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<mods:title id="2A67AF32FA54DDA91B2896736650BFA8">Taxonomy of the freshwater crabs of Costa Rica, with a revision of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Pseudothelphusidae)</mods:title>
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<treatment id="710D87BB2913160ED6BDFAA9FB7BFE00" ID-DOI="http://doi.org/10.5281/zenodo.5491508" ID-GBIF-Taxon="119590161" ID-Zenodo-Dep="5491508" LSID="urn:lsid:plazi:treatment:710D87BB2913160ED6BDFAA9FB7BFE00" httpUri="http://treatment.plazi.org/id/710D87BB2913160ED6BDFAA9FB7BFE00" lastPageId="6" lastPageNumber="307" pageId="5" pageNumber="306">
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<paragraph id="F91B36AD2913160DD6BDFAA9FD71FAF5" blockId="5.[151,672,1290,1350]" box="[151,672,1290,1317]" pageId="5" pageNumber="306">
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<taxonomicName id="3EA44D2E2913160DD6BDFAA9FD71FAF5" ID-CoL="78N74" authority="Pretzmann, 1965" authorityName="Pretzmann" authorityYear="1965" box="[151,672,1290,1317]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="cocleensis">
<emphasis id="CBD0EABF2913160DD6BDFAA9FD71FAF5" bold="true" box="[151,672,1290,1317]" pageId="5" pageNumber="306">
<emphasis id="CBD0EABF2913160DD6BDFAA9FE14FAF4" bold="true" box="[151,453,1290,1316]" italics="true" pageId="5" pageNumber="306">Ptychophallus cocleensis</emphasis>
<bibRefCitation id="9D354B5C2913160DD7E6FAA8FD71FAF5" author="Pretzmann" box="[460,672,1291,1317]" pageId="5" pageNumber="306" refString="Pretzmann, G. (1965) Vorlaufiger Bericht uber die Familie Pseudothelphusidae. Sitzungsberichte der Mathematisch- Naturwissenschaftlichen Klasse der Osterreichischen Akademie der Wissenschaften, Wien, 1, 1 - 11." type="journal article" year="1965">Pretzmann, 1965</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="F91B36AD2913160DD6BDFA8EFEEAFA96" blockId="5.[151,672,1290,1350]" box="[151,315,1325,1350]" pageId="5" pageNumber="306">
(
<figureCitation id="619F2A282913160DD6B5FA8EFEDBFA96" box="[159,266,1325,1350]" captionStart="FIGURES 1 9" captionStartId="4.[151,264,1764,1787]" captionTargetBox="[152,1431,193,1735]" captionTargetId="figure@4.[152,1436,193,1743]" captionTargetPageId="4" captionText="FIGURES 1 9. Achlidon agrestis (Rathbun, 1898): 1, left first gonopod (partially damaged), holotype, USNM 19487, mesial view; 2, same data, lateral view of distal part; 3, left first gonopod, NHMW 21827, cephalic view of distal part; 4, same data, apical view of apex; 5, same data, latero-cephalic view. Achlidon puntarenas (Hobbs, 1991): 6, left first gonopod, holotype, USNM 250555, caudo-mesial view; 7, same data, latero-cephalic view of distal part. Ptychophallus cocleensis Pretzmann, 1965: 8, left first gonopod, holotype, USNM 119869, lateral view of distal part; 9, same data, mesial view (dr, disto-caudal ridge; fs, field of apical spines; lp, lateral process; me, mesial process; mp, marginal process; ms, marginal suture). Scale bars = 1 mm." httpUri="https://zenodo.org/record/235128/files/figure.png" pageId="5" pageNumber="306">Figs. 8, 9</figureCitation>
,
<figureCitation id="619F2A282913160DD73CFA8DFEE2FA96" box="[278,307,1326,1350]" captionStart="FIGURE 78" captionStartId="32.[151,250,1218,1240]" captionTargetBox="[156,1432,201,1194]" captionTargetId="figure@32.[151,1436,193,1196]" captionTargetPageId="32" captionText="FIGURE 78. Distribution map of Ptychophallus cocleensis Pretzmann, 1965, P. lavallensis Pretzmann, 1978, and P. micracanthus Rodríguez, 1994." httpUri="https://zenodo.org/record/235136/files/figure.png" pageId="5" pageNumber="306">78</figureCitation>
)
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<taxonomicName id="3EA44D2E2913160DD6BDFAD6FCA4FA5B" ID-CoL="78N74" authority="Pretzmann, 1965: 5" authorityName="Pretzmann" authorityPageNumber="5" authorityYear="1965" box="[151,885,1397,1419]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="306" phylum="Arthropoda" rank="subSpecies" species="montanus" subGenus="Ptychophallus" subSpecies="cocleensis">
<emphasis id="CBD0EABF2913160DD6BDFAD6FEF9FA5B" box="[151,296,1397,1419]" italics="true" pageId="5" pageNumber="306">Ptychophallus</emphasis>
(
<emphasis id="CBD0EABF2913160DD71CFAD6FE16FA5B" box="[310,455,1397,1419]" italics="true" pageId="5" pageNumber="306">Ptychophallus</emphasis>
)
<emphasis id="CBD0EABF2913160DD7FCFAD5FD77FA5B" box="[470,678,1397,1419]" italics="true" pageId="5" pageNumber="306">montanus cocleensis</emphasis>
<treatmentCitation id="780510BC2913160DD487FAD5FCA4FA5B" author="Pretzmann" box="[685,885,1397,1419]" page="5" pageId="5" pageNumber="306" year="1965">
<bibRefCitation id="9D354B5C2913160DD487FAD5FC8DFA5B" author="Pretzmann" box="[685,860,1397,1419]" pageId="5" pageNumber="306" refString="Pretzmann, G. (1965) Vorlaufiger Bericht uber die Familie Pseudothelphusidae. Sitzungsberichte der Mathematisch- Naturwissenschaftlichen Klasse der Osterreichischen Akademie der Wissenschaften, Wien, 1, 1 - 11." type="journal article" year="1965">Pretzmann, 1965</bibRefCitation>
: 5
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.
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</paragraph>
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<treatmentCitationGroup id="D9B411832913160DD6BDFA37FAC2FA7A" box="[151,1299,1428,1450]" pageId="5" pageNumber="306">
<taxonomicName id="3EA44D2E2913160DD6BDFA37FDEFFA7A" ID-CoL="78N74" box="[151,574,1428,1450]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="cocleensis" subGenus="Ptychophallus">
<emphasis id="CBD0EABF2913160DD6BDFA37FEF9FA7A" box="[151,296,1428,1450]" italics="true" pageId="5" pageNumber="306">Ptychophallus</emphasis>
(
<emphasis id="CBD0EABF2913160DD71CFA37FE16FA7A" box="[310,455,1428,1450]" italics="true" pageId="5" pageNumber="306">Ptychophallus</emphasis>
)
<emphasis id="CBD0EABF2913160DD7FCFA37FDEFFA7A" box="[470,574,1428,1450]" italics="true" pageId="5" pageNumber="306">cocleensis</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC2913160DD472FA36FCF0FA7A" author="Pretzmann" box="[600,801,1428,1450]" page="21" pageId="5" pageNumber="306" year="1971">
<bibRefCitation id="9D354B5C2913160DD472FA36FCD1FA7A" author="Pretzmann" box="[600,768,1428,1450]" pageId="5" pageNumber="306" refString="Pretzmann, G. (1971) Fortschritte in der Klassifizierung der Pseudothelphusidae. Anzeiger der Mathematisch Naturwissenschaftliche Klasse der Osterreichischen Akademie der Wissenschaften, Series 1, 179 (1 - 4), 12 - 24." type="journal article" year="1971">Pretzmann 1971</bibRefCitation>
: 21
</treatmentCitation>
.—
<treatmentCitation id="780510BC2913160DD514FA37FBC1FA7A" author="Pretzmann" box="[830,1040,1428,1450]" page="88" pageId="5" pageNumber="306" year="1972">
<bibRefCitation id="9D354B5C2913160DD514FA37FC3AFA7A" author="Pretzmann" box="[830,1003,1428,1450]" pageId="5" pageNumber="306" refString="Pretzmann, G. (1972) Die Pseudothelphusidae (Crustacea Brachyura). Zoologica, 42 (120), pt. 1, 1 - 182." type="journal article" year="1972">Pretzmann 1972</bibRefCitation>
: 88
</treatmentCitation>
, figs. 505507, 534, 535.
</treatmentCitationGroup>
</paragraph>
<paragraph id="F91B36AD2913160DD6BDFA11FD25F9D6" blockId="5.[151,1436,1397,1572]" pageId="5" pageNumber="306">
<treatmentCitationGroup id="D9B411832913160DD6BDFA11FD25F9D6" pageId="5" pageNumber="306">
<taxonomicName id="3EA44D2E2913160DD6BDFA11FE47FA18" ID-CoL="78N74" box="[151,406,1458,1480]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="cocleensis">
<emphasis id="CBD0EABF2913160DD6BDFA11FE47FA18" box="[151,406,1458,1480]" italics="true" pageId="5" pageNumber="306">Ptychophallus cocleensis</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC2913160DD79AFA11FDA8FA18" author="Rodriguez" box="[432,633,1458,1480]" page="86" pageId="5" pageNumber="306" year="1982">
<bibRefCitation id="9D354B5C2913160DD79AFA11FD87FA18" author="Rodriguez" box="[432,598,1458,1480]" pageId="5" pageNumber="306" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Familie des Pseudothelphusidae. ORSTOM, Paris, 223 pp. [Faune Tropicale 22]" type="book" year="1982">Rodriguez 1982</bibRefCitation>
: 86
</treatmentCitation>
.—
<treatmentCitation id="780510BC2913160DD4BDFA11FCBEFA18" author="Villalobos" box="[663,879,1458,1480]" page="220" pageId="5" pageNumber="306" year="1982">
<bibRefCitation id="9D354B5C2913160DD4BDFA11FCEEFA18" author="Villalobos" box="[663,831,1458,1480]" pageId="5" pageNumber="306" refString="Villalobos, A. (1982) Decapoda. In: Hurlbert, S. H. &amp; Villalobos, A. (Eds.), Aquatic Biota of Mexico, Central America and the West Indies. San Diego State University, San Diego, pp. 215 - 239." type="book chapter" year="1982">Villalobos 1982</bibRefCitation>
: 220
</treatmentCitation>
(in list).—
<treatmentCitation id="780510BC2913160DD5F7FA11FB6BFA18" author="Rodriguez" box="[989,1210,1458,1480]" page="302" pageId="5" pageNumber="306" year="1994">
<bibRefCitation id="9D354B5C2913160DD5F7FA11FB56FA18" author="Rodriguez" box="[989,1159,1458,1480]" pageId="5" pageNumber="306" refString="Rodriguez, G. (1994) A revision of the type material of some species of Hypolobocera and Ptychophallus (Crustacea: Decapoda: Pseudothelphusidae) in the National Museum of Natural History, Washington, D. C., with descriptions of a new species and a new subspecies. Proceedings of the Biological Society of Washington, 107 (2), 296 - 307." type="journal article" year="1994">Rodríguez 1994</bibRefCitation>
: 302
</treatmentCitation>
, fig. 3D, E.—
<treatmentCitation id="780510BC2913160DD36DFA11FE12FA37" author="Campos" page="560" pageId="5" pageNumber="306" year="1999">
<bibRefCitation id="9D354B5C2913160DD36DFA11FE5BFA37" author="Campos" pageId="5" pageNumber="306" refString="Campos, M. R. &amp; Lemaitre, R. (1999) Two new freshwater crabs of the genus Ptychophallus Smalley, 1964 (Crustacea: Decapoda: Brachyura: Pseudothelphusidae) from Panama, with notes on the distribution of the genus. Proceedings of the Biological Society of Washington, 112 (3), 553 - 561." type="journal article" year="1999">Campos &amp; Lemaitre 1999</bibRefCitation>
: 560
</treatmentCitation>
(map).—
<treatmentCitation id="780510BC2913160DD400FA72FC7CFA37" author="Rodriguez" box="[554,941,1489,1511]" page="423" pageId="5" pageNumber="306" year="2000">
<bibRefCitation id="9D354B5C2913160DD400FA72FCA2FA37" author="Rodriguez" box="[554,883,1489,1511]" pageId="5" pageNumber="306" refString="Rodriguez, G. &amp; Hedstrom, I. (2000) The freswater crabs of the Barbilla National Park, Costa Rica (Crustacea: Brachyura: Pseudothelphusidae), with notes on the evolution of structures for spermatophore retention. Proceedings of the Biological Society of Washington, 113 (2), 420 - 425." type="journal article" year="2000">Rodríguez &amp; Hedström 2000</bibRefCitation>
: 423
</treatmentCitation>
, 424, fig. 2D, I.—
<treatmentCitation id="780510BC2913160DD2A2FA71FAA0FA37" author="Ng" box="[1160,1393,1489,1511]" page="176" pageId="5" pageNumber="306" year="2008">
<bibRefCitation id="9D354B5C2913160DD2A2FA71FAE9FA37" author="Ng" box="[1160,1336,1489,1511]" pageId="5" pageNumber="306" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology Supplement, 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="CBD0EABF2913160DD29DFA71FB25FA37" box="[1207,1268,1489,1511]" italics="true" pageId="5" pageNumber="306">et al.</emphasis>
2008
</bibRefCitation>
: 176
</treatmentCitation>
(in list).—
<treatmentCitation id="780510BC2913160DD721FA53FD4DF9D6" author="Villalobos" box="[267,668,1520,1542]" page="297" pageId="5" pageNumber="306" year="2008">
<bibRefCitation id="9D354B5C2913160DD721FA53FDBDF9D6" author="Villalobos" box="[267,620,1520,1542]" pageId="5" pageNumber="306" refString="Villalobos Hiriart, J. L. &amp; Alvarez, F. (2008) Los cangrejos de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico, con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F. &amp; Rodriguez- Almaraz, G. (Eds.), Crustaceos de Mexico: Estado actual de su Conocimiento. Universidad Autonoma de Nuevo Leon, Monterrey, Nuevo Leon, pp. 239 - 300. [Mexico]" type="book chapter" year="2008">Villalobos Hiriart &amp; Álvarez 2008</bibRefCitation>
: 297
</treatmentCitation>
(in list).
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</paragraph>
<paragraph id="F91B36AD2913160DD6BDF9ADFC9FF9F4" blockId="5.[151,1436,1397,1572]" box="[151,846,1550,1572]" pageId="5" pageNumber="306">
<treatmentCitationGroup id="D9B411832913160DD6BDF9ADFC9FF9F4" box="[151,846,1550,1572]" pageId="5" pageNumber="306">
?
<taxonomicName id="3EA44D2E2913160DD682F9ADFE7AF9F4" ID-CoL="6WN77" box="[168,427,1550,1572]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="goldmanni">
<emphasis id="CBD0EABF2913160DD682F9ADFE7AF9F4" box="[168,427,1550,1572]" italics="true" pageId="5" pageNumber="306">Ptychophallus goldmanni</emphasis>
</taxonomicName>
<treatmentCitation id="780510BC2913160DD7ECF9ADFD7FF9F4" author="Abele" box="[454,686,1550,1572]" page="39" pageId="5" pageNumber="306" year="1989">
<bibRefCitation id="9D354B5C2913160DD7ECF9ADFD5AF9F4" author="Abele" box="[454,651,1550,1572]" pageId="5" pageNumber="306" refString="Abele, L. G. &amp; Kim, W. (1989) The decapod crustaceans of the Panama Canal. Smithsonian Contributions to Zoology, 482, 1 - 50. http: // dx. doi. org / 10.5479 / si. 00810282.482" type="journal article" year="1989">Abele &amp; Kim 1989</bibRefCitation>
: 39
</treatmentCitation>
(see Remarks).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
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<emphasis id="CBD0EABF2913160DD6BDF9F2FEACF9BA" bold="true" box="[151,381,1617,1642]" pageId="5" pageNumber="306">Material examined</emphasis>
.
<collectingCountry id="81B3763D2913160DD7A4F9F1FDD8F9BA" box="[398,521,1618,1642]" name="Panama" pageId="5" pageNumber="306">PANAMA</collectingCountry>
. Colón Province, Atlantic drainage: male,
<typeStatus id="261F880F2913160DD5D0F9F2FB8EF9BA" box="[1018,1119,1617,1642]" pageId="5" pageNumber="306" type="holotype">holotype</typeStatus>
,
<collectionCode id="9FB5AE682913160DD244F9F1FB6EF9BA" LSID="urn:lsid:biocol.org:col:34871" box="[1134,1215,1618,1642]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="306">USNM</collectionCode>
119869, Coclé del Norte River, 1951, M.W. Stirling leg.—
<collectingCountry id="81B3763D2913160DD440F9D6FD17F95E" box="[618,710,1653,1678]" name="Panama" pageId="5" pageNumber="306">Panamá</collectingCountry>
Province:
<specimenCount id="EFA2FD242913160DD562F9D5FC4AF95E" box="[840,923,1653,1678]" pageId="5" pageNumber="306" type="male">1 male</specimenCount>
,
<specimenCount id="EFA2FD242913160DD586F9D5FBC4F95E" box="[940,1045,1653,1678]" pageId="5" pageNumber="306" type="female">1 female</specimenCount>
,
<collectionCode id="9FB5AE682913160DD20FF9D5FBA7F95E" LSID="urn:lsid:biocol.org:col:34871" box="[1061,1142,1654,1678]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="306">USNM</collectionCode>
230097, Barro Colorado Island, Wheeler trail,
<date id="8D1A106D2913160DD7A0F93AFE2FF962" box="[394,510,1689,1714]" pageId="5" pageNumber="306" value="1969-07-09">9.vii.1969</date>
, R. Foster leg.
</paragraph>
</subSubSection>
<subSubSection id="B1BE65262913160DD6EDF91EFBC4F87E" pageId="5" pageNumber="306" type="description">
<paragraph id="F91B36AD2913160DD6EDF91EFBC4F87E" blockId="5.[151,1437,1617,2038]" pageId="5" pageNumber="306">
<emphasis id="CBD0EABF2913160DD6EDF91EFE3CF906" bold="true" box="[199,493,1725,1750]" pageId="5" pageNumber="306">Description of gonopod</emphasis>
. Straight in mesiocaudal view, apex bent approximately 90° in laterocephalic direction. Marginal suture on mesial side, straight; marginal process subtriangular, not produced beyond the caudal border of apex. Mesial process subtriangular, slightly downturned, narrow. Lateral process strongly divided into 2 lobes by deep median notch; proximal lobe smaller than distal lobe, digitiform, curved in distal direction, with few scattered, minute setae; distal lobe broadly rounded, swollen, projected beyond distal end of caudal ridge. Distocaudal ridge short, narrow. Apex oblong, narrow; caudal border slightly convex, with small notch near lateral end. Field of apical spines well developed, directed towards laterocephalic side.
</paragraph>
</subSubSection>
<subSubSection id="B1BE65262913160DD6EDF81AFE69F826" pageId="5" pageNumber="306" type="materials_examined">
<paragraph id="F91B36AD2913160DD6EDF81AFE69F826" blockId="5.[151,1437,1617,2038]" pageId="5" pageNumber="306">
<emphasis id="CBD0EABF2913160DD6EDF81AFE2CF802" bold="true" box="[199,509,1977,2002]" pageId="5" pageNumber="306">
<typeStatus id="261F880F2913160DD6EDF81AFEE5F802" box="[199,308,1977,2002]" pageId="5" pageNumber="306" type="holotype">Holotype</typeStatus>
and
<typeStatus id="261F880F2913160DD744F81AFE70F802" box="[366,417,1977,2002]" pageId="5" pageNumber="306">type</typeStatus>
locality
</emphasis>
. Male, cw 26.4, cl 15.8,
<collectionCode id="9FB5AE682913160DD53BF819FCB3F802" LSID="urn:lsid:biocol.org:col:34871" box="[785,866,1978,2002]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="306">USNM</collectionCode>
119869.
<collectingCountry id="81B3763D2913160DD5E3F819FBF4F801" box="[969,1061,1978,2001]" name="Panama" pageId="5" pageNumber="306">Panama</collectingCountry>
, Colón Province, Coclé del Norte River (Atlantic drainage).
</paragraph>
</subSubSection>
<subSubSection id="B1BE65262910160ED6EDFF34FC9EFF04" pageId="6" pageNumber="307" type="distribution">
<paragraph id="F91B36AD2910160ED6EDFF34FC9EFF04" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="307">
<emphasis id="CBD0EABF2910160ED6EDFF34FE88FF60" bold="true" box="[199,345,151,176]" pageId="6" pageNumber="307">Distribution</emphasis>
. The only two known records suggest a distribution in central
<collectingCountry id="81B3763D2910160ED203FF3BFB54FF7F" box="[1065,1157,152,175]" name="Panama" pageId="6" pageNumber="307">Panama</collectingCountry>
, provinces of Colón and
<collectingCountry id="81B3763D2910160ED6BDFF1FFF22FF04" box="[151,243,188,212]" name="Panama" pageId="6" pageNumber="307">Panamá</collectingCountry>
, probably restricted to the Atlantic drainage (
<figureCitation id="619F2A282910160ED4C4FF1FFC93FF04" box="[750,834,188,213]" captionStart="FIGURE 78" captionStartId="32.[151,250,1218,1240]" captionTargetBox="[156,1432,201,1194]" captionTargetId="figure@32.[151,1436,193,1196]" captionTargetPageId="32" captionText="FIGURE 78. Distribution map of Ptychophallus cocleensis Pretzmann, 1965, P. lavallensis Pretzmann, 1978, and P. micracanthus Rodríguez, 1994." httpUri="https://zenodo.org/record/235136/files/figure.png" pageId="6" pageNumber="307">Fig. 78</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="B1BE65262910160ED6EDFF7CFB7BFE00" pageId="6" pageNumber="307" type="discussion">
<paragraph id="F91B36AD2910160ED6EDFF7CFB7BFE00" blockId="6.[151,1437,151,464]" pageId="6" pageNumber="307">
<emphasis id="CBD0EABF2910160ED6EDFF7CFEE5FF28" bold="true" box="[199,308,223,248]" pageId="6" pageNumber="307">Remarks</emphasis>
. The record of two males of
<taxonomicName id="3EA44D2E2910160ED4A0FF7CFC7DFF28" box="[650,940,223,248]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="goldmanni">
<emphasis id="CBD0EABF2910160ED4A0FF7CFC7DFF28" box="[650,940,223,248]" italics="true" pageId="6" pageNumber="307">Ptychophallus goldmanni</emphasis>
</taxonomicName>
from Barro Colorado Island, in the former
<collectingCountry id="81B3763D2910160ED6BDFEA6FF21FECC" box="[151,240,261,284]" name="Panama" pageId="6" pageNumber="307">Panama</collectingCountry>
Canal Zone made by
<bibRefCitation id="9D354B5C2910160ED7C1FEA7FCD5FECC" author="Abele" box="[491,772,260,285]" pageId="6" pageNumber="307" refString="Abele, L. G. &amp; Kim, W. (1989) The decapod crustaceans of the Panama Canal. Smithsonian Contributions to Zoology, 482, 1 - 50. http: // dx. doi. org / 10.5479 / si. 00810282.482" type="journal article" year="1989">Abele &amp; Kim (1989: 39)</bibRefCitation>
possibly refers to
<taxonomicName id="3EA44D2E2910160ED5FCFEA6FBB7FECD" box="[982,1126,260,285]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="cocleensis">
<emphasis id="CBD0EABF2910160ED5FCFEA6FBB7FECD" box="[982,1126,260,285]" italics="true" pageId="6" pageNumber="307">P. cocleensis</emphasis>
</taxonomicName>
. One of us (CM) could not find these specimens in the USNM holdings. A male and female specimen collected during the same period (
<date id="8D1A106D2910160ED346FE84FF01FEB5" pageId="6" pageNumber="307" value="1969-07">July 1969</date>
) and nearly at the same place (Barro Colorado Island, Wheeler trail) (USNM 230097), however, was positively identified as
<taxonomicName id="3EA44D2E2910160ED7B6FED2FDFBFE58" box="[412,554,367,392]" class="Malacostraca" family="Pseudothelphusidae" genus="Ptychophallus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="cocleensis">
<emphasis id="CBD0EABF2910160ED7B6FED2FDFBFE58" box="[412,554,367,392]" italics="true" pageId="6" pageNumber="307">P. cocleensis</emphasis>
</taxonomicName>
. Although
<bibRefCitation id="9D354B5C2910160ED482FECCFC40FE58" author="Abele" box="[680,913,367,392]" pageId="6" pageNumber="307" refString="Abele, L. G. &amp; Kim, W. (1989) The decapod crustaceans of the Panama Canal. Smithsonian Contributions to Zoology, 482, 1 - 50. http: // dx. doi. org / 10.5479 / si. 00810282.482" type="journal article" year="1989">Abele &amp; Kim (1989)</bibRefCitation>
mentioned that their specimens were collected by T. Zaret at the Zetek trail, the lot USNM 230099 has two small labels in the respective jar stating &quot;
<taxonomicName id="3EA44D2E2910160ED688FE14FEB0FE00" box="[162,353,439,464]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="307" phylum="Arthropoda" rank="genus">
<emphasis id="CBD0EABF2910160ED688FE14FEB0FE00" box="[162,353,439,464]" italics="true" pageId="6" pageNumber="307">Pseudothelphusa</emphasis>
</taxonomicName>
sp. illustrated by Abele&quot;, and &quot;male illustrated in decapod
<collectingCountry id="81B3763D2910160ED5DCFE1BFB9EFE1F" box="[1014,1103,440,463]" name="Panama" pageId="6" pageNumber="307">Panama</collectingCountry>
Canal&quot;.
</paragraph>
</subSubSection>
</treatment>
</document>