treatments-xml/data/6B/86/51/6B865165A8335B55AE6657BAE6F0C62F.xml

<document ID-DOI="http://dx.doi.org/10.3897/BDJ.11.e107751" ID-Pensoft-Pub="1314-2828-11-e107751" ID-Pensoft-UUID="473F14BBE43C56218CA054FFDF2B55A1" ID-ZooBank="29C26A7941324AE0B685C1D30B9266B2" ModsDocID="1314-2828-11-e107751" checkinTime="1692714683841" checkinUser="pensoft" docAuthor="Yang, Weicheng, He, Lianfeng, Yu, Hao &amp; Lin, Yucheng" docDate="2023" docId="6B865165A8335B55AE6657BAE6F0C62F" docLanguage="en" docName="BiodivDatJour 11: e107751" docOrigin="Biodiversity Data Journal 11" docPubDate="2023-08-21" docSource="http://dx.doi.org/10.3897/BDJ.11.e107751" docTitle="Speleoticus hei Yu &amp; Lin 2023, sp. nov." docType="treatment" docVersion="1" id="473F14BBE43C56218CA054FFDF2B55A1" lastPageNumber="107751" masterDocId="473F14BBE43C56218CA054FFDF2B55A1" masterDocTitle="A new cave-dwelling spider of the genus Speleoticus (Araneae, Nesticidae) from Sichuan, China" masterLastPageNumber="107751" masterPageNumber="107751" pageNumber="107751" updateTime="1692714683841" updateUser="pensoft">
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<mods:title id="039537EA35D9117F02696744BF1D8F9E">A new cave-dwelling spider of the genus Speleoticus (Araneae, Nesticidae) from Sichuan, China</mods:title>
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<mods:namePart id="3C826E2F5DB99DD9C278FB626A3F1199">Yang, Weicheng</mods:namePart>
<mods:affiliation id="4CBDB10B0BE11B4C2838A15E2498810C">School of Life Sciences, Guizhou Normal University, Guiyang, China</mods:affiliation>
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<mods:namePart id="E83F950748F0F1EE0C845DF37974F26F">He, Lianfeng</mods:namePart>
<mods:affiliation id="5E7BFB6AEA07A1A6CDD439D0F4203C30">School of Life Sciences, Guizhou Normal University, Guiyang, China</mods:affiliation>
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<mods:namePart id="87AD1C703210F8C9ACF79A7E3B62D954">Yu, Hao</mods:namePart>
<mods:affiliation id="82BD9E63BBE7D0A862D2DD116D5471B1">School of Life Sciences, Guizhou Normal University, Guiyang, China</mods:affiliation>
<mods:nameIdentifier id="075BE403907618A7C5411AFFB6BB7C55" type="email">insect1986@126.com</mods:nameIdentifier>
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<mods:namePart id="38E66A1400C7BE957C228A8121132F52">Lin, Yucheng</mods:namePart>
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<mods:affiliation id="14932C808A6B4FEFD0E3ED3392781A9F">Key Laboratory Bio-resources and Eco-environment (Ministry of Education), College of Life Sciences, Sichuan University, Chengdu, China</mods:affiliation>
<mods:nameIdentifier id="3CB0A738AF4CB3F5D69382C754E41CEC" type="email">linyucheng@scu.edu.cn</mods:nameIdentifier>
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<treatment id="6B865165A8335B55AE6657BAE6F0C62F" LSID="urn:lsid:plazi:treatment:6B865165A8335B55AE6657BAE6F0C62F" httpUri="http://treatment.plazi.org/id/6B865165A8335B55AE6657BAE6F0C62F" lastPageNumber="107751" pageId="0" pageNumber="107751">
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<taxonomicName id="1AC30CDA1E39EB21746B89539C99AEF4" LSID="6B865165-A833-5B55-AE66-57BAE6F0C62F" authority="Yu &amp; Lin" authorityName="Yu &amp; Lin" authorityYear="2023" class="Arachnida" family="Nesticidae" genus="Speleoticus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Speleoticus hei" order="Araneae" pageId="0" pageNumber="107751" phylum="Arthropoda" rank="species" species="hei" status="sp. nov.">Speleoticus hei Yu &amp; Lin</taxonomicName>
<taxonomicNameLabel id="02362101BA8B34AC95F7F918A79B24A2" pageId="0" pageNumber="107751">sp. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection id="204A8E13A54B4FD8A6AA14960C8F86FC" pageId="0" pageNumber="107751" type="materials_examined">
<paragraph id="6F2E8FD9C50361D44E3FD36F7695AA51" pageId="0" pageNumber="107751">Materials</paragraph>
<paragraph id="A32F8F0499674DA1D5FD0C9B31A63B90" pageId="0" pageNumber="107751">
<materialsCitation id="4BD607120586A55A886F60D05D7592E8" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectingMethod="by hand" collectorName="Li He, Yu, Lin" country="China" determinerName="Hao Yu and Yucheng Lin" latitude="32.577297" location="cave Hanwang Dong" longitude="106.10698" pageId="0" pageNumber="107751" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="Holotype">
<emphasis id="F993472AF627E80F874BC716D1DD8575" bold="true" pageId="0" pageNumber="107751">Type status:</emphasis>
<materialsCitation id="897F1B9D3F53BD38FA8FCAB2822F1699" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectingMethod="by hand" collectorName="Li He, Yu, Lin" country="China" determinerName="Hao Yu and Yucheng Lin" latitude="32.577297" location="cave Hanwang Dong" longitude="106.10698" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="Holotype">
<typeStatus id="E77B28BF661EF9C13624CAAC1DA8333F" pageId="0" pageNumber="107751">Holotype</typeStatus>
. 
<emphasis id="EE27375CEF6C3EA0EF30F35B8BC28125" bold="true" pageId="0" pageNumber="107751">Occurrence:</emphasis>
recordedBy: 
<collectorName id="471EFA75447119C85DA01A94CB2379AA" pageId="0" pageNumber="107751">Li He</collectorName>
; individualCount: 
<specimenCount id="726AAF1360A28F0A074CA0E44E1582D4" pageId="0" pageNumber="107751" type="generic">1</specimenCount>
; sex: 
<specimenType id="623AC4B4FC626E9145B290B4B58A3270" pageId="0" pageNumber="107751">male</specimenType>
; lifeStage: 
<specimenType id="EAB7DDC4D408C0612C97BC4EA5B58D86" pageId="0" pageNumber="107751">adult</specimenType>
; behavior: weaver; preparations: whole animal (ETOH); occurrenceID: 
<uuid id="5D367F2E736FF9027708E9F62D9503B8">4F2FECF6-4539-5B2E-88BA-A9B6C98A6BDC</uuid>
; 
<emphasis id="5667E96E9E341D05A2B8CE9E311EAAC2" bold="true" pageId="0" pageNumber="107751">Taxon:</emphasis>
order: Araneae; family: Nesticidae; genus: Speleoticus; specificEpithet: hei; scientificNameAuthorship: 
<collectorName id="7737575BF54B3A6CDB716169F3E9B988">Yu</collectorName>
&amp; 
<collectorName id="10B01709385D98624B4F0396BC9F3CE6">Lin</collectorName>
; 
<emphasis id="E9C329CB21EF3FF13F5F17359FD0031F" bold="true" pageId="0" pageNumber="107751">Location:</emphasis>
continent: Asia; country: 
<collectingCountry id="12E89E97B93A676255D39372B8723721" name="China" pageId="0" pageNumber="107751">China</collectingCountry>
; countryCode: CHN; stateProvince: 
<collectingRegion id="E6221B0E30F476E9046D799E8D6542A3" country="China" name="Sichuan">Sichuan</collectingRegion>
; county: Guangyuan; locality: 
<location id="FC7959EDB10F04C7F5E657E4D958B262" LSID="urn:lsid:plazi:treatment:6B865165A8335B55AE6657BAE6F0C62F:FC7959EDB10F04C7F5E657E4D958B262" country="China" latitude="32.577297" longitude="106.10698" name="cave Hanwang Dong" pageId="0" pageNumber="107751" stateProvince="Sichuan">cave Hanwang Dong</location>
; decimalLatitude: 
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; decimalLongitude: 
<geoCoordinate id="67395E68A0CFCEF3529F6DD294BDA7E8" orientation="longitude" pageId="0" pageNumber="107751" value="106.106979">106.106979</geoCoordinate>
; 
<emphasis id="4ECDFA2EEAF4EA51647FCC24D3CCC93E" bold="true" pageId="0" pageNumber="107751">Identification:</emphasis>
identifiedBy: 
<determinerName id="F580C69C669CF77F6F1A67F6EEE56708" pageId="0" pageNumber="107751">Hao Yu and Yucheng Lin</determinerName>
; dateIdentified: 2022-03; identificationReferences: Lin et al. 2016; 
<emphasis id="2F657CFACEB5270B2C8D65CB5A8DEA8D" bold="true" pageId="0" pageNumber="107751">Event:</emphasis>
samplingProtocol: 
<collectingMethod id="3CEF304BA74066120C89C9F48C716FB5" pageId="0" pageNumber="107751">by hand</collectingMethod>
; samplingEffort: 
<quantity id="C9F584F0D1D4BD030EF2FE51EF3399A5" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
by foot; year: 2018; month: 4; day: 6; 
<emphasis id="C9134AA7FAB1E88A30FD937A93466C08" bold="true" pageId="0" pageNumber="107751">Record Level:</emphasis>
basisOfRecord: PreservedSpecimen 
<materialsCitation id="F4B73D62553F81519AF5BD1658403650" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectingMethod="by hand" collectorName="Li He, Yu, Lin" country="China" determinerName="Hao Yu and Yucheng Lin" latitude="32.577297" location="cave Hanwang Dong" longitude="106.10698" pageId="0" pageNumber="107751" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Sichuan" typeStatus="Paratype">
<emphasis id="5AC028C65D703C6B8729BA906A629DAF" bold="true" pageId="0" pageNumber="107751">Type status:</emphasis>
<materialsCitation id="8523AB3BEEC4937BC9BEB7777FA51C9D" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectingMethod="by hand" collectorName="Li He, Yu, Lin" country="China" determinerName="Hao Yu and Yucheng Lin" latitude="32.577297" location="cave Hanwang Dong" longitude="106.10698" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Sichuan" typeStatus="Paratype">
<typeStatus id="4270DD625D332EEBA806BA202C3AF26B" pageId="0" pageNumber="107751">Paratype</typeStatus>
. 
<emphasis id="87F51E124B55974F21268AAAFCE23DC3" bold="true" pageId="0" pageNumber="107751">Occurrence:</emphasis>
recordedBy: 
<collectorName id="1D45F75569333861E68388F17D55E986" pageId="0" pageNumber="107751">Li He</collectorName>
; individualCount: 
<specimenCount id="67A6559354E24FE067EC5F156367C357" pageId="0" pageNumber="107751" type="generic">5</specimenCount>
; sex: 
<specimenCount id="42ABDDE94E9C2186625CDA91EBE16FE0" type="male">2 males</specimenCount>
, 
<specimenCount id="ACAFC82E3248723518A954802B258BBD" type="female">3 females</specimenCount>
; lifeStage: 
<specimenType id="12D74DD5797F1600EFA3198F9E0BB45D" pageId="0" pageNumber="107751">adult</specimenType>
; behavior: weaver; preparations: whole animal (ETOH); occurrenceID: 
<uuid id="04925C115AD0DBDE9EA1F6A97E6660B5">F3ACEB49-BC9E-5D14-8965-7D441B18B1AA</uuid>
; 
<emphasis id="A6EB52CCA690FD829D9DC5A399617055" bold="true" pageId="0" pageNumber="107751">Taxon:</emphasis>
order: Araneae; family: Nesticidae; genus: Speleoticus; specificEpithet: hei; scientificNameAuthorship: 
<collectorName id="94BF7C11F4B8F4209200744CE8C4ADF8">Yu</collectorName>
&amp; 
<collectorName id="580EC936688180F68A6410EB30B72234">Lin</collectorName>
; 
<emphasis id="74308131DFCEDB301712735E59F1CC19" bold="true" pageId="0" pageNumber="107751">Location:</emphasis>
continent: Asia; country: 
<collectingCountry id="66643DE6743C2FFF6569013943CCDC7E" name="China" pageId="0" pageNumber="107751">China</collectingCountry>
; countryCode: CHN; stateProvince: 
<collectingRegion id="B5CEAA5C3A4DBD80470A58988023964A" country="China" name="Sichuan">Sichuan</collectingRegion>
; county: Guangyuan; locality: 
<location id="596CF42B7C75E129A33401BF89A6F719" LSID="urn:lsid:plazi:treatment:6B865165A8335B55AE6657BAE6F0C62F:596CF42B7C75E129A33401BF89A6F719" country="China" latitude="32.577297" longitude="106.10698" name="cave Hanwang Dong" pageId="0" pageNumber="107751" stateProvince="Sichuan">cave Hanwang Dong</location>
; decimalLatitude: 
<geoCoordinate id="C76E3E1AB6790677C8685215A94705AE" orientation="latitude" pageId="0" pageNumber="107751" value="32.577297">32.577297</geoCoordinate>
; decimalLongitude: 
<geoCoordinate id="07B556E5507AE3ADA7EE6A9C09386992" orientation="longitude" pageId="0" pageNumber="107751" value="106.106979">106.106979</geoCoordinate>
; 
<emphasis id="3EF647E559282798673AB04D23326837" bold="true" pageId="0" pageNumber="107751">Identification:</emphasis>
identifiedBy: 
<determinerName id="D3D9A69F281892884EEF86A9C722553B" pageId="0" pageNumber="107751">Hao Yu and Yucheng Lin</determinerName>
; dateIdentified: 2022-03; identificationReferences: Lin et al. 2016; 
<emphasis id="F4F18743F552D544F1D74355EAE211B3" bold="true" pageId="0" pageNumber="107751">Event:</emphasis>
samplingProtocol: 
<collectingMethod id="7F6F7C5B0320D0C44DAC6DC22AE90395" pageId="0" pageNumber="107751">by hand</collectingMethod>
; samplingEffort: 
<quantity id="196BB881605E9D8BFA682598961A0F91" metricMagnitude="4" metricUnit="m" metricValue="1.0" unit="km" value="10.0">10 km</quantity>
by foot; year: 2018; month: 4; day: 6; 
<emphasis id="605219AF31A45E37B777E66F98875B69" bold="true" pageId="0" pageNumber="107751">Record Level:</emphasis>
basisOfRecord: PreservedSpecimen
</materialsCitation>
</materialsCitation>
</materialsCitation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5BD575828FD86E191B85AAADB28D59AF" pageId="0" pageNumber="107751" type="description">
<paragraph id="015A90BC81F39C755730EAF26D5948A5" pageId="0" pageNumber="107751">Description</paragraph>
<paragraph id="741B63A861614F86390E1322FFB50A53" pageId="0" pageNumber="107751">
<emphasis id="9B558A440EC31559022E355EC2066E08" bold="true" pageId="0" pageNumber="107751">Male holotype.</emphasis>
Habitus as in Fig. 
<figureCitation id="3ECB3065CAA23C23197E0135CDA4FBB3" captionStart="Figure 3" captionStartId="F9886411" captionText="Figure 3. Speleoticus hei Yu &amp; Lin, sp. n., female paratype and male holotype. A, B Macerated epigyne, ventral and dorsal view; C, D Epigyne, macerated and embedded in Arabic gum, ventral and dorsal view; E, F Male, habitus, ventral and dorsal view; G, H Female, habitus, ventral and dorsal view. Abbreviations: Co = copulatory opening; Ms = median septum; S = spermatheca; Vp = vulval pocket. Scale bars: 0.2 mm (equal for A-D); 2 mm (equal for E-F, equal for G-H)." figureDoi="10.3897/BDJ.11.e107751.figure3" httpUri="https://binary.pensoft.net/fig/885493" pageId="0" pageNumber="107751">3</figureCitation>
E and F. Total length 3.15. Carapace length 1.45, width 1.39, pale yellow to beige, with faint dark areas around the cervical furrow and fovea. Cervical groove and fovea distinct. Eyes (Fig. 
<figureCitation id="8EB6EB22CDD49068008E1251326C86C7" captionStart="Figure 3" captionStartId="F9886411" captionText="Figure 3. Speleoticus hei Yu &amp; Lin, sp. n., female paratype and male holotype. A, B Macerated epigyne, ventral and dorsal view; C, D Epigyne, macerated and embedded in Arabic gum, ventral and dorsal view; E, F Male, habitus, ventral and dorsal view; G, H Female, habitus, ventral and dorsal view. Abbreviations: Co = copulatory opening; Ms = median septum; S = spermatheca; Vp = vulval pocket. Scale bars: 0.2 mm (equal for A-D); 2 mm (equal for E-F, equal for G-H)." figureDoi="10.3897/BDJ.11.e107751.figure3" httpUri="https://binary.pensoft.net/fig/885493" pageId="0" pageNumber="107751">3</figureCitation>
E): in dorsal view, AER slightly recurved, PER almost straight and slightly wider than AER. Eye sizes and interdistances: AME 0.09, ALE 0.09, PME 0.09, PLE 0.10; AME-AME 0.02, AME-ALE 0.05, PME-PME 0.11, PME-PLE 0.06; MOA 0.20 long, anterior width 0.19, posterior width 0.29. Mouthparts: chelicerae light orange, with three teeth on promargin; endite length 0.55, width 0.29, depressed posteriorly, slightly convergent anteriorly, with dense setae on inner margin; labium nearly rectangular, length 0.18, width 0.14. Sternum yellowish with sparse setae, length 0.75, width 0.85. Leg uniformly yellowish. Leg measurements: I 9.44 (0.43, 0.24, 2.37, 2.93, 2.33, 1.14); II 7.82 (0.48, 0.27, 2.08, 2.30, 1.80, 0.89); III 5.92 (0.43, 0.24, 1.70, 1.62, 1.26, 0.67); IV 7.96 (0.47, 0.26, 2.28, 2.38, 1.74, 0.83). Opisthosoma ovoid, 1.70 long, 1.07 wide, grey, dorsum with setae, with two indistinctly coloured chevron stripes posteriorly.
</paragraph>
<paragraph id="989BA23C86B5C32D7F37EB0688E9A583" pageId="0" pageNumber="107751">
<emphasis id="3494898B14B239D069E460405F8D1CE3" bold="true" italics="true" pageId="0" pageNumber="107751">Palp</emphasis>
(Fig. 
<figureCitation id="5C9753F32E46A599425EEB0F60CCE94E" captionStart="Figure 4" captionStartId="F9886409" captionText="Figure 4. Male left palp of the holotype of Speleoticus hei Yu &amp; Lin, sp. n. A Ventral view; B Dorsal view; C Prolateral view; D Retrolateral view. Abbreviations: Cp (I-III) = processes of the conductor (I-III); Dp (I-II) = distal processes of the paracymbium (I-II); E = embolus; Eb = embolic base; Ma = median apophysis; P = paracymbium; St = subtegulum; T = tegulum; Ta = terminal apophysis; Va = ventral apophysis of the paracymbium. Scale bar: 0.2 mm (equal for A-D)." figureDoi="10.3897/BDJ.11.e107751.figure4" httpUri="https://binary.pensoft.net/fig/885573" pageId="0" pageNumber="107751">4</figureCitation>
A-D): Paracymbium long, with three branches or apophyses; ventral apophysis very small, shaped like a quadrahedron; Dp-I about 1/2 of Dp-II length, harpoon-shaped and with a subapical barb; Dp-II broad and long, well-developed, generally shaped like a sickle. Conductor complex, distally with three apophyses: Cp-I long and heavily-sclerotised, blade-shaped; Cp-II small, papilliform; Cp-III laminar, ending with a blunt tip. Terminal apophysis nearly triangular in prolateral view, protruding ventrally. Median apophysis heavily sclerotised, strongly expanded, directed ventrally, concave with two branches; proximal branch small, papilliform; distal branch relatively long, finger-like. Embolus slender and filiform, arising on the retrolateral flank (approximately 4 
<normalizedToken id="437A9B38DB2FD01E967E2DC149CD3525" originalValue="o’">o'</normalizedToken>
clock position), surrounding the base, terminating at approximately 12 
<normalizedToken id="D54E55292BE9500D260AAC270FB9947F" originalValue="o’">o'</normalizedToken>
clock position, its tip filiform and hidden behind the conductor.
</paragraph>
<paragraph id="91994CE305DEC2AF89646149F2FB4091" pageId="0" pageNumber="107751">
<emphasis id="4E39418ABC6F48916D92167B3D71D10D" bold="true" pageId="0" pageNumber="107751">Female.</emphasis>
Colour of the living female light brown, abdomen dorsally with two ˄-shaped stripes and 3-4 pairs black spots (Fig. 
<figureCitation id="89A6E2A265379D55DF98684E53A10209" captionStart="Figure 2" captionStartId="F9886407" captionText="Figure 2. Speleoticus hei Yu &amp; Lin, sp. n., female paratype, live specimens. Photographs by Li He (Chengdu, Sichuan)." figureDoi="10.3897/BDJ.11.e107751.figure2" httpUri="https://binary.pensoft.net/fig/861451" pageId="0" pageNumber="107751">2</figureCitation>
). Habitus in alcohol as in Fig. 
<figureCitation id="EE7A994BEE9F3BE827E734540768A448" captionStart="Figure 3" captionStartId="F9886411" captionText="Figure 3. Speleoticus hei Yu &amp; Lin, sp. n., female paratype and male holotype. A, B Macerated epigyne, ventral and dorsal view; C, D Epigyne, macerated and embedded in Arabic gum, ventral and dorsal view; E, F Male, habitus, ventral and dorsal view; G, H Female, habitus, ventral and dorsal view. Abbreviations: Co = copulatory opening; Ms = median septum; S = spermatheca; Vp = vulval pocket. Scale bars: 0.2 mm (equal for A-D); 2 mm (equal for E-F, equal for G-H)." figureDoi="10.3897/BDJ.11.e107751.figure3" httpUri="https://binary.pensoft.net/fig/885493" pageId="0" pageNumber="107751">3</figureCitation>
G and H. As in male, except as noted. Total length 3.29. length 1.31, width 1.21, pyriform. Eye diameters: AME 0.09, ALE 0.09, PME 0.09, PLE 0.07; interdistances: AME-AME 0.02, AME-ALE 0.25, PME-PME 0.12, PME-PLE 0.07; MOA 0.22 long, anterior width 0.19, posterior width 0.29. Chelicerae with three promarginal teeth. Opisthosoma: length 1.99, width 1.40, dorsum anteriorly with two ˄-shaped stripes, posteriorly with paired marks, partially fused each other in the posterior side. Leg measurements: I 8.34 (0.45, 0.25, 2.45, 2.39, 1.88, 0.93); II 6.60 (0.41, 0.23, 1.79, 1.99, 1.41, 0.77); III 5.62 (0.39, 0.22, 1.56, 1.59, 1.46, 0.41); IV 7.28 (0.45, 0.25, 2.16, 2.19, 1.53, 0.70).
</paragraph>
<paragraph id="4629D8B94F5D233F8DB373F3B11C73C4" pageId="0" pageNumber="107751">
<emphasis id="EAEB5CA1D678607DF7AB2391704F044B" bold="true" italics="true" pageId="0" pageNumber="107751">Epigyne</emphasis>
(Fig. 
<figureCitation id="0B26E29B637136E746C81530F1C11586" captionStart="Figure 3" captionStartId="F9886411" captionText="Figure 3. Speleoticus hei Yu &amp; Lin, sp. n., female paratype and male holotype. A, B Macerated epigyne, ventral and dorsal view; C, D Epigyne, macerated and embedded in Arabic gum, ventral and dorsal view; E, F Male, habitus, ventral and dorsal view; G, H Female, habitus, ventral and dorsal view. Abbreviations: Co = copulatory opening; Ms = median septum; S = spermatheca; Vp = vulval pocket. Scale bars: 0.2 mm (equal for A-D); 2 mm (equal for E-F, equal for G-H)." figureDoi="10.3897/BDJ.11.e107751.figure3" httpUri="https://binary.pensoft.net/fig/885493" pageId="0" pageNumber="107751">3</figureCitation>
A-D): Epigynal plate simple, disc-shaped, scape indistinct; lateral and posterior margins indistinct, spermathecae and bursae distinctly visible through integument after dissection. Median septum short, slightly protruding, shaped as a tongue. Copulatory openings indistinct, slit like, located on the anterior margin of the cleft situated on the posterior margin of the epigynal plate. Vulva centrally with a ˄-shaped structure. Vulval pocket global, situated anteriorly, widely separated from each other by ca. 1.5 
<normalizedToken id="C5646A43F0909E397AC4A937787E0CB9" originalValue="×">x</normalizedToken>
diameters. Spermathecae situated posteriorly to vulval pocket, relatively large, oval, ca. 1.6 
<normalizedToken id="3C232FD9D31291F232CD779F97922B23" originalValue="×">x</normalizedToken>
longer than wide, the two spermathecae are separated by 0.6 lengths.
</paragraph>
</subSubSection>
<subSubSection id="296B012938DD0FDD90E5BED4C9CECA69" pageId="0" pageNumber="107751" type="diagnosis">
<paragraph id="09AAF0C0CE7D012A161C93F6E1E3E277" pageId="0" pageNumber="107751">Diagnosis</paragraph>
<paragraph id="E1EE502EEA269E60EF7587043063F4F1" pageId="0" pageNumber="107751">
Males of 
<taxonomicName id="FD2C32E325AEADDD711D5C18D472E5FD" lsidName="S. hei" pageId="0" pageNumber="107751" rank="species" species="hei">
<emphasis id="9511DE657EB803237B7B2BF50D3D9489" italics="true" pageId="0" pageNumber="107751">S. hei</emphasis>
</taxonomicName>
Yu &amp; Lin, sp. n. can be distinguished from those of all other congeners by the direction of the terminal apophysis (MA in 
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), which is pointed ventrally (Fig. 
<figureCitation id="8D88FBDA502ECE7279A519E927DC7E58" captionStart="Figure 4" captionStartId="F9886409" captionText="Figure 4. Male left palp of the holotype of Speleoticus hei Yu &amp; Lin, sp. n. A Ventral view; B Dorsal view; C Prolateral view; D Retrolateral view. Abbreviations: Cp (I-III) = processes of the conductor (I-III); Dp (I-II) = distal processes of the paracymbium (I-II); E = embolus; Eb = embolic base; Ma = median apophysis; P = paracymbium; St = subtegulum; T = tegulum; Ta = terminal apophysis; Va = ventral apophysis of the paracymbium. Scale bar: 0.2 mm (equal for A-D)." figureDoi="10.3897/BDJ.11.e107751.figure4" httpUri="https://binary.pensoft.net/fig/885573" pageId="0" pageNumber="107751">4</figureCitation>
A, C and D) (vs. pointed prolaterally) and by the presence of a distinct and heavily-sclerotised median apophysis (Fig. 
<figureCitation id="0D71FC226C6BF262E829A56CD581FCFD" captionStart="Figure 4" captionStartId="F9886409" captionText="Figure 4. Male left palp of the holotype of Speleoticus hei Yu &amp; Lin, sp. n. A Ventral view; B Dorsal view; C Prolateral view; D Retrolateral view. Abbreviations: Cp (I-III) = processes of the conductor (I-III); Dp (I-II) = distal processes of the paracymbium (I-II); E = embolus; Eb = embolic base; Ma = median apophysis; P = paracymbium; St = subtegulum; T = tegulum; Ta = terminal apophysis; Va = ventral apophysis of the paracymbium. Scale bar: 0.2 mm (equal for A-D)." figureDoi="10.3897/BDJ.11.e107751.figure4" httpUri="https://binary.pensoft.net/fig/885573" pageId="0" pageNumber="107751">4</figureCitation>
A and D) (vs. median apophysis absent). Amongst the species of genus 
<taxonomicName id="861EA01C83E30921C49B97E48D9604B9" authorityName="Ballarin &amp; Li" authorityYear="2016" class="Arachnida" family="Nesticidae" genus="Speleoticus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Speleoticus" order="Araneae" pageId="0" pageNumber="107751" phylum="Arthropoda" rank="genus">
<emphasis id="A0F4667F283276848D7B879602AB58D6" italics="true" pageId="0" pageNumber="107751">Speleoticus</emphasis>
</taxonomicName>
, the female of the new species resembles 
<taxonomicName id="2DACBD2D9A99072C004536596876217D" lsidName="S. globosus" pageId="0" pageNumber="107751" rank="species" species="globosus">
<emphasis id="373A0433E0F450BF1F1555D7B5E78E44" italics="true" pageId="0" pageNumber="107751">S. globosus</emphasis>
</taxonomicName>
in having a similarly-shaped epigynum, but can be distinguished from the latter by: (1) copulatory openings indistinct (vs. distinct) (cf. Fig. 
<figureCitation id="535A00611F208C11FFFB84FFE25D2225" captionStart="Figure 3" captionStartId="F9886411" captionText="Figure 3. Speleoticus hei Yu &amp; Lin, sp. n., female paratype and male holotype. A, B Macerated epigyne, ventral and dorsal view; C, D Epigyne, macerated and embedded in Arabic gum, ventral and dorsal view; E, F Male, habitus, ventral and dorsal view; G, H Female, habitus, ventral and dorsal view. Abbreviations: Co = copulatory opening; Ms = median septum; S = spermatheca; Vp = vulval pocket. Scale bars: 0.2 mm (equal for A-D); 2 mm (equal for E-F, equal for G-H)." figureDoi="10.3897/BDJ.11.e107751.figure3" httpUri="https://binary.pensoft.net/fig/885493" pageId="0" pageNumber="107751">3</figureCitation>
A and C and 
<bibRefCitation id="C0106CE7F8D2DD406C7DA6B302F3AF13" author="Liu, J." journalOrPublisher="Zootaxa" pageId="0" pageNumber="107751" pagination="501 - 547" refId="B9886365" refString="Liu, J., Li, S. Q., 2013. New cave-dwelling spiders of the family Nesticidae (Arachnida, Araneae) from China. Zootaxa 3613: 501 - 547" title="New cave-dwelling spiders of the family Nesticidae (Arachnida, Araneae) from China" volume="3613" year="2013">Liu and Li (2013)</bibRefCitation>
: figs. 32D and 33B); (2) vulval pocket slightly smaller than spermatheca (vs. vulval pocket distinctly larger) (cf. Fig. 
<figureCitation id="0459046198DC81D0C28C0E2A529B9221" captionStart="Figure 3" captionStartId="F9886411" captionText="Figure 3. Speleoticus hei Yu &amp; Lin, sp. n., female paratype and male holotype. A, B Macerated epigyne, ventral and dorsal view; C, D Epigyne, macerated and embedded in Arabic gum, ventral and dorsal view; E, F Male, habitus, ventral and dorsal view; G, H Female, habitus, ventral and dorsal view. Abbreviations: Co = copulatory opening; Ms = median septum; S = spermatheca; Vp = vulval pocket. Scale bars: 0.2 mm (equal for A-D); 2 mm (equal for E-F, equal for G-H)." figureDoi="10.3897/BDJ.11.e107751.figure3" httpUri="https://binary.pensoft.net/fig/885493" pageId="0" pageNumber="107751">3</figureCitation>
B and D and 
<bibRefCitation id="DE571BE9C5F97A16F624617EA0B6E13B" author="Liu, J." journalOrPublisher="Zootaxa" pageId="0" pageNumber="107751" pagination="501 - 547" refId="B9886365" refString="Liu, J., Li, S. Q., 2013. New cave-dwelling spiders of the family Nesticidae (Arachnida, Araneae) from China. Zootaxa 3613: 501 - 547" title="New cave-dwelling spiders of the family Nesticidae (Arachnida, Araneae) from China" volume="3613" year="2013">Liu and Li (2013)</bibRefCitation>
: figs. 32B, D and 34A); (3) the presence of all eight eyes (vs. eyes absent) (cf. Fig. 
<figureCitation id="5BDC77E4750DDC502D4C2BE7C172EB2D" captionStart="Figure 3" captionStartId="F9886411" captionText="Figure 3. Speleoticus hei Yu &amp; Lin, sp. n., female paratype and male holotype. A, B Macerated epigyne, ventral and dorsal view; C, D Epigyne, macerated and embedded in Arabic gum, ventral and dorsal view; E, F Male, habitus, ventral and dorsal view; G, H Female, habitus, ventral and dorsal view. Abbreviations: Co = copulatory opening; Ms = median septum; S = spermatheca; Vp = vulval pocket. Scale bars: 0.2 mm (equal for A-D); 2 mm (equal for E-F, equal for G-H)." figureDoi="10.3897/BDJ.11.e107751.figure3" httpUri="https://binary.pensoft.net/fig/885493" pageId="0" pageNumber="107751">3</figureCitation>
and 
<bibRefCitation id="23E10102EFCEDC500BF9FD05BC3386A6" author="Liu, J." journalOrPublisher="Zootaxa" pageId="0" pageNumber="107751" pagination="501 - 547" refId="B9886365" refString="Liu, J., Li, S. Q., 2013. New cave-dwelling spiders of the family Nesticidae (Arachnida, Araneae) from China. Zootaxa 3613: 501 - 547" title="New cave-dwelling spiders of the family Nesticidae (Arachnida, Araneae) from China" volume="3613" year="2013">Liu and Li (2013)</bibRefCitation>
: fig. 34D).
</paragraph>
</subSubSection>
<subSubSection id="B6C36440ABA892EA391A0FD70153792B" pageId="0" pageNumber="107751" type="etymology">
<paragraph id="47A0CD823EFB5C4C434BAAA88669B7E9" pageId="0" pageNumber="107751">Etymology</paragraph>
<paragraph id="FB7569E4B73483E499FC7AA080BBD77F" pageId="0" pageNumber="107751">This species is a patronymic named after Mr. Li He (Chengdu City, China), collector of the types, who has greatly helped us in our research.</paragraph>
</subSubSection>
<subSubSection id="8CE643D9AE737B9665FE26BE15ADBB3A" pageId="0" pageNumber="107751" type="distribution">
<paragraph id="8DCB58AAE39F0B46AACB3768C1357766" pageId="0" pageNumber="107751">Distribution</paragraph>
<paragraph id="C1E57C0BC434BD053FF238DD62B1659B" pageId="0" pageNumber="107751">
Known only from the type locality, Hanwang Dong Cave, Guangyuan, Sichuan, China (Fig. 
<figureCitation id="5864BBB54B234971E088B029D8A69D39" captionStart="Figure 1" captionStartId="F9886405" captionText="Figure 1. Hanwang Dong Cave, type locality of Speleoticus hei Yu &amp; Lin, sp. n. A Location of Hanwang Dong Cave; B, C Entrance of Hanwang Dong Cave; D Habitat inside the cave; E Collection inside the cave." figureDoi="10.3897/BDJ.11.e107751.figure1" httpUri="https://binary.pensoft.net/fig/861449" pageId="0" pageNumber="107751">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="1C127A46D489EF01C4831376D44C5D77" pageId="0" pageNumber="107751" type="biology">
<paragraph id="0C5865EC0A276E60F65177D315DA79A9" pageId="0" pageNumber="107751">Biology</paragraph>
<paragraph id="336BC47DD8CCDD02DD310FC3930B83B8" pageId="0" pageNumber="107751">
The types of 
<taxonomicName id="9DA3E420C2B29D43726F830A1D83084D" lsidName="S. hei" pageId="0" pageNumber="107751" rank="species" species="hei">
<emphasis id="F8534C0973279EA042FAE745214E43E6" italics="true" pageId="0" pageNumber="107751">S. hei</emphasis>
</taxonomicName>
Yu &amp; Lin, sp. n. were collected under stones in the moist area about 50-500 m from the entrance in Hanwang Dong Cave.
</paragraph>
</subSubSection>
</treatment>
</document>