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<document id="3E736CE59A113342FDF80E35F9FB8793" ID-DOI="10.5281/zenodo.11205625" ID-ISSN="2189-6720" ID-Zenodo-Dep="11205625" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1714415462846" checkinUser="felipe" docAuthor="Kyohei Watanabe" docDate="2019" docId="60171F75FFF36615FC70FE44FABAFA64" docLanguage="en" docName="BullKanagawaPrefectMus.48.81-113.pdf" docOrigin="Bulletin of the Kanagawa Prefectural Museum (Natural Science) 48" docStyle="DocumentStyle:020B98DC1285EBBD02620724ECB44EDE.1:BullKanagawaPrefectMus.2013-.journal_article.english" docStyleId="020B98DC1285EBBD02620724ECB44EDE" docStyleName="BullKanagawaPrefectMus.2013-.journal_article.english" docStyleVersion="1" docTitle="Dihelus Townes 1970" docType="treatment" docVersion="4" lastPageNumber="89" masterDocId="9C2E670DFFFB661DFFA2FF97FF8CFF98" masterDocTitle="Taxonomic and Zoogeographical Notes of Japanese Cryptinae (Hymenoptera, Ichneumonidae), with Description of Five New Species" masterLastPageNumber="113" masterPageNumber="81" pageNumber="89" updateTime="1715891727401" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="4DE3F4DABA27E3D0B3BB19EC2B7024E5">Taxonomic and Zoogeographical Notes of Japanese Cryptinae (Hymenoptera, Ichneumonidae), with Description of Five New Species</mods:title>
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<mods:namePart id="54D97CF9844FFDA6998890D694292100">Kyohei Watanabe</mods:namePart>
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<subSubSection id="A0A4FDE8FFF36615FC70FE44FA97FE73" box="[978,1307,465,491]" pageId="8" pageNumber="89" type="nomenclature">
<paragraph id="E801AE63FFF36615FC70FE44FA97FE73" blockId="8.[850,1435,465,559]" box="[978,1307,465,491]" pageId="8" pageNumber="89">
<heading id="B349190FFFF36615FC70FE44FA97FE73" box="[978,1307,465,491]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="89" reason="2">
Genus
<taxonomicName id="2FBED5E0FFF36615FBBDFE46FA97FE73" ID-CoL="45F6" ID-ENA="540368" authority="Townes, 1970" authorityName="Townes" authorityYear="1970" box="[1055,1307,465,491]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="DACA7271FFF36615FBBDFE46FBF2FE73" bold="true" box="[1055,1150,465,491]" italics="true" pageId="8" pageNumber="89">Dihelus</emphasis>
<bibRefCitation id="8C2FD392FFF36615FB21FE44FA97FE73" author="Townes, H." box="[1155,1307,467,491]" pageId="8" pageNumber="89" pagination="1 - 537" refId="ref25408" refString="Townes, H., 1970. The genera of Ichneumonidae, Part 2. Memoirs of the American Entomological Institute, 12: 1 - 537." type="journal article" year="1970">Townes, 1970</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A0A4FDE8FFF36615FCF0FE62FAD3FDB7" pageId="8" pageNumber="89" type="reference_group">
<paragraph id="E801AE63FFF36615FCF0FE62FAD3FDB7" blockId="8.[850,1435,465,559]" pageId="8" pageNumber="89">
<treatmentCitationGroup id="C8AE894DFFF36615FCF0FE62FAD3FDB7" pageId="8" pageNumber="89">
<treatmentCitation id="691F8872FFF36615FCF0FE62FB1EFD94" author="Townes, H." box="[850,1170,501,524]" page="170" pageId="8" pageNumber="89" year="1970">
<taxonomicName id="2FBED5E0FFF36615FCF0FE62FB1EFD94" ID-CoL="45F6" authority="Townes, 1970: 170" authorityName="Townes" authorityPageNumber="170" authorityYear="1970" box="[850,1170,501,524]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="genus">
<emphasis id="DACA7271FFF36615FCF0FE62FC25FD94" box="[850,937,501,524]" italics="true" pageId="8" pageNumber="89">Dihelus</emphasis>
<bibRefCitation id="8C2FD392FFF36615FC16FE62FB1EFD94" author="Townes, H." box="[948,1170,501,524]" pageId="8" pageNumber="89" pagination="1 - 537" refId="ref25408" refString="Townes, H., 1970. The genera of Ichneumonidae, Part 2. Memoirs of the American Entomological Institute, 12: 1 - 537." type="journal article" year="1970">Townes, 1970: 170</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="370510C1FFF36615FB02FE62FB5BFD94" box="[1184,1239,501,524]" pageId="8" pageNumber="89">Type</typeStatus>
species:
<taxonomicName id="2FBED5E0FFF36615FAE6FE62FBF9FDB7" ID-CoL="364QD" authority="Townes, 1970" authorityName="Townes" authorityYear="1970" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="rufipleuris">
<emphasis id="DACA7271FFF36615FAE6FE62FC54FDB7" italics="true" pageId="8" pageNumber="89">Dihelus rufipleuris</emphasis>
<bibRefCitation id="8C2FD392FFF36615FC7CFD8FFBF9FDB7" author="Townes, H." box="[990,1141,536,559]" pageId="8" pageNumber="89" pagination="1 - 537" refId="ref25408" refString="Townes, H., 1970. The genera of Ichneumonidae, Part 2. Memoirs of the American Entomological Institute, 12: 1 - 537." type="journal article" year="1970">Townes, 1970</bibRefCitation>
</taxonomicName>
. Original designation.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="A0A4FDE8FFF36615FCC8FDF2FABAFA64" pageId="8" pageNumber="89" type="discussion">
<paragraph id="E801AE63FFF36615FCC8FDF2FABAFA64" blockId="8.[850,1437,612,1533]" pageId="8" pageNumber="89">
In
<collectingCountry id="90A9EEF3FFF36615FC2AFDF2FC49FDE4" box="[904,965,613,636]" name="Japan" pageId="8" pageNumber="89">Japan</collectingCountry>
, two species,
<taxonomicName id="2FBED5E0FFF36615FBF4FDF3FA17FDE4" ID-CoL="364Q7" authority="(Momoi, 1966)" baseAuthorityName="Momoi" baseAuthorityYear="1966" box="[1110,1435,612,638]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="hylaevorus">
<emphasis id="DACA7271FFF36615FBF4FDF3FB79FDE6" box="[1110,1269,612,638]" italics="true" pageId="8" pageNumber="89">D. hylaevorus</emphasis>
(
<bibRefCitation id="8C2FD392FFF36615FAA7FDF2FA18FDE4" author="Momoi, S." box="[1285,1428,613,636]" pageId="8" pageNumber="89" pagination="158 - 167" refId="ref24974" refString="Momoi, S., 1966. Descriptions of seven new species and a new genus of Mesostenini from Japan (Hymenoptera: Ichneumonidae). Kontyu, Tokyo, 34: 158 - 167." type="journal article" year="1966">Momoi, 1966</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="2FBED5E0FFF36615FCDFFD1DFB46FD3B" ID-CoL="6D67L" authority="Gupta &amp; Gupta (1978)" authorityName="Gupta &amp; Gupta" authorityYear="1978" box="[893,1226,650,676]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="DACA7271FFF36615FCDFFD1DFC56FD3C" box="[893,986,650,676]" italics="true" pageId="8" pageNumber="89">D. niger</emphasis>
<bibRefCitation id="8C2FD392FFF36615FC40FD1BFB46FD3B" author="Gupta, V. K. &amp; S. Gupta" box="[994,1226,652,675]" pageId="8" pageNumber="89" pagination="557 - 574" refId="ref24672" refString="Gupta, V. K. &amp; S. Gupta, 1978. Studies on the genus Dihelus from the oriental and eastern Palaearctic regions (Hymenoptera: Ichneumonidae). Oriental Insects, 12: 557 - 574." type="journal article" year="1978">Gupta &amp; Gupta (1978)</bibRefCitation>
</taxonomicName>
have been recorded. The latter species was described based on a single female (
<typeStatus id="370510C1FFF36615FCFBFD4DFC3FFD69" box="[857,947,730,753]" pageId="8" pageNumber="89" type="holotype">holotype</typeStatus>
) and a single male (
<typeStatus id="370510C1FFF36615FBDAFD4DFB47FD69" box="[1144,1227,730,753]" pageId="8" pageNumber="89" type="allotype">allotype</typeStatus>
) and
<bibRefCitation id="8C2FD392FFF36615FAA3FD4DFC1AFC80" author="Gupta, V. K. &amp; S. Gupta" pageId="8" pageNumber="89" pagination="557 - 574" refId="ref24672" refString="Gupta, V. K. &amp; S. Gupta, 1978. Studies on the genus Dihelus from the oriental and eastern Palaearctic regions (Hymenoptera: Ichneumonidae). Oriental Insects, 12: 557 - 574." type="journal article" year="1978">Gupta &amp; Gupta (1978)</bibRefCitation>
noted that this species can be distinguished from
<taxonomicName id="2FBED5E0FFF36615FCF0FCB1FC61FCD8" ID-CoL="364Q7" baseAuthorityName="Momoi" baseAuthorityYear="1966" box="[850,1005,806,832]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="hylaevorus">
<emphasis id="DACA7271FFF36615FCF0FCB1FC61FCD8" box="[850,1005,806,832]" italics="true" pageId="8" pageNumber="89">D. hylaevorus</emphasis>
</taxonomicName>
by the surface sculpture of face, frons and propodeum and the coloration of T1T4. Among them, the surface sculptures on face and propodeum are subjective and I could not recognize the differences between
<taxonomicName id="2FBED5E0FFF36615FADDFC0CFC47FC44" ID-CoL="364Q7" baseAuthorityName="Momoi" baseAuthorityYear="1966" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="hylaevorus">
<emphasis id="DACA7271FFF36615FADDFC0CFC47FC44" italics="true" pageId="8" pageNumber="89">D. hylaevorus</emphasis>
</taxonomicName>
and
<taxonomicName id="2FBED5E0FFF36615FC5CFC55FBD0FC44" ID-CoL="6D67L" authorityName="Gupta &amp; Gupta" authorityYear="1978" box="[1022,1116,962,988]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="DACA7271FFF36615FC5CFC55FBD0FC44" box="[1022,1116,962,988]" italics="true" pageId="8" pageNumber="89">D. niger</emphasis>
</taxonomicName>
. The surface sculpture of frons is somewhat objective, i.e., frons with (
<taxonomicName id="2FBED5E0FFF36615FB50FC7EFA1FFB9B" ID-CoL="364Q7" baseAuthorityName="Momoi" baseAuthorityYear="1966" box="[1266,1427,1001,1027]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="hylaevorus">
<emphasis id="DACA7271FFF36615FB50FC7EFA1FFB9B" box="[1266,1427,1001,1027]" italics="true" pageId="8" pageNumber="89">D. hylaevorus</emphasis>
</taxonomicName>
) or without (
<taxonomicName id="2FBED5E0FFF36615FC68FB87FBA4FBB2" ID-CoL="6D67L" authorityName="Gupta &amp; Gupta" authorityYear="1978" box="[970,1064,1040,1066]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="DACA7271FFF36615FC68FB87FBA4FBB2" box="[970,1064,1040,1066]" italics="true" pageId="8" pageNumber="89">D. niger</emphasis>
</taxonomicName>
) rugae below anterior ocellus, while this character states gradually changed in size (
<figureCitation id="7085B2E6FFF36615FA95FBAEFCEFFBEE" captionStart="Fig" captionStartId="2.[188,222,1328,1349]" captionTargetBox="[194,1433,158,1310]" captionTargetId="figure-238@2.[188,1436,151,1318]" captionTargetPageId="2" captionText="Fig. 1. Giraudia japonica sp. nov. (A, KPM-NK 69946; G, KPM-NK 69947), Apsilops scotinus (Tosquinet, 1903) (B, H, KPM-NK 69931), Hedycryptus ohshimensis (Uchida, 1930) (C, J, NIAES), Dihelus hylaevorus (Momoi, 1966) (D, MNHAH), Ischnus splendens sp. nov. (D, E, K, KPM-NK 69406), Nematopodius (Nematopodius) montanus sp. nov. (F, KPM-NK 69583) and Xoridesopus amamiensis sp. nov. (L, KPM-NK 69529), female ― AF, head, frontal (A, D), lateral (B, C, E) and posterolateral (F) view; G, H, IL, areolet of right wing; J, areolet and second recurrent vein of right wing." figureDoi="http://doi.org/10.5281/zenodo.11205628" httpUri="https://zenodo.org/record/11205628/files/figure.png" pageId="8" pageNumber="89">Figs. 1C F</figureCitation>
).
<bibRefCitation id="8C2FD392FFF36615FCD5FBC8FB81FBEE" author="Momoi, S." box="[887,1037,1119,1142]" pageId="8" pageNumber="89" pagination="158 - 167" refId="ref24974" refString="Momoi, S., 1966. Descriptions of seven new species and a new genus of Mesostenini from Japan (Hymenoptera: Ichneumonidae). Kontyu, Tokyo, 34: 158 - 167." type="journal article" year="1966">Momoi (1966)</bibRefCitation>
described the coloration of metasomal tergites as “tergites 1 through 4, and an apicomedian large spot on tergite 8, white.” but
<typeStatus id="370510C1FFF36615FB36FB3AFB76FB5C" box="[1172,1274,1197,1220]" pageId="8" pageNumber="89" type="paratype">paratypes</typeStatus>
and additional materials of
<taxonomicName id="2FBED5E0FFF36615FC7BFB45FBF2FB74" ID-CoL="364Q7" baseAuthorityName="Momoi" baseAuthorityYear="1966" box="[985,1150,1234,1260]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="hylaevorus">
<emphasis id="DACA7271FFF36615FC7BFB45FBF2FB74" box="[985,1150,1234,1260]" italics="true" pageId="8" pageNumber="89">D. hylaevorus</emphasis>
</taxonomicName>
indicate that this sentence may be “posterior margin of tergites 1 through 4...”.
<bibRefCitation id="8C2FD392FFF36615FCF0FAB5FBB7FAA1" author="Gupta, V. K. &amp; S. Gupta" box="[850,1083,1314,1337]" pageId="8" pageNumber="89" pagination="557 - 574" refId="ref24672" refString="Gupta, V. K. &amp; S. Gupta, 1978. Studies on the genus Dihelus from the oriental and eastern Palaearctic regions (Hymenoptera: Ichneumonidae). Oriental Insects, 12: 557 - 574." type="journal article" year="1978">Gupta &amp; Gupta (1978)</bibRefCitation>
referred this character states of
<taxonomicName id="2FBED5E0FFF36615FADDFAB7FC43FAF9" ID-CoL="364Q7" baseAuthorityName="Momoi" baseAuthorityYear="1966" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="hylaevorus">
<emphasis id="DACA7271FFF36615FADDFAB7FC43FAF9" italics="true" pageId="8" pageNumber="89">D. hylaevorus</emphasis>
</taxonomicName>
without correction and thus the differences of T1T4 between
<taxonomicName id="2FBED5E0FFF36615FB8AFAF9FB5EFA10" ID-CoL="364Q7" baseAuthorityName="Momoi" baseAuthorityYear="1966" box="[1064,1234,1390,1416]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="hylaevorus">
<emphasis id="DACA7271FFF36615FB8AFAF9FB5EFA10" box="[1064,1234,1390,1416]" italics="true" pageId="8" pageNumber="89">D. hylaevorus</emphasis>
</taxonomicName>
and
<taxonomicName id="2FBED5E0FFF36615FAAEFAF9FAFEFA10" ID-CoL="6D67L" authorityName="Gupta &amp; Gupta" authorityYear="1978" box="[1292,1394,1390,1416]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="DACA7271FFF36615FAAEFAF9FAFEFA10" box="[1292,1394,1390,1416]" italics="true" pageId="8" pageNumber="89">D. niger</emphasis>
</taxonomicName>
are based on incorrect information. By the above reasons, I conclude that both species are same and
<taxonomicName id="2FBED5E0FFF36615FB47FA2BFACDFA4E" ID-CoL="6D67L" authorityName="Gupta &amp; Gupta" authorityYear="1978" box="[1253,1345,1468,1494]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="DACA7271FFF36615FB47FA2BFACDFA4E" box="[1253,1345,1468,1494]" italics="true" pageId="8" pageNumber="89">D. niger</emphasis>
</taxonomicName>
is newly synonymized under
<taxonomicName id="2FBED5E0FFF36615FBB9FA74FB39FA65" ID-CoL="364Q7" baseAuthorityName="Momoi" baseAuthorityYear="1966" box="[1051,1205,1507,1533]" class="Insecta" family="Ichneumonidae" genus="Dihelus" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="89" phylum="Arthropoda" rank="species" species="hylaevorus">
<emphasis id="DACA7271FFF36615FBB9FA74FB39FA65" box="[1051,1205,1507,1533]" italics="true" pageId="8" pageNumber="89">D. hylaevorus</emphasis>
</taxonomicName>
(
<emphasis id="DACA7271FFF36615FB61FA74FAA4FA65" bold="true" box="[1219,1320,1507,1533]" pageId="8" pageNumber="89">syn. nov.</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>