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<document ID-DOI="http://doi.org/10.5281/zenodo.5670176" ID-GBIF-Dataset="5c1b56c8-dfa2-478a-833a-8c93d0f6b04a" ID-GBIF-Taxon="127638180" ID-ISSN="1464-5262" ID-Zenodo-Dep="5670176" checkinTime="1463679414422" checkinUser="plazi" docAuthor="Mariano C. Michat &amp; Patricia L. M. Torres" docDate="2016" docId="5D459B65FFA7FFBF059FAC39240257C8" docLanguage="en" docName="jNatHist.50.25.6.pdf" docOrigin="Journal of Natural History 50 (25)" docStyle="DocumentStyle{}" docTitle="Thermonectus tremouillesi" docType="treatment" docVersion="6" lastPageId="15" lastPageNumber="1646" masterDocId="A17CE31DFFA4FFB00423A94620745724" masterDocTitle="Thermonectus tremouillesi sp. nov. (Coleoptera: Dytiscidae: Aciliini): description of the adults and larvae and comparisons with other species of the genus" masterLastPageNumber="1648" masterPageNumber="1633" pageId="3" pageNumber="1635" updateTime="1636575875680" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Thermonectus tremouillesi sp. nov. (Coleoptera: Dytiscidae: Aciliini): description of the adults and larvae and comparisons with other species of the genus</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mariano C. Michat</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Patricia L. M. Torres</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>50</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>25</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1633</mods:start>
<mods:end>1648</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2016.1145274</mods:identifier>
<mods:identifier type="GBIF-Dataset">5c1b56c8-dfa2-478a-833a-8c93d0f6b04a</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">269894</mods:identifier>
<mods:identifier type="ZooBank">B3603E04-ADE4-4772-A12F-38EB77AC5951</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.5670176" ID-GBIF-Taxon="127638180" ID-Zenodo-Dep="5670176" LSID="urn:lsid:plazi:treatment:5D459B65FFA7FFBF059FAC39240257C8" httpUri="http://treatment.plazi.org/id/5D459B65FFA7FFBF059FAC39240257C8" lastPageId="15" lastPageNumber="1646" pageId="3" pageNumber="1635">
<subSubSection box="[444,872,1407,1432]" pageId="3" pageNumber="1635" type="nomenclature">
<paragraph blockId="3.[444,872,1407,1466]" box="[444,872,1407,1432]" pageId="3" pageNumber="1635">
<heading box="[444,872,1407,1432]" centered="true" fontSize="11" level="3" pageId="3" pageNumber="1635" reason="1">
<taxonomicName box="[444,770,1407,1432]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1633" phylum="Arthropoda" rank="species" species="tremouillesi" status="sp. nov.">
<emphasis bold="true" box="[444,770,1407,1432]" italics="true" pageId="3" pageNumber="1635">Thermonectus tremouIllesI</emphasis>
</taxonomicName>
sp. nov.
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[581,734,1441,1466]" pageId="3" pageNumber="1635" type="description">
<paragraph blockId="3.[444,872,1407,1466]" box="[581,734,1441,1466]" pageId="3" pageNumber="1635">
<heading box="[581,734,1441,1466]" centered="true" fontSize="11" level="3" pageId="3" pageNumber="1635" reason="1">
(
<figureCitation box="[589,726,1441,1466]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStart-3="Figure 4" captionStartId-0="6.[160,225,1584,1607]" captionStartId-1="7.[160,225,1580,1603]" captionStartId-2="10.[160,225,1592,1615]" captionStartId-3="11.[160,225,1587,1610]" captionTargetBox-0="[9,1150,142,1552]" captionTargetBox-1="[9,1152,142,1547]" captionTargetBox-2="[9,1152,142,1560]" captionTargetBox-3="[9,1152,142,1555]" captionTargetId-0="figure@6.[163,1153,142,1553]" captionTargetId-1="figure@7.[163,1153,142,1549]" captionTargetId-2="figure@10.[163,1153,142,1561]" captionTargetId-3="figure@11.[163,1153,142,1556]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="10" captionTargetPageId-3="11" captionText-0="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." captionText-1="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." captionText-2="Figure 3. Line drawings of instar I larvae of Thermonectus tremouillesi. (A) Cephalic capsule, dorsal aspect; (B) cephalic capsule, ventral aspect; (C) left antenna, dorsal aspect; (D) right antenna, ventral aspect; (E) left maxilla, dorsal aspect; (F) right maxilla, ventral aspect; (G) left mandible, dorsal aspect; (H) labium, dorsal aspect; (I) labium, ventral aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AN: antenna; EB: egg burster; FR: frontoclypeus; LA: labium; MN: mandible; MX: maxilla; PA: parietal; Sp: spinula; TP: tentorial pit." captionText-3="Figure 4. Line drawings of larvae of Thermonectus tremouillesi. (A) Left prothoracic leg of instar I, anterior aspect; (B) right prothoracic leg of instar I, posterior aspect; (C) abdominal segment VIII of instar I, dorsal aspect; (D) abdominal segment VIII of instar I, ventral aspect; (E) right urogomphus of instar I, dorsal aspect; (F) head of instar III, dorsal aspect, showing colour pattern on cephalic capsule (chaetotaxy omitted); (G) labium of instar III, dorsal aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AB: abdominal segment VIII; CO: coxa; FE: femur; PT: pretarsus; TA: tarsus; TI: tibia; TR: trochanter; UR: urogomphus." httpUri-0="https://zenodo.org/record/269895/files/figure.png" httpUri-1="https://zenodo.org/record/269896/files/figure.png" httpUri-2="https://zenodo.org/record/269897/files/figure.png" httpUri-3="https://zenodo.org/record/269898/files/figure.png" pageId="3" pageNumber="1635">
Figures
<number box="[684,726,1441,1466]" pageId="3" pageNumber="1635" value="2.5" valueMax="4.0" valueMin="1.0">
1
<emphasis bold="true" box="[698,712,1441,1466]" italics="true" pageId="3" pageNumber="1635"></emphasis>
4
</number>
</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="1637" pageId="3" pageNumber="1635" type="materials_examined">
<paragraph blockId="3.[160,671,1524,1583]" box="[160,311,1524,1549]" pageId="3" pageNumber="1635">
<heading bold="true" box="[160,311,1524,1549]" fontSize="11" level="2" pageId="3" pageNumber="1635" reason="0">
<emphasis bold="true" box="[160,311,1524,1549]" italics="true" pageId="3" pageNumber="1635">
<typeStatus box="[160,216,1524,1549]" pageId="3" pageNumber="1635">Type</typeStatus>
locality
</emphasis>
</heading>
</paragraph>
<paragraph blockId="3.[160,671,1524,1583]" box="[160,671,1559,1583]" pageId="3" pageNumber="1635">
<collectingCountry box="[160,272,1559,1583]" name="Argentina" pageId="3" pageNumber="1635">Argentina</collectingCountry>
, Entre Ríos Province, El Palmar NP.
</paragraph>
<paragraph blockId="3.[160,1156,1633,1727]" box="[160,420,1633,1658]" pageId="3" pageNumber="1635">
<heading bold="true" box="[160,420,1633,1658]" fontSize="11" level="2" pageId="3" pageNumber="1635" reason="0">
<emphasis bold="true" box="[160,420,1633,1658]" italics="true" pageId="3" pageNumber="1635">
<typeStatus box="[160,216,1633,1658]" pageId="3" pageNumber="1635">Type</typeStatus>
material (labels)
</emphasis>
</heading>
</paragraph>
<paragraph blockId="3.[160,1156,1633,1727]" lastBlockId="4.[160,1156,1320,1552]" lastPageId="4" lastPageNumber="1636" pageId="3" pageNumber="1635">
<typeStatus box="[160,265,1669,1693]" pageId="3" pageNumber="1635">Holotype</typeStatus>
male (
<collectionCode box="[346,423,1669,1693]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="3" pageNumber="1635">MACN</collectionCode>
):
<emphasis bold="true" box="[442,448,1668,1692]" italics="true" pageId="3" pageNumber="1635"></emphasis>
<collectingCountry box="[448,563,1669,1693]" name="Argentina" pageId="3" pageNumber="1635">Argentina</collectingCountry>
, Entre Ríos Prov., El Palmar NP,
<date box="[938,1065,1668,1693]" pageId="3" pageNumber="1635" value="2002-02-26">
<number box="[938,966,1669,1693]" pageId="3" pageNumber="1635" value="26.0">26</number>
<emphasis bold="true" box="[966,979,1668,1692]" italics="true" pageId="3" pageNumber="1635"></emphasis>
II
<emphasis bold="true" box="[993,1006,1668,1692]" italics="true" pageId="3" pageNumber="1635"></emphasis>
<number box="[1007,1065,1669,1693]" pageId="3" pageNumber="1635" value="2002.0">2002</number>
</date>
, Michat &amp; Torres col.
<emphasis bold="true" box="[305,311,1703,1727]" italics="true" pageId="3" pageNumber="1635"></emphasis>
[white, rectangular, printed label],
<emphasis bold="true" box="[724,730,1703,1727]" italics="true" pageId="3" pageNumber="1635"></emphasis>
<typeStatus box="[729,853,1703,1727]" pageId="3" pageNumber="1635">HOLOTYPE</typeStatus>
<taxonomicName authority="Michat &amp; Torres" authorityName="Michat &amp; Torres" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" lastPageId="4" lastPageNumber="1636" order="Coleoptera" pageId="3" pageNumber="1635" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[862,1155,1703,1727]" italics="true" pageId="3" pageNumber="1635">Thermonectus tremouillesi</emphasis>
Michat &amp; Torres
</taxonomicName>
<emphasis bold="true" box="[349,355,1320,1344]" italics="true" pageId="4" pageNumber="1636"></emphasis>
[red, rectangular, printed label].
<typeStatus box="[757,868,1320,1344]" pageId="4" pageNumber="1636">Paratypes</typeStatus>
(
<number box="[888,916,1320,1344]" pageId="4" pageNumber="1636" value="28.0">28</number>
exs.): same data as
<typeStatus box="[160,262,1355,1379]" pageId="4" pageNumber="1636">holotype</typeStatus>
[white, rectangular, printed label],
<emphasis bold="true" box="[714,720,1355,1379]" italics="true" pageId="4" pageNumber="1636"></emphasis>
<typeStatus box="[719,837,1355,1379]" pageId="4" pageNumber="1636">PARATYPE</typeStatus>
<taxonomicName authority="Michat &amp; Torres" authorityName="Michat &amp; Torres" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1636" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[854,1155,1355,1379]" italics="true" pageId="4" pageNumber="1636">Thermonectus tremouillesi</emphasis>
Michat &amp; Torres
</taxonomicName>
<emphasis bold="true" box="[346,352,1389,1413]" italics="true" pageId="4" pageNumber="1636"></emphasis>
[red, rectangular, printed label] (
<specimenCount box="[747,853,1390,1414]" pageId="4" pageNumber="1636" type="male">
<number box="[747,774,1390,1414]" pageId="4" pageNumber="1636" value="13.0">13</number>
males
</specimenCount>
, five females,
<collectionCode box="[1024,1098,1390,1414]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="4" pageNumber="1636">MACN</collectionCode>
; one male, one female
<collectionCode box="[387,459,1424,1448]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="4" pageNumber="1636">NHML</collectionCode>
; one male, one female
<collectionCode box="[764,842,1424,1448]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="1636">MNHN</collectionCode>
); same data as
<typeStatus box="[1047,1151,1424,1448]" pageId="4" pageNumber="1636">holotype</typeStatus>
, except
<emphasis bold="true" box="[249,255,1459,1483]" italics="true" pageId="4" pageNumber="1636"></emphasis>
<date box="[255,379,1459,1483]" pageId="4" pageNumber="1636" value="2004-02-26">
<number box="[255,283,1459,1483]" pageId="4" pageNumber="1636" value="26.0">26</number>
<emphasis bold="true" box="[282,295,1459,1483]" italics="true" pageId="4" pageNumber="1636"></emphasis>
II
<emphasis bold="true" box="[309,322,1459,1483]" italics="true" pageId="4" pageNumber="1636"></emphasis>
<number box="[323,379,1459,1483]" pageId="4" pageNumber="1636" value="2004.0">2004</number>
</date>
<emphasis bold="true" box="[379,385,1459,1483]" italics="true" pageId="4" pageNumber="1636"></emphasis>
[white, rectangular, printed label],
<emphasis bold="true" box="[813,819,1459,1483]" italics="true" pageId="4" pageNumber="1636"></emphasis>
<typeStatus box="[819,937,1459,1483]" pageId="4" pageNumber="1636">PARATYPE</typeStatus>
<taxonomicName authority="Michat &amp; Torres" authorityName="Michat &amp; Torres" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1636" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis italics="true" pageId="4" pageNumber="1636">Thermonectus tremouillesi</emphasis>
Michat &amp; Torres
</taxonomicName>
<emphasis bold="true" box="[462,468,1493,1517]" italics="true" pageId="4" pageNumber="1636"></emphasis>
[red, rectangular, printed label] (five males, one female,
<collectionCode box="[160,236,1528,1552]" country="Argentina" httpUri="http://grbio.org/cool/pyng-6456" name="Museo Argentino de Ciencias Naturales Bernardino Rivadavia" pageId="4" pageNumber="1636">MACN</collectionCode>
).
</paragraph>
<caption ID-Table-UUID="81937AFBFFA0FFB404D3A9C823885781" box="[240,1020,142,165]" httpUri="http://table.plazi.org/id/81937AFBFFA0FFB404D3A9C823885781" pageId="4" pageNumber="1636" targetBox="[250,1064,178,1069]" targetIsTable="true" targetPageId="4">
<paragraph blockId="4.[240,1077,142,1222]" box="[240,1020,142,165]" pageId="4" pageNumber="1636">
<emphasis bold="true" box="[240,323,142,165]" pageId="4" pageNumber="1636">Table 1.</emphasis>
Measurements and ratios for the larvae of
<taxonomicName box="[763,1015,142,165]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1636" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[763,1015,142,165]" italics="true" pageId="4" pageNumber="1636">Thermonectus tremouillesi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="4.[240,1077,142,1222]" pageId="4" pageNumber="1636">
<table box="[250,1064,178,1069]" gridcols="4" gridrows="37" pageId="4" pageNumber="1636">
<tr box="[250,1064,178,197]" gridrow="0" pageId="4" pageNumber="1636">
<th box="[250,344,178,197]" gridcol="0" gridrow="0" pageId="4" pageNumber="1636">Measure</th>
<th box="[454,579,178,197]" gridcol="1" gridrow="0" pageId="4" pageNumber="1636">Instar I (n = 2)</th>
<th box="[690,820,178,197]" gridcol="2" gridrow="0" pageId="4" pageNumber="1636">Instar II (n = 1)</th>
<th box="[930,1064,178,197]" gridcol="3" gridrow="0" pageId="4" pageNumber="1636">Instar III (n = 5)</th>
</tr>
<tr box="[250,1064,210,229]" gridrow="1" pageId="4" pageNumber="1636">
<th box="[250,344,210,229]" gridcol="0" gridrow="1" pageId="4" pageNumber="1636">TL (mm)</th>
<td box="[454,579,210,229]" gridcol="1" gridrow="1" pageId="4" pageNumber="1636">8.008.30</td>
<td box="[690,820,210,229]" gridcol="2" gridrow="1" pageId="4" pageNumber="1636">13.10</td>
<td box="[930,1064,210,229]" gridcol="3" gridrow="1" pageId="4" pageNumber="1636">15.2023.60</td>
</tr>
<tr box="[250,1064,234,253]" gridrow="2" pageId="4" pageNumber="1636">
<th box="[250,344,234,253]" gridcol="0" gridrow="2" pageId="4" pageNumber="1636">MW (mm)</th>
<td box="[454,579,234,253]" gridcol="1" gridrow="2" pageId="4" pageNumber="1636">0.801.20</td>
<td box="[690,820,234,253]" gridcol="2" gridrow="2" pageId="4" pageNumber="1636">1.60</td>
<td box="[930,1064,234,253]" gridcol="3" gridrow="2" pageId="4" pageNumber="1636">2.102.90</td>
</tr>
<tr box="[250,1064,258,277]" gridrow="3" pageId="4" pageNumber="1636">
<th box="[250,344,258,277]" gridcol="0" gridrow="3" pageId="4" pageNumber="1636">HL (mm)</th>
<td box="[454,579,258,277]" gridcol="1" gridrow="3" pageId="4" pageNumber="1636">1.40</td>
<td box="[690,820,258,277]" gridcol="2" gridrow="3" pageId="4" pageNumber="1636">1.88</td>
<td box="[930,1064,258,277]" gridcol="3" gridrow="3" pageId="4" pageNumber="1636">2.702.83</td>
</tr>
<tr box="[250,1064,282,301]" gridrow="4" pageId="4" pageNumber="1636">
<th box="[250,344,282,301]" gridcol="0" gridrow="4" pageId="4" pageNumber="1636">HW (mm)</th>
<td box="[454,579,282,301]" gridcol="1" gridrow="4" pageId="4" pageNumber="1636">1.15</td>
<td box="[690,820,282,301]" gridcol="2" gridrow="4" pageId="4" pageNumber="1636">1.56</td>
<td box="[930,1064,282,301]" gridcol="3" gridrow="4" pageId="4" pageNumber="1636">2.162.21</td>
</tr>
<tr box="[250,1064,306,325]" gridrow="5" pageId="4" pageNumber="1636">
<th box="[250,344,306,325]" gridcol="0" gridrow="5" pageId="4" pageNumber="1636">FRL (mm)</th>
<td box="[454,579,306,325]" gridcol="1" gridrow="5" pageId="4" pageNumber="1636">0.640.65</td>
<td box="[690,820,306,325]" gridcol="2" gridrow="5" pageId="4" pageNumber="1636">0.84</td>
<td box="[930,1064,306,325]" gridcol="3" gridrow="5" pageId="4" pageNumber="1636">1.161.22</td>
</tr>
<tr box="[250,1064,330,349]" gridrow="6" pageId="4" pageNumber="1636">
<th box="[250,344,330,349]" gridcol="0" gridrow="6" pageId="4" pageNumber="1636">OCW (mm)</th>
<td box="[454,579,330,349]" gridcol="1" gridrow="6" pageId="4" pageNumber="1636">0.43</td>
<td box="[690,820,330,349]" gridcol="2" gridrow="6" pageId="4" pageNumber="1636">0.56</td>
<td box="[930,1064,330,349]" gridcol="3" gridrow="6" pageId="4" pageNumber="1636">0.920.99</td>
</tr>
<tr box="[250,1064,354,373]" gridrow="7" pageId="4" pageNumber="1636">
<th box="[250,344,354,373]" gridcol="0" gridrow="7" pageId="4" pageNumber="1636">HL/HW</th>
<td box="[454,579,354,373]" gridcol="1" gridrow="7" pageId="4" pageNumber="1636">1.22</td>
<td box="[690,820,354,373]" gridcol="2" gridrow="7" pageId="4" pageNumber="1636">1.20</td>
<td box="[930,1064,354,373]" gridcol="3" gridrow="7" pageId="4" pageNumber="1636">1.231.28</td>
</tr>
<tr box="[250,1064,378,397]" gridrow="8" pageId="4" pageNumber="1636">
<th box="[250,344,378,397]" gridcol="0" gridrow="8" pageId="4" pageNumber="1636">HW/OCW</th>
<td box="[454,579,378,397]" gridcol="1" gridrow="8" pageId="4" pageNumber="1636">2.71</td>
<td box="[690,820,378,397]" gridcol="2" gridrow="8" pageId="4" pageNumber="1636">2.78</td>
<td box="[930,1064,378,397]" gridcol="3" gridrow="8" pageId="4" pageNumber="1636">2.222.38</td>
</tr>
<tr box="[250,1064,402,421]" gridrow="9" pageId="4" pageNumber="1636">
<th box="[250,344,402,421]" gridcol="0" gridrow="9" pageId="4" pageNumber="1636">COL/HL</th>
<td box="[454,579,402,421]" gridcol="1" gridrow="9" pageId="4" pageNumber="1636">0.54</td>
<td box="[690,820,402,421]" gridcol="2" gridrow="9" pageId="4" pageNumber="1636">0.55</td>
<td box="[930,1064,402,421]" gridcol="3" gridrow="9" pageId="4" pageNumber="1636">0.560.57</td>
</tr>
<tr box="[250,1064,426,445]" gridrow="10" pageId="4" pageNumber="1636">
<th box="[250,344,426,445]" gridcol="0" gridrow="10" pageId="4" pageNumber="1636">FRL/HL</th>
<td box="[454,579,426,445]" gridcol="1" gridrow="10" pageId="4" pageNumber="1636">0.46</td>
<td box="[690,820,426,445]" gridcol="2" gridrow="10" pageId="4" pageNumber="1636">0.45</td>
<td box="[930,1064,426,445]" gridcol="3" gridrow="10" pageId="4" pageNumber="1636">0.430.44</td>
</tr>
<tr box="[250,1064,450,469]" gridrow="11" pageId="4" pageNumber="1636">
<th box="[250,344,450,469]" gridcol="0" gridrow="11" pageId="4" pageNumber="1636">A/HW</th>
<td box="[454,579,450,469]" gridcol="1" gridrow="11" pageId="4" pageNumber="1636">0.50</td>
<td box="[690,820,450,469]" gridcol="2" gridrow="11" pageId="4" pageNumber="1636">0.44</td>
<td box="[930,1064,450,469]" gridcol="3" gridrow="11" pageId="4" pageNumber="1636">0.370.42</td>
</tr>
<tr box="[250,1064,474,493]" gridrow="12" pageId="4" pageNumber="1636">
<th box="[250,344,474,493]" gridcol="0" gridrow="12" pageId="4" pageNumber="1636">A1/A3</th>
<td box="[454,579,474,493]" gridcol="1" gridrow="12" pageId="4" pageNumber="1636">0.79</td>
<td box="[690,820,474,493]" gridcol="2" gridrow="12" pageId="4" pageNumber="1636">0.79</td>
<td box="[930,1064,474,493]" gridcol="3" gridrow="12" pageId="4" pageNumber="1636">0.710.86</td>
</tr>
<tr box="[250,1064,498,517]" gridrow="13" pageId="4" pageNumber="1636">
<th box="[250,344,498,517]" gridcol="0" gridrow="13" pageId="4" pageNumber="1636">A2/A3</th>
<td box="[454,579,498,517]" gridcol="1" gridrow="13" pageId="4" pageNumber="1636">0.790.82</td>
<td box="[690,820,498,517]" gridcol="2" gridrow="13" pageId="4" pageNumber="1636">0.88</td>
<td box="[930,1064,498,517]" gridcol="3" gridrow="13" pageId="4" pageNumber="1636">0.810.93</td>
</tr>
<tr box="[250,1064,522,541]" gridrow="14" pageId="4" pageNumber="1636">
<th box="[250,344,522,541]" gridcol="0" gridrow="14" pageId="4" pageNumber="1636">A4/A3</th>
<td box="[454,579,522,541]" gridcol="1" gridrow="14" pageId="4" pageNumber="1636">0.330.37</td>
<td box="[690,820,522,541]" gridcol="2" gridrow="14" pageId="4" pageNumber="1636">0.25</td>
<td box="[930,1064,522,541]" gridcol="3" gridrow="14" pageId="4" pageNumber="1636">0.160.20</td>
</tr>
<tr box="[250,1064,546,565]" gridrow="15" pageId="4" pageNumber="1636">
<th box="[250,344,546,565]" gridcol="0" gridrow="15" pageId="4" pageNumber="1636">MNL/MNW</th>
<td box="[454,579,546,565]" gridcol="1" gridrow="15" pageId="4" pageNumber="1636">2.742.89</td>
<td box="[690,820,546,565]" gridcol="2" gridrow="15" pageId="4" pageNumber="1636">3.04</td>
<td box="[930,1064,546,565]" gridcol="3" gridrow="15" pageId="4" pageNumber="1636">3.133.30</td>
</tr>
<tr box="[250,1064,570,589]" gridrow="16" pageId="4" pageNumber="1636">
<th box="[250,344,570,589]" gridcol="0" gridrow="16" pageId="4" pageNumber="1636">MNL/HL</th>
<td box="[454,579,570,589]" gridcol="1" gridrow="16" pageId="4" pageNumber="1636">0.46</td>
<td box="[690,820,570,589]" gridcol="2" gridrow="16" pageId="4" pageNumber="1636">0.47</td>
<td box="[930,1064,570,589]" gridcol="3" gridrow="16" pageId="4" pageNumber="1636">0.440.46</td>
</tr>
<tr box="[250,1064,594,613]" gridrow="17" pageId="4" pageNumber="1636">
<th box="[250,344,594,613]" gridcol="0" gridrow="17" pageId="4" pageNumber="1636">PPF/MP1</th>
<td box="[454,579,594,613]" gridcol="1" gridrow="17" pageId="4" pageNumber="1636">0.360.40</td>
<td box="[690,820,594,613]" gridcol="2" gridrow="17" pageId="4" pageNumber="1636">0.22</td>
<td box="[930,1064,594,613]" gridcol="3" gridrow="17" pageId="4" pageNumber="1636">0.200.25</td>
</tr>
<tr box="[250,1064,618,637]" gridrow="18" pageId="4" pageNumber="1636">
<th box="[250,344,618,637]" gridcol="0" gridrow="18" pageId="4" pageNumber="1636">A/MP</th>
<td box="[454,579,618,637]" gridcol="1" gridrow="18" pageId="4" pageNumber="1636">1.771.81</td>
<td box="[690,820,618,637]" gridcol="2" gridrow="18" pageId="4" pageNumber="1636">1.79</td>
<td box="[930,1064,618,637]" gridcol="3" gridrow="18" pageId="4" pageNumber="1636">1.581.76</td>
</tr>
<tr box="[250,1064,642,661]" gridrow="19" pageId="4" pageNumber="1636">
<th box="[250,344,642,661]" gridcol="0" gridrow="19" pageId="4" pageNumber="1636">MP1/MP2</th>
<td box="[454,579,642,661]" gridcol="1" gridrow="19" pageId="4" pageNumber="1636">0.560.58</td>
<td box="[690,820,642,661]" gridcol="2" gridrow="19" pageId="4" pageNumber="1636">0.75</td>
<td box="[930,1064,642,661]" gridcol="3" gridrow="19" pageId="4" pageNumber="1636">0.760.91</td>
</tr>
<tr box="[250,1064,666,685]" gridrow="20" pageId="4" pageNumber="1636">
<th box="[250,344,666,685]" gridcol="0" gridrow="20" pageId="4" pageNumber="1636">MP3/MP2</th>
<td box="[454,579,666,685]" gridcol="1" gridrow="20" pageId="4" pageNumber="1636">1.841.89</td>
<td box="[690,820,666,685]" gridcol="2" gridrow="20" pageId="4" pageNumber="1636">1.50</td>
<td box="[930,1064,666,685]" gridcol="3" gridrow="20" pageId="4" pageNumber="1636">1.291.45</td>
</tr>
<tr box="[250,1064,690,709]" gridrow="21" pageId="4" pageNumber="1636">
<th box="[250,344,690,709]" gridcol="0" gridrow="21" pageId="4" pageNumber="1636">GA/MP1</th>
<td box="[454,579,690,709]" gridcol="1" gridrow="21" pageId="4" pageNumber="1636">2.913.20</td>
<td box="[690,820,690,709]" gridcol="2" gridrow="21" pageId="4" pageNumber="1636">2.61</td>
<td box="[930,1064,690,709]" gridcol="3" gridrow="21" pageId="4" pageNumber="1636">2.062.48</td>
</tr>
<tr box="[250,1064,714,733]" gridrow="22" pageId="4" pageNumber="1636">
<th box="[250,344,714,733]" gridcol="0" gridrow="22" pageId="4" pageNumber="1636">MP/LP</th>
<td box="[454,579,714,733]" gridcol="1" gridrow="22" pageId="4" pageNumber="1636">0.880.89</td>
<td box="[690,820,714,733]" gridcol="2" gridrow="22" pageId="4" pageNumber="1636">0.88</td>
<td box="[930,1064,714,733]" gridcol="3" gridrow="22" pageId="4" pageNumber="1636">0.820.87</td>
</tr>
<tr box="[250,1064,738,757]" gridrow="23" pageId="4" pageNumber="1636">
<th box="[250,344,738,757]" gridcol="0" gridrow="23" pageId="4" pageNumber="1636">LP2/LP1</th>
<td box="[454,579,738,757]" gridcol="1" gridrow="23" pageId="4" pageNumber="1636">0.940.95</td>
<td box="[690,820,738,757]" gridcol="2" gridrow="23" pageId="4" pageNumber="1636">0.82</td>
<td box="[930,1064,738,757]" gridcol="3" gridrow="23" pageId="4" pageNumber="1636">0.640.72</td>
</tr>
<tr box="[250,1064,762,781]" gridrow="24" pageId="4" pageNumber="1636">
<th box="[250,344,762,781]" gridcol="0" gridrow="24" pageId="4" pageNumber="1636">L3 (mm)</th>
<td box="[454,579,762,781]" gridcol="1" gridrow="24" pageId="4" pageNumber="1636">2.862.98</td>
<td box="[690,820,762,781]" gridcol="2" gridrow="24" pageId="4" pageNumber="1636">4.03</td>
<td box="[930,1064,762,781]" gridcol="3" gridrow="24" pageId="4" pageNumber="1636">5.816.25</td>
</tr>
<tr box="[250,1064,786,805]" gridrow="25" pageId="4" pageNumber="1636">
<th box="[250,344,786,805]" gridcol="0" gridrow="25" pageId="4" pageNumber="1636">L3/L1</th>
<td box="[454,579,786,805]" gridcol="1" gridrow="25" pageId="4" pageNumber="1636">1.001.03</td>
<td box="[690,820,786,805]" gridcol="2" gridrow="25" pageId="4" pageNumber="1636">1.07</td>
<td box="[930,1064,786,805]" gridcol="3" gridrow="25" pageId="4" pageNumber="1636">1.111.12</td>
</tr>
<tr box="[250,1064,810,829]" gridrow="26" pageId="4" pageNumber="1636">
<th box="[250,344,810,829]" gridcol="0" gridrow="26" pageId="4" pageNumber="1636">L3/L2</th>
<td box="[454,579,810,829]" gridcol="1" gridrow="26" pageId="4" pageNumber="1636">0.950.97</td>
<td box="[690,820,810,829]" gridcol="2" gridrow="26" pageId="4" pageNumber="1636">0.99</td>
<td box="[930,1064,810,829]" gridcol="3" gridrow="26" pageId="4" pageNumber="1636">1.011.02</td>
</tr>
<tr box="[250,1064,834,853]" gridrow="27" pageId="4" pageNumber="1636">
<th box="[250,344,834,853]" gridcol="0" gridrow="27" pageId="4" pageNumber="1636">L3/HW</th>
<td box="[454,579,834,853]" gridcol="1" gridrow="27" pageId="4" pageNumber="1636">2.59</td>
<td box="[690,820,834,853]" gridcol="2" gridrow="27" pageId="4" pageNumber="1636">2.58</td>
<td box="[930,1064,834,853]" gridcol="3" gridrow="27" pageId="4" pageNumber="1636">2.682.82</td>
</tr>
<tr box="[250,1064,858,877]" gridrow="28" pageId="4" pageNumber="1636">
<th box="[250,344,858,877]" gridcol="0" gridrow="28" pageId="4" pageNumber="1636">L3 (CO/FE)</th>
<td box="[454,579,858,877]" gridcol="1" gridrow="28" pageId="4" pageNumber="1636">0.740.81</td>
<td box="[690,820,858,877]" gridcol="2" gridrow="28" pageId="4" pageNumber="1636">0.76</td>
<td box="[930,1064,858,877]" gridcol="3" gridrow="28" pageId="4" pageNumber="1636">0.810.86</td>
</tr>
<tr box="[250,1064,882,901]" gridrow="29" pageId="4" pageNumber="1636">
<th box="[250,344,882,901]" gridcol="0" gridrow="29" pageId="4" pageNumber="1636">L3 (TI/FE)</th>
<td box="[454,579,882,901]" gridcol="1" gridrow="29" pageId="4" pageNumber="1636">0.760.77</td>
<td box="[690,820,882,901]" gridcol="2" gridrow="29" pageId="4" pageNumber="1636">0.72</td>
<td box="[930,1064,882,901]" gridcol="3" gridrow="29" pageId="4" pageNumber="1636">0.650.70</td>
</tr>
<tr box="[250,1064,906,925]" gridrow="30" pageId="4" pageNumber="1636">
<th box="[250,344,906,925]" gridcol="0" gridrow="30" pageId="4" pageNumber="1636">L3 (TA/FE)</th>
<td box="[454,579,906,925]" gridcol="1" gridrow="30" pageId="4" pageNumber="1636">0.580.59</td>
<td box="[690,820,906,925]" gridcol="2" gridrow="30" pageId="4" pageNumber="1636">0.52</td>
<td box="[930,1064,906,925]" gridcol="3" gridrow="30" pageId="4" pageNumber="1636">0.450.48</td>
</tr>
<tr box="[250,1064,930,949]" gridrow="31" pageId="4" pageNumber="1636">
<th box="[250,344,930,949]" gridcol="0" gridrow="31" pageId="4" pageNumber="1636">L3 (CL/TA)</th>
<td box="[454,579,930,949]" gridcol="1" gridrow="31" pageId="4" pageNumber="1636">0.360.37</td>
<td box="[690,820,930,949]" gridcol="2" gridrow="31" pageId="4" pageNumber="1636">0.25</td>
<td box="[930,1064,930,949]" gridcol="3" gridrow="31" pageId="4" pageNumber="1636">0.230.24</td>
</tr>
<tr box="[250,1064,954,973]" gridrow="32" pageId="4" pageNumber="1636">
<th box="[250,344,954,973]" gridcol="0" gridrow="32" pageId="4" pageNumber="1636">LAS (mm)</th>
<td box="[454,579,954,973]" gridcol="1" gridrow="32" pageId="4" pageNumber="1636">1.551.63</td>
<td box="[690,820,954,973]" gridcol="2" gridrow="32" pageId="4" pageNumber="1636">1.93</td>
<td box="[930,1064,954,973]" gridcol="3" gridrow="32" pageId="4" pageNumber="1636">2.683.00</td>
</tr>
<tr box="[250,1064,978,997]" gridrow="33" pageId="4" pageNumber="1636">
<th box="[250,344,978,997]" gridcol="0" gridrow="33" pageId="4" pageNumber="1636">LAS/HW</th>
<td box="[454,579,978,997]" gridcol="1" gridrow="33" pageId="4" pageNumber="1636">1.41</td>
<td box="[690,820,978,997]" gridcol="2" gridrow="33" pageId="4" pageNumber="1636">1.23</td>
<td box="[930,1064,978,997]" gridcol="3" gridrow="33" pageId="4" pageNumber="1636">1.221.33</td>
</tr>
<tr box="[250,1064,1002,1021]" gridrow="34" pageId="4" pageNumber="1636">
<th box="[250,344,1002,1021]" gridcol="0" gridrow="34" pageId="4" pageNumber="1636">U (mm)</th>
<td box="[454,579,1002,1021]" gridcol="1" gridrow="34" pageId="4" pageNumber="1636">0.60</td>
<td box="[690,820,1002,1021]" gridcol="2" gridrow="34" pageId="4" pageNumber="1636">0.75</td>
<td box="[930,1064,1002,1021]" gridcol="3" gridrow="34" pageId="4" pageNumber="1636">1.051.34</td>
</tr>
<tr box="[250,1064,1026,1045]" gridrow="35" pageId="4" pageNumber="1636">
<th box="[250,344,1026,1045]" gridcol="0" gridrow="35" pageId="4" pageNumber="1636">U/LAS</th>
<td box="[454,579,1026,1045]" gridcol="1" gridrow="35" pageId="4" pageNumber="1636">0.370.39</td>
<td box="[690,820,1026,1045]" gridcol="2" gridrow="35" pageId="4" pageNumber="1636">0.39</td>
<td box="[930,1064,1026,1045]" gridcol="3" gridrow="35" pageId="4" pageNumber="1636">0.390.45</td>
</tr>
<tr box="[250,1064,1050,1069]" gridrow="36" pageId="4" pageNumber="1636">
<th box="[250,344,1050,1069]" gridcol="0" gridrow="36" pageId="4" pageNumber="1636">U/HW</th>
<td box="[454,579,1050,1069]" gridcol="1" gridrow="36" pageId="4" pageNumber="1636">0.52</td>
<td box="[690,820,1050,1069]" gridcol="2" gridrow="36" pageId="4" pageNumber="1636">0.48</td>
<td box="[930,1064,1050,1069]" gridcol="3" gridrow="36" pageId="4" pageNumber="1636">0.490.59</td>
</tr>
</table>
</paragraph>
<tableNote pageId="4" pageNumber="1636" targetBox="[250,1064,178,1069]" targetPageId="4">
<paragraph blockId="4.[240,1077,142,1222]" box="[240,1077,1083,1102]" pageId="4" pageNumber="1636">A: antenna; A1A4: antennomeres 14; CL: longest claw; CO: coxa; COL: coronal line length; FE:</paragraph>
<paragraph blockId="4.[240,1077,142,1222]" box="[240,1075,1107,1126]" pageId="4" pageNumber="1636">femur; FRL: frontoclypeus length; GA: galea; HL: head length; HW: head width; LAS: length of last</paragraph>
<paragraph blockId="4.[240,1077,142,1222]" box="[240,1077,1131,1150]" pageId="4" pageNumber="1636">abdominal segment; LP: labial palpus; LP1LP2: labial palpomeres 12; L1L3: legs 13; MNL:</paragraph>
<paragraph blockId="4.[240,1077,142,1222]" box="[240,1075,1155,1175]" pageId="4" pageNumber="1636">mandible length; MNW: mandible width; MP: maxillary palpus; MP1MP3: maxillary palpomeres</paragraph>
<paragraph blockId="4.[240,1077,142,1222]" box="[240,1077,1179,1198]" pageId="4" pageNumber="1636">13; MW: maximum body width; OCW: occipital foramen width; PPF: palpifer; TA: tarsus; TI: tibia; TL:</paragraph>
<paragraph blockId="4.[240,1077,142,1222]" box="[240,795,1203,1222]" pageId="4" pageNumber="1636">total body length (excluding urogomphi); U: urogomphus length.</paragraph>
</tableNote>
<paragraph blockId="4.[160,1155,1618,1712]" box="[160,517,1618,1643]" pageId="4" pageNumber="1636">
<heading bold="true" box="[160,517,1618,1643]" fontSize="11" level="2" pageId="4" pageNumber="1636" reason="0">
<emphasis bold="true" box="[160,517,1618,1643]" italics="true" pageId="4" pageNumber="1636">Additional material examined</emphasis>
</heading>
</paragraph>
<paragraph blockId="4.[160,1155,1618,1712]" lastBlockId="5.[160,1155,916,975]" lastPageId="5" lastPageNumber="1637" pageId="4" pageNumber="1636">
These are damaged or parts of specimens that we prefer not to include in the
<typeStatus box="[1105,1155,1654,1678]" pageId="4" pageNumber="1636">type</typeStatus>
series:
<collectingCountry box="[238,350,1688,1712]" name="Argentina" pageId="4" pageNumber="1636">Argentina</collectingCountry>
, Entre Ríos Province, same data as
<typeStatus box="[759,859,1688,1712]" pageId="4" pageNumber="1636">holotype</typeStatus>
(six males, three females,
<collectionCode box="[160,237,916,940]" country="Mexico" httpUri="http://grbio.org/cool/654h-avhc" name="Museo de Historia Natural de la Ciudad de Mexico" pageId="5" pageNumber="1637">MCMC</collectionCode>
); Misiones Province, Iguazú NP,
<number box="[605,632,916,940]" pageId="5" pageNumber="1637" value="12.0">12</number>
/
<date box="[642,823,916,940]" pageId="5" pageNumber="1637" value="2000-08-19">
<number box="[642,669,916,940]" pageId="5" pageNumber="1637" value="19.0">19</number>
August
<number box="[768,823,916,940]" pageId="5" pageNumber="1637" value="2000.0">2000</number>
</date>
(one male,
<collectionCode box="[961,1038,916,940]" country="Mexico" httpUri="http://grbio.org/cool/654h-avhc" name="Museo de Historia Natural de la Ciudad de Mexico" pageId="5" pageNumber="1637">MCMC</collectionCode>
); Misiones Province, Iguazú NP, San Martín Island,
<date box="[612,811,951,975]" pageId="5" pageNumber="1637" value="2002-02-15">
<number box="[612,639,951,975]" pageId="5" pageNumber="1637" value="15.0">15</number>
February
<number box="[756,811,951,975]" pageId="5" pageNumber="1637" value="2002.0">2002</number>
</date>
(
<specimenCount box="[830,927,951,975]" pageId="5" pageNumber="1637" type="female">
<number box="[830,841,951,975]" pageId="5" pageNumber="1637" value="1.0">1</number>
female
</specimenCount>
,
<collectionCode box="[940,1018,951,975]" country="Mexico" httpUri="http://grbio.org/cool/654h-avhc" name="Museo de Historia Natural de la Ciudad de Mexico" pageId="5" pageNumber="1637">MCMC</collectionCode>
).
</paragraph>
</subSubSection>
<caption box="[160,1155,142,165]" pageId="5" pageNumber="1637">
<paragraph blockId="5.[160,1157,142,822]" box="[160,1155,142,165]" pageId="5" pageNumber="1637">
<emphasis bold="true" box="[160,242,142,165]" pageId="5" pageNumber="1637">Table 2.</emphasis>
Number and position of secondary setae on the legs of larvae of
<taxonomicName box="[899,1150,142,165]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1637" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[899,1150,142,165]" italics="true" pageId="5" pageNumber="1637">Thermonectus tremouillesi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph blockId="5.[160,1157,142,822]" pageId="5" pageNumber="1637">
<table box="[171,1144,178,741]" gridcols="4" gridrows="22" pageId="5" pageNumber="1637">
<tr box="[171,1144,178,197]" gridrow="0" pageId="5" pageNumber="1637">
<th box="[171,261,178,197]" gridcol="0" gridrow="0" pageId="5" pageNumber="1637">Segment</th>
<th box="[421,487,178,197]" gridcol="1" gridrow="0" pageId="5" pageNumber="1637">Position</th>
<th box="[648,816,178,197]" gridcol="2" gridrow="0" pageId="5" pageNumber="1637">Instar II (n = 1)</th>
<th box="[977,1144,178,197]" gridcol="3" gridrow="0" pageId="5" pageNumber="1637">Instar III (n = 3)</th>
</tr>
<tr box="[171,1144,210,229]" gridrow="1" pageId="5" pageNumber="1637">
<th box="[171,261,210,229]" gridcol="0" gridrow="1" pageId="5" pageNumber="1637">Coxa</th>
<td box="[421,487,210,229]" gridcol="1" gridrow="1" pageId="5" pageNumber="1637">P</td>
<td box="[648,816,210,229]" gridcol="2" gridrow="1" pageId="5" pageNumber="1637">2/45/12</td>
<td box="[977,1144,210,229]" gridcol="3" gridrow="1" pageId="5" pageNumber="1637">916/517/321</td>
</tr>
<tr box="[171,1144,234,253]" gridrow="2" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,234,253]" gridcol="1" gridrow="2" pageId="5" pageNumber="1637">V</td>
<td box="[648,816,234,253]" gridcol="2" gridrow="2" pageId="5" pageNumber="1637">57/45/4</td>
<td box="[977,1144,234,253]" gridcol="3" gridrow="2" pageId="5" pageNumber="1637">1117/720/1122</td>
</tr>
<tr box="[171,1144,258,277]" gridrow="3" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,258,277]" gridcol="1" gridrow="3" pageId="5" pageNumber="1637">Total</td>
<td box="[648,816,258,277]" gridcol="2" gridrow="3" pageId="5" pageNumber="1637">79/810/56</td>
<td box="[977,1144,258,277]" gridcol="3" gridrow="3" pageId="5" pageNumber="1637">2333/1233/2139</td>
</tr>
<tr box="[171,1144,290,309]" gridrow="4" pageId="5" pageNumber="1637">
<th box="[171,261,290,309]" gridcol="0" gridrow="4" pageId="5" pageNumber="1637">Trochanter</th>
<td box="[421,487,290,309]" gridcol="1" gridrow="4" pageId="5" pageNumber="1637">Di</td>
<td box="[648,816,290,309]" gridcol="2" gridrow="4" pageId="5" pageNumber="1637">0/0/01</td>
<td box="[977,1144,290,309]" gridcol="3" gridrow="4" pageId="5" pageNumber="1637">01/01/02</td>
</tr>
<tr box="[171,1144,314,333]" gridrow="5" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,314,333]" gridcol="1" gridrow="5" pageId="5" pageNumber="1637">Pr</td>
<td box="[648,816,314,333]" gridcol="2" gridrow="5" pageId="5" pageNumber="1637">2/2/2</td>
<td box="[977,1144,314,333]" gridcol="3" gridrow="5" pageId="5" pageNumber="1637">49/47/49</td>
</tr>
<tr box="[171,1144,338,357]" gridrow="6" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,338,357]" gridcol="1" gridrow="6" pageId="5" pageNumber="1637">Total</td>
<td box="[648,816,338,357]" gridcol="2" gridrow="6" pageId="5" pageNumber="1637">2/2/23</td>
<td box="[977,1144,338,357]" gridcol="3" gridrow="6" pageId="5" pageNumber="1637">59/57/59</td>
</tr>
<tr box="[171,1144,370,389]" gridrow="7" pageId="5" pageNumber="1637">
<th box="[171,261,370,389]" gridcol="0" gridrow="7" pageId="5" pageNumber="1637">Femur</th>
<td box="[421,487,370,389]" gridcol="1" gridrow="7" pageId="5" pageNumber="1637">A</td>
<td box="[648,816,370,389]" gridcol="2" gridrow="7" pageId="5" pageNumber="1637">14/8/20</td>
<td box="[977,1144,370,389]" gridcol="3" gridrow="7" pageId="5" pageNumber="1637">913/1631/1640</td>
</tr>
<tr box="[171,1144,394,413]" gridrow="8" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,394,413]" gridcol="1" gridrow="8" pageId="5" pageNumber="1637">ADi</td>
<td box="[648,816,394,413]" gridcol="2" gridrow="8" pageId="5" pageNumber="1637">12/23/4</td>
<td box="[977,1144,394,413]" gridcol="3" gridrow="8" pageId="5" pageNumber="1637">12/45/26</td>
</tr>
<tr box="[171,1144,418,437]" gridrow="9" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,418,437]" gridcol="1" gridrow="9" pageId="5" pageNumber="1637">D</td>
<td box="[648,816,418,437]" gridcol="2" gridrow="9" pageId="5" pageNumber="1637">68/67/1012</td>
<td box="[977,1144,418,437]" gridcol="3" gridrow="9" pageId="5" pageNumber="1637">1015/1824/1628</td>
</tr>
<tr box="[171,1144,442,461]" gridrow="10" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,442,461]" gridcol="1" gridrow="10" pageId="5" pageNumber="1637">PDi</td>
<td box="[648,816,442,461]" gridcol="2" gridrow="10" pageId="5" pageNumber="1637">1/1/1</td>
<td box="[977,1144,442,461]" gridcol="3" gridrow="10" pageId="5" pageNumber="1637">1/1/1</td>
</tr>
<tr box="[171,1144,466,485]" gridrow="11" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,466,485]" gridcol="1" gridrow="11" pageId="5" pageNumber="1637">PV</td>
<td box="[648,816,466,485]" gridcol="2" gridrow="11" pageId="5" pageNumber="1637">1617/1820/2224</td>
<td box="[977,1144,466,485]" gridcol="3" gridrow="11" pageId="5" pageNumber="1637">2224/2430/2228</td>
</tr>
<tr box="[171,1144,490,509]" gridrow="12" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,490,509]" gridcol="1" gridrow="12" pageId="5" pageNumber="1637">Total</td>
<td box="[648,816,490,509]" gridcol="2" gridrow="12" pageId="5" pageNumber="1637">2730/37/5761</td>
<td box="[977,1144,490,509]" gridcol="3" gridrow="12" pageId="5" pageNumber="1637">4554/6391/6299</td>
</tr>
<tr box="[171,1144,522,541]" gridrow="13" pageId="5" pageNumber="1637">
<th box="[171,261,522,541]" gridcol="0" gridrow="13" pageId="5" pageNumber="1637">Tibia</th>
<td box="[421,487,522,541]" gridcol="1" gridrow="13" pageId="5" pageNumber="1637">A</td>
<td box="[648,816,522,541]" gridcol="2" gridrow="13" pageId="5" pageNumber="1637">12/810/24</td>
<td box="[977,1144,522,541]" gridcol="3" gridrow="13" pageId="5" pageNumber="1637">69/1335/1439</td>
</tr>
<tr box="[171,1144,546,565]" gridrow="14" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,546,565]" gridcol="1" gridrow="14" pageId="5" pageNumber="1637">ADi</td>
<td box="[648,816,546,565]" gridcol="2" gridrow="14" pageId="5" pageNumber="1637">2/1/2</td>
<td box="[977,1144,546,565]" gridcol="3" gridrow="14" pageId="5" pageNumber="1637">2/14/23</td>
</tr>
<tr box="[171,1144,570,589]" gridrow="15" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,570,589]" gridcol="1" gridrow="15" pageId="5" pageNumber="1637">D</td>
<td box="[648,816,570,589]" gridcol="2" gridrow="15" pageId="5" pageNumber="1637">1112/1112/14</td>
<td box="[977,1144,570,589]" gridcol="3" gridrow="15" pageId="5" pageNumber="1637">1113/1625/1521</td>
</tr>
<tr box="[171,1144,594,613]" gridrow="16" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,594,613]" gridcol="1" gridrow="16" pageId="5" pageNumber="1637">PDi</td>
<td box="[648,816,594,613]" gridcol="2" gridrow="16" pageId="5" pageNumber="1637">1/1/1</td>
<td box="[977,1144,594,613]" gridcol="3" gridrow="16" pageId="5" pageNumber="1637">2/1/1</td>
</tr>
<tr box="[171,1144,618,637]" gridrow="17" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,618,637]" gridcol="1" gridrow="17" pageId="5" pageNumber="1637">PV</td>
<td box="[648,816,618,637]" gridcol="2" gridrow="17" pageId="5" pageNumber="1637">0/0/0</td>
<td box="[977,1144,618,637]" gridcol="3" gridrow="17" pageId="5" pageNumber="1637">69/710/612</td>
</tr>
<tr box="[171,1144,642,661]" gridrow="18" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,642,661]" gridcol="1" gridrow="18" pageId="5" pageNumber="1637">Total</td>
<td box="[648,816,642,661]" gridcol="2" gridrow="18" pageId="5" pageNumber="1637">16/2124/41</td>
<td box="[977,1144,642,661]" gridcol="3" gridrow="18" pageId="5" pageNumber="1637">2735/4073/4376</td>
</tr>
<tr box="[171,1144,674,693]" gridrow="19" pageId="5" pageNumber="1637">
<th box="[171,261,674,693]" gridcol="0" gridrow="19" pageId="5" pageNumber="1637">Tarsus</th>
<td box="[421,487,674,693]" gridcol="1" gridrow="19" pageId="5" pageNumber="1637">D</td>
<td box="[648,816,674,693]" gridcol="2" gridrow="19" pageId="5" pageNumber="1637">78/78/5</td>
<td box="[977,1144,674,693]" gridcol="3" gridrow="19" pageId="5" pageNumber="1637">68/79/8</td>
</tr>
<tr box="[171,1144,698,717]" gridrow="20" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,698,717]" gridcol="1" gridrow="20" pageId="5" pageNumber="1637">V</td>
<td box="[648,816,698,717]" gridcol="2" gridrow="20" pageId="5" pageNumber="1637">89/610/11</td>
<td box="[977,1144,698,717]" gridcol="3" gridrow="20" pageId="5" pageNumber="1637">1012/913/1012</td>
</tr>
<tr box="[171,1144,722,741]" gridrow="21" pageId="5" pageNumber="1637" rowspan-0="1">
<td box="[421,487,722,741]" gridcol="1" gridrow="21" pageId="5" pageNumber="1637">Total</td>
<td box="[648,816,722,741]" gridcol="2" gridrow="21" pageId="5" pageNumber="1637">16/1417/16</td>
<td box="[977,1144,722,741]" gridcol="3" gridrow="21" pageId="5" pageNumber="1637">1719/1622/1820</td>
</tr>
</table>
</paragraph>
<paragraph blockId="5.[160,1157,142,822]" pageId="5" pageNumber="1637">
<tableNote pageId="5" pageNumber="1637" targetBox="[171,1144,178,741]" targetPageId="5">Numbers between slashes refer to pro-, meso- and metathoracic leg, respectively. A: anterior, D: dorsal, Di: distal, P: posterior, Pr: proximal, V: ventral, Total: total number of secondary setae on the segment (excluding primary and natatory setae).</tableNote>
</paragraph>
<subSubSection pageId="5" pageNumber="1637" type="diagnosis">
<paragraph blockId="5.[160,1156,1027,1226]" box="[160,276,1027,1052]" pageId="5" pageNumber="1637">
<heading bold="true" box="[160,276,1027,1052]" fontSize="11" level="2" pageId="5" pageNumber="1637" reason="0">
<emphasis bold="true" box="[160,276,1027,1052]" italics="true" pageId="5" pageNumber="1637">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[160,1156,1027,1226]" pageId="5" pageNumber="1637">
Medium-sized species; colour pattern on head, pronotum and elytra as in
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Figure
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B, D, E; sexual sculpture of female elytra well visible, covering basal fourth to basal half (
<figureCitation box="[167,257,1132,1156]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="5" pageNumber="1637">
Figure
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D); male protibia narrow, dorsal margin slightly prominent (
<figureCitation box="[947,1040,1132,1156]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="5" pageNumber="1637">
Figure
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D); tergite IX of male with four posterior projections (
<figureCitation box="[664,758,1167,1191]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="5" pageNumber="1637">
Figure
<number box="[746,758,1167,1191]" pageId="5" pageNumber="1637" value="2.0">2</number>
</figureCitation>
F); median lobe of aedeagus bifid apically (
<figureCitation box="[261,352,1202,1226]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="5" pageNumber="1637">
Figure
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E); parameres with a terminal spur (
<figureCitation box="[763,855,1202,1226]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="5" pageNumber="1637">
Figure
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E).
</paragraph>
</subSubSection>
<subSubSection lastPageId="9" lastPageNumber="1641" pageId="5" pageNumber="1637" type="description">
<paragraph blockId="5.[160,1155,1278,1476]" box="[160,294,1278,1303]" pageId="5" pageNumber="1637">
<heading bold="true" box="[160,294,1278,1303]" fontSize="11" level="2" pageId="5" pageNumber="1637" reason="0">
<emphasis bold="true" box="[160,294,1278,1303]" italics="true" pageId="5" pageNumber="1637">Description</emphasis>
</heading>
</paragraph>
<paragraph blockId="5.[160,1155,1278,1476]" pageId="5" pageNumber="1637">
<emphasis box="[160,243,1314,1338]" italics="true" pageId="5" pageNumber="1637">Habitus</emphasis>
(
<figureCitation box="[263,356,1314,1338]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="5" pageNumber="1637">
Figure
<number box="[347,356,1314,1338]" pageId="5" pageNumber="1637" value="1.0">1</number>
</figureCitation>
B, C). Oval, greatest width at posterior third, moderately narrowed anteriorly, regularly curved on margins, dorsal surface markedly convex; females somewhat larger than males; male: TL =
<quantity box="[560,718,1383,1407]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" metricValueMax="1.26" metricValueMin="1.18" pageId="5" pageNumber="1637" unit="mm" value="12.2" valueMax="12.6" valueMin="11.8">
11.8
<emphasis bold="true" box="[606,619,1383,1407]" italics="true" pageId="5" pageNumber="1637"></emphasis>
12.6 mm
</quantity>
(mean =
<quantity box="[828,929,1383,1407]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="5" pageNumber="1637" unit="mm" value="12.1">12.1 mm</quantity>
); MW =
<quantity box="[1024,1155,1383,1407]" metricMagnitude="-3" metricUnit="m" metricValue="7.05" metricValueMax="7.4" metricValueMin="6.7" pageId="5" pageNumber="1637" unit="mm" value="7.05" valueMax="7.4" valueMin="6.7">
6.7
<emphasis bold="true" box="[1057,1070,1383,1407]" italics="true" pageId="5" pageNumber="1637"></emphasis>
7.4 mm
</quantity>
(mean =
<quantity box="[264,350,1418,1442]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="5" pageNumber="1637" unit="mm" value="7.2">7.2 mm</quantity>
) (N =
<number box="[424,451,1418,1442]" pageId="5" pageNumber="1637" value="18.0">18</number>
) (
<typeStatus box="[475,577,1418,1442]" pageId="5" pageNumber="1637">holotype</typeStatus>
: TL =
<quantity box="[648,749,1418,1442]" metricMagnitude="-2" metricUnit="m" metricValue="1.24" pageId="5" pageNumber="1637" unit="mm" value="12.4">12.4 mm</quantity>
; MW =
<quantity box="[839,926,1418,1442]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="5" pageNumber="1637" unit="mm" value="7.3">7.3 mm</quantity>
); female: TL =
<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.27" metricValueMax="1.3199999999999998" metricValueMin="1.22" pageId="5" pageNumber="1637" unit="mm" value="12.7" valueMax="13.2" valueMin="12.2">
12.2
<emphasis bold="true" box="[1142,1155,1417,1441]" italics="true" pageId="5" pageNumber="1637"></emphasis>
13.2 mm
</quantity>
(mean =
<quantity box="[373,474,1452,1476]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="5" pageNumber="1637" unit="mm" value="12.5">12.5 mm</quantity>
); MW = 7.0
<emphasis bold="true" box="[605,618,1452,1476]" italics="true" pageId="5" pageNumber="1637"></emphasis>
<quantity box="[618,704,1452,1476]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="5" pageNumber="1637" unit="mm" value="7.7">7.7 mm</quantity>
(mean =
<quantity box="[818,904,1452,1476]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="5" pageNumber="1637" unit="mm" value="7.3">7.3 mm</quantity>
) (N =
<number box="[979,994,1452,1476]" pageId="5" pageNumber="1637" value="6.0">6</number>
).
</paragraph>
<paragraph blockId="5.[160,1155,1529,1727]" lastBlockId="8.[160,1156,142,651]" lastPageId="8" lastPageNumber="1640" pageId="5" pageNumber="1637">
<emphasis bold="true" box="[160,244,1529,1554]" italics="true" pageId="5" pageNumber="1637">Colour.</emphasis>
Dorsal surface (
<figureCitation box="[440,530,1530,1554]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="5" pageNumber="1637">
Figure
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B, D, E): yellowish to orangish background with black markings as follows: head with postocular band all along posterior margin, projected forward on internal ocular margin to approximately half of eye length, and isolated V-shaped macula in middle of interocular space (
<figureCitation box="[759,853,1634,1658]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="5" pageNumber="1637">
Figure
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E), sometimes coalescent with ocular markings (
<figureCitation box="[417,506,1669,1693]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="5" pageNumber="1637">
Figure
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</figureCitation>
B) [
<typeStatus box="[549,654,1669,1693]" pageId="5" pageNumber="1637">holotype</typeStatus>
: V-shaped macula isolated]; pronotum with transverse band on anterior margin, its width slightly greater than head width, continued posterolaterally in short projections (
<figureCitation box="[721,813,142,166]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
B) that sometimes appear as isolated maculae (
<figureCitation box="[376,468,177,201]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
E) [
<typeStatus box="[512,618,177,201]" pageId="8" pageNumber="1640">holotype</typeStatus>
: anterior band and posterolateral projections connected], and transverse band on posterior margin, a bit narrower and about twice thicker than anterior band, with anterior border indented medially and laterals irregularly rounded and continued to pronotal edge as a fine marginal band; elytra irrorate, irroration absent or reduced on humeral angles, external and internal margins, and two submarginal areas, one just posterior to mid-length, the other subapical; irroration interrupted by an irregular transverse black fascia on distal fourth in some specimens (
<figureCitation box="[384,482,420,444]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
D), in other specimens only an irregular submarginal macula is evident on each elytron (
<figureCitation box="[576,667,454,478]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
B) [
<typeStatus box="[710,815,454,478]" pageId="8" pageNumber="1640">holotype</typeStatus>
: transverse fascia well developed]; internal margin of elytra with a fine black band from scutellum to apex. Ventral surface (
<figureCitation box="[258,347,524,548]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
C): ferrugineous to piceus black except antennae, maxillary and labial palpi, pro- and mesolegs, ventral surface of pronotum, and epipleura yellowish, and base of prosternal process, metatrochantera and distal portion of metafemora orangish.
</paragraph>
<caption httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="6" pageNumber="1638" targetBox="[9,1150,142,1552]" targetPageId="6">
<paragraph blockId="6.[160,1157,1584,1695]" pageId="6" pageNumber="1638">
<emphasis bold="true" box="[160,251,1584,1607]" pageId="6" pageNumber="1638">Figure 1.</emphasis>
Photographs of adults of
<taxonomicName box="[514,648,1584,1607]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1638" phylum="Arthropoda" rank="genus">
<emphasis box="[514,648,1584,1607]" italics="true" pageId="6" pageNumber="1638">Thermonectus</emphasis>
</taxonomicName>
. (A)
<taxonomicName box="[696,779,1584,1607]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1638" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[696,779,1584,1607]" italics="true" pageId="6" pageNumber="1638">T. alfredi</emphasis>
</taxonomicName>
, male, habitus, dorsal aspect; (BE)
<taxonomicName class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="1638" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis italics="true" pageId="6" pageNumber="1638">T. tremouillesi</emphasis>
</taxonomicName>
: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect.
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="7" pageNumber="1639" targetBox="[9,1152,142,1547]" targetPageId="7">
<paragraph blockId="7.[160,1156,1579,1691]" pageId="7" pageNumber="1639">
<emphasis bold="true" box="[160,254,1580,1603]" pageId="7" pageNumber="1639">Figure 2.</emphasis>
SEM images of male adults of
<taxonomicName box="[586,720,1580,1603]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1639" phylum="Arthropoda" rank="genus">
<emphasis box="[586,720,1580,1603]" italics="true" pageId="7" pageNumber="1639">Thermonectus</emphasis>
</taxonomicName>
. (AF)
<taxonomicName box="[798,935,1580,1603]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1639" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[798,935,1580,1603]" italics="true" pageId="7" pageNumber="1639">T. tremouillesi</emphasis>
</taxonomicName>
: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G)
<taxonomicName box="[653,737,1668,1691]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="1639" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[653,737,1668,1691]" italics="true" pageId="7" pageNumber="1639">T. alfredi</emphasis>
</taxonomicName>
, tergite IX, ventral aspect.
</paragraph>
</caption>
<paragraph blockId="8.[160,1155,696,894]" pageId="8" pageNumber="1640">
<emphasis bold="true" box="[160,468,696,721]" italics="true" pageId="8" pageNumber="1640">Sculpture and punctation.</emphasis>
Smooth, shiny; microreticulation minute, of regular meshes; head, pronotum and to somewhat lesser degree elytra with very fine, closely distributed punctures combined with larger, sparsely distributed punctures (
<figureCitation box="[898,987,766,790]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
D, SEM detail); each elytron with four longitudinal rows of widely spaced setiferous punctures, one contiguous to elytral commissure, the other three on disc; ventral side with fine, irregularly distributed punctation.
</paragraph>
<paragraph blockId="8.[160,1156,939,1137]" pageId="8" pageNumber="1640">
<emphasis bold="true" box="[160,274,939,964]" italics="true" pageId="8" pageNumber="1640">Structure.</emphasis>
Head deeply set into pronotum, ratio head length/maximum head width about
<number box="[237,271,975,999]" pageId="8" pageNumber="1640" value="0.4">0.4</number>
; pronotum deeply notched to hold head, ratio length of pronotum at midpoint/greatest width of pronotum about
<number box="[635,646,1009,1033]" pageId="8" pageNumber="1640" value="1.0">1</number>
/
<number box="[657,672,1009,1033]" pageId="8" pageNumber="1640" value="3.0">3</number>
, ratio width on anterior margin of pronotum/width on posterior margin of pronotum about
<number box="[761,795,1044,1068]" pageId="8" pageNumber="1640" value="0.6">0.6</number>
; prosternal process moderately widened on distal third, rounded apically (
<figureCitation box="[682,780,1079,1103]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
C); posterior apex of scutellum rounded; elytra widely rounded on distal third (
<figureCitation box="[706,795,1113,1137]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
B, D).
</paragraph>
<paragraph blockId="8.[160,1155,1182,1380]" pageId="8" pageNumber="1640">
<emphasis bold="true" box="[160,224,1182,1207]" italics="true" pageId="8" pageNumber="1640">Male.</emphasis>
Protibia narrow, dorsal margin slightly prominent (
<figureCitation box="[856,955,1183,1207]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
D); tarsal palette bearing
<number box="[260,287,1217,1241]" pageId="8" pageNumber="1640" value="18.0">18</number>
(occasionally
<number box="[459,486,1217,1241]" pageId="8" pageNumber="1640" value="19.0">19</number>
) sucker-like setae, three of the basal ones a bit larger (among these, one about
<number box="[460,488,1252,1276]" pageId="8" pageNumber="1640" value="20.0">20</number>
% larger than the other two) and two of the distal ones a bit smaller than the others (
<figureCitation box="[516,614,1287,1311]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="8" pageNumber="1640">
Figure
<number box="[602,614,1287,1311]" pageId="8" pageNumber="1640" value="2.0">2</number>
</figureCitation>
B); tergite IX with four posterior projections (
<figureCitation box="[167,260,1321,1345]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="8" pageNumber="1640">
Figure
<number box="[248,260,1321,1345]" pageId="8" pageNumber="1640" value="2.0">2</number>
</figureCitation>
F); median lobe of aedeagus bifid apically (
<figureCitation box="[758,850,1321,1345]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="8" pageNumber="1640">
Figure
<number box="[838,850,1321,1345]" pageId="8" pageNumber="1640" value="2.0">2</number>
</figureCitation>
E); parameres symmetrical, with a minute, partly concealed apical spur (
<figureCitation box="[670,762,1356,1380]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
E, detail).
</paragraph>
<paragraph blockId="8.[160,1156,1425,1484]" pageId="8" pageNumber="1640">
<emphasis bold="true" box="[160,252,1425,1450]" italics="true" pageId="8" pageNumber="1640">Female.</emphasis>
Protibia not distinctly modified, dorsal margin not prominent; elytra bearing well-visible, closely spaced longitudinal sculpture on basal fourth to basal half (
<figureCitation box="[1035,1120,1460,1484]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
D).
</paragraph>
<paragraph blockId="8.[160,1155,1529,1727]" box="[160,270,1529,1554]" pageId="8" pageNumber="1640">
<heading bold="true" box="[160,270,1529,1554]" fontSize="11" level="2" pageId="8" pageNumber="1640" reason="0">
<emphasis bold="true" box="[160,270,1529,1554]" italics="true" pageId="8" pageNumber="1640">Variation</emphasis>
</heading>
</paragraph>
<paragraph blockId="8.[160,1155,1529,1727]" lastBlockId="9.[160,1155,142,270]" lastPageId="9" lastPageNumber="1641" pageId="8" pageNumber="1640">
The V-shaped macula of head may be more or less developed; in extreme cases it reaches the ocular markings (
<figureCitation box="[533,631,1599,1623]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
B, E). The pronotal central maculae show different degree of development, from small and isolated (
<figureCitation box="[869,963,1634,1658]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="8" pageNumber="1640">
Figure
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</figureCitation>
E) to somewhat larger and widely connected with the anterior transverse band of the pronotum. The transverse black fascia on distal fourth of the elytra varies greatly in extent; in most specimens it is well developed (
<figureCitation box="[537,628,142,166]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="9" pageNumber="1641">
Figure
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</figureCitation>
D), but in some specimens it is reduced and appears as an irregular submarginal macula on each elytron (
<figureCitation box="[890,982,177,201]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="9" pageNumber="1641">
Figure
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</figureCitation>
B). The female sexual sculpture varies in extent, covering basal fourth (
<figureCitation box="[836,929,212,236]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="9" pageNumber="1641">
Figure
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D) to basal half of elytra.
</paragraph>
<paragraph blockId="9.[160,1156,315,1449]" box="[160,240,315,340]" pageId="9" pageNumber="1641">
<heading bold="true" box="[160,240,315,340]" fontSize="11" level="2" pageId="9" pageNumber="1641" reason="0">
<emphasis bold="true" box="[160,240,315,340]" italics="true" pageId="9" pageNumber="1641">Larvae</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="1644" pageId="9" pageNumber="1641" type="diagnosis">
<paragraph blockId="9.[160,1156,315,1449]" pageId="9" pageNumber="1641">
<emphasis bold="true" box="[160,282,350,375]" italics="true" pageId="9" pageNumber="1641">Diagnosis.</emphasis>
Instar I (
<figureCitation box="[394,496,351,375]" captionStart="Figure 3" captionStartId="10.[160,225,1592,1615]" captionTargetBox="[9,1152,142,1560]" captionTargetId="figure@10.[163,1153,142,1561]" captionTargetPageId="10" captionText="Figure 3. Line drawings of instar I larvae of Thermonectus tremouillesi. (A) Cephalic capsule, dorsal aspect; (B) cephalic capsule, ventral aspect; (C) left antenna, dorsal aspect; (D) right antenna, ventral aspect; (E) left maxilla, dorsal aspect; (F) right maxilla, ventral aspect; (G) left mandible, dorsal aspect; (H) labium, dorsal aspect; (I) labium, ventral aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AN: antenna; EB: egg burster; FR: frontoclypeus; LA: labium; MN: mandible; MX: maxilla; PA: parietal; Sp: spinula; TP: tentorial pit." httpUri="https://zenodo.org/record/269897/files/figure.png" pageId="9" pageNumber="1641">
Figures
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A
<emphasis bold="true" box="[515,528,351,375]" italics="true" pageId="9" pageNumber="1641"></emphasis>
I,
<number box="[550,562,351,375]" pageId="9" pageNumber="1641" value="4.0">4</number>
A
<emphasis bold="true" box="[580,593,351,375]" italics="true" pageId="9" pageNumber="1641"></emphasis>
E): head length:
<quantity box="[783,885,351,375]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="9" pageNumber="1641" unit="mm" value="1.4">1.40 mm</quantity>
; nasale rounded, markedly projected forward, bearing about
<number box="[620,702,385,409]" pageId="9" pageNumber="1641" value="90.0" valueMax="100.0" valueMin="80.0">
80
<emphasis bold="true" box="[647,660,385,409]" italics="true" pageId="9" pageNumber="1641"></emphasis>
100
</number>
lamellae clypeales on anterior margin (
<figureCitation box="[167,256,420,444]" captionStart="Figure 3" captionStartId="10.[160,225,1592,1615]" captionTargetBox="[9,1152,142,1560]" captionTargetId="figure@10.[163,1153,142,1561]" captionTargetPageId="10" captionText="Figure 3. Line drawings of instar I larvae of Thermonectus tremouillesi. (A) Cephalic capsule, dorsal aspect; (B) cephalic capsule, ventral aspect; (C) left antenna, dorsal aspect; (D) right antenna, ventral aspect; (E) left maxilla, dorsal aspect; (F) right maxilla, ventral aspect; (G) left mandible, dorsal aspect; (H) labium, dorsal aspect; (I) labium, ventral aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AN: antenna; EB: egg burster; FR: frontoclypeus; LA: labium; MN: mandible; MX: maxilla; PA: parietal; Sp: spinula; TP: tentorial pit." httpUri="https://zenodo.org/record/269897/files/figure.png" pageId="9" pageNumber="1641">
Figure
<number box="[245,256,420,444]" pageId="9" pageNumber="1641" value="3.0">3</number>
</figureCitation>
A); ratio length of maxillary palpus/length of labial palpus:
<number box="[915,1023,420,444]" pageId="9" pageNumber="1641" value="0.885" valueMax="0.89" valueMin="0.88">
0.88
<emphasis bold="true" box="[962,975,420,444]" italics="true" pageId="9" pageNumber="1641"></emphasis>
0.89
</number>
; stipes with a row of
<number box="[259,327,454,479]" pageId="9" pageNumber="1641" value="17.0" valueMax="19.0" valueMin="15.0">
15
<emphasis bold="true" box="[287,300,454,478]" italics="true" pageId="9" pageNumber="1641"></emphasis>
19
</number>
additional spine-like setae on dorsal surface (
<figureCitation box="[840,929,455,479]" captionStart="Figure 3" captionStartId="10.[160,225,1592,1615]" captionTargetBox="[9,1152,142,1560]" captionTargetId="figure@10.[163,1153,142,1561]" captionTargetPageId="10" captionText="Figure 3. Line drawings of instar I larvae of Thermonectus tremouillesi. (A) Cephalic capsule, dorsal aspect; (B) cephalic capsule, ventral aspect; (C) left antenna, dorsal aspect; (D) right antenna, ventral aspect; (E) left maxilla, dorsal aspect; (F) right maxilla, ventral aspect; (G) left mandible, dorsal aspect; (H) labium, dorsal aspect; (I) labium, ventral aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AN: antenna; EB: egg burster; FR: frontoclypeus; LA: labium; MN: mandible; MX: maxilla; PA: parietal; Sp: spinula; TP: tentorial pit." httpUri="https://zenodo.org/record/269897/files/figure.png" pageId="9" pageNumber="1641">
Figure
<number box="[918,929,455,479]" pageId="9" pageNumber="1641" value="3.0">3</number>
</figureCitation>
E); prementum with three additional spine-like setae on each side of anterodorsal surface (
<figureCitation box="[970,1061,489,513]" captionStart="Figure 3" captionStartId="10.[160,225,1592,1615]" captionTargetBox="[9,1152,142,1560]" captionTargetId="figure@10.[163,1153,142,1561]" captionTargetPageId="10" captionText="Figure 3. Line drawings of instar I larvae of Thermonectus tremouillesi. (A) Cephalic capsule, dorsal aspect; (B) cephalic capsule, ventral aspect; (C) left antenna, dorsal aspect; (D) right antenna, ventral aspect; (E) left maxilla, dorsal aspect; (F) right maxilla, ventral aspect; (G) left mandible, dorsal aspect; (H) labium, dorsal aspect; (I) labium, ventral aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AN: antenna; EB: egg burster; FR: frontoclypeus; LA: labium; MN: mandible; MX: maxilla; PA: parietal; Sp: spinula; TP: tentorial pit." httpUri="https://zenodo.org/record/269897/files/figure.png" pageId="9" pageNumber="1641">
Figure
<number box="[1050,1061,489,513]" pageId="9" pageNumber="1641" value="3.0">3</number>
</figureCitation>
H); ratio length of metathoracic leg/head width:
<number box="[616,664,524,548]" pageId="9" pageNumber="1641" value="2.59">2.59</number>
; pro-, meso- and metafemur with one,
<number box="[1115,1155,524,548]" pageId="9" pageNumber="1641" value="1.5" valueMax="2.0" valueMin="1.0">
1
<emphasis bold="true" box="[1128,1141,524,548]" italics="true" pageId="9" pageNumber="1641"></emphasis>
2
</number>
and
<number box="[211,252,558,583]" pageId="9" pageNumber="1641" value="2.5" valueMax="3.0" valueMin="2.0">
2
<emphasis bold="true" box="[224,237,558,582]" italics="true" pageId="9" pageNumber="1641"></emphasis>
3
</number>
additional anterodistal setae respectively (
<figureCitation box="[738,829,559,583]" captionStart="Figure 4" captionStartId="11.[160,225,1587,1610]" captionTargetBox="[9,1152,142,1555]" captionTargetId="figure@11.[163,1153,142,1556]" captionTargetPageId="11" captionText="Figure 4. Line drawings of larvae of Thermonectus tremouillesi. (A) Left prothoracic leg of instar I, anterior aspect; (B) right prothoracic leg of instar I, posterior aspect; (C) abdominal segment VIII of instar I, dorsal aspect; (D) abdominal segment VIII of instar I, ventral aspect; (E) right urogomphus of instar I, dorsal aspect; (F) head of instar III, dorsal aspect, showing colour pattern on cephalic capsule (chaetotaxy omitted); (G) labium of instar III, dorsal aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AB: abdominal segment VIII; CO: coxa; FE: femur; PT: pretarsus; TA: tarsus; TI: tibia; TR: trochanter; UR: urogomphus." httpUri="https://zenodo.org/record/269898/files/figure.png" pageId="9" pageNumber="1641">
Figure
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A); additional measurements and ratios that characterise the body shape are shown in
<tableCitation box="[824,907,593,617]" captionStart="Table 1" captionStartId="4.[240,296,142,165]" captionTargetBox="[250,1064,178,1069]" captionTargetPageId="4" captionText="Table 1. Measurements and ratios for the larvae of Thermonectus tremouillesi." httpUri="http://table.plazi.org/id/81937AFBFFA0FFB404D3A9C823885781" pageId="9" pageNumber="1641" tableUuid="81937AFBFFA0FFB404D3A9C823885781">
Table
<number box="[893,907,593,617]" pageId="9" pageNumber="1641" value="1.0">1</number>
</tableCitation>
.
</paragraph>
<paragraph blockId="9.[160,1156,315,1449]" pageId="9" pageNumber="1641">
Instar II: head length:
<quantity box="[439,540,628,652]" metricMagnitude="-3" metricUnit="m" metricValue="1.88" pageId="9" pageNumber="1641" unit="mm" value="1.88">1.88 mm</quantity>
; nasale rounded, markedly projected forward, bearing about
<number box="[234,329,662,687]" pageId="9" pageNumber="1641" value="120.0" valueMax="130.0" valueMin="110.0">
110
<emphasis bold="true" box="[275,288,662,686]" italics="true" pageId="9" pageNumber="1641"></emphasis>
130
</number>
lamellae clypeales on anterior margin; parietal with
<number box="[920,947,663,687]" pageId="9" pageNumber="1641" value="11.0">11</number>
temporal and
<number box="[1115,1155,662,687]" pageId="9" pageNumber="1641" value="8.5" valueMax="9.0" valueMin="8.0">
8
<emphasis bold="true" box="[1128,1141,662,686]" italics="true" pageId="9" pageNumber="1641"></emphasis>
9
</number>
ventral spine-like setae on each side; antennomere
<number box="[744,758,697,721]" pageId="9" pageNumber="1641" value="1.0">1</number>
with
<number box="[823,864,697,721]" pageId="9" pageNumber="1641" value="6.0" valueMax="7.0" valueMin="5.0">
5
<emphasis bold="true" box="[836,849,697,721]" italics="true" pageId="9" pageNumber="1641"></emphasis>
7
</number>
secondary hair-like setae; mandible with a row of
<number box="[444,512,732,756]" pageId="9" pageNumber="1641" value="31.0" valueMax="32.0" valueMin="30.0">
30
<emphasis bold="true" box="[472,485,732,756]" italics="true" pageId="9" pageNumber="1641"></emphasis>
32
</number>
secondary hair-like setae on basoexternal margin; ratio length of maxillary palpus/length of labial palpus:
<number box="[745,793,767,791]" pageId="9" pageNumber="1641" value="0.88">0.88</number>
; stipes with a row of
<number box="[1050,1077,767,791]" pageId="9" pageNumber="1641" value="28.0">28</number>
dorsal secondary spine-like setae and
<number box="[516,584,801,825]" pageId="9" pageNumber="1641" value="17.0" valueMax="18.0" valueMin="16.0">
16
<emphasis bold="true" box="[544,557,801,825]" italics="true" pageId="9" pageNumber="1641"></emphasis>
18
</number>
external and
<number box="[743,810,801,825]" pageId="9" pageNumber="1641" value="37.5" valueMax="40.0" valueMin="35.0">
35
<emphasis bold="true" box="[770,783,801,825]" italics="true" pageId="9" pageNumber="1641"></emphasis>
40
</number>
dorsoexternal secondary hairlike setae; prementum with
<number box="[474,528,836,860]" pageId="9" pageNumber="1641" value="9.5" valueMax="10.0" valueMin="9.0">
9
<emphasis bold="true" box="[487,500,836,860]" italics="true" pageId="9" pageNumber="1641"></emphasis>
10
</number>
secondary spine-like setae on each side of anterodorsal surface, bearing dorsal spinulae on basal half and on margins; ratio length of metathoracic leg/head width:
<number box="[397,445,905,929]" pageId="9" pageNumber="1641" value="2.58">2.58</number>
; posterior surface of coxa with one secondary pore; additional measurements and ratios that characterise the body shape are shown in
<tableCitation box="[1061,1150,940,964]" captionStart="Table 1" captionStartId="4.[240,296,142,165]" captionTargetBox="[250,1064,178,1069]" captionTargetPageId="4" captionText="Table 1. Measurements and ratios for the larvae of Thermonectus tremouillesi." httpUri="http://table.plazi.org/id/81937AFBFFA0FFB404D3A9C823885781" pageId="9" pageNumber="1641" tableUuid="81937AFBFFA0FFB404D3A9C823885781">
Table
<number box="[1136,1150,940,964]" pageId="9" pageNumber="1641" value="1.0">1</number>
</tableCitation>
; detailed secondary leg setation is given in
<tableCitation box="[652,735,975,999]" captionStart="Table 2" captionStartId="5.[160,216,142,165]" captionText="Table 2. Number and position of secondary setae on the legs of larvae of Thermonectus tremouillesi." pageId="9" pageNumber="1641">
Table
<number box="[721,735,975,999]" pageId="9" pageNumber="1641" value="2.0">2</number>
</tableCitation>
.
</paragraph>
<paragraph blockId="9.[160,1156,315,1449]" pageId="9" pageNumber="1641">
Instar III (
<figureCitation box="[293,382,1009,1033]" captionStart="Figure 4" captionStartId="11.[160,225,1587,1610]" captionTargetBox="[9,1152,142,1555]" captionTargetId="figure@11.[163,1153,142,1556]" captionTargetPageId="11" captionText="Figure 4. Line drawings of larvae of Thermonectus tremouillesi. (A) Left prothoracic leg of instar I, anterior aspect; (B) right prothoracic leg of instar I, posterior aspect; (C) abdominal segment VIII of instar I, dorsal aspect; (D) abdominal segment VIII of instar I, ventral aspect; (E) right urogomphus of instar I, dorsal aspect; (F) head of instar III, dorsal aspect, showing colour pattern on cephalic capsule (chaetotaxy omitted); (G) labium of instar III, dorsal aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AB: abdominal segment VIII; CO: coxa; FE: femur; PT: pretarsus; TA: tarsus; TI: tibia; TR: trochanter; UR: urogomphus." httpUri="https://zenodo.org/record/269898/files/figure.png" pageId="9" pageNumber="1641">
Figure
<number box="[370,382,1009,1033]" pageId="9" pageNumber="1641" value="4.0">4</number>
</figureCitation>
F
<emphasis bold="true" box="[396,409,1009,1033]" italics="true" pageId="9" pageNumber="1641"></emphasis>
G): head length:
<quantity box="[593,747,1009,1033]" metricMagnitude="-3" metricUnit="m" metricValue="2.765" metricValueMax="2.83" metricValueMin="2.7" pageId="9" pageNumber="1641" unit="mm" value="2.765" valueMax="2.83" valueMin="2.7">
2.70
<emphasis bold="true" box="[637,650,1009,1033]" italics="true" pageId="9" pageNumber="1641"></emphasis>
2.83 mm
</quantity>
; nasale rounded, markedly projected forward, bearing about
<number box="[415,507,1044,1068]" pageId="9" pageNumber="1641" value="200.0" valueMax="210.0" valueMin="190.0">
190
<emphasis bold="true" box="[455,468,1044,1068]" italics="true" pageId="9" pageNumber="1641"></emphasis>
210
</number>
lamellae clypeales on anterior margin (
<figureCitation box="[931,1018,1044,1068]" captionStart="Figure 4" captionStartId="11.[160,225,1587,1610]" captionTargetBox="[9,1152,142,1555]" captionTargetId="figure@11.[163,1153,142,1556]" captionTargetPageId="11" captionText="Figure 4. Line drawings of larvae of Thermonectus tremouillesi. (A) Left prothoracic leg of instar I, anterior aspect; (B) right prothoracic leg of instar I, posterior aspect; (C) abdominal segment VIII of instar I, dorsal aspect; (D) abdominal segment VIII of instar I, ventral aspect; (E) right urogomphus of instar I, dorsal aspect; (F) head of instar III, dorsal aspect, showing colour pattern on cephalic capsule (chaetotaxy omitted); (G) labium of instar III, dorsal aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AB: abdominal segment VIII; CO: coxa; FE: femur; PT: pretarsus; TA: tarsus; TI: tibia; TR: trochanter; UR: urogomphus." httpUri="https://zenodo.org/record/269898/files/figure.png" pageId="9" pageNumber="1641">
Figure
<number box="[1006,1018,1044,1068]" pageId="9" pageNumber="1641" value="4.0">4</number>
</figureCitation>
F); anterolateral lobes of frontoclypeus not projecting beyond nasale (
<figureCitation box="[814,904,1079,1103]" captionStart="Figure 4" captionStartId="11.[160,225,1587,1610]" captionTargetBox="[9,1152,142,1555]" captionTargetId="figure@11.[163,1153,142,1556]" captionTargetPageId="11" captionText="Figure 4. Line drawings of larvae of Thermonectus tremouillesi. (A) Left prothoracic leg of instar I, anterior aspect; (B) right prothoracic leg of instar I, posterior aspect; (C) abdominal segment VIII of instar I, dorsal aspect; (D) abdominal segment VIII of instar I, ventral aspect; (E) right urogomphus of instar I, dorsal aspect; (F) head of instar III, dorsal aspect, showing colour pattern on cephalic capsule (chaetotaxy omitted); (G) labium of instar III, dorsal aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AB: abdominal segment VIII; CO: coxa; FE: femur; PT: pretarsus; TA: tarsus; TI: tibia; TR: trochanter; UR: urogomphus." httpUri="https://zenodo.org/record/269898/files/figure.png" pageId="9" pageNumber="1641">
Figure
<number box="[892,904,1079,1103]" pageId="9" pageNumber="1641" value="4.0">4</number>
</figureCitation>
F); parietal with
<number box="[1089,1154,1078,1103]" pageId="9" pageNumber="1641" value="12.0" valueMax="13.0" valueMin="11.0">
11
<emphasis bold="true" box="[1116,1129,1078,1102]" italics="true" pageId="9" pageNumber="1641"></emphasis>
13
</number>
temporal and
<number box="[311,363,1113,1137]" pageId="9" pageNumber="1641" value="12.0" valueMax="15.0" valueMin="9.0">
9
<emphasis bold="true" box="[324,337,1113,1137]" italics="true" pageId="9" pageNumber="1641"></emphasis>
15
</number>
ventral spine-like setae on each side; antennomere
<number box="[915,928,1113,1137]" pageId="9" pageNumber="1641" value="1.0">1</number>
with
<number box="[987,1039,1113,1137]" pageId="9" pageNumber="1641" value="9.5" valueMax="11.0" valueMin="8.0">
8
<emphasis bold="true" box="[1000,1013,1113,1137]" italics="true" pageId="9" pageNumber="1641"></emphasis>
11
</number>
secondary hair-like setae; mandible with a row of
<number box="[613,678,1148,1172]" pageId="9" pageNumber="1641" value="33.0" valueMax="38.0" valueMin="28.0">
28
<emphasis bold="true" box="[640,653,1148,1172]" italics="true" pageId="9" pageNumber="1641"></emphasis>
38
</number>
secondary hair-like setae on basoexternal margin, and several minute secondary setae on external margin; ratio length of maxillary palpus/length of labial palpus:
<number box="[493,597,1217,1241]" pageId="9" pageNumber="1641" value="0.845" valueMax="0.87" valueMin="0.82">
0.82
<emphasis bold="true" box="[538,551,1217,1241]" italics="true" pageId="9" pageNumber="1641"></emphasis>
0.87
</number>
; stipes with a row of
<number box="[828,893,1217,1241]" pageId="9" pageNumber="1641" value="30.0" valueMax="35.0" valueMin="25.0">
25
<emphasis bold="true" box="[854,867,1217,1241]" italics="true" pageId="9" pageNumber="1641"></emphasis>
35
</number>
dorsal secondary spinelike setae,
<number box="[276,342,1252,1276]" pageId="9" pageNumber="1641" value="18.5" valueMax="20.0" valueMin="17.0">
17
<emphasis bold="true" box="[303,316,1252,1276]" italics="true" pageId="9" pageNumber="1641"></emphasis>
20
</number>
external and
<number box="[498,563,1252,1276]" pageId="9" pageNumber="1641" value="41.0" valueMax="50.0" valueMin="32.0">
32
<emphasis bold="true" box="[524,537,1252,1276]" italics="true" pageId="9" pageNumber="1641"></emphasis>
50
</number>
dorsoexternal secondary hair-like setae, and few secondary, minute, ventroexternal setae; prementum with
<number box="[762,828,1286,1311]" pageId="9" pageNumber="1641" value="12.0" valueMax="14.0" valueMin="10.0">
10
<emphasis bold="true" box="[789,802,1286,1310]" italics="true" pageId="9" pageNumber="1641"></emphasis>
14
</number>
secondary spine-like setae on each side of anterodorsal surface, bearing dorsal spinulae on basal half and on margins (
<figureCitation box="[167,259,1356,1380]" captionStart="Figure 4" captionStartId="11.[160,225,1587,1610]" captionTargetBox="[9,1152,142,1555]" captionTargetId="figure@11.[163,1153,142,1556]" captionTargetPageId="11" captionText="Figure 4. Line drawings of larvae of Thermonectus tremouillesi. (A) Left prothoracic leg of instar I, anterior aspect; (B) right prothoracic leg of instar I, posterior aspect; (C) abdominal segment VIII of instar I, dorsal aspect; (D) abdominal segment VIII of instar I, ventral aspect; (E) right urogomphus of instar I, dorsal aspect; (F) head of instar III, dorsal aspect, showing colour pattern on cephalic capsule (chaetotaxy omitted); (G) labium of instar III, dorsal aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AB: abdominal segment VIII; CO: coxa; FE: femur; PT: pretarsus; TA: tarsus; TI: tibia; TR: trochanter; UR: urogomphus." httpUri="https://zenodo.org/record/269898/files/figure.png" pageId="9" pageNumber="1641">
Figure
<number box="[247,259,1356,1380]" pageId="9" pageNumber="1641" value="4.0">4</number>
</figureCitation>
G); ratio length of metathoracic leg/head width:
<number box="[818,921,1356,1380]" pageId="9" pageNumber="1641" value="2.75" valueMax="2.82" valueMin="2.68">
2.68
<emphasis bold="true" box="[862,875,1356,1380]" italics="true" pageId="9" pageNumber="1641"></emphasis>
2.82
</number>
; posterior surface of coxa with
<number box="[277,316,1390,1415]" pageId="9" pageNumber="1641" value="2.0" valueMax="3.0" valueMin="1.0">
1
<emphasis bold="true" box="[290,303,1390,1414]" italics="true" pageId="9" pageNumber="1641"></emphasis>
3
</number>
secondary pores; additional measurements and ratios that characterise the body shape are shown in
<tableCitation box="[440,519,1425,1449]" captionStart="Table 1" captionStartId="4.[240,296,142,165]" captionTargetBox="[250,1064,178,1069]" captionTargetPageId="4" captionText="Table 1. Measurements and ratios for the larvae of Thermonectus tremouillesi." httpUri="http://table.plazi.org/id/81937AFBFFA0FFB404D3A9C823885781" pageId="9" pageNumber="1641" tableUuid="81937AFBFFA0FFB404D3A9C823885781">
Table
<number box="[506,519,1425,1449]" pageId="9" pageNumber="1641" value="1.0">1</number>
</tableCitation>
; detailed secondary leg setation is given in
<tableCitation box="[992,1071,1425,1449]" captionStart="Table 2" captionStartId="5.[160,216,142,165]" captionText="Table 2. Number and position of secondary setae on the legs of larvae of Thermonectus tremouillesi." pageId="9" pageNumber="1641">
Table
<number box="[1058,1071,1425,1449]" pageId="9" pageNumber="1641" value="2.0">2</number>
</tableCitation>
.
</paragraph>
<paragraph blockId="9.[160,1156,1494,1727]" pageId="9" pageNumber="1641">
<emphasis bold="true" box="[160,244,1494,1519]" italics="true" pageId="9" pageNumber="1641">Colour.</emphasis>
Based on ethanol preserved specimens; therefore, colour pattern might be altered. Instar I: Background colour of cephalic capsule testaceous, stemmatal areas with more or less extended light brown macula; head appendages testaceous, distal half of mandible light brown; thoracic tergites light brown with diffuse testaceous maculae on laterals; legs testaceous to light brown; abdominal sclerites light brown; membranous parts creamy white; urogomphus brown with a narrow ring-like testaceous band at mid length.
</paragraph>
<caption httpUri="https://zenodo.org/record/269897/files/figure.png" pageId="10" pageNumber="1642" targetBox="[9,1152,142,1560]" targetPageId="10">
<paragraph blockId="10.[160,1157,1592,1762]" pageId="10" pageNumber="1642">
<emphasis bold="true" box="[160,252,1592,1615]" pageId="10" pageNumber="1642">Figure 3.</emphasis>
Line drawings of instar I larvae of
<taxonomicName box="[612,865,1592,1615]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="1642" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[612,865,1592,1615]" italics="true" pageId="10" pageNumber="1642">Thermonectus tremouillesi</emphasis>
</taxonomicName>
. (A) Cephalic capsule, dorsal aspect; (B) cephalic capsule, ventral aspect; (C) left antenna, dorsal aspect; (D) right antenna, ventral aspect; (E) left maxilla, dorsal aspect; (F) right maxilla, ventral aspect; (G) left mandible, dorsal aspect; (H) labium, dorsal aspect; (I) labium, ventral aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AN: antenna; EB: egg burster; FR: frontoclypeus; LA: labium; MN: mandible; MX: maxilla; PA: parietal; Sp: spinula; TP: tentorial pit.
</paragraph>
</caption>
<caption httpUri="https://zenodo.org/record/269898/files/figure.png" pageId="11" pageNumber="1643" targetBox="[9,1152,142,1555]" targetPageId="11">
<paragraph blockId="11.[160,1156,1587,1786]" pageId="11" pageNumber="1643">
<emphasis bold="true" box="[160,253,1587,1610]" pageId="11" pageNumber="1643">Figure 4.</emphasis>
Line drawings of larvae of
<taxonomicName box="[538,791,1587,1610]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1643" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[538,791,1587,1610]" italics="true" pageId="11" pageNumber="1643">
<taxonomicName box="[538,672,1587,1610]" class="Insecta" family="Dytiscidae" genus="Thermonectus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="1643" phylum="Arthropoda" rank="genus">Thermonectus</taxonomicName>
tremouillesi
</emphasis>
</taxonomicName>
. (A) Left prothoracic leg of instar I, anterior aspect; (B) right prothoracic leg of instar I, posterior aspect; (C) abdominal segment VIII of instar I, dorsal aspect; (D) abdominal segment VIII of instar I, ventral aspect; (E) right urogomphus of instar I, dorsal aspect; (F) head of instar III, dorsal aspect, showing colour pattern on cephalic capsule (chaetotaxy omitted); (G) labium of instar III, dorsal aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AB: abdominal segment VIII; CO: coxa; FE: femur; PT: pretarsus; TA: tarsus; TI: tibia; TR: trochanter; UR: urogomphus.
</paragraph>
</caption>
<paragraph blockId="12.[160,1155,142,409]" pageId="12" pageNumber="1644">Instar II. As for instar I except: light brown stemmatal macula restricted to area surrounding anterolateral stemma; lateral margin of parietal with diffuse light brown area from posterior (smallest) stemma to occipital suture; urogomphus light brown, lacking ringlike testaceous band.</paragraph>
<paragraph blockId="12.[160,1155,142,409]" pageId="12" pageNumber="1644">
Instar III (
<figureCitation box="[299,391,281,305]" captionStart="Figure 4" captionStartId="11.[160,225,1587,1610]" captionTargetBox="[9,1152,142,1555]" captionTargetId="figure@11.[163,1153,142,1556]" captionTargetPageId="11" captionText="Figure 4. Line drawings of larvae of Thermonectus tremouillesi. (A) Left prothoracic leg of instar I, anterior aspect; (B) right prothoracic leg of instar I, posterior aspect; (C) abdominal segment VIII of instar I, dorsal aspect; (D) abdominal segment VIII of instar I, ventral aspect; (E) right urogomphus of instar I, dorsal aspect; (F) head of instar III, dorsal aspect, showing colour pattern on cephalic capsule (chaetotaxy omitted); (G) labium of instar III, dorsal aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AB: abdominal segment VIII; CO: coxa; FE: femur; PT: pretarsus; TA: tarsus; TI: tibia; TR: trochanter; UR: urogomphus." httpUri="https://zenodo.org/record/269898/files/figure.png" pageId="12" pageNumber="1644">
Figure
<number box="[379,391,281,305]" pageId="12" pageNumber="1644" value="4.0">4</number>
</figureCitation>
F). As for instar II except: somewhat darker in general; background colour of cephalic capsule light brown; light brown area on lateral margin of parietal more or less developed (sometimes absent); one small lighter macula contiguous to largest stemma; legs light brown.
</paragraph>
</subSubSection>
<subSubSection box="[160,679,453,513]" pageId="12" pageNumber="1644" type="distribution">
<paragraph blockId="12.[160,679,453,513]" box="[160,300,453,478]" pageId="12" pageNumber="1644">
<heading bold="true" box="[160,300,453,478]" fontSize="11" level="2" pageId="12" pageNumber="1644" reason="0">
<emphasis bold="true" box="[160,300,453,478]" italics="true" pageId="12" pageNumber="1644">Distribution</emphasis>
</heading>
</paragraph>
<paragraph blockId="12.[160,679,453,513]" box="[160,679,489,513]" pageId="12" pageNumber="1644">
<collectingCountry box="[160,272,489,513]" name="Argentina" pageId="12" pageNumber="1644">Argentina</collectingCountry>
, Entre Ríos and Misiones Provinces.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="1644" type="biology_ecology">
<paragraph blockId="12.[160,1156,558,1068]" box="[160,249,558,583]" pageId="12" pageNumber="1644">
<heading bold="true" box="[160,249,558,583]" fontSize="11" level="2" pageId="12" pageNumber="1644" reason="0">
<emphasis bold="true" box="[160,249,558,583]" italics="true" pageId="12" pageNumber="1644">Habitat</emphasis>
</heading>
</paragraph>
<paragraph blockId="12.[160,1156,558,1068]" pageId="12" pageNumber="1644">
<taxonomicName box="[160,448,593,617]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1644" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[160,448,593,617]" italics="true" pageId="12" pageNumber="1644">Thermonectus tremouillesi</emphasis>
</taxonomicName>
apparently exploits a particular habitat. It was found only in lentic water bodies in the forest, shaded with large- and medium-sized trees, with muddy bottom and dark cold water with abundant organic matter. In Iguazú NP (
<date pageId="12" pageNumber="1644" value="2002-02-15">
<number box="[1129,1156,663,687]" pageId="12" pageNumber="1644" value="15.0">15</number>
February
<number box="[267,323,697,721]" pageId="12" pageNumber="1644" value="2002.0">2002</number>
</date>
), nine larvae (one of them reared to the adult stage) were collected from a relatively small pond under a bridge, almost completely devoid of aquatic vegetation, in association with adults of
<taxonomicName authority="Solier, 1834" authorityName="Solier" authorityYear="1834" box="[511,792,767,791]" class="Insecta" family="Hydrophilidae" genus="Tropisternus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1644" phylum="Arthropoda" rank="genus">
<emphasis box="[511,643,767,790]" italics="true" pageId="12" pageNumber="1644">Tropisternus</emphasis>
<bibRefCitation author="Solier AJJ." box="[656,792,767,791]" journalOrPublisher="An Soc Entomol Fr" pageId="12" pageNumber="1644" pagination="299 - 318" part="3" refString="Solier AJJ. 1834. Observations sur la tribu des Hydrophiliens, et principalement sur le genre Hydrophilus de Fabricius. An Soc Entomol Fr. 3: 299 - 318." title="Observations sur la tribu des Hydrophiliens, et principalement sur le genre Hydrophilus de Fabricius" type="journal article" year="1834">
Solier,
<number box="[737,792,767,791]" pageId="12" pageNumber="1644" value="1834.0">1834</number>
</bibRefCitation>
</taxonomicName>
,
<taxonomicName authority="Erichson, 1832" authorityName="Erichson" authorityYear="1832" box="[811,1100,767,791]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1644" phylum="Arthropoda" rank="genus">
<emphasis box="[811,918,767,791]" italics="true" pageId="12" pageNumber="1644">Copelatus</emphasis>
<bibRefCitation author="Erichson WF." box="[931,1100,767,791]" journalOrPublisher="Berolini: Nietackianis" pageId="12" pageNumber="1644" refString="Erichson WF. 1832. Genera Dyticeorum. Berolini: Nietackianis." title="Genera Dyticeorum" type="book" year="1832">
Erichson,
<number box="[1045,1100,767,791]" pageId="12" pageNumber="1644" value="1832.0">1832</number>
</bibRefCitation>
</taxonomicName>
and
<taxonomicName authority="Mulsant, 1844" authorityName="Mulsant" authorityYear="1844" box="[160,459,801,825]" class="Insecta" family="Hydrophilidae" genus="Helochares" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1644" phylum="Arthropoda" rank="genus">
<emphasis box="[160,279,801,825]" italics="true" pageId="12" pageNumber="1644">Helochares</emphasis>
<bibRefCitation author="Mulsant E." box="[294,459,801,825]" journalOrPublisher="Paris: L. Maison." pageId="12" pageNumber="1644" refString="Mulsant E. 1844. Histoire naturelle des Coleopteres de France. Palpicornies. Paris: L. Maison." title="Histoire naturelle des Coleopteres de France. Palpicornies" type="book" year="1844">
Mulsant,
<number box="[404,459,801,825]" pageId="12" pageNumber="1644" value="1844.0">1844</number>
</bibRefCitation>
</taxonomicName>
, and larvae of
<taxonomicName box="[656,788,802,825]" class="Insecta" family="Hydrophilidae" genus="Tropisternus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1644" phylum="Arthropoda" rank="genus">
<emphasis box="[656,788,802,825]" italics="true" pageId="12" pageNumber="1644">Tropisternus</emphasis>
</taxonomicName>
and
<taxonomicName authority="Aube, 1838" authorityName="Aube" authorityYear="1838" class="Insecta" family="Dytiscidae" genus="Hydaticus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1644" phylum="Arthropoda" rank="species" species="palliatus">
<emphasis box="[861,1075,801,825]" italics="true" pageId="12" pageNumber="1644">Hydaticus palliatus</emphasis>
<bibRefCitation author="Aube C." editor="Dejean PFMA" journalOrPublisher="Paris: Mequignon Pere et Fils" pageId="12" pageNumber="1644" refString="Aube C. 1838. Hydrocanthares et gyriniens. In: Dejean PFMA, editor. Species general des coleopteres de la collection de M. le Comte Dejean. Vol. 6. Paris: Mequignon Pere et Fils." title="Hydrocanthares et gyriniens" type="book" volumeTitle="Species general des coleopteres de la collection de M. le Comte Dejean. Vol. 6" year="1838">
Aubé,
<number box="[160,215,836,860]" pageId="12" pageNumber="1644" value="1838.0">1838</number>
</bibRefCitation>
</taxonomicName>
. In El Palmar NP (
<date box="[442,650,836,860]" pageId="12" pageNumber="1644" value="2002-02-26">
<number box="[442,469,836,860]" pageId="12" pageNumber="1644" value="26.0">26</number>
February
<number box="[594,650,836,860]" pageId="12" pageNumber="1644" value="2002.0">2002</number>
</date>
), several adults and one larva were taken from a small pond
<quantity box="[393,441,871,895]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="1644" unit="m" value="3.0">3 m</quantity>
long,
<quantity box="[521,569,871,895]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="1644" unit="m" value="2.0">2 m</quantity>
wide and
<quantity box="[703,776,871,895]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="1644" unit="cm" value="20.0">20 cm</quantity>
in depth, completely devoid of aquatic vegetation, in association with adults of
<taxonomicName authority="Enochrus Thomson, 1859" authorityName="Enochrus Thomson" authorityYear="1859" box="[720,1150,905,929]" class="Insecta" family="Hydrophilidae" genus="Tropisternus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1644" phylum="Arthropoda" rank="genus">
<emphasis box="[720,966,905,929]" italics="true" pageId="12" pageNumber="1644">Tropisternus, Enochrus</emphasis>
<bibRefCitation author="Thomson CG." box="[975,1150,905,929]" journalOrPublisher="Vol. 1. Lund: Berlingska Boktryckeriet" pageId="12" pageNumber="1644" refString="Thomson CG. 1859. Skandinaviens Coleoptera. Vol. 1. Lund: Berlingska Boktryckeriet." title="Skandinaviens Coleoptera" type="book" year="1859">
Thomson,
<number box="[1095,1150,905,929]" pageId="12" pageNumber="1644" value="1859.0">1859</number>
</bibRefCitation>
</taxonomicName>
,
<taxonomicName box="[160,267,940,964]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1644" phylum="Arthropoda" rank="genus">
<emphasis box="[160,267,940,964]" italics="true" pageId="12" pageNumber="1644">Copelatus</emphasis>
</taxonomicName>
and
<taxonomicName authority="Fabricius, 1792" authorityName="Fabricius" authorityYear="1792" box="[335,713,940,964]" class="Insecta" family="Dytiscidae" genus="Rhantus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1644" phylum="Arthropoda" rank="species" species="calidus">
<emphasis box="[335,511,940,964]" italics="true" pageId="12" pageNumber="1644">Rhantus calidus</emphasis>
(Fabricius,
<number box="[649,706,940,964]" pageId="12" pageNumber="1644" value="1792.0">1792</number>
)
</taxonomicName>
. In El Palmar NP (
<date box="[936,1143,940,964]" pageId="12" pageNumber="1644" value="2004-02-26">
<number box="[936,963,940,964]" pageId="12" pageNumber="1644" value="26.0">26</number>
February
<number box="[1087,1143,940,964]" pageId="12" pageNumber="1644" value="2004.0">2004</number>
</date>
), several adults were taken from a large pond with similar characteristics to the previous sites except for the presence of floating vegetation (
<figureCitation box="[744,836,1009,1033]" captionStart="Figure 5" captionStartId="13.[160,225,890,913]" captionTargetBox="[178,1137,142,859]" captionTargetId="figure@13.[178,1138,142,859]" captionTargetPageId="13" captionText="Figure 5. Habitat of Thermonectus tremouillesi: shaded, lentic water body at El Palmar NP, Entre Ríos Province, Argentina." httpUri="https://zenodo.org/record/269899/files/figure.png" pageId="12" pageNumber="1644">
Figure
<number box="[822,836,1009,1033]" pageId="12" pageNumber="1644" value="5.0">5</number>
</figureCitation>
), placed contiguously to the small pond sampled in
<number box="[430,486,1044,1068]" pageId="12" pageNumber="1644" value="2002.0">2002</number>
, which by that time was dry.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="1644" type="etymology">
<paragraph blockId="12.[160,1155,1113,1241]" box="[160,285,1113,1138]" pageId="12" pageNumber="1644">
<heading bold="true" box="[160,285,1113,1138]" fontSize="11" level="2" pageId="12" pageNumber="1644" reason="0">
<emphasis bold="true" box="[160,285,1113,1138]" italics="true" pageId="12" pageNumber="1644">Etymology</emphasis>
</heading>
</paragraph>
<paragraph blockId="12.[160,1155,1113,1241]" pageId="12" pageNumber="1644">
Dedicated to Prof. Edgardo R. Trémouilles, in recognition of his valuable contributions to the knowledge of the dytiscid fauna of South
<collectingCountry box="[685,779,1183,1207]" name="United States of America" pageId="12" pageNumber="1644">America</collectingCountry>
. The specific epithet is a noun in the genitive case.
</paragraph>
</subSubSection>
<subSubSection lastPageId="15" lastPageNumber="1647" pageId="12" pageNumber="1644" type="discussion">
<paragraph blockId="12.[160,299,1303,1329]" box="[160,299,1303,1329]" pageId="12" pageNumber="1644">
<heading bold="true" box="[160,299,1303,1329]" fontSize="11" level="2" pageId="12" pageNumber="1644" reason="0">
<emphasis bold="true" box="[160,299,1303,1329]" pageId="12" pageNumber="1644">Discussion</emphasis>
</heading>
</paragraph>
<paragraph blockId="12.[160,1156,1357,1727]" lastBlockId="13.[160,1156,1026,1709]" lastPageId="13" lastPageNumber="1645" pageId="12" pageNumber="1644">
The identity of many insect species described a long time ago is based on succinct, undetailed and unillustrated original descriptions. Although, by the time they appeared, these descriptions served their purpose, the substantial increase in known diversity during the last few centuries has undoubtedly rendered many old descriptions insufficiently documented from a comparative point of view. This is particularly problematic in the cases in which the
<typeStatus box="[377,486,1530,1554]" pageId="12" pageNumber="1644">holotypes</typeStatus>
are lost, where species identity might be severely compromised. In the case documented in this paper, after gathering convincing information that the
<typeStatus box="[207,305,1599,1623]" pageId="12" pageNumber="1644">holotype</typeStatus>
of
<taxonomicName box="[348,445,1599,1623]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1644" phylum="Arthropoda" rank="species" species="laporti">
<emphasis box="[348,445,1599,1623]" italics="true" pageId="12" pageNumber="1644">T. laporti</emphasis>
</taxonomicName>
is lost, we considered two possibilities of how to proceed: to consider that our specimens belong to
<taxonomicName box="[592,685,1634,1658]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1644" phylum="Arthropoda" rank="species" species="laporti">
<emphasis box="[592,685,1634,1658]" italics="true" pageId="12" pageNumber="1644">T. laporti</emphasis>
</taxonomicName>
and possibly to designate a
<typeStatus box="[999,1091,1634,1658]" pageId="12" pageNumber="1644">neotype</typeStatus>
, or to describe them as a new species. After several discussions with colleagues, the first option appeared more problematic because if we described our material as
<taxonomicName box="[921,1015,1703,1727]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="1644" phylum="Arthropoda" rank="species" species="laporti">
<emphasis box="[921,1015,1703,1727]" italics="true" pageId="12" pageNumber="1644">T. laporti</emphasis>
</taxonomicName>
and the lost
<typeStatus box="[160,259,1026,1050]" pageId="13" pageNumber="1645">holotype</typeStatus>
eventually appeared, there could be some confusion. Also, some presumed differences in dorsal colour pattern (see below) might argue against the recognition of our specimens as
<taxonomicName box="[352,444,1096,1120]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="laporti">
<emphasis box="[352,444,1096,1120]" italics="true" pageId="13" pageNumber="1645">T. laporti</emphasis>
</taxonomicName>
. On the other hand, if we described our material as a new species, and if the
<typeStatus box="[270,368,1130,1154]" pageId="13" pageNumber="1645">holotype</typeStatus>
of
<taxonomicName box="[402,495,1130,1154]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="laporti">
<emphasis box="[402,495,1130,1154]" italics="true" pageId="13" pageNumber="1645">T. laporti</emphasis>
</taxonomicName>
appeared, two scenarios are possible: (
<number box="[923,933,1130,1154]" pageId="13" pageNumber="1645" value="1.0">1</number>
) both
<typeStatus box="[1006,1115,1130,1154]" pageId="13" pageNumber="1645">holotypes</typeStatus>
are identical implying a new synonymy, which is no major problem; and (
<number box="[920,933,1165,1189]" pageId="13" pageNumber="1645" value="2.0">2</number>
) both
<typeStatus box="[1006,1115,1165,1189]" pageId="13" pageNumber="1645">holotypes</typeStatus>
are different species which means there is no conflict. A careful consideration of these arguments underpinned our decision to describe our material as a new species.
</paragraph>
<caption httpUri="https://zenodo.org/record/269899/files/figure.png" pageId="13" pageNumber="1645" targetBox="[178,1137,142,859]" targetPageId="13">
<paragraph blockId="13.[160,1155,890,943]" pageId="13" pageNumber="1645">
<emphasis bold="true" box="[160,251,890,913]" pageId="13" pageNumber="1645">Figure 5.</emphasis>
Habitat of
<taxonomicName box="[365,616,890,913]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[365,616,890,913]" italics="true" pageId="13" pageNumber="1645">
<taxonomicName box="[365,499,890,913]" class="Insecta" family="Dytiscidae" genus="Thermonectus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="genus">Thermonectus</taxonomicName>
tremouillesi
</emphasis>
</taxonomicName>
: shaded, lentic water body at El Palmar NP, Entre Ríos Province, Argentina.
</paragraph>
</caption>
<paragraph blockId="13.[160,1156,1026,1709]" lastBlockId="14.[160,1156,142,1726]" lastPageId="14" lastPageNumber="1646" pageId="13" pageNumber="1645">
Body size, dorsal colour pattern, and sexual and genital characters have proven to be the most useful in separating species within
<taxonomicName box="[658,811,1304,1328]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="genus">
<emphasis box="[658,811,1304,1328]" italics="true" pageId="13" pageNumber="1645">Thermonectus</emphasis>
</taxonomicName>
(
<bibRefCitation author="McWilliams KL." bookContentInfo="226 p." box="[824,1014,1304,1328]" journalOrPublisher="Indiana University, Bloomington, Indiana" pageId="13" pageNumber="1645" refString="McWilliams KL. 1968. A taxonomic revision of the North American species of the genus Thermonectus Dejean (Coleoptera: Dytiscidae) [Doctoral thesis]. Indiana University, Bloomington, Indiana; 226 p." title="A taxonomic revision of the North American species of the genus Thermonectus Dejean (Coleoptera: Dytiscidae) [Doctoral thesis]" type="book" year="1968">
McWilliams
<number box="[959,1014,1304,1328]" pageId="13" pageNumber="1645" value="1968.0">1968</number>
</bibRefCitation>
;
<bibRefCitation author="Tremouilles ER." journalOrPublisher="Rev Soc Entomol Argentina" pageId="13" pageNumber="1645" pagination="95 - 115" part="46" refString="Tremouilles ER. 1989. Contribucion para el conocimiento del genero Thermonectus Dejean en la Argentina y areas limitrofes (Coleoptera, Dytiscidae). Rev Soc Entomol Argentina. 46: 95 - 115." title="Contribucion para el conocimiento del genero Thermonectus Dejean en la Argentina y areas limitrofes (Coleoptera, Dytiscidae)" type="journal article" year="1989">
Trémouilles
<number box="[160,215,1338,1362]" pageId="13" pageNumber="1645" value="1989.0">1989</number>
</bibRefCitation>
). The distinct dorsal colour pattern of
<taxonomicName box="[680,835,1338,1362]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[680,835,1338,1362]" italics="true" pageId="13" pageNumber="1645">T. tremouillesi</emphasis>
</taxonomicName>
differentiates it from most species of the genus. Only four species (
<emphasis box="[632,875,1373,1397]" italics="true" pageId="13" pageNumber="1645">
<taxonomicName box="[632,731,1373,1397]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="alfredi">T. alfredi</taxonomicName>
,
<taxonomicName authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[745,1021,1373,1397]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="simulator">
T. simulator
<bibRefCitation author="Sharp D." box="[886,1021,1373,1397]" journalOrPublisher="Sci T Roy Dublin Soc" pageId="13" pageNumber="1645" pagination="179 - 1003" part="2" refString="Sharp D. 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Sci T Roy Dublin Soc. 2: 179 - 1003." title="On aquatic carnivorous Coleoptera or Dytiscidae" type="journal article" year="1882">
Sharp,
<number box="[966,1021,1373,1397]" pageId="13" pageNumber="1645" value="1882.0">1882</number>
</bibRefCitation>
</taxonomicName>
</emphasis>
,
<taxonomicName authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="depictus">
<emphasis box="[1037,1155,1373,1397]" italics="true" pageId="13" pageNumber="1645">T. depictus</emphasis>
<bibRefCitation author="Sharp D." box="[160,296,1408,1432]" journalOrPublisher="Sci T Roy Dublin Soc" pageId="13" pageNumber="1645" pagination="179 - 1003" part="2" refString="Sharp D. 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Sci T Roy Dublin Soc. 2: 179 - 1003." title="On aquatic carnivorous Coleoptera or Dytiscidae" type="journal article" year="1882">
Sharp,
<number box="[241,296,1408,1432]" pageId="13" pageNumber="1645" value="1882.0">1882</number>
</bibRefCitation>
</taxonomicName>
and
<taxonomicName box="[360,460,1408,1432]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="laporti">
<emphasis box="[360,460,1408,1432]" italics="true" pageId="13" pageNumber="1645">T. laporti</emphasis>
</taxonomicName>
) exhibit rather similar colouration, mostly on the head and pronotum.
<taxonomicName box="[297,536,1442,1466]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[297,536,1442,1466]" italics="true" pageId="13" pageNumber="1645">Thermonectus alfredi</emphasis>
</taxonomicName>
and
<taxonomicName box="[614,752,1442,1466]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="simulator">
<emphasis box="[614,752,1442,1466]" italics="true" pageId="13" pageNumber="1645">T. simulator</emphasis>
</taxonomicName>
are two closely related species (
<bibRefCitation author="Tremouilles ER." box="[167,361,1477,1501]" journalOrPublisher="Rev Soc Entomol Argentina" pageId="13" pageNumber="1645" pagination="95 - 115" part="46" refString="Tremouilles ER. 1989. Contribucion para el conocimiento del genero Thermonectus Dejean en la Argentina y areas limitrofes (Coleoptera, Dytiscidae). Rev Soc Entomol Argentina. 46: 95 - 115." title="Contribucion para el conocimiento del genero Thermonectus Dejean en la Argentina y areas limitrofes (Coleoptera, Dytiscidae)" type="journal article" year="1989">
Trémouilles
<number box="[306,361,1477,1501]" pageId="13" pageNumber="1645" value="1989.0">1989</number>
</bibRefCitation>
), undoubtedly close to
<taxonomicName box="[636,789,1477,1501]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[636,789,1477,1501]" italics="true" pageId="13" pageNumber="1645">T. tremouillesi</emphasis>
</taxonomicName>
in aedeagal structure, although visibly larger and bearing completely (or sometimes almost completely in
<taxonomicName box="[1048,1147,1512,1536]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[1048,1147,1512,1536]" italics="true" pageId="13" pageNumber="1645">T. alfredi</emphasis>
</taxonomicName>
) irrorate elytra (
<figureCitation box="[336,427,1546,1570]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="13" pageNumber="1645">
Figure
<number box="[418,427,1546,1570]" pageId="13" pageNumber="1645" value="1.0">1</number>
</figureCitation>
A). The elytra in
<taxonomicName box="[634,789,1546,1570]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[634,789,1546,1570]" italics="true" pageId="13" pageNumber="1645">T. tremouillesi</emphasis>
</taxonomicName>
are mostly irrorate but exhibit premarginal yellowish to orangish areas and a more or less extended transverse black fascia (
<figureCitation box="[238,327,1616,1640]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="13" pageNumber="1645">
Figure
<number box="[318,327,1616,1640]" pageId="13" pageNumber="1645" value="1.0">1</number>
</figureCitation>
B, D) which is never present in the other species. The shape of the male tergite IX of
<taxonomicName box="[306,458,1650,1674]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[306,458,1650,1674]" italics="true" pageId="13" pageNumber="1645">T. tremouillesi</emphasis>
</taxonomicName>
, bearing four posterior projections (
<figureCitation box="[875,968,1650,1674]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="13" pageNumber="1645">
Figure
<number box="[956,968,1650,1674]" pageId="13" pageNumber="1645" value="2.0">2</number>
</figureCitation>
F), clearly differs from that of
<taxonomicName box="[306,400,1685,1709]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[306,400,1685,1709]" italics="true" pageId="13" pageNumber="1645">T. alfredi</emphasis>
</taxonomicName>
(
<figureCitation box="[416,507,1685,1709]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="13" pageNumber="1645">
Figure
<number box="[495,507,1685,1709]" pageId="13" pageNumber="1645" value="2.0">2</number>
</figureCitation>
G) but is rather similar to that of
<taxonomicName box="[887,1015,1685,1709]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="1645" phylum="Arthropoda" rank="species" species="simulator">
<emphasis box="[887,1015,1685,1709]" italics="true" pageId="13" pageNumber="1645">T. simulator</emphasis>
</taxonomicName>
. The female sexual sculpture, however, is well marked and closely spaced in
<taxonomicName box="[993,1155,142,166]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[993,1155,142,166]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
(
<figureCitation box="[167,260,177,201]" captionStart="Figure 1" captionStartId="6.[160,225,1584,1607]" captionTargetBox="[9,1150,142,1552]" captionTargetId="figure@6.[163,1153,142,1553]" captionTargetPageId="6" captionText="Figure 1. Photographs of adults of Thermonectus. (A) T. alfredi, male, habitus, dorsal aspect; (B E) T. tremouillesi: (B) male, habitus, dorsal aspect; (C) male, habitus, ventral aspect; (D) female, left elytron, dorsal aspect, including details of punctation and sexual sculpture (right detail from SEM); (E) female, head and pronotum, dorsal aspect." httpUri="https://zenodo.org/record/269895/files/figure.png" pageId="14" pageNumber="1646">
Figure
<number box="[251,260,177,201]" pageId="14" pageNumber="1646" value="1.0">1</number>
</figureCitation>
D), whereas it is very subtle and more sparsely distributed in
<taxonomicName box="[1016,1149,177,201]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="simulator">
<emphasis box="[1016,1149,177,201]" italics="true" pageId="14" pageNumber="1646">T. simulator</emphasis>
</taxonomicName>
. Also, the male protibia is narrow, with the dorsal margin slightly prominent in
<taxonomicName box="[160,314,246,270]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[160,314,246,270]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
(
<figureCitation box="[333,428,246,270]" captionStart="Figure 2" captionStartId="7.[160,225,1580,1603]" captionTargetBox="[9,1152,142,1547]" captionTargetId="figure@7.[163,1153,142,1549]" captionTargetPageId="7" captionText="Figure 2. SEM images of male adults of Thermonectus. (A F) T. tremouillesi: (A) anterior portion, ventral aspect; (B) right protarsal palette, ventral aspect; (C) prosternal process, ventral aspect; (D) right tibia, anterior aspect (line drawing); (E) aedeagus, ventral aspect, including detail of apex of right paramere; (F) tergite IX, ventral aspect; (G) T. alfredi, tergite IX, ventral aspect." httpUri="https://zenodo.org/record/269896/files/figure.png" pageId="14" pageNumber="1646">
Figure
<number box="[415,428,246,270]" pageId="14" pageNumber="1646" value="2.0">2</number>
</figureCitation>
D), whereas in
<taxonomicName box="[606,738,246,270]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="simulator">
<emphasis box="[606,738,246,270]" italics="true" pageId="14" pageNumber="1646">T. simulator</emphasis>
</taxonomicName>
and
<taxonomicName box="[803,900,246,270]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[803,900,246,270]" italics="true" pageId="14" pageNumber="1646">T. alfredi</emphasis>
</taxonomicName>
it is broader and the dorsal margin is more prominent, giving the tibia a rather triangular appearance (
<bibRefCitation author="Tremouilles ER." box="[167,363,316,340]" journalOrPublisher="Rev Soc Entomol Argentina" pageId="14" pageNumber="1646" pagination="95 - 115" part="46" refString="Tremouilles ER. 1989. Contribucion para el conocimiento del genero Thermonectus Dejean en la Argentina y areas limitrofes (Coleoptera, Dytiscidae). Rev Soc Entomol Argentina. 46: 95 - 115." title="Contribucion para el conocimiento del genero Thermonectus Dejean en la Argentina y areas limitrofes (Coleoptera, Dytiscidae)" type="journal article" year="1989">
Trémouilles
<number box="[308,363,316,340]" pageId="14" pageNumber="1646" value="1989.0">1989</number>
</bibRefCitation>
). On the other hand, in
<taxonomicName box="[653,772,316,340]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="depictus">
<emphasis box="[653,772,316,340]" italics="true" pageId="14" pageNumber="1646">T. depictus</emphasis>
</taxonomicName>
, the anterior pronotal transverse band has pointed ends, and the posterolateral projections (or isolated central maculae) are absent, whereas the tergite IX of the male bears only two posterior projections (
<bibRefCitation author="Tremouilles ER." box="[167,360,420,444]" journalOrPublisher="Rev Soc Entomol Argentina" pageId="14" pageNumber="1646" pagination="95 - 115" part="46" refString="Tremouilles ER. 1989. Contribucion para el conocimiento del genero Thermonectus Dejean en la Argentina y areas limitrofes (Coleoptera, Dytiscidae). Rev Soc Entomol Argentina. 46: 95 - 115." title="Contribucion para el conocimiento del genero Thermonectus Dejean en la Argentina y areas limitrofes (Coleoptera, Dytiscidae)" type="journal article" year="1989">
Trémouilles
<number box="[305,360,420,444]" pageId="14" pageNumber="1646" value="1989.0">1989</number>
</bibRefCitation>
). As already mentioned, the separation of
<taxonomicName box="[847,998,420,444]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[847,998,420,444]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
and
<taxonomicName box="[1058,1155,420,444]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="laporti">
<emphasis box="[1058,1155,420,444]" italics="true" pageId="14" pageNumber="1646">T. laporti</emphasis>
</taxonomicName>
is more difficult based on available information. Both species have similar size and general colouration, although the following differences can be marked after comparison of our specimens with the original description of
<taxonomicName authority="Aube 1838" authorityName="Aube" authorityYear="1838" box="[732,979,524,548]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="laporti">
<emphasis box="[732,830,524,548]" italics="true" pageId="14" pageNumber="1646">T. laporti</emphasis>
(
<bibRefCitation author="Aube C." box="[846,970,524,548]" editor="Dejean PFMA" journalOrPublisher="Paris: Mequignon Pere et Fils" pageId="14" pageNumber="1646" refString="Aube C. 1838. Hydrocanthares et gyriniens. In: Dejean PFMA, editor. Species general des coleopteres de la collection de M. le Comte Dejean. Vol. 6. Paris: Mequignon Pere et Fils." title="Hydrocanthares et gyriniens" type="book" volumeTitle="Species general des coleopteres de la collection de M. le Comte Dejean. Vol. 6" year="1838">
Aubé
<number box="[915,970,524,548]" pageId="14" pageNumber="1646" value="1838.0">1838</number>
</bibRefCitation>
)
</taxonomicName>
: (
<number box="[1002,1012,524,548]" pageId="14" pageNumber="1646" value="1.0">1</number>
) the projection forward (following the internal ocular margins to approximately half of the eye length) of the postocular black band, which is present in
<taxonomicName box="[800,950,593,617]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[800,950,593,617]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
, is not mentioned for
<taxonomicName box="[198,294,627,651]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="laporti">
<emphasis box="[198,294,627,651]" italics="true" pageId="14" pageNumber="1646">T. laporti</emphasis>
</taxonomicName>
; this is a rather conspicuous feature, and its omission from the description of the latter species may indicate an absence; (
<number box="[656,669,662,686]" pageId="14" pageNumber="1646" value="2.0">2</number>
) the posterior transverse black band of the pronotum is
<number box="[306,339,697,721]" pageId="14" pageNumber="1646" value="2.5">2.5</number>
times broader than long in
<taxonomicName box="[657,754,697,721]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="laporti">
<emphasis box="[657,754,697,721]" italics="true" pageId="14" pageNumber="1646">T. laporti</emphasis>
</taxonomicName>
, whereas it is more than four times broader than long in
<taxonomicName box="[408,561,731,755]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[408,561,731,755]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
; also, the laterals of this band are continued to the pronotal edge as a fine marginal band in
<taxonomicName box="[661,815,766,790]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[661,815,766,790]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
, which is not mentioned for
<taxonomicName box="[160,262,801,825]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="laporti">
<emphasis box="[160,262,801,825]" italics="true" pageId="14" pageNumber="1646">T. laporti</emphasis>
</taxonomicName>
; (
<number box="[288,300,801,825]" pageId="14" pageNumber="1646" value="3.0">3</number>
) the elytra of
<taxonomicName box="[480,582,801,825]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="laporti">
<emphasis box="[480,582,801,825]" italics="true" pageId="14" pageNumber="1646">T. laporti</emphasis>
</taxonomicName>
are described as being black and covered with yellowish rounded maculae; in
<taxonomicName box="[531,687,835,859]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[531,687,835,859]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
the background colour of the elytra is yellowish to orangish and covered with black irroration. This last character, however, should be taken with care as it is just a point of view which colour (black or yellow) to choose as the background.
<taxonomicName box="[481,768,939,963]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[481,768,939,963]" italics="true" pageId="14" pageNumber="1646">Thermonectus tremouillesi</emphasis>
</taxonomicName>
keys to couplet
<quantity box="[965,1010,939,963]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" pageId="14" pageNumber="1646" unit="in" value="6.0">6 in</quantity>
<bibRefCitation author="Tremouilles ER." journalOrPublisher="Rev Soc Entomol Argentina" pageId="14" pageNumber="1646" pagination="95 - 115" part="46" refString="Tremouilles ER. 1989. Contribucion para el conocimiento del genero Thermonectus Dejean en la Argentina y areas limitrofes (Coleoptera, Dytiscidae). Rev Soc Entomol Argentina. 46: 95 - 115." title="Contribucion para el conocimiento del genero Thermonectus Dejean en la Argentina y areas limitrofes (Coleoptera, Dytiscidae)" type="journal article" year="1989">
Tremouilles
<emphasis bold="true" box="[1149,1155,939,963]" italics="true" pageId="14" pageNumber="1646"></emphasis>
(
<number box="[167,222,974,998]" pageId="14" pageNumber="1646" value="1989.0">1989</number>
)
</bibRefCitation>
key on South American
<taxonomicName box="[511,664,974,998]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="genus">
<emphasis box="[511,664,974,998]" italics="true" pageId="14" pageNumber="1646">Thermonectus</emphasis>
</taxonomicName>
. This couplet is formed by
<taxonomicName box="[975,1104,974,998]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="simulator">
<emphasis box="[975,1104,974,998]" italics="true" pageId="14" pageNumber="1646">T. simulator</emphasis>
</taxonomicName>
and
<taxonomicName box="[160,255,1009,1033]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[160,255,1009,1033]" italics="true" pageId="14" pageNumber="1646">T. alfredi</emphasis>
</taxonomicName>
, both species differing from
<taxonomicName box="[588,740,1009,1033]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[588,740,1009,1033]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
in the characters mentioned above.
</paragraph>
<paragraph blockId="14.[160,1156,142,1726]" lastBlockId="15.[160,1155,142,236]" lastPageId="15" lastPageNumber="1647" pageId="14" pageNumber="1646">
Despite great morphological similarity in larval morphology among
<taxonomicName box="[1002,1155,1043,1067]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="genus">
<emphasis box="[1002,1155,1043,1067]" italics="true" pageId="14" pageNumber="1646">Thermonectus</emphasis>
</taxonomicName>
species, the larvae of
<taxonomicName box="[425,581,1078,1102]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[425,581,1078,1102]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
look more similar to those of
<taxonomicName authority="Michat 2013" authorityName="Michat" authorityYear="2013" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[959,1058,1078,1102]" italics="true" pageId="14" pageNumber="1646">T. alfredi</emphasis>
(
<bibRefCitation author="Michat MC." journalOrPublisher="Kol Rdsch" pageId="14" pageNumber="1646" pagination="7 - 15" part="83" refString="Michat MC. 2013. Description of the larvae of Thermonectus alfredi Griffini, 1898 (Coleoptera: Dytiscidae). Kol Rdsch. 83: 7 - 15." title="Description of the larvae of Thermonectus alfredi Griffini, 1898 (Coleoptera: Dytiscidae)" type="journal article" year="2013">
Michat
<number box="[160,215,1113,1137]" pageId="14" pageNumber="1646" value="2013.0">2013</number>
</bibRefCitation>
)
</taxonomicName>
than to those of
<taxonomicName authority="Michat and Torres 2005" authorityName="Michat and Torres" authorityYear="2005" box="[437,881,1113,1137]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="succinctus">
<emphasis box="[437,577,1113,1137]" italics="true" pageId="14" pageNumber="1646">T. succinctus</emphasis>
(
<bibRefCitation author="Michat MC &amp; Torres PLM." box="[595,872,1113,1137]" journalOrPublisher="Aquat Insects" pageId="14" pageNumber="1646" pagination="281 - 292" part="27" refString="Michat MC, Torres PLM. 2005. Larval morphology of Thermonectus succinctus (Aube 1838) (Coleoptera: Dytiscidae: Dytiscinae), with biological notes and chaetotaxic analysis. Aquat Insects. 27: 281 - 292." title="Larval morphology of Thermonectus succinctus (Aube 1838) (Coleoptera: Dytiscidae: Dytiscinae), with biological notes and chaetotaxic analysis" type="journal article" year="2005">
Michat and Torres
<number box="[817,872,1113,1137]" pageId="14" pageNumber="1646" value="2005.0">2005</number>
</bibRefCitation>
)
</taxonomicName>
. The nasale is rounded and somewhat more projected forward in all instars of
<taxonomicName box="[831,987,1147,1171]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[831,987,1147,1171]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
and
<taxonomicName box="[1056,1155,1147,1171]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[1056,1155,1147,1171]" italics="true" pageId="14" pageNumber="1646">T. alfredi</emphasis>
</taxonomicName>
(
<figureCitation box="[167,270,1182,1206]" captionStart="Figure 3" captionStartId="10.[160,225,1592,1615]" captionTargetBox="[9,1152,142,1560]" captionTargetId="figure@10.[163,1153,142,1561]" captionTargetPageId="10" captionText="Figure 3. Line drawings of instar I larvae of Thermonectus tremouillesi. (A) Cephalic capsule, dorsal aspect; (B) cephalic capsule, ventral aspect; (C) left antenna, dorsal aspect; (D) right antenna, ventral aspect; (E) left maxilla, dorsal aspect; (F) right maxilla, ventral aspect; (G) left mandible, dorsal aspect; (H) labium, dorsal aspect; (I) labium, ventral aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AN: antenna; EB: egg burster; FR: frontoclypeus; LA: labium; MN: mandible; MX: maxilla; PA: parietal; Sp: spinula; TP: tentorial pit." httpUri="https://zenodo.org/record/269897/files/figure.png" pageId="14" pageNumber="1646">
Figures
<number box="[259,270,1182,1206]" pageId="14" pageNumber="1646" value="3.0">3</number>
</figureCitation>
A,
<number box="[305,317,1182,1206]" pageId="14" pageNumber="1646" value="4.0">4</number>
F). In
<taxonomicName box="[386,524,1183,1206]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="succinctus">
<emphasis box="[386,524,1183,1206]" italics="true" pageId="14" pageNumber="1646">T. succinctus</emphasis>
</taxonomicName>
, the nasale is more flattened and less prominent. The first instars of
<taxonomicName box="[324,476,1217,1241]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[324,476,1217,1241]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
and
<taxonomicName box="[536,631,1217,1241]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[536,631,1217,1241]" italics="true" pageId="14" pageNumber="1646">T. alfredi</emphasis>
</taxonomicName>
have a larger number of additional spine-like setae on the dorsal surface of the stipes (
<number box="[637,704,1251,1275]" pageId="14" pageNumber="1646" value="17.0" valueMax="19.0" valueMin="15.0">
15
<emphasis bold="true" box="[664,677,1251,1275]" italics="true" pageId="14" pageNumber="1646"></emphasis>
19
</number>
and
<number box="[765,832,1251,1275]" pageId="14" pageNumber="1646" value="19.0" valueMax="21.0" valueMin="17.0">
17
<emphasis bold="true" box="[792,805,1251,1275]" italics="true" pageId="14" pageNumber="1646"></emphasis>
21
</number>
respectively,
<figureCitation box="[989,1080,1251,1275]" captionStart="Figure 3" captionStartId="10.[160,225,1592,1615]" captionTargetBox="[9,1152,142,1560]" captionTargetId="figure@10.[163,1153,142,1561]" captionTargetPageId="10" captionText="Figure 3. Line drawings of instar I larvae of Thermonectus tremouillesi. (A) Cephalic capsule, dorsal aspect; (B) cephalic capsule, ventral aspect; (C) left antenna, dorsal aspect; (D) right antenna, ventral aspect; (E) left maxilla, dorsal aspect; (F) right maxilla, ventral aspect; (G) left mandible, dorsal aspect; (H) labium, dorsal aspect; (I) labium, ventral aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AN: antenna; EB: egg burster; FR: frontoclypeus; LA: labium; MN: mandible; MX: maxilla; PA: parietal; Sp: spinula; TP: tentorial pit." httpUri="https://zenodo.org/record/269897/files/figure.png" pageId="14" pageNumber="1646">
Figure
<number box="[1069,1080,1251,1275]" pageId="14" pageNumber="1646" value="3.0">3</number>
</figureCitation>
E) and bear additional anterodistal setae on the femur (
<figureCitation box="[711,802,1286,1310]" captionStart="Figure 4" captionStartId="11.[160,225,1587,1610]" captionTargetBox="[9,1152,142,1555]" captionTargetId="figure@11.[163,1153,142,1556]" captionTargetPageId="11" captionText="Figure 4. Line drawings of larvae of Thermonectus tremouillesi. (A) Left prothoracic leg of instar I, anterior aspect; (B) right prothoracic leg of instar I, posterior aspect; (C) abdominal segment VIII of instar I, dorsal aspect; (D) abdominal segment VIII of instar I, ventral aspect; (E) right urogomphus of instar I, dorsal aspect; (F) head of instar III, dorsal aspect, showing colour pattern on cephalic capsule (chaetotaxy omitted); (G) labium of instar III, dorsal aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AB: abdominal segment VIII; CO: coxa; FE: femur; PT: pretarsus; TA: tarsus; TI: tibia; TR: trochanter; UR: urogomphus." httpUri="https://zenodo.org/record/269898/files/figure.png" pageId="14" pageNumber="1646">
Figure
<number box="[790,802,1286,1310]" pageId="14" pageNumber="1646" value="4.0">4</number>
</figureCitation>
A).
<taxonomicName box="[841,1112,1286,1310]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="succinctus">
<emphasis box="[841,1112,1286,1310]" italics="true" pageId="14" pageNumber="1646">Thermonectus succinctus</emphasis>
</taxonomicName>
, on the other hand, has fewer (
<number box="[488,541,1321,1345]" pageId="14" pageNumber="1646" value="9.0" valueMax="12.0" valueMin="6.0">
6
<emphasis bold="true" box="[501,514,1321,1345]" italics="true" pageId="14" pageNumber="1646"></emphasis>
12
</number>
) additional setae on the stipes and lacks additional anterodistal setae on the femur. The second and third instars of
<taxonomicName box="[914,1067,1355,1379]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[914,1067,1355,1379]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
can be distinguished from those of
<taxonomicName box="[478,571,1390,1414]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[478,571,1390,1414]" italics="true" pageId="14" pageNumber="1646">T. alfredi</emphasis>
</taxonomicName>
and
<taxonomicName box="[628,763,1391,1414]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="succinctus">
<emphasis box="[628,763,1391,1414]" italics="true" pageId="14" pageNumber="1646">T. succinctus</emphasis>
</taxonomicName>
in the larger number of secondary spine-like setae on the anterodorsal surface of the prementum. Also, the number of secondary hair-like setae on the basoexternal margin of the mandible is larger in
<taxonomicName box="[160,315,1494,1518]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[160,315,1494,1518]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
and
<taxonomicName box="[382,480,1494,1518]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[382,480,1494,1518]" italics="true" pageId="14" pageNumber="1646">T. alfredi</emphasis>
</taxonomicName>
than in
<taxonomicName box="[588,728,1495,1518]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="succinctus">
<emphasis box="[588,728,1495,1518]" italics="true" pageId="14" pageNumber="1646">T. succinctus</emphasis>
</taxonomicName>
. With respect to the third instar, in
<taxonomicName box="[160,312,1529,1553]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[160,312,1529,1553]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
and
<taxonomicName box="[371,465,1529,1553]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[371,465,1529,1553]" italics="true" pageId="14" pageNumber="1646">T. alfredi</emphasis>
</taxonomicName>
, the basal one-half to two-thirds of the dorsal surface of the prementum is covered with spinulae (
<figureCitation box="[597,689,1563,1587]" captionStart="Figure 4" captionStartId="11.[160,225,1587,1610]" captionTargetBox="[9,1152,142,1555]" captionTargetId="figure@11.[163,1153,142,1556]" captionTargetPageId="11" captionText="Figure 4. Line drawings of larvae of Thermonectus tremouillesi. (A) Left prothoracic leg of instar I, anterior aspect; (B) right prothoracic leg of instar I, posterior aspect; (C) abdominal segment VIII of instar I, dorsal aspect; (D) abdominal segment VIII of instar I, ventral aspect; (E) right urogomphus of instar I, dorsal aspect; (F) head of instar III, dorsal aspect, showing colour pattern on cephalic capsule (chaetotaxy omitted); (G) labium of instar III, dorsal aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AB: abdominal segment VIII; CO: coxa; FE: femur; PT: pretarsus; TA: tarsus; TI: tibia; TR: trochanter; UR: urogomphus." httpUri="https://zenodo.org/record/269898/files/figure.png" pageId="14" pageNumber="1646">
Figure
<number box="[677,689,1563,1587]" pageId="14" pageNumber="1646" value="4.0">4</number>
</figureCitation>
G), whereas in
<taxonomicName box="[862,1000,1564,1587]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="succinctus">
<emphasis box="[862,1000,1564,1587]" italics="true" pageId="14" pageNumber="1646">T. succinctus</emphasis>
</taxonomicName>
, the spinulae are present only on the basal third. The dorsal colour pattern of the cephalic capsule in
<taxonomicName box="[160,310,1633,1657]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[160,310,1633,1657]" italics="true" pageId="14" pageNumber="1646">T. tremouillesi</emphasis>
</taxonomicName>
and
<taxonomicName box="[367,502,1633,1657]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="succinctus">
<emphasis box="[367,502,1633,1657]" italics="true" pageId="14" pageNumber="1646">T. succinctus</emphasis>
</taxonomicName>
lacks light maculae on each side of the frontoclypeus and on the parietals (close to the joint of frontal and coronal sutures), thus differing from
<taxonomicName box="[160,258,1702,1726]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="1646" phylum="Arthropoda" rank="species" species="alfredi">
<emphasis box="[160,258,1702,1726]" italics="true" pageId="14" pageNumber="1646">T. alfredi</emphasis>
</taxonomicName>
. The absence of dark maculae on the frontoclypeus of the mature larva of
<taxonomicName box="[160,313,142,166]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1647" phylum="Arthropoda" rank="species" species="tremouillesi">
<emphasis box="[160,313,142,166]" italics="true" pageId="15" pageNumber="1647">T. tremouillesi</emphasis>
</taxonomicName>
(
<figureCitation box="[330,422,142,166]" captionStart="Figure 4" captionStartId="11.[160,225,1587,1610]" captionTargetBox="[9,1152,142,1555]" captionTargetId="figure@11.[163,1153,142,1556]" captionTargetPageId="11" captionText="Figure 4. Line drawings of larvae of Thermonectus tremouillesi. (A) Left prothoracic leg of instar I, anterior aspect; (B) right prothoracic leg of instar I, posterior aspect; (C) abdominal segment VIII of instar I, dorsal aspect; (D) abdominal segment VIII of instar I, ventral aspect; (E) right urogomphus of instar I, dorsal aspect; (F) head of instar III, dorsal aspect, showing colour pattern on cephalic capsule (chaetotaxy omitted); (G) labium of instar III, dorsal aspect. Numbers and lowercase letters indicate ancestral setae and pores, respectively; additional sensilla not labelled. AB: abdominal segment VIII; CO: coxa; FE: femur; PT: pretarsus; TA: tarsus; TI: tibia; TR: trochanter; UR: urogomphus." httpUri="https://zenodo.org/record/269898/files/figure.png" pageId="15" pageNumber="1647">
Figure
<number box="[410,422,142,166]" pageId="15" pageNumber="1647" value="4.0">4</number>
</figureCitation>
F) also distinguishes it from the mainly North American species
<taxonomicName authority="Harris, 1829" authorityName="Harris" authorityYear="1829" box="[160,446,177,201]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1647" phylum="Arthropoda" rank="species" species="basillaris">
<emphasis box="[160,284,177,201]" italics="true" pageId="15" pageNumber="1647">T. basillaris</emphasis>
(
<bibRefCitation author="Harris TW." box="[302,439,177,201]" journalOrPublisher="Contributions to entomology &quot;. New England Farmer" pageId="15" pageNumber="1647" pagination="1 - 2" part="8" refString="Harris TW. 1829. Corrections and additions for the &quot; Contributions to entomology &quot;. New England Farmer 8: 1 - 2." title="Corrections and additions for the" type="journal article" year="1829">
Harris,
<number box="[384,439,177,201]" pageId="15" pageNumber="1647" value="1829.0">1829</number>
</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName authority="Aube 1838" authorityName="Aube" authorityYear="1838" box="[511,973,177,201]" class="Insecta" family="Dytiscidae" genus="Thermonectus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="1647" phylum="Arthropoda" rank="subSpecies" species="nigrofasciatus" subSpecies="ornaticollis">
<emphasis box="[511,821,177,201]" italics="true" pageId="15" pageNumber="1647">T. nigrofasciatus ornaticollis</emphasis>
(
<bibRefCitation author="Aube C." box="[839,965,177,201]" editor="Dejean PFMA" journalOrPublisher="Paris: Mequignon Pere et Fils" pageId="15" pageNumber="1647" refString="Aube C. 1838. Hydrocanthares et gyriniens. In: Dejean PFMA, editor. Species general des coleopteres de la collection de M. le Comte Dejean. Vol. 6. Paris: Mequignon Pere et Fils." title="Hydrocanthares et gyriniens" type="book" volumeTitle="Species general des coleopteres de la collection de M. le Comte Dejean. Vol. 6" year="1838">
Aubé
<number box="[910,965,177,201]" pageId="15" pageNumber="1647" value="1838.0">1838</number>
</bibRefCitation>
)
</taxonomicName>
, which bear, in different degrees, frontoclypeal dark maculae (
<bibRefCitation author="Wilson CB." box="[693,832,212,236]" journalOrPublisher="Bull Bur Fish" pageId="15" pageNumber="1647" pagination="232 - 345" part="39" refString="Wilson CB. 1923. Water beetles in relation to pondfish culture, with life histories of those found in fishponds at Fairport, Iowa. Bull Bur Fish. 39: 232 - 345." title="Water beetles in relation to pondfish culture, with life histories of those found in fishponds at Fairport, Iowa" type="journal article" year="1923">
Wilson
<number box="[777,832,212,236]" pageId="15" pageNumber="1647" value="1923.0">1923</number>
</bibRefCitation>
;
<bibRefCitation author="Carroll K &amp; Barman EH." box="[846,1129,212,236]" journalOrPublisher="Georgia J Sci" pageId="15" pageNumber="1647" pagination="171 - 178" part="62" refString="Carroll K, Barman EH. 2004. A redescription of the mature larva of Thermonectus basillaris Harris (Coleoptera: Dytiscidae: Dytiscinae). Georgia J Sci. 62: 171 - 178." title="A redescription of the mature larva of Thermonectus basillaris Harris (Coleoptera: Dytiscidae: Dytiscinae)" type="journal article" year="2004">
Carroll and Barman
<number box="[1074,1129,212,236]" pageId="15" pageNumber="1647" value="2004.0">2004</number>
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>