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<document id="4D00C22237D4CE044237079E49C7BF88" ID-CLB-Dataset="29089" ID-DOI="10.11646/zootaxa.4455.3.2" ID-GBIF-Dataset="839210a1-c9d7-4833-893d-24d9f29c72d2" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457427" ID-ZooBank="A01F20BE-F0C4-4B45-97C5-E996A2C84EE1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539262663038" checkinUser="plazi" docAuthor="Yamana, Yusuke &amp; Kohtsuka, Hisanori" docDate="2018" docId="6D72879EFFAB0C60D7A2F906E3FB8CF0" docLanguage="en" docName="zootaxa.4455.3.2.pdf" docOrigin="Zootaxa 4455 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lipotrapeza purpurata Yamana &amp; Kohtsuka 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="445" masterDocId="914BFFE6FFA00C70D735FFC4E0238F70" masterDocTitle="Dendrochirotid holothurians (Echinodermata: Holothuroidea: Dendrochirotida) including four new species, from off Misaki, Japan" masterLastPageNumber="453" masterPageNumber="429" pageNumber="440" updateTime="1698575945573" updateUser="ExternalLinkService">
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<mods:title id="5061EE182EF355A0F20B5DFDCB67D399">Dendrochirotid holothurians (Echinodermata: Holothuroidea: Dendrochirotida) including four new species, from off Misaki, Japan</mods:title>
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<mods:namePart id="759F421ACD9C177218D2D2E2A0903A4F">Yamana, Yusuke</mods:namePart>
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<mods:namePart id="B5D5042C967A929843FD834CD45CE83B">Kohtsuka, Hisanori</mods:namePart>
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<treatment id="6D72879EFFAB0C60D7A2F906E3FB8CF0" ID-DOI="http://doi.org/10.5281/zenodo.5951766" ID-GBIF-Taxon="148691376" ID-Zenodo-Dep="5951766" LSID="urn:lsid:plazi:treatment:6D72879EFFAB0C60D7A2F906E3FB8CF0" httpUri="http://treatment.plazi.org/id/6D72879EFFAB0C60D7A2F906E3FB8CF0" lastPageId="16" lastPageNumber="445" pageId="11" pageNumber="440">
<subSubSection id="ADC16503FFAB0C7BD7A2F906E2C1898E" pageId="11" pageNumber="440" type="nomenclature">
<paragraph id="E5643688FFAB0C7BD7A2F906E22F89AD" blockId="11.[151,738,1730,1826]" box="[151,524,1730,1757]" pageId="11" pageNumber="440">
<heading id="BE2C81E4FFAB0C7BD7A2F906E22F89AD" bold="true" box="[151,524,1730,1757]" fontSize="11" level="1" pageId="11" pageNumber="440" reason="1">
<emphasis id="D7AFEA9AFFAB0C7BD7A2F906E22F89AD" bold="true" box="[151,524,1730,1757]" pageId="11" pageNumber="440">
<taxonomicName id="22DB4D0BFFAB0C7BD7A2F906E18489AC" ID-CoL="3VBF9" authority="Yamana &amp; Kohtsuka, 2018" authorityName="Yamana &amp; Kohtsuka" authorityYear="2018" box="[151,423,1730,1756]" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="11" pageNumber="440" phylum="Echinodermata" rank="species" species="purpurata" status="sp. nov.">
<emphasis id="D7AFEA9AFFAB0C7BD7A2F906E18489AC" bold="true" box="[151,423,1730,1756]" italics="true" pageId="11" pageNumber="440">Lipotrapeza purpurata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9C57E1FFAB0C7BD69BF907E22F89AD" box="[430,524,1731,1757]" pageId="11" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E5643688FFAB0C7BD7A2F922E2C1898E" blockId="11.[151,738,1730,1826]" box="[151,738,1765,1790]" pageId="11" pageNumber="440">[New Japanese name: Murasaki-honenashi-namako]</paragraph>
</subSubSection>
<subSubSection id="ADC16503FFAB0C7BD7A2F8CDE1588852" box="[151,379,1801,1826]" pageId="11" pageNumber="440" type="description">
<paragraph id="E5643688FFAB0C7BD7A2F8CDE1588852" blockId="11.[151,738,1730,1826]" box="[151,379,1801,1826]" pageId="11" pageNumber="440">
(
<figureCitation id="7DE02A0DFFAB0C7BD7AAF8CDE0DB8852" box="[159,248,1801,1826]" captionStart="FIGURE 1" captionStartId="4.[151,250,1855,1877]" captionTargetBox="[199,1397,199,1836]" captionTargetId="figure@4.[193,1400,195,1838]" captionTargetPageId="4" captionText="FIGURE 1. Lateral views of the specimens, dorsal upper, anterior left. A, Amphicyclus japonicus Bell, 1884, (a) WMNH-INV- 2015-13, (b) -2015-14, (c) -2015-311, (d) -2015-312; B, Neocucumis misakiensis sp. nov., (e) WMNH-INV-2016-208, (f) - 2016-209; C, Pseudocolochirus misakiensis sp. nov., (g) WMNH-INV-2015-348, (h) -2015-349, (i) -2015-350; D, Hemiocnus tegulatus (Augustin, 1908) comb. nov., (j) WMNH-INV-2015-378, (k) -2015-379; E, Lipotrapeza purpurata sp. nov., (l) WMNH-INV-2015-38, (m) -2015-39, (n) -2015-40; F, Pentamera misakiensis sp. nov., (o) WMNH-INV-2015-34, (p) -2015- 35, (q) -2015-36; G, Pseudoplacothuria ohshimai (Liao, 1997) gen. nov., comb. nov., (r) WMNH-INV-2015-9, (s) -2015-10." httpUri="https://zenodo.org/record/1457429/files/figure.png" pageId="11" pageNumber="440">Figs 1E</figureCitation>
,
<figureCitation id="7DE02A0DFFAB0C7BD631F8CEE1058852" box="[260,294,1802,1826]" captionStart="FIGURE 2" captionStartId="5.[151,250,1810,1832]" captionTargetBox="[240,1366,199,1750]" captionTargetId="figure@5.[212,1376,193,1795]" captionTargetPageId="5" captionText="FIGURE 2. Elements of calcareous rings, viewed from body cavity, anterior upper. A, Amphicyclus japonicus Bell, 1884 (WMNH-INV-2015-14); B, Neocucumis misakiensis sp. nov. (WMNH-INV-2016-208); C, Pseudocolochirus misakiensis sp. nov. (WMNH-INV-2015-350); D, Hemiocnus tegulatus (Augustin, 1908) comb. nov. (WMNH-INV-2015-379); E, Lipotrapeza purpurata sp. nov. (WMNH-INV-2015-39); Pentamera misakiensis sp. nov. (WMNH-INV-2015-34); Pseudoplacothuria ohshimai (Liao, 1997) gen. nov., comb. nov. (WMNH-INV-2015-9). Abbreviations: ld, left-dorsal part; lv, left-ventral part; md, mid-dorsal part; mv, mid-ventral part; rd, right-dorsal part; rv, right-ventral part; -ir, inter radial element; -r, radial element." httpUri="https://zenodo.org/record/1457431/files/figure.png" pageId="11" pageNumber="440">2E</figureCitation>
,
<figureCitation id="7DE02A0DFFAB0C7BD607F8CEE1508852" box="[306,371,1802,1826]" captionStart="FIGURE 7" captionStartId="14.[151,250,1878,1900]" captionTargetBox="[187,1400,196,1864]" captionTargetId="figure@14.[184,1404,193,1867]" captionTargetPageId="14" captionText="FIGURE 7. SEM image of ossicles of Lipotrapeza purpurata sp. nov. (AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla." httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="11" pageNumber="440">7AP</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="ADC16503FFAB0C7FD7A2F895E3C58E6C" lastPageId="15" lastPageNumber="444" pageId="11" pageNumber="440" type="materials_examined">
<paragraph id="E5643688FFAB0C7BD7A2F895E43F88A6" blockId="11.[151,1437,1873,2006]" pageId="11" pageNumber="440">
<emphasis id="D7AFEA9AFFAB0C7BD7A2F895E1A5881A" bold="true" box="[151,390,1873,1898]" pageId="11" pageNumber="440">Material examined.</emphasis>
<emphasis id="D7AFEA9AFFAB0C7BD6A6F895E1DA8819" box="[403,505,1873,1897]" italics="true" pageId="11" pageNumber="440">Holotype</emphasis>
, WMNH-INV-
<date id="91651048FFAB0C7BD585F895E32B881A" box="[688,776,1873,1898]" pageId="11" pageNumber="440" value="2015" valueMax="2039">2015-39</date>
(St.
<date id="91651048FFAB0C7BD47CF896E3C6881A" box="[841,997,1873,1898]" pageId="11" pageNumber="440" value="1926-08-01">1, 26 August</date>
2014, length
<quantity id="22239B6DFFAB0C7BD3A5F896E4C48819" box="[1168,1255,1874,1898]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="11" pageNumber="440" unit="mm" value="21.0">21 mm</quantity>
, width
<quantity id="22239B6DFFAB0C7BD270F895E5B3881A" box="[1349,1424,1873,1898]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="11" pageNumber="440" unit="mm" value="5.0">5 mm</quantity>
).
<emphasis id="D7AFEA9AFFAB0C7BD7A2F8B2E12A88FD" box="[151,265,1910,1933]" italics="true" pageId="11" pageNumber="440">Paratypes</emphasis>
: WMNH-INV-
<date id="91651048FFAB0C7BD68AF8B1E23488FE" box="[447,535,1909,1934]" pageId="11" pageNumber="440" value="2015" valueMax="2038">2015-38</date>
(St.
<date id="91651048FFAB0C7BD563F8B2E2D388FE" box="[598,752,1909,1934]" pageId="11" pageNumber="440" value="1926-08-01">1, 26 August</date>
2014, length
<quantity id="22239B6DFFAB0C7BD4ACF8B2E3CC88FD" box="[921,1007,1910,1934]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="11" pageNumber="440" unit="mm" value="28.0">28 mm</quantity>
, width
<quantity id="22239B6DFFAB0C7BD37EF8B2E4B088FD" box="[1099,1171,1910,1934]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="11" pageNumber="440" unit="mm" value="9.0">9 mm</quantity>
, partially eviscerated); WMNH-INV-
<date id="91651048FFAB0C7BD60DF85DE1B188C2" box="[312,402,1945,1970]" pageId="11" pageNumber="440" value="2015" valueMax="2040">2015-40</date>
(St.
<date id="91651048FFAB0C7BD6FFF85EE27E88C2" box="[458,605,1945,1970]" pageId="11" pageNumber="440" value="1926-08-01">1, 26 August</date>
2014, length
<quantity id="22239B6DFFAB0C7BD5CEF85EE36E88C1" box="[763,845,1946,1970]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="11" pageNumber="440" unit="mm" value="32.0">32 mm</quantity>
, width
<quantity id="22239B6DFFAB0C7BD496F85EE3D788C1" box="[931,1012,1946,1970]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="11" pageNumber="440" unit="mm" value="11.0">11 mm</quantity>
, extremely expanded); WMNH-INV- 2016-4 (St.
<date id="91651048FFAB0C7BD62EF87AE18888A6" box="[283,427,1981,2006]" pageId="11" pageNumber="440" value="1926-08-01">1, 26 August</date>
2014, length
<quantity id="22239B6DFFAB0C7BD570F87AE2AB88A5" box="[581,648,1982,2006]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="11" pageNumber="440" unit="mm" value="8.0">8 mm</quantity>
, width
<quantity id="22239B6DFFAB0C7BD5EEF87AE33E88A5" box="[731,797,1982,2006]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="440" unit="mm" value="3.0">3 mm</quantity>
, partially eviscerated).
</paragraph>
<caption id="B1A46600FFAC0C7CD7A2F8B2E2FC8898" httpUri="https://zenodo.org/record/1457439/files/figure.png" pageId="12" pageNumber="441" startId="12.[151,250,1910,1932]" targetBox="[178,1412,197,1897]" targetPageId="12">
<paragraph id="E5643688FFAC0C7CD7A2F8B2E2FC8898" blockId="12.[151,1436,1910,2024]" pageId="12" pageNumber="441">
<emphasis id="D7AFEA9AFFAC0C7CD7A2F8B2E13788FB" bold="true" box="[151,276,1910,1932]" pageId="12" pageNumber="441">FIGURE 5.</emphasis>
SEM image of ossicles of
<taxonomicName id="22DB4D0BFFAC0C7CD51AF8B2E34188FC" authority="Yamana &amp; Kohtsuka, 2018" authorityName="Yamana &amp; Kohtsuka" authorityYear="2018" box="[559,866,1910,1932]" class="Holothuroidea" family="Cucumariidae" genus="Pseudocolochirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="12" pageNumber="441" phylum="Echinodermata" rank="species" species="misakiensis" status="sp. nov.">
<emphasis id="D7AFEA9AFFAC0C7CD51AF8B2E34188FC" box="[559,866,1910,1932]" italics="true" pageId="12" pageNumber="441">Pseudocolochirus misakiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9C57E1FFAC0C7CD45CF8B2E39888FB" box="[873,955,1910,1931]" pageId="12" pageNumber="441" rank="species">
<emphasis id="D7AFEA9AFFAC0C7CD45CF8B2E39888FB" bold="true" box="[873,955,1910,1931]" pageId="12" pageNumber="441">sp. nov.</emphasis>
</taxonomicNameLabel>
(AG, WMNH-INV-2015-350). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral skin of introvert; E the pedicel of the abdominal side; F and G, the middle part of the body wall on the abdominal side, and on the dorsal side, respectively; H, the ventrolateral anal papilla.
</paragraph>
</caption>
<caption id="B1A46600FFAD0C7DD7A2F8B1E5BF8897" httpUri="https://zenodo.org/record/1457441/files/figure.png" pageId="13" pageNumber="442" startId="13.[151,250,1909,1931]" targetBox="[180,1416,196,1896]" targetPageId="13">
<paragraph id="E5643688FFAD0C7DD7A2F8B1E5BF8897" blockId="13.[151,1436,1909,2023]" pageId="13" pageNumber="442">
<emphasis id="D7AFEA9AFFAD0C7DD7A2F8B1E13688FA" bold="true" box="[151,277,1909,1931]" pageId="13" pageNumber="442">FIGURE 6.</emphasis>
SEM image of ossicles of
<taxonomicName id="22DB4D0BFFAD0C7DD507F8B2E39488FB" authority="(Augustin, 1908)" authorityName="Yamana &amp; Kohtsuka" authorityYear="2018" baseAuthorityName="Augustin" baseAuthorityYear="1908" box="[562,951,1909,1931]" class="Holothuroidea" family="Cucumariidae" genus="Hemiocnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="13" pageNumber="442" phylum="Echinodermata" rank="species" species="tegulata" status="comb. nov.">
<emphasis id="D7AFEA9AFFAD0C7DD507F8B2E2DC88FB" box="[562,767,1909,1931]" italics="true" pageId="13" pageNumber="442">Hemiocnus tegulata</emphasis>
(Augustin, 1908)
</taxonomicName>
<taxonomicNameLabel id="CC9C57E1FFAD0C7DD48AF8B2E41688FB" box="[959,1077,1910,1931]" pageId="13" pageNumber="442">
<emphasis id="D7AFEA9AFFAD0C7DD48AF8B2E41688FB" bold="true" box="[959,1077,1910,1931]" pageId="13" pageNumber="442">comb. nov.</emphasis>
</taxonomicNameLabel>
(AI, WMNH-INV-2015-378). A, ossicles of the ventrolateral tentacle; B, the ventrolateral skin of peri-oral; C, the ventrolateral skin of pharynx; D, the ventrolateral skin of introvert; E the pedicel of the abdominal side; F and G, the middle part of the body wall on the abdominal side, and on the dorsal side, respectively; H, extra large ossicles of body wall on the dorsal side; I, the ventrolateral anal papilla.
</paragraph>
</caption>
<caption id="B1A46600FFAE0C7ED7A2F892E0C08883" httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="14" pageNumber="443" startId="14.[151,250,1878,1900]" targetBox="[187,1400,196,1864]" targetPageId="14">
<paragraph id="E5643688FFAE0C7ED7A2F892E0C08883" blockId="14.[151,1436,1878,2035]" pageId="14" pageNumber="443">
<emphasis id="D7AFEA9AFFAE0C7ED7A2F892E130881C" bold="true" box="[151,275,1878,1900]" pageId="14" pageNumber="443">FIGURE 7.</emphasis>
SEM image of ossicles of
<taxonomicName id="22DB4D0BFFAE0C7ED512F893E32C881C" authority="Yamana &amp; Kohtsuka, 2018" authorityName="Yamana &amp; Kohtsuka" authorityYear="2018" box="[551,783,1879,1900]" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="14" pageNumber="443" phylum="Echinodermata" rank="species" species="purpurata" status="sp. nov.">
<emphasis id="D7AFEA9AFFAE0C7ED512F893E32C881C" box="[551,783,1879,1900]" italics="true" pageId="14" pageNumber="443">Lipotrapeza purpurata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9C57E1FFAE0C7ED420F893E345881C" box="[789,870,1879,1900]" pageId="14" pageNumber="443" rank="species">
<emphasis id="D7AFEA9AFFAE0C7ED420F893E345881C" bold="true" box="[789,870,1879,1900]" pageId="14" pageNumber="443">sp. nov.</emphasis>
</taxonomicNameLabel>
(AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla.
</paragraph>
</caption>
<paragraph id="E5643688FFAF0C7FD7F2FF5CE3C58E6C" blockId="15.[151,1437,151,1112]" pageId="15" pageNumber="444">
<emphasis id="D7AFEA9AFFAF0C7FD7F2FF5CE1568FC0" box="[199,373,152,176]" italics="true" pageId="15" pageNumber="444">Other material</emphasis>
: 1 specimen, WMNH-INV-2015-8 (St.
<date id="91651048FFAF0C7FD463FF5CE3D78FC0" box="[854,1012,151,176]" pageId="15" pageNumber="444" value="1926-08-01">1, 26 August</date>
2014, length
<quantity id="22239B6DFFAF0C7FD396FF5CE4CE8FDF" box="[1187,1261,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="15" pageNumber="444" unit="mm" value="9.0">9 mm</quantity>
, width
<quantity id="22239B6DFFAF0C7FD278FF5CE5BB8FDF" box="[1357,1432,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="444" unit="mm" value="3.0">3 mm</quantity>
, eviscerating); 1 specimen, WMNH-INV-
<date id="91651048FFAF0C7FD550FF78E29E8FA5" box="[613,701,188,213]" pageId="15" pageNumber="444" value="2015" valueMax="2037">2015-37</date>
(St.
<date id="91651048FFAF0C7FD5C7FF78E3A18FA4" box="[754,898,188,213]" pageId="15" pageNumber="444" value="1926-08-01">1, 26 August</date>
2014, length
<quantity id="22239B6DFFAF0C7FD32EFF78E44F8FA4" box="[1051,1132,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="444" unit="mm" value="13.0">13 mm</quantity>
, width
<quantity id="22239B6DFFAF0C7FD38BFF78E5208FA4" box="[1214,1283,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="444" unit="mm" value="6.0">6 mm</quantity>
); 1 specimen, WMNH-INV-
<date id="91651048FFAF0C7FD60FFF1BE1B18F88" box="[314,402,223,248]" pageId="15" pageNumber="444" value="2015" valueMax="2041">2015-41</date>
(St.
<date id="91651048FFAF0C7FD6FBFF24E2458F88" box="[462,614,223,248]" pageId="15" pageNumber="444" value="1926-08-01">1, 26 August</date>
2014, length
<quantity id="22239B6DFFAF0C7FD439FF24E3438F87" box="[780,864,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="15" pageNumber="444" unit="mm" value="21.0">21 mm</quantity>
, width
<quantity id="22239B6DFFAF0C7FD48FFF1BE4228F87" box="[954,1025,223,248]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="444" unit="mm" value="6.0">6 mm</quantity>
, partially eviscerated); 1 specimen, WMNH-INV-
<date id="91651048FFAF0C7FD60FFEC0E1B18E6C" box="[314,402,260,284]" pageId="15" pageNumber="444" value="2017" valueMax="2041">2017-41</date>
(St.
<date id="91651048FFAF0C7FD6F2FEC0E2748E6C" box="[455,599,260,285]" pageId="15" pageNumber="444" value="1926-08-01">1, 26 August</date>
2014, length
<quantity id="22239B6DFFAF0C7FD5C4FEC0E3618E6C" box="[753,834,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="15" pageNumber="444" unit="mm" value="13.0">13 mm</quantity>
, width
<quantity id="22239B6DFFAF0C7FD4A1FEC0E3FA8E6C" box="[916,985,260,285]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="15" pageNumber="444" unit="mm" value="6.0">6 mm</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="ADC16503FFAF0C60D7F2FEE3E18E8E85" lastPageId="16" lastPageNumber="445" pageId="15" pageNumber="444" type="description">
<paragraph id="E5643688FFAF0C7FD7F2FEE3E2008DF5" blockId="15.[151,1437,151,1112]" pageId="15" pageNumber="444">
<emphasis id="D7AFEA9AFFAF0C7FD7F2FEE3E1D38E30" bold="true" box="[199,496,295,320]" pageId="15" pageNumber="444">
Description of
<typeStatus id="3A60882AFFAF0C7FD6B6FEE3E1CF8E30" box="[387,492,295,320]" pageId="15" pageNumber="444" type="holotype">holotype</typeStatus>
.
</emphasis>
Body fusiform, straight or curved, with both ends tapered and turned slightly upwards (
<figureCitation id="7DE02A0DFFAF0C7FD63FFE88E1478E14" box="[266,356,332,357]" captionStart="FIGURE 1" captionStartId="4.[151,250,1855,1877]" captionTargetBox="[199,1397,199,1836]" captionTargetId="figure@4.[193,1400,195,1838]" captionTargetPageId="4" captionText="FIGURE 1. Lateral views of the specimens, dorsal upper, anterior left. A, Amphicyclus japonicus Bell, 1884, (a) WMNH-INV- 2015-13, (b) -2015-14, (c) -2015-311, (d) -2015-312; B, Neocucumis misakiensis sp. nov., (e) WMNH-INV-2016-208, (f) - 2016-209; C, Pseudocolochirus misakiensis sp. nov., (g) WMNH-INV-2015-348, (h) -2015-349, (i) -2015-350; D, Hemiocnus tegulatus (Augustin, 1908) comb. nov., (j) WMNH-INV-2015-378, (k) -2015-379; E, Lipotrapeza purpurata sp. nov., (l) WMNH-INV-2015-38, (m) -2015-39, (n) -2015-40; F, Pentamera misakiensis sp. nov., (o) WMNH-INV-2015-34, (p) -2015- 35, (q) -2015-36; G, Pseudoplacothuria ohshimai (Liao, 1997) gen. nov., comb. nov., (r) WMNH-INV-2015-9, (s) -2015-10." httpUri="https://zenodo.org/record/1457429/files/figure.png" pageId="15" pageNumber="444">Fig. 1E</figureCitation>
); body wall thick and soft. Body color dark purplish brown or brownish white (living and preserved specimens), inside of body cavity purplish white and inside of pedicel dark purple (preserved specimens). Tentacles 20, arranged in double circle (15 + 5), including five pairs of large inter radial tentacles and five single small radial tentacles in outer circle and five small radial tentacles in inner circle. Color of tentacles and introvert, cream or whitish-brown (living and preserved material). Several rows of villi-like projections surrounding oral opening, short and wide, dark purplish brown (living and preserving). Pedicels non-retractile or partially retractile, scattered over entire body, lacking on introvert, becoming gradually smaller anteriorly and posteriorly. Color of pedicels same as body wall, occasionally tips tinged with dark purplish color. Ten anal papillae and five anal teeth in radii.
</paragraph>
<paragraph id="E5643688FFAF0C7FD7F2FD4BE5BE8C65" blockId="15.[151,1437,151,1112]" pageId="15" pageNumber="444">
Calcareous ring long (
<figureCitation id="7DE02A0DFFAF0C7FD6E1FD4BE2118DD8" box="[468,562,655,680]" captionStart="FIGURE 2" captionStartId="5.[151,250,1810,1832]" captionTargetBox="[240,1366,199,1750]" captionTargetId="figure@5.[212,1376,193,1795]" captionTargetPageId="5" captionText="FIGURE 2. Elements of calcareous rings, viewed from body cavity, anterior upper. A, Amphicyclus japonicus Bell, 1884 (WMNH-INV-2015-14); B, Neocucumis misakiensis sp. nov. (WMNH-INV-2016-208); C, Pseudocolochirus misakiensis sp. nov. (WMNH-INV-2015-350); D, Hemiocnus tegulatus (Augustin, 1908) comb. nov. (WMNH-INV-2015-379); E, Lipotrapeza purpurata sp. nov. (WMNH-INV-2015-39); Pentamera misakiensis sp. nov. (WMNH-INV-2015-34); Pseudoplacothuria ohshimai (Liao, 1997) gen. nov., comb. nov. (WMNH-INV-2015-9). Abbreviations: ld, left-dorsal part; lv, left-ventral part; md, mid-dorsal part; mv, mid-ventral part; rd, right-dorsal part; rv, right-ventral part; -ir, inter radial element; -r, radial element." httpUri="https://zenodo.org/record/1457431/files/figure.png" pageId="15" pageNumber="444">Fig. 2E</figureCitation>
), radial and most of interradial elements not fragmented. Radials entire, anteriorly notched, carrying paired, fragmented, posterior prolongations. Interradials plates sagittate, sharply pointed anteriorly, divided into two fragments, lacking posterior prolongations. Polian vesicle and stone canal single. Gonad situated in mid-body, in two clusters, one on each side of dorsal mesentery, most tubules unbranched.
</paragraph>
<paragraph id="E5643688FFAF0C7FD7F2FCDBE4648B28" blockId="15.[151,1437,151,1112]" pageId="15" pageNumber="444">
The
<typeStatus id="3A60882AFFAF0C7FD7CCFCDBE1788C48" box="[249,347,799,824]" pageId="15" pageNumber="444" type="holotype">holotype</typeStatus>
possesses rosettes in the body wall (
<figureCitation id="7DE02A0DFFAF0C7FD5C0FCDBE3458C48" box="[757,870,799,824]" captionStart="FIGURE 7" captionStartId="14.[151,250,1878,1900]" captionTargetBox="[187,1400,196,1864]" captionTargetId="figure@14.[184,1404,193,1867]" captionTargetPageId="14" captionText="FIGURE 7. SEM image of ossicles of Lipotrapeza purpurata sp. nov. (AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla." httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="15" pageNumber="444">Fig. 7F, G</figureCitation>
,
<tableCitation id="A8590333FFAF0C7FD444FCDBE3E68C48" box="[881,965,799,824]" captionStart="TABLE 5" captionStartId="15.[151,239,1160,1183]" captionText="TABLE 5. Measurements (µm) of ossicles from four specimens of Lipotrapeza purpurata sp. nov." pageId="15" pageNumber="444">Table 5</tableCitation>
), typical of the open
<typeStatus id="3A60882AFFAF0C7FD387FCE4E4C68C48" box="[1202,1253,800,824]" pageId="15" pageNumber="444">type</typeStatus>
, branches rarely overlapping. Anal papilla with rosettes, rods, tables, and plates (
<figureCitation id="7DE02A0DFFAF0C7FD45CFC80E3E18C2C" box="[873,962,836,861]" captionStart="FIGURE 7" captionStartId="14.[151,250,1878,1900]" captionTargetBox="[187,1400,196,1864]" captionTargetId="figure@14.[184,1404,193,1867]" captionTargetPageId="14" captionText="FIGURE 7. SEM image of ossicles of Lipotrapeza purpurata sp. nov. (AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla." httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="15" pageNumber="444">Fig. 7H</figureCitation>
,
<tableCitation id="A8590333FFAF0C7FD4FAFC80E4078C2D" box="[975,1060,836,861]" captionStart="TABLE 5" captionStartId="15.[151,239,1160,1183]" captionText="TABLE 5. Measurements (µm) of ossicles from four specimens of Lipotrapeza purpurata sp. nov." pageId="15" pageNumber="444">Table 5</tableCitation>
). Tentacle ossicles comprise rods and rosettes (
<figureCitation id="7DE02A0DFFAF0C7FD603FCA3E1B68CF0" box="[310,405,871,896]" captionStart="FIGURE 7" captionStartId="14.[151,250,1878,1900]" captionTargetBox="[187,1400,196,1864]" captionTargetId="figure@14.[184,1404,193,1867]" captionTargetPageId="14" captionText="FIGURE 7. SEM image of ossicles of Lipotrapeza purpurata sp. nov. (AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla." httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="15" pageNumber="444">Fig. 7A</figureCitation>
,
<tableCitation id="A8590333FFAF0C7FD693FCA3E2238CF0" box="[422,512,871,896]" captionStart="TABLE 5" captionStartId="15.[151,239,1160,1183]" captionText="TABLE 5. Measurements (µm) of ossicles from four specimens of Lipotrapeza purpurata sp. nov." pageId="15" pageNumber="444">Table 5</tableCitation>
); rods, short, simple, with small numbers of distal branches and a few distal perforations. Peri-oral skin, pharyngeal villi and introvert with rosettes (
<figureCitation id="7DE02A0DFFAF0C7FD489FC48E4168CD4" box="[956,1077,908,933]" captionStart="FIGURE 7" captionStartId="14.[151,250,1878,1900]" captionTargetBox="[187,1400,196,1864]" captionTargetId="figure@14.[184,1404,193,1867]" captionTargetPageId="14" captionText="FIGURE 7. SEM image of ossicles of Lipotrapeza purpurata sp. nov. (AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla." httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="15" pageNumber="444">Fig. 7BD</figureCitation>
,
<tableCitation id="A8590333FFAF0C7FD375FC48E4B68CD5" box="[1088,1173,908,933]" captionStart="TABLE 5" captionStartId="15.[151,239,1160,1183]" captionText="TABLE 5. Measurements (µm) of ossicles from four specimens of Lipotrapeza purpurata sp. nov." pageId="15" pageNumber="444">Table 5</tableCitation>
). Pedicels with circular endplate and other supporting plates (
<figureCitation id="7DE02A0DFFAF0C7FD571FC6BE2BF8CB7" box="[580,668,943,968]" captionStart="FIGURE 7" captionStartId="14.[151,250,1878,1900]" captionTargetBox="[187,1400,196,1864]" captionTargetId="figure@14.[184,1404,193,1867]" captionTargetPageId="14" captionText="FIGURE 7. SEM image of ossicles of Lipotrapeza purpurata sp. nov. (AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla." httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="15" pageNumber="444">Fig. 7E</figureCitation>
,
<tableCitation id="A8590333FFAF0C7FD59CFC6BE3238CB8" box="[681,768,943,968]" captionStart="TABLE 5" captionStartId="15.[151,239,1160,1183]" captionText="TABLE 5. Measurements (µm) of ossicles from four specimens of Lipotrapeza purpurata sp. nov." pageId="15" pageNumber="444">Table 5</tableCitation>
). Endplate with approximately uniform-sized perforations; perforations of supporting plates mostly arranged in single or double rows medially and three or more rows distally. Anal papillae with mostly simple rods distally, other rods present in middle to basal part, and simple-shaped tables and rosettes present only in basal part of anal papilla and anal periphery. Most tables with low spire with four pillars and eight teeth, disc with 8-10 marginal perforations. Gonad lacking ossicles.
</paragraph>
<caption id="B1A46600FFAF0C7FD7A2FB4CE4968BEF" box="[151,1205,1160,1183]" pageId="15" pageNumber="444">
<paragraph id="E5643688FFAF0C7FD7A2FB4CE4968BEF" blockId="15.[151,1205,1160,1183]" box="[151,1205,1160,1183]" pageId="15" pageNumber="444">
<emphasis id="D7AFEA9AFFAF0C7FD7A2FB4CE1298BEF" bold="true" box="[151,266,1160,1183]" pageId="15" pageNumber="444">TABLE 5.</emphasis>
Measurements (µm) of ossicles from four specimens of
<taxonomicName id="22DB4D0BFFAF0C7FD451FB4CE47B8BEF" authority="Yamana &amp; Kohtsuka, 2018" authorityName="Yamana &amp; Kohtsuka" authorityYear="2018" box="[868,1112,1160,1183]" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="15" pageNumber="444" phylum="Echinodermata" rank="species" species="purpurata" status="sp. nov.">
<emphasis id="D7AFEA9AFFAF0C7FD451FB4CE47B8BEF" box="[868,1112,1160,1183]" italics="true" pageId="15" pageNumber="444">Lipotrapeza purpurata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CC9C57E1FFAF0C7FD36AFB4CE4968BEF" box="[1119,1205,1160,1183]" pageId="15" pageNumber="444" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="E5643688FFAF0C7FD6BCFB7EE55C8890" pageId="15" pageNumber="444">
<table id="97DBC428FFAFF38FD7AAFB7EE5AF8890" box="[159,1420,1210,2016]" gridcols="13" gridrows="17" pageId="15" pageNumber="444">
<tr id="5BEB34CAFFAFF38FD7AAFB7EE5AF8BA0" box="[159,1420,1210,1232]" gridrow="0" pageId="15" pageNumber="444" rowspan-0="1">
<th id="183A5DB6FFAFF38FD6BCFB7EE2A18BA0" box="[393,642,1210,1232]" colspan="3" colspanRight="2" gridcol="1" gridrow="0" pageId="15" pageNumber="444">WMNH-INV-2015-38</th>
<th id="183A5DB6FFAFF38FD5A6FB7EE3AA8BA0" box="[659,905,1210,1232]" colspan="3" colspanRight="2" gridcol="4" gridrow="0" pageId="15" pageNumber="444">WMNH-INV-2015-39</th>
<th id="183A5DB6FFAFF38FD4AFFB7EE4B28BA0" box="[922,1169,1210,1232]" colspan="3" colspanRight="2" gridcol="7" gridrow="0" pageId="15" pageNumber="444">WMNH-INV-2015-40</th>
<th id="183A5DB6FFAFF38FD397FB7EE5AF8BA0" box="[1186,1420,1210,1232]" colspan="3" colspanRight="2" gridcol="10" gridrow="0" pageId="15" pageNumber="444">WMNH-INV-2016-4</th>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAFB20E5AF8B8A" box="[159,1420,1252,1274]" gridrow="1" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAFB20E15A8B8A" box="[159,377,1252,1274]" gridcol="0" gridrow="1" pageId="15" pageNumber="444">Ossicle type</th>
<td id="183A5DB6FFAFF38FD6BCFB20E1808B8A" box="[393,419,1252,1274]" gridcol="1" gridrow="1" pageId="15" pageNumber="444">n</td>
<td id="183A5DB6FFAFF38FD68DFB20E2358B8A" box="[440,534,1252,1274]" gridcol="2" gridrow="1" pageId="15" pageNumber="444">Mean±sd</td>
<td id="183A5DB6FFAFF38FD51FFB20E2A18B8A" box="[554,642,1252,1274]" gridcol="3" gridrow="1" pageId="15" pageNumber="444">Range</td>
<td id="183A5DB6FFAFF38FD5A6FB20E28E8B8A" box="[659,685,1252,1274]" gridcol="4" gridrow="1" pageId="15" pageNumber="444">n</td>
<td id="183A5DB6FFAFF38FD58BFB20E3AA8B8A" box="[702,905,1252,1274]" colspan="2" colspanRight="1" gridcol="5" gridrow="1" pageId="15" pageNumber="444">Mean±sd Range</td>
<td id="183A5DB6FFAFF38FD4AFFB20E3978B8A" box="[922,948,1252,1274]" gridcol="7" gridrow="1" pageId="15" pageNumber="444">n</td>
<td id="183A5DB6FFAFF38FD4F3FB20E4078B8A" box="[966,1060,1252,1274]" gridcol="8" gridrow="1" pageId="15" pageNumber="444">Mean±sd</td>
<td id="183A5DB6FFAFF38FD30DFB20E4B28B8A" box="[1080,1169,1252,1274]" gridcol="9" gridrow="1" pageId="15" pageNumber="444">Range</td>
<td id="183A5DB6FFAFF38FD397FB20E4988B8A" box="[1186,1211,1252,1274]" gridcol="10" gridrow="1" pageId="15" pageNumber="444">n</td>
<td id="183A5DB6FFAFF38FD3FBFB20E5AF8B8A" box="[1230,1420,1252,1274]" colspan="2" colspanRight="1" gridcol="11" gridrow="1" pageId="15" pageNumber="444">Mean±sd Range</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAFAC9E5AF8A53" box="[159,1420,1293,1315]" gridrow="2" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAFAC9E15A8A53" box="[159,377,1293,1315]" gridcol="0" gridrow="2" pageId="15" pageNumber="444">Tentacle rod</th>
<td id="183A5DB6FFAFF38FD6BCFAC9E1808A53" box="[393,419,1293,1315]" gridcol="1" gridrow="2" pageId="15" pageNumber="444">21</td>
<td id="183A5DB6FFAFF38FD68DFAC9E2358A53" box="[440,534,1293,1315]" gridcol="2" gridrow="2" pageId="15" pageNumber="444">91±26</td>
<td id="183A5DB6FFAFF38FD51FFAC9E2A18A53" box="[554,642,1293,1315]" gridcol="3" gridrow="2" pageId="15" pageNumber="444">50132</td>
<td id="183A5DB6FFAFF38FD5A6FAC9E28E8A53" box="[659,685,1293,1315]" gridcol="4" gridrow="2" pageId="15" pageNumber="444">12</td>
<td id="183A5DB6FFAFF38FD58BFAC9E32F8A53" box="[702,780,1293,1315]" gridcol="5" gridrow="2" pageId="15" pageNumber="444">81±14</td>
<td id="183A5DB6FFAFF38FD405FAC9E3AA8A53" box="[816,905,1293,1315]" gridcol="6" gridrow="2" pageId="15" pageNumber="444">65115</td>
<td id="183A5DB6FFAFF38FD4AFFAC9E3978A53" box="[922,948,1293,1315]" gridcol="7" gridrow="2" pageId="15" pageNumber="444">28</td>
<td id="183A5DB6FFAFF38FD4F3FAC9E4078A53" box="[966,1060,1293,1315]" gridcol="8" gridrow="2" pageId="15" pageNumber="444">112±56</td>
<td id="183A5DB6FFAFF38FD30DFAC9E4B28A53" box="[1080,1169,1293,1315]" gridcol="9" gridrow="2" pageId="15" pageNumber="444">50246</td>
<td id="183A5DB6FFAFF38FD397FAC9E4988A53" box="[1186,1211,1293,1315]" gridcol="10" gridrow="2" pageId="15" pageNumber="444">22</td>
<td id="183A5DB6FFAFF38FD3FBFAC9E53F8A53" box="[1230,1308,1293,1315]" gridcol="11" gridrow="2" pageId="15" pageNumber="444">71±9</td>
<td id="183A5DB6FFAFF38FD275FAC9E5AF8A53" box="[1344,1420,1293,1315]" gridcol="12" gridrow="2" pageId="15" pageNumber="444">5593</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAFAF2E5AF8A3C" box="[159,1420,1334,1356]" gridrow="3" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAFAF2E15A8A3C" box="[159,377,1334,1356]" gridcol="0" gridrow="3" pageId="15" pageNumber="444">Tentacle rosette</th>
<td id="183A5DB6FFAFF38FD6BCFAF2E1808A3C" box="[393,419,1334,1356]" gridcol="1" gridrow="3" pageId="15" pageNumber="444">15</td>
<td id="183A5DB6FFAFF38FD68DFAF2E2358A3C" box="[440,534,1334,1356]" gridcol="2" gridrow="3" pageId="15" pageNumber="444">55±8</td>
<td id="183A5DB6FFAFF38FD51FFAF2E2A18A3C" box="[554,642,1334,1356]" gridcol="3" gridrow="3" pageId="15" pageNumber="444">3770</td>
<td id="183A5DB6FFAFF38FD5A6FAF2E28E8A3C" box="[659,685,1334,1356]" gridcol="4" gridrow="3" pageId="15" pageNumber="444">16</td>
<td id="183A5DB6FFAFF38FD58BFAF2E32F8A3C" box="[702,780,1334,1356]" gridcol="5" gridrow="3" pageId="15" pageNumber="444">56±12</td>
<td id="183A5DB6FFAFF38FD405FAF2E3AA8A3C" box="[816,905,1334,1356]" gridcol="6" gridrow="3" pageId="15" pageNumber="444">3878</td>
<td id="183A5DB6FFAFF38FD4AFFAF2E3978A3C" box="[922,948,1334,1356]" gridcol="7" gridrow="3" pageId="15" pageNumber="444">0</td>
<td id="183A5DB6FFAFF38FD4F3FAF2E4078A3C" box="[966,1060,1334,1356]" gridcol="8" gridrow="3" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD30DFAF2E4B28A3C" box="[1080,1169,1334,1356]" gridcol="9" gridrow="3" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD397FAF2E4988A3C" box="[1186,1211,1334,1356]" gridcol="10" gridrow="3" pageId="15" pageNumber="444">32</td>
<td id="183A5DB6FFAFF38FD3FBFAF2E53F8A3C" box="[1230,1308,1334,1356]" gridcol="11" gridrow="3" pageId="15" pageNumber="444">49±9</td>
<td id="183A5DB6FFAFF38FD275FAF2E5AF8A3C" box="[1344,1420,1334,1356]" gridcol="12" gridrow="3" pageId="15" pageNumber="444">3269</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAFAA4E5AF8A06" box="[159,1420,1376,1398]" gridrow="4" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAFAA4E15A8A06" box="[159,377,1376,1398]" gridcol="0" gridrow="4" pageId="15" pageNumber="444">Peri-oral rosette</th>
<td id="183A5DB6FFAFF38FD6BCFAA4E1808A06" box="[393,419,1376,1398]" gridcol="1" gridrow="4" pageId="15" pageNumber="444">41</td>
<td id="183A5DB6FFAFF38FD68DFAA4E2358A06" box="[440,534,1376,1398]" gridcol="2" gridrow="4" pageId="15" pageNumber="444">33±6</td>
<td id="183A5DB6FFAFF38FD51FFAA4E2A18A06" box="[554,642,1376,1398]" gridcol="3" gridrow="4" pageId="15" pageNumber="444">2446</td>
<td id="183A5DB6FFAFF38FD5A6FAA4E28E8A06" box="[659,685,1376,1398]" gridcol="4" gridrow="4" pageId="15" pageNumber="444">20</td>
<td id="183A5DB6FFAFF38FD58BFAA4E32F8A06" box="[702,780,1376,1398]" gridcol="5" gridrow="4" pageId="15" pageNumber="444">46±10</td>
<td id="183A5DB6FFAFF38FD405FAA4E3AA8A06" box="[816,905,1376,1398]" gridcol="6" gridrow="4" pageId="15" pageNumber="444">3274</td>
<td id="183A5DB6FFAFF38FD4AFFAA4E3978A06" box="[922,948,1376,1398]" gridcol="7" gridrow="4" pageId="15" pageNumber="444">7</td>
<td id="183A5DB6FFAFF38FD4F3FAA4E4078A06" box="[966,1060,1376,1398]" gridcol="8" gridrow="4" pageId="15" pageNumber="444">58±15</td>
<td id="183A5DB6FFAFF38FD30DFAA4E4B28A06" box="[1080,1169,1376,1398]" gridcol="9" gridrow="4" pageId="15" pageNumber="444">4187</td>
<td id="183A5DB6FFAFF38FD397FAA4E4988A06" box="[1186,1211,1376,1398]" gridcol="10" gridrow="4" pageId="15" pageNumber="444">44</td>
<td id="183A5DB6FFAFF38FD3FBFAA4E53F8A06" box="[1230,1308,1376,1398]" gridcol="11" gridrow="4" pageId="15" pageNumber="444">30±5</td>
<td id="183A5DB6FFAFF38FD275FAA4E5AF8A06" box="[1344,1420,1376,1398]" gridcol="12" gridrow="4" pageId="15" pageNumber="444">2139</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAFA4DE5AF8AEF" box="[159,1420,1417,1439]" gridrow="5" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAFA4DE15A8AEF" box="[159,377,1417,1439]" gridcol="0" gridrow="5" pageId="15" pageNumber="444">Pharyngeal rosette</th>
<td id="183A5DB6FFAFF38FD6BCFA4DE1808AEF" box="[393,419,1417,1439]" gridcol="1" gridrow="5" pageId="15" pageNumber="444">43</td>
<td id="183A5DB6FFAFF38FD68DFA4DE2358AEF" box="[440,534,1417,1439]" gridcol="2" gridrow="5" pageId="15" pageNumber="444">30±5</td>
<td id="183A5DB6FFAFF38FD51FFA4DE2A18AEF" box="[554,642,1417,1439]" gridcol="3" gridrow="5" pageId="15" pageNumber="444">2242</td>
<td id="183A5DB6FFAFF38FD5A6FA4DE28E8AEF" box="[659,685,1417,1439]" gridcol="4" gridrow="5" pageId="15" pageNumber="444">10</td>
<td id="183A5DB6FFAFF38FD58BFA4DE32F8AEF" box="[702,780,1417,1439]" gridcol="5" gridrow="5" pageId="15" pageNumber="444">37±6</td>
<td id="183A5DB6FFAFF38FD405FA4DE3AA8AEF" box="[816,905,1417,1439]" gridcol="6" gridrow="5" pageId="15" pageNumber="444">2646</td>
<td id="183A5DB6FFAFF38FD4AFFA4DE3978AEF" box="[922,948,1417,1439]" gridcol="7" gridrow="5" pageId="15" pageNumber="444">13</td>
<td id="183A5DB6FFAFF38FD4F3FA4DE4078AEF" box="[966,1060,1417,1439]" gridcol="8" gridrow="5" pageId="15" pageNumber="444">38±9</td>
<td id="183A5DB6FFAFF38FD30DFA4DE4B28AEF" box="[1080,1169,1417,1439]" gridcol="9" gridrow="5" pageId="15" pageNumber="444">2859</td>
<td id="183A5DB6FFAFF38FD397FA4DE4988AEF" box="[1186,1211,1417,1439]" gridcol="10" gridrow="5" pageId="15" pageNumber="444">38</td>
<td id="183A5DB6FFAFF38FD3FBFA4DE53F8AEF" box="[1230,1308,1417,1439]" gridcol="11" gridrow="5" pageId="15" pageNumber="444">30±5</td>
<td id="183A5DB6FFAFF38FD275FA4DE5AF8AEF" box="[1344,1420,1417,1439]" gridcol="12" gridrow="5" pageId="15" pageNumber="444">1841</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAFA76E5AF8A97" box="[159,1420,1458,1511]" gridrow="6" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAFA76E15A8A97" box="[159,377,1458,1511]" gridcol="0" gridrow="6" pageId="15" pageNumber="444">Introvert rosette &amp; plate</th>
<td id="183A5DB6FFAFF38FD6BCFA76E1808A97" box="[393,419,1458,1511]" gridcol="1" gridrow="6" pageId="15" pageNumber="444">32</td>
<td id="183A5DB6FFAFF38FD68DFA76E2358A97" box="[440,534,1458,1511]" gridcol="2" gridrow="6" pageId="15" pageNumber="444">43±8</td>
<td id="183A5DB6FFAFF38FD51FFA76E2A18A97" box="[554,642,1458,1511]" gridcol="3" gridrow="6" pageId="15" pageNumber="444">3371</td>
<td id="183A5DB6FFAFF38FD5A6FA76E28E8A97" box="[659,685,1458,1511]" gridcol="4" gridrow="6" pageId="15" pageNumber="444">9</td>
<td id="183A5DB6FFAFF38FD58BFA76E32F8A97" box="[702,780,1458,1511]" gridcol="5" gridrow="6" pageId="15" pageNumber="444">50±8</td>
<td id="183A5DB6FFAFF38FD405FA76E3AA8A97" box="[816,905,1458,1511]" gridcol="6" gridrow="6" pageId="15" pageNumber="444">3559</td>
<td id="183A5DB6FFAFF38FD4AFFA76E3978A97" box="[922,948,1458,1511]" gridcol="7" gridrow="6" pageId="15" pageNumber="444">20</td>
<td id="183A5DB6FFAFF38FD4F3FA76E4078A97" box="[966,1060,1458,1511]" gridcol="8" gridrow="6" pageId="15" pageNumber="444">47±19</td>
<td id="183A5DB6FFAFF38FD30DFA76E4B28A97" box="[1080,1169,1458,1511]" gridcol="9" gridrow="6" pageId="15" pageNumber="444">27118</td>
<td id="183A5DB6FFAFF38FD397FA76E4988A97" box="[1186,1211,1458,1511]" gridcol="10" gridrow="6" pageId="15" pageNumber="444">25</td>
<td id="183A5DB6FFAFF38FD3FBFA76E53F8A97" box="[1230,1308,1458,1511]" gridcol="11" gridrow="6" pageId="15" pageNumber="444">36±4</td>
<td id="183A5DB6FFAFF38FD275FA76E5AF8A97" box="[1344,1420,1458,1511]" gridcol="12" gridrow="6" pageId="15" pageNumber="444">2847</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAFA3EE5AF8960" box="[159,1420,1530,1552]" gridrow="7" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAFA3EE15A8960" box="[159,377,1530,1552]" gridcol="0" gridrow="7" pageId="15" pageNumber="444">Pedicel endplate</th>
<td id="183A5DB6FFAFF38FD6BCFA3EE1808960" box="[393,419,1530,1552]" gridcol="1" gridrow="7" pageId="15" pageNumber="444">8</td>
<td id="183A5DB6FFAFF38FD68DFA3EE2358960" box="[440,534,1530,1552]" gridcol="2" gridrow="7" pageId="15" pageNumber="444">224±11</td>
<td id="183A5DB6FFAFF38FD51FFA3EE2A18960" box="[554,642,1530,1552]" gridcol="3" gridrow="7" pageId="15" pageNumber="444">210238</td>
<td id="183A5DB6FFAFF38FD5A6FA3EE28E8960" box="[659,685,1530,1552]" gridcol="4" gridrow="7" pageId="15" pageNumber="444">3</td>
<td id="183A5DB6FFAFF38FD58BFA3EE32F8960" box="[702,780,1530,1552]" gridcol="5" gridrow="7" pageId="15" pageNumber="444">208±21</td>
<td id="183A5DB6FFAFF38FD405FA3EE3AA8960" box="[816,905,1530,1552]" gridcol="6" gridrow="7" pageId="15" pageNumber="444">185227</td>
<td id="183A5DB6FFAFF38FD4AFFA3EE3978960" box="[922,948,1530,1552]" gridcol="7" gridrow="7" pageId="15" pageNumber="444">3</td>
<td id="183A5DB6FFAFF38FD4F3FA3EE4078960" box="[966,1060,1530,1552]" gridcol="8" gridrow="7" pageId="15" pageNumber="444">215±15</td>
<td id="183A5DB6FFAFF38FD30DFA3EE4B28960" box="[1080,1169,1530,1552]" gridcol="9" gridrow="7" pageId="15" pageNumber="444">200229</td>
<td id="183A5DB6FFAFF38FD397FA3EE4988960" box="[1186,1211,1530,1552]" gridcol="10" gridrow="7" pageId="15" pageNumber="444">6</td>
<td id="183A5DB6FFAFF38FD3FBFA3EE53F8960" box="[1230,1308,1530,1552]" gridcol="11" gridrow="7" pageId="15" pageNumber="444">169±42</td>
<td id="183A5DB6FFAFF38FD275FA3EE5AF8960" box="[1344,1420,1530,1552]" gridcol="12" gridrow="7" pageId="15" pageNumber="444">85194</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAF9E0E5AF8928" box="[159,1420,1572,1624]" gridrow="8" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAF9E0E15A8928" box="[159,377,1572,1624]" gridcol="0" gridrow="8" pageId="15" pageNumber="444">Pedicel supporting plate</th>
<td id="183A5DB6FFAFF38FD6BCF9E0E1808928" box="[393,419,1572,1624]" gridcol="1" gridrow="8" pageId="15" pageNumber="444">8</td>
<td id="183A5DB6FFAFF38FD68DF9E0E2358928" box="[440,534,1572,1624]" gridcol="2" gridrow="8" pageId="15" pageNumber="444">131±30</td>
<td id="183A5DB6FFAFF38FD51FF9E0E2A18928" box="[554,642,1572,1624]" gridcol="3" gridrow="8" pageId="15" pageNumber="444">75169</td>
<td id="183A5DB6FFAFF38FD5A6F9E0E28E8928" box="[659,685,1572,1624]" gridcol="4" gridrow="8" pageId="15" pageNumber="444">5</td>
<td id="183A5DB6FFAFF38FD58BF9E0E32F8928" box="[702,780,1572,1624]" gridcol="5" gridrow="8" pageId="15" pageNumber="444">149±21</td>
<td id="183A5DB6FFAFF38FD405F9E0E3AA8928" box="[816,905,1572,1624]" gridcol="6" gridrow="8" pageId="15" pageNumber="444">133184</td>
<td id="183A5DB6FFAFF38FD4AFF9E0E3978928" box="[922,948,1572,1624]" gridcol="7" gridrow="8" pageId="15" pageNumber="444">5</td>
<td id="183A5DB6FFAFF38FD4F3F9E0E4078928" box="[966,1060,1572,1624]" gridcol="8" gridrow="8" pageId="15" pageNumber="444">153±27</td>
<td id="183A5DB6FFAFF38FD30DF9E0E4B28928" box="[1080,1169,1572,1624]" gridcol="9" gridrow="8" pageId="15" pageNumber="444">125195</td>
<td id="183A5DB6FFAFF38FD397F9E0E4988928" box="[1186,1211,1572,1624]" gridcol="10" gridrow="8" pageId="15" pageNumber="444">11</td>
<td id="183A5DB6FFAFF38FD3FBF9E0E53F8928" box="[1230,1308,1572,1624]" gridcol="11" gridrow="8" pageId="15" pageNumber="444">126±21</td>
<td id="183A5DB6FFAFF38FD275F9E0E5AF8928" box="[1344,1420,1572,1624]" gridcol="12" gridrow="8" pageId="15" pageNumber="444">81165</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAF9A8E5AF89D0" box="[159,1420,1644,1696]" gridrow="9" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAF9A8E15A89D0" box="[159,377,1644,1696]" gridcol="0" gridrow="9" pageId="15" pageNumber="444">Body rosette (abdominal)</th>
<td id="183A5DB6FFAFF38FD6BCF9A8E18089D0" box="[393,419,1644,1696]" gridcol="1" gridrow="9" pageId="15" pageNumber="444">12</td>
<td id="183A5DB6FFAFF38FD68DF9A8E23589D0" box="[440,534,1644,1696]" gridcol="2" gridrow="9" pageId="15" pageNumber="444">45±5</td>
<td id="183A5DB6FFAFF38FD51FF9A8E2A189D0" box="[554,642,1644,1696]" gridcol="3" gridrow="9" pageId="15" pageNumber="444">3856</td>
<td id="183A5DB6FFAFF38FD5A6F9A8E28E89D0" box="[659,685,1644,1696]" gridcol="4" gridrow="9" pageId="15" pageNumber="444">8</td>
<td id="183A5DB6FFAFF38FD58BF9A8E32F89D0" box="[702,780,1644,1696]" gridcol="5" gridrow="9" pageId="15" pageNumber="444">43±4</td>
<td id="183A5DB6FFAFF38FD405F9A8E3AA89D0" box="[816,905,1644,1696]" gridcol="6" gridrow="9" pageId="15" pageNumber="444">3851</td>
<td id="183A5DB6FFAFF38FD4AFF9A8E39789D0" box="[922,948,1644,1696]" gridcol="7" gridrow="9" pageId="15" pageNumber="444">0</td>
<td id="183A5DB6FFAFF38FD4F3F9A8E40789D0" box="[966,1060,1644,1696]" gridcol="8" gridrow="9" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD30DF9A8E4B289D0" box="[1080,1169,1644,1696]" gridcol="9" gridrow="9" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD397F9A8E49889D0" box="[1186,1211,1644,1696]" gridcol="10" gridrow="9" pageId="15" pageNumber="444">0</td>
<td id="183A5DB6FFAFF38FD3FBF9A8E53F89D0" box="[1230,1308,1644,1696]" gridcol="11" gridrow="9" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD275F9A8E5AF89D0" box="[1344,1420,1644,1696]" gridcol="12" gridrow="9" pageId="15" pageNumber="444">-</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAF970E5AF8998" box="[159,1420,1716,1768]" gridrow="10" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAF970E15A8998" box="[159,377,1716,1768]" gridcol="0" gridrow="10" pageId="15" pageNumber="444">Body table (abdominal)</th>
<td id="183A5DB6FFAFF38FD6BCF970E1808998" box="[393,419,1716,1768]" gridcol="1" gridrow="10" pageId="15" pageNumber="444">3</td>
<td id="183A5DB6FFAFF38FD68DF970E2358998" box="[440,534,1716,1768]" gridcol="2" gridrow="10" pageId="15" pageNumber="444">55±8</td>
<td id="183A5DB6FFAFF38FD51FF970E2A18998" box="[554,642,1716,1768]" gridcol="3" gridrow="10" pageId="15" pageNumber="444">4661</td>
<td id="183A5DB6FFAFF38FD5A6F970E28E8998" box="[659,685,1716,1768]" gridcol="4" gridrow="10" pageId="15" pageNumber="444">0</td>
<td id="183A5DB6FFAFF38FD58BF970E32F8998" box="[702,780,1716,1768]" gridcol="5" gridrow="10" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD405F970E3AA8998" box="[816,905,1716,1768]" gridcol="6" gridrow="10" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD4AFF970E3978998" box="[922,948,1716,1768]" gridcol="7" gridrow="10" pageId="15" pageNumber="444">3</td>
<td id="183A5DB6FFAFF38FD4F3F970E4078998" box="[966,1060,1716,1768]" gridcol="8" gridrow="10" pageId="15" pageNumber="444">74±15</td>
<td id="183A5DB6FFAFF38FD30DF970E4B28998" box="[1080,1169,1716,1768]" gridcol="9" gridrow="10" pageId="15" pageNumber="444">5783</td>
<td id="183A5DB6FFAFF38FD397F970E4988998" box="[1186,1211,1716,1768]" gridcol="10" gridrow="10" pageId="15" pageNumber="444">11</td>
<td id="183A5DB6FFAFF38FD3FBF970E53F8998" box="[1230,1308,1716,1768]" gridcol="11" gridrow="10" pageId="15" pageNumber="444">68±6</td>
<td id="183A5DB6FFAFF38FD275F970E5AF8998" box="[1344,1420,1716,1768]" gridcol="12" gridrow="10" pageId="15" pageNumber="444">5678</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAF938E5AF8862" box="[159,1420,1788,1810]" gridrow="11" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAF938E15A8862" box="[159,377,1788,1810]" gridcol="0" gridrow="11" pageId="15" pageNumber="444">Body rosette (dorsal)</th>
<td id="183A5DB6FFAFF38FD6BCF938E1808862" box="[393,419,1788,1810]" gridcol="1" gridrow="11" pageId="15" pageNumber="444">12</td>
<td id="183A5DB6FFAFF38FD68DF938E2358862" box="[440,534,1788,1810]" gridcol="2" gridrow="11" pageId="15" pageNumber="444">46±6</td>
<td id="183A5DB6FFAFF38FD51FF938E2A18862" box="[554,642,1788,1810]" gridcol="3" gridrow="11" pageId="15" pageNumber="444">3857</td>
<td id="183A5DB6FFAFF38FD5A6F938E28E8862" box="[659,685,1788,1810]" gridcol="4" gridrow="11" pageId="15" pageNumber="444">9</td>
<td id="183A5DB6FFAFF38FD58BF938E32F8862" box="[702,780,1788,1810]" gridcol="5" gridrow="11" pageId="15" pageNumber="444">42±5</td>
<td id="183A5DB6FFAFF38FD405F938E3AA8862" box="[816,905,1788,1810]" gridcol="6" gridrow="11" pageId="15" pageNumber="444">3149</td>
<td id="183A5DB6FFAFF38FD4AFF938E3978862" box="[922,948,1788,1810]" gridcol="7" gridrow="11" pageId="15" pageNumber="444">0</td>
<td id="183A5DB6FFAFF38FD4F3F938E4078862" box="[966,1060,1788,1810]" gridcol="8" gridrow="11" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD30DF938E4B28862" box="[1080,1169,1788,1810]" gridcol="9" gridrow="11" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD397F938E4988862" box="[1186,1211,1788,1810]" gridcol="10" gridrow="11" pageId="15" pageNumber="444">0</td>
<td id="183A5DB6FFAFF38FD3FBF938E53F8862" box="[1230,1308,1788,1810]" gridcol="11" gridrow="11" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD275F938E5AF8862" box="[1344,1420,1788,1810]" gridcol="12" gridrow="11" pageId="15" pageNumber="444">-</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAF8E1E5AF884B" box="[159,1420,1829,1851]" gridrow="12" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAF8E1E15A884B" box="[159,377,1829,1851]" gridcol="0" gridrow="12" pageId="15" pageNumber="444">Body table (dorsal)</th>
<td id="183A5DB6FFAFF38FD6BCF8E1E180884B" box="[393,419,1829,1851]" gridcol="1" gridrow="12" pageId="15" pageNumber="444">0</td>
<td id="183A5DB6FFAFF38FD68DF8E1E235884B" box="[440,534,1829,1851]" gridcol="2" gridrow="12" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD51FF8E1E2A1884B" box="[554,642,1829,1851]" gridcol="3" gridrow="12" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD5A6F8E1E28E884B" box="[659,685,1829,1851]" gridcol="4" gridrow="12" pageId="15" pageNumber="444">0</td>
<td id="183A5DB6FFAFF38FD58BF8E1E32F884B" box="[702,780,1829,1851]" gridcol="5" gridrow="12" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD405F8E1E3AA884B" box="[816,905,1829,1851]" gridcol="6" gridrow="12" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD4AFF8E1E397884B" box="[922,948,1829,1851]" gridcol="7" gridrow="12" pageId="15" pageNumber="444">6</td>
<td id="183A5DB6FFAFF38FD4F3F8E1E407884B" box="[966,1060,1829,1851]" gridcol="8" gridrow="12" pageId="15" pageNumber="444">72±10</td>
<td id="183A5DB6FFAFF38FD30DF8E1E4B2884B" box="[1080,1169,1829,1851]" gridcol="9" gridrow="12" pageId="15" pageNumber="444">5882</td>
<td id="183A5DB6FFAFF38FD397F8E1E498884B" box="[1186,1211,1829,1851]" gridcol="10" gridrow="12" pageId="15" pageNumber="444">9</td>
<td id="183A5DB6FFAFF38FD3FBF8E1E53F884B" box="[1230,1308,1829,1851]" gridcol="11" gridrow="12" pageId="15" pageNumber="444">67±8</td>
<td id="183A5DB6FFAFF38FD275F8E1E5AF884B" box="[1344,1420,1829,1851]" gridcol="12" gridrow="12" pageId="15" pageNumber="444">5782</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAF88AE5AF8814" box="[159,1420,1870,1892]" gridrow="13" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAF88AE15A8814" box="[159,377,1870,1892]" gridcol="0" gridrow="13" pageId="15" pageNumber="444">Anal rod</th>
<td id="183A5DB6FFAFF38FD6BCF88AE1808814" box="[393,419,1870,1892]" gridcol="1" gridrow="13" pageId="15" pageNumber="444">26</td>
<td id="183A5DB6FFAFF38FD68DF88AE2358814" box="[440,534,1870,1892]" gridcol="2" gridrow="13" pageId="15" pageNumber="444">83±27</td>
<td id="183A5DB6FFAFF38FD51FF88AE2A18814" box="[554,642,1870,1892]" gridcol="3" gridrow="13" pageId="15" pageNumber="444">20115</td>
<td id="183A5DB6FFAFF38FD5A6F88AE28E8814" box="[659,685,1870,1892]" gridcol="4" gridrow="13" pageId="15" pageNumber="444">7</td>
<td id="183A5DB6FFAFF38FD58BF88AE32F8814" box="[702,780,1870,1892]" gridcol="5" gridrow="13" pageId="15" pageNumber="444">101±21</td>
<td id="183A5DB6FFAFF38FD405F88AE3AA8814" box="[816,905,1870,1892]" gridcol="6" gridrow="13" pageId="15" pageNumber="444">62124</td>
<td id="183A5DB6FFAFF38FD4AFF88AE3978814" box="[922,948,1870,1892]" gridcol="7" gridrow="13" pageId="15" pageNumber="444">2</td>
<td id="183A5DB6FFAFF38FD4F3F88AE4078814" box="[966,1060,1870,1892]" gridcol="8" gridrow="13" pageId="15" pageNumber="444">59</td>
<td id="183A5DB6FFAFF38FD30DF88AE4B28814" box="[1080,1169,1870,1892]" gridcol="9" gridrow="13" pageId="15" pageNumber="444">5266</td>
<td id="183A5DB6FFAFF38FD397F88AE4988814" box="[1186,1211,1870,1892]" gridcol="10" gridrow="13" pageId="15" pageNumber="444">21</td>
<td id="183A5DB6FFAFF38FD3FBF88AE53F8814" box="[1230,1308,1870,1892]" gridcol="11" gridrow="13" pageId="15" pageNumber="444">76±23</td>
<td id="183A5DB6FFAFF38FD275F88AE5AF8814" box="[1344,1420,1870,1892]" gridcol="12" gridrow="13" pageId="15" pageNumber="444">23113</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAF8BCE5AF88FE" box="[159,1420,1912,1934]" gridrow="14" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAF8BCE15A88FE" box="[159,377,1912,1934]" gridcol="0" gridrow="14" pageId="15" pageNumber="444">Anal rosette</th>
<td id="183A5DB6FFAFF38FD6BCF8BCE18088FE" box="[393,419,1912,1934]" gridcol="1" gridrow="14" pageId="15" pageNumber="444">22</td>
<td id="183A5DB6FFAFF38FD68DF8BCE23588FE" box="[440,534,1912,1934]" gridcol="2" gridrow="14" pageId="15" pageNumber="444">37±10</td>
<td id="183A5DB6FFAFF38FD51FF8BCE2A188FE" box="[554,642,1912,1934]" gridcol="3" gridrow="14" pageId="15" pageNumber="444">2167</td>
<td id="183A5DB6FFAFF38FD5A6F8BCE28E88FE" box="[659,685,1912,1934]" gridcol="4" gridrow="14" pageId="15" pageNumber="444">9</td>
<td id="183A5DB6FFAFF38FD58BF8BCE32F88FE" box="[702,780,1912,1934]" gridcol="5" gridrow="14" pageId="15" pageNumber="444">38±10</td>
<td id="183A5DB6FFAFF38FD405F8BCE3AA88FE" box="[816,905,1912,1934]" gridcol="6" gridrow="14" pageId="15" pageNumber="444">2754</td>
<td id="183A5DB6FFAFF38FD4AFF8BCE39788FE" box="[922,948,1912,1934]" gridcol="7" gridrow="14" pageId="15" pageNumber="444">2</td>
<td id="183A5DB6FFAFF38FD4F3F8BCE40788FE" box="[966,1060,1912,1934]" gridcol="8" gridrow="14" pageId="15" pageNumber="444">40</td>
<td id="183A5DB6FFAFF38FD30DF8BCE4B288FE" box="[1080,1169,1912,1934]" gridcol="9" gridrow="14" pageId="15" pageNumber="444">3148</td>
<td id="183A5DB6FFAFF38FD397F8BCE49888FE" box="[1186,1211,1912,1934]" gridcol="10" gridrow="14" pageId="15" pageNumber="444">1</td>
<td id="183A5DB6FFAFF38FD3FBF8BCE53F88FE" box="[1230,1308,1912,1934]" gridcol="11" gridrow="14" pageId="15" pageNumber="444">35</td>
<td id="183A5DB6FFAFF38FD275F8BCE5AF88FE" box="[1344,1420,1912,1934]" gridcol="12" gridrow="14" pageId="15" pageNumber="444">-</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAF865E5AF88C7" box="[159,1420,1953,1975]" gridrow="15" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAF865E15A88C7" box="[159,377,1953,1975]" gridcol="0" gridrow="15" pageId="15" pageNumber="444">Anal supporting table</th>
<td id="183A5DB6FFAFF38FD6BCF865E18088C7" box="[393,419,1953,1975]" gridcol="1" gridrow="15" pageId="15" pageNumber="444">8</td>
<td id="183A5DB6FFAFF38FD68DF865E23588C7" box="[440,534,1953,1975]" gridcol="2" gridrow="15" pageId="15" pageNumber="444">87±17</td>
<td id="183A5DB6FFAFF38FD51FF865E2A188C7" box="[554,642,1953,1975]" gridcol="3" gridrow="15" pageId="15" pageNumber="444">71112</td>
<td id="183A5DB6FFAFF38FD5A6F865E28E88C7" box="[659,685,1953,1975]" gridcol="4" gridrow="15" pageId="15" pageNumber="444">1</td>
<td id="183A5DB6FFAFF38FD58BF865E32F88C7" box="[702,780,1953,1975]" gridcol="5" gridrow="15" pageId="15" pageNumber="444">89</td>
<td id="183A5DB6FFAFF38FD405F865E3AA88C7" box="[816,905,1953,1975]" gridcol="6" gridrow="15" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD4AFF865E39788C7" box="[922,948,1953,1975]" gridcol="7" gridrow="15" pageId="15" pageNumber="444">8</td>
<td id="183A5DB6FFAFF38FD4F3F865E40788C7" box="[966,1060,1953,1975]" gridcol="8" gridrow="15" pageId="15" pageNumber="444">101±19</td>
<td id="183A5DB6FFAFF38FD30DF865E4B288C7" box="[1080,1169,1953,1975]" gridcol="9" gridrow="15" pageId="15" pageNumber="444">75138</td>
<td id="183A5DB6FFAFF38FD397F865E49888C7" box="[1186,1211,1953,1975]" gridcol="10" gridrow="15" pageId="15" pageNumber="444">0</td>
<td id="183A5DB6FFAFF38FD3FBF865E53F88C7" box="[1230,1308,1953,1975]" gridcol="11" gridrow="15" pageId="15" pageNumber="444">-</td>
<td id="183A5DB6FFAFF38FD275F865E5AF88C7" box="[1344,1420,1953,1975]" gridcol="12" gridrow="15" pageId="15" pageNumber="444">-</td>
</tr>
<tr id="5BEB34CAFFAFF38FD7AAF80EE5AF8890" box="[159,1420,1994,2016]" gridrow="16" pageId="15" pageNumber="444">
<th id="183A5DB6FFAFF38FD7AAF80EE15A8890" box="[159,377,1994,2016]" gridcol="0" gridrow="16" pageId="15" pageNumber="444">Anal table</th>
<td id="183A5DB6FFAFF38FD6BCF80EE1808890" box="[393,419,1994,2016]" gridcol="1" gridrow="16" pageId="15" pageNumber="444">4</td>
<td id="183A5DB6FFAFF38FD68DF80EE2358890" box="[440,534,1994,2016]" gridcol="2" gridrow="16" pageId="15" pageNumber="444">61±10</td>
<td id="183A5DB6FFAFF38FD51FF80EE2A18890" box="[554,642,1994,2016]" gridcol="3" gridrow="16" pageId="15" pageNumber="444">5173</td>
<td id="183A5DB6FFAFF38FD5A6F80EE28E8890" box="[659,685,1994,2016]" gridcol="4" gridrow="16" pageId="15" pageNumber="444">2</td>
<td id="183A5DB6FFAFF38FD58BF80EE32F8890" box="[702,780,1994,2016]" gridcol="5" gridrow="16" pageId="15" pageNumber="444">52</td>
<td id="183A5DB6FFAFF38FD405F80EE3AA8890" box="[816,905,1994,2016]" gridcol="6" gridrow="16" pageId="15" pageNumber="444">5053</td>
<td id="183A5DB6FFAFF38FD4AFF80EE3978890" box="[922,948,1994,2016]" gridcol="7" gridrow="16" pageId="15" pageNumber="444">5</td>
<td id="183A5DB6FFAFF38FD4F3F80EE4078890" box="[966,1060,1994,2016]" gridcol="8" gridrow="16" pageId="15" pageNumber="444">54±6</td>
<td id="183A5DB6FFAFF38FD30DF80EE4B28890" box="[1080,1169,1994,2016]" gridcol="9" gridrow="16" pageId="15" pageNumber="444">4763</td>
<td id="183A5DB6FFAFF38FD397F80EE4988890" box="[1186,1211,1994,2016]" gridcol="10" gridrow="16" pageId="15" pageNumber="444">5</td>
<td id="183A5DB6FFAFF38FD3FBF80EE53F8890" box="[1230,1308,1994,2016]" gridcol="11" gridrow="16" pageId="15" pageNumber="444">59±8</td>
<td id="183A5DB6FFAFF38FD275F80EE5AF8890" box="[1344,1420,1994,2016]" gridcol="12" gridrow="16" pageId="15" pageNumber="444">4870</td>
</tr>
</table>
</paragraph>
<paragraph id="E5643688FFB00C60D7F2FF53E2F08E6D" blockId="16.[151,1437,151,896]" pageId="16" pageNumber="445">
<emphasis id="D7AFEA9AFFB00C60D7F2FF53E1DB8FC0" bold="true" box="[199,504,151,176]" pageId="16" pageNumber="445">
Description of
<typeStatus id="3A60882AFFB00C60D64EFF53E1D08FC0" box="[379,499,151,176]" pageId="16" pageNumber="445" type="paratype">paratypes</typeStatus>
.
</emphasis>
Body color varies among specimens from purplish to light brown; some swollen specimens with thin transparent purplish skin (
<figureCitation id="7DE02A0DFFB00C60D597FF78E2DB8FA4" box="[674,760,188,213]" captionStart="FIGURE 1" captionStartId="4.[151,250,1855,1877]" captionTargetBox="[199,1397,199,1836]" captionTargetId="figure@4.[193,1400,195,1838]" captionTargetPageId="4" captionText="FIGURE 1. Lateral views of the specimens, dorsal upper, anterior left. A, Amphicyclus japonicus Bell, 1884, (a) WMNH-INV- 2015-13, (b) -2015-14, (c) -2015-311, (d) -2015-312; B, Neocucumis misakiensis sp. nov., (e) WMNH-INV-2016-208, (f) - 2016-209; C, Pseudocolochirus misakiensis sp. nov., (g) WMNH-INV-2015-348, (h) -2015-349, (i) -2015-350; D, Hemiocnus tegulatus (Augustin, 1908) comb. nov., (j) WMNH-INV-2015-378, (k) -2015-379; E, Lipotrapeza purpurata sp. nov., (l) WMNH-INV-2015-38, (m) -2015-39, (n) -2015-40; F, Pentamera misakiensis sp. nov., (o) WMNH-INV-2015-34, (p) -2015- 35, (q) -2015-36; G, Pseudoplacothuria ohshimai (Liao, 1997) gen. nov., comb. nov., (r) WMNH-INV-2015-9, (s) -2015-10." httpUri="https://zenodo.org/record/1457429/files/figure.png" pageId="16" pageNumber="445">Fig. 1E</figureCitation>
). Color of body cavity, oral opening, and inside of body and pedicel wall are uniform. In small specimens, large pedicels form two longitudinal rows along each radius, and small pedicels sparsley distributed in the interradii.
</paragraph>
<paragraph id="E5643688FFB00C60D7F2FEECE18E8E85" blockId="16.[151,1437,151,896]" pageId="16" pageNumber="445">
In a swollen, immature paratype (WMNH-INV-
<date id="91651048FFB00C60D5D3FEE3E3628E30" box="[742,833,295,320]" pageId="16" pageNumber="445" value="2015" valueMax="2040">2015-40</date>
), tentacle with swollen rods (
<figureCitation id="7DE02A0DFFB00C60D3BFFEE3E4FA8E30" box="[1162,1241,295,320]" captionStart="FIGURE 7" captionStartId="14.[151,250,1878,1900]" captionTargetBox="[187,1400,196,1864]" captionTargetId="figure@14.[184,1404,193,1867]" captionTargetPageId="14" captionText="FIGURE 7. SEM image of ossicles of Lipotrapeza purpurata sp. nov. (AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla." httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="16" pageNumber="445">
Fig.
<date id="91651048FFB00C60D38AFEECE4FA8E30" box="[1215,1241,296,320]" pageId="16" pageNumber="445">7I</date>
</figureCitation>
), and without the usual rosettes. Peri-oral skin and pharyngeal villi with the usual rosettes (
<figureCitation id="7DE02A0DFFB00C60D4E1FE88E46B8E14" box="[980,1096,332,357]" captionStart="FIGURE 7" captionStartId="14.[151,250,1878,1900]" captionTargetBox="[187,1400,196,1864]" captionTargetId="figure@14.[184,1404,193,1867]" captionTargetPageId="14" captionText="FIGURE 7. SEM image of ossicles of Lipotrapeza purpurata sp. nov. (AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla." httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="16" pageNumber="445">Fig. 7J, K</figureCitation>
), similar to those of holotype. Introvert with usual rosettes and additionally with table-like ossicles and large, rosette-derived plates (
<figureCitation id="7DE02A0DFFB00C60D200FEABE5AC8EF8" box="[1333,1423,367,392]" captionStart="FIGURE 7" captionStartId="14.[151,250,1878,1900]" captionTargetBox="[187,1400,196,1864]" captionTargetId="figure@14.[184,1404,193,1867]" captionTargetPageId="14" captionText="FIGURE 7. SEM image of ossicles of Lipotrapeza purpurata sp. nov. (AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla." httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="16" pageNumber="445">Fig. 7L</figureCitation>
). Pedicel with normal endplate and supporting plates (
<figureCitation id="7DE02A0DFFB00C60D5D0FE50E3678EDC" box="[741,836,404,429]" captionStart="FIGURE 7" captionStartId="14.[151,250,1878,1900]" captionTargetBox="[187,1400,196,1864]" captionTargetId="figure@14.[184,1404,193,1867]" captionTargetPageId="14" captionText="FIGURE 7. SEM image of ossicles of Lipotrapeza purpurata sp. nov. (AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla." httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="16" pageNumber="445">Fig. 7M</figureCitation>
), similar to those of holotype. Mid-body wall lacking rosettes but tables present (
<figureCitation id="7DE02A0DFFB00C60D6FDFE73E2678EA0" box="[456,580,439,464]" captionStart="FIGURE 7" captionStartId="14.[151,250,1878,1900]" captionTargetBox="[187,1400,196,1864]" captionTargetId="figure@14.[184,1404,193,1867]" captionTargetPageId="14" captionText="FIGURE 7. SEM image of ossicles of Lipotrapeza purpurata sp. nov. (AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla." httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="16" pageNumber="445">Fig. 7N, O</figureCitation>
). Ossicles of anal papilla exhibit rosettes, rods, tables, and supporting tables (
<figureCitation id="7DE02A0DFFB00C60D7AAFE18E0D78E84" box="[159,244,476,501]" captionStart="FIGURE 7" captionStartId="14.[151,250,1878,1900]" captionTargetBox="[187,1400,196,1864]" captionTargetId="figure@14.[184,1404,193,1867]" captionTargetPageId="14" captionText="FIGURE 7. SEM image of ossicles of Lipotrapeza purpurata sp. nov. (AH, WMNH-INV-2015-39; IP, WMNH-INV-2015- 40). A and I, ossicles of the ventrolateral tentacle; B and J, the ventrolateral skin of peri-oral; C and K, the ventrolateral skin of pharynx; D and L, the ventrolateral skin of introvert; E and M, the pedicel of the abdominal side; F and N, the middle part of the body wall on the abdominal side; G and O, the middle part of the body wall on the dorsal side; H and P, ventrolateral anal papilla." httpUri="https://zenodo.org/record/1457443/files/figure.png" pageId="16" pageNumber="445">Fig. 7P</figureCitation>
), as in holotype.
</paragraph>
</subSubSection>
<subSubSection id="ADC16503FFB00C60D7F2FE3BE2DA8C64" pageId="16" pageNumber="445" type="discussion">
<paragraph id="E5643688FFB00C60D7F2FE3BE2DA8C64" blockId="16.[151,1437,151,896]" pageId="16" pageNumber="445">
<emphasis id="D7AFEA9AFFB00C60D7F2FE3BE1188D68" bold="true" box="[199,315,511,536]" pageId="16" pageNumber="445">Remarks.</emphasis>
This species has 20 tentacles, a peculiar calcareous ring, and rosette ossicles in the introvert and body wall. This combination of features is typical of the genus
<taxonomicName id="22DB4D0BFFB00C60D4A5FDE1E4348D4C" authorityName="Yamana &amp; Kohtsuka" authorityYear="2018" box="[912,1047,549,572]" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="445" phylum="Echinodermata" rank="genus">
<emphasis id="D7AFEA9AFFB00C60D4A5FDE1E4348D4C" box="[912,1047,549,572]" italics="true" pageId="16" pageNumber="445">Lipotrapeza</emphasis>
</taxonomicName>
. Among the
<taxonomicName id="22DB4D0BFFB00C60D38DFDE1E51C8D4C" authorityName="Yamana &amp; Kohtsuka" authorityYear="2018" box="[1208,1343,549,572]" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="445" phylum="Echinodermata" rank="genus">
<emphasis id="D7AFEA9AFFB00C60D38DFDE1E51C8D4C" box="[1208,1343,549,572]" italics="true" pageId="16" pageNumber="445">Lipotrapeza</emphasis>
</taxonomicName>
, a long calcareous ring had been only observed in two nominal species,
<taxonomicName id="22DB4D0BFFB00C60D4BBFD8DE5608D10" authority="Heding &amp; Panning, 1954" authorityName="Heding &amp; Panning" authorityYear="1954" box="[910,1347,583,608]" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="445" phylum="Echinodermata" rank="species" species="japonica">
<emphasis id="D7AFEA9AFFB00C60D4BBFD8DE4318D2F" box="[910,1042,584,608]" italics="true" pageId="16" pageNumber="445">L. japonica</emphasis>
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</taxonomicName>
, and
<taxonomicName id="22DB4D0BFFB00C60D2B0FD8DE21B8DF4" authority="Yamana &amp; Kohtsuka, 2017" authorityName="Yamana &amp; Kohtsuka" authorityYear="2017" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="445" phylum="Echinodermata" rank="species" species="littoralis">
<emphasis id="D7AFEA9AFFB00C60D2B0FD8DE0DB8DF4" italics="true" pageId="16" pageNumber="445">L. littoralis</emphasis>
<bibRefCitation id="814A4B79FFB00C60D635FDA9E21B8DF4" box="[256,568,620,645]" pageId="16" pageNumber="445" refId="ref12551" refString="Yamana, Y. &amp; Kohtsuka, H. (2017) A new species of Lipotrapeza (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae) from Japan, with a partial Redescription of the syntypes of L. japonica. Plankton and Benthos Research, 12, 151 - 163. https: // doi. org / 10.3800 / pbr. 12.151" type="journal article">Yamana &amp; Kohtsuka, 2017</bibRefCitation>
</taxonomicName>
. However, the number of anal papillae of the present species is significantly different from
<taxonomicName id="22DB4D0BFFB00C60D609FD55E19E8DD7" authorityName="Heding &amp; Panning" authorityYear="1954" box="[316,445,656,680]" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="445" phylum="Echinodermata" rank="species" species="japonica">
<emphasis id="D7AFEA9AFFB00C60D609FD55E19E8DD7" box="[316,445,656,680]" italics="true" pageId="16" pageNumber="445">L. japonica</emphasis>
</taxonomicName>
, in which only five papillae are present (
<bibRefCitation id="814A4B79FFB00C60D4B9FD4BE4878DD8" box="[908,1188,655,680]" pageId="16" pageNumber="445" refId="ref10776" refString="Heding, S. G. &amp; Panning, A. (1954) Phyllophoridae. Spolia Zoologica Musei Hauniensis, 13, 1 - 209." type="journal article">Heding &amp; Panning 1954</bibRefCitation>
;
<bibRefCitation id="814A4B79FFB00C60D384FD54E0F78DBC" pageId="16" pageNumber="445" refId="ref12551" refString="Yamana, Y. &amp; Kohtsuka, H. (2017) A new species of Lipotrapeza (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae) from Japan, with a partial Redescription of the syntypes of L. japonica. Plankton and Benthos Research, 12, 151 - 163. https: // doi. org / 10.3800 / pbr. 12.151" type="journal article">Yamana &amp; Kohtsuka 2017</bibRefCitation>
). The present species is also distinguishable from both
<taxonomicName id="22DB4D0BFFB00C60D40CFD71E39B8DBC" authorityName="Heding &amp; Panning" authorityYear="1954" box="[825,952,693,716]" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="445" phylum="Echinodermata" rank="species" species="japonica">
<emphasis id="D7AFEA9AFFB00C60D40CFD71E39B8DBC" box="[825,952,693,716]" italics="true" pageId="16" pageNumber="445">L. japonica</emphasis>
</taxonomicName>
and
<taxonomicName id="22DB4D0BFFB00C60D4DBFD71E4488DBC" authorityName="Yamana &amp; Kohtsuka" authorityYear="2017" box="[1006,1131,692,716]" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="445" phylum="Echinodermata" rank="species" species="littoralis">
<emphasis id="D7AFEA9AFFB00C60D4DBFD71E4488DBC" box="[1006,1131,692,716]" italics="true" pageId="16" pageNumber="445">L. littoralis</emphasis>
</taxonomicName>
, by the existence of rosette ossicles and/or tables in the body wall, while no ossicles are present in the body wall of
<taxonomicName id="22DB4D0BFFB00C60D35FFD1DE4CA8D9F" authorityName="Heding &amp; Panning" authorityYear="1954" box="[1130,1257,728,752]" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="445" phylum="Echinodermata" rank="species" species="japonica">
<emphasis id="D7AFEA9AFFB00C60D35FFD1DE4CA8D9F" box="[1130,1257,728,752]" italics="true" pageId="16" pageNumber="445">L. japonica</emphasis>
</taxonomicName>
and
<taxonomicName id="22DB4D0BFFB00C60D22BFD1DE5BF8D80" authorityName="Yamana &amp; Kohtsuka" authorityYear="2017" box="[1310,1436,728,752]" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="445" phylum="Echinodermata" rank="species" species="littoralis">
<emphasis id="D7AFEA9AFFB00C60D22BFD1DE5BF8D80" box="[1310,1436,728,752]" italics="true" pageId="16" pageNumber="445">L. littoralis</emphasis>
</taxonomicName>
(
<bibRefCitation id="814A4B79FFB00C60D7ABFD38E1968C65" box="[158,437,764,789]" pageId="16" pageNumber="445" refId="ref10776" refString="Heding, S. G. &amp; Panning, A. (1954) Phyllophoridae. Spolia Zoologica Musei Hauniensis, 13, 1 - 209." type="journal article">Heding &amp; Panning 1954</bibRefCitation>
;
<bibRefCitation id="814A4B79FFB00C60D6F5FD39E2CE8C64" box="[448,749,764,789]" pageId="16" pageNumber="445" refId="ref12551" refString="Yamana, Y. &amp; Kohtsuka, H. (2017) A new species of Lipotrapeza (Echinodermata: Holothuroidea: Dendrochirotida: Phyllophoridae) from Japan, with a partial Redescription of the syntypes of L. japonica. Plankton and Benthos Research, 12, 151 - 163. https: // doi. org / 10.3800 / pbr. 12.151" type="journal article">Yamana &amp; Kohtsuka 2017</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="ADC16503FFB00C60D7F2FCDBE3448C2C" pageId="16" pageNumber="445" type="distribution">
<paragraph id="E5643688FFB00C60D7F2FCDBE3448C2C" blockId="16.[151,1437,151,896]" pageId="16" pageNumber="445">
<emphasis id="D7AFEA9AFFB00C60D7F2FCDBE1438C48" bold="true" box="[199,352,799,824]" pageId="16" pageNumber="445">Distribution.</emphasis>
So far known only from the
<typeStatus id="3A60882AFFB00C60D5F1FCE4E2D78C48" box="[708,756,800,824]" pageId="16" pageNumber="445">type</typeStatus>
locality, off Jyogashima Island, south coast of
<collectingRegion id="271FF86AFFB00C60D213FCE4E12E8C2D" country="Japan" name="Kanagawa" pageId="16" pageNumber="445">Kanagawa Prefecture</collectingRegion>
, sand bottom,
<quantity id="22239B6DFFB00C60D69AFC80E2368C2C" box="[431,533,836,860]" metricMagnitude="1" metricUnit="m" metricValue="8.75" metricValueMax="8.8" metricValueMin="8.7" pageId="16" pageNumber="445" unit="m" value="87.5" valueMax="88.0" valueMin="87.0">8788 m</quantity>
, Pacific Ocean, middle
<collectingCountry id="9DCC7618FFB00C60D415FC81E3408C2C" box="[800,867,837,860]" name="Japan" pageId="16" pageNumber="445">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="ADC16503FFB00C60D7F2FCA3E3FB8CF0" box="[199,984,871,896]" pageId="16" pageNumber="445" type="etymology">
<paragraph id="E5643688FFB00C60D7F2FCA3E3FB8CF0" blockId="16.[151,1437,151,896]" box="[199,984,871,896]" pageId="16" pageNumber="445">
<emphasis id="D7AFEA9AFFB00C60D7F2FCA3E16D8CF0" bold="true" box="[199,334,871,896]" pageId="16" pageNumber="445">Etymology.</emphasis>
The specific name
<taxonomicName id="22DB4D0BFFB00C60D51CFCADE2B88CF0" authorityName="Yamana &amp; Kohtsuka" authorityYear="2018" box="[553,667,873,896]" class="Holothuroidea" family="Phyllophoridae" genus="Lipotrapeza" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dendrochirotida" pageId="16" pageNumber="445" phylum="Echinodermata" rank="species" species="purpurata">
<emphasis id="D7AFEA9AFFB00C60D51CFCADE2B88CF0" box="[553,667,873,896]" italics="true" pageId="16" pageNumber="445">purpurata</emphasis>
</taxonomicName>
alludes to its purplish color.
</paragraph>
</subSubSection>
</treatment>
</document>