251 lines
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251 lines
22 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.960.55007" ID-GBIF-Dataset="d33b7f20-b953-459e-bf96-129919db2fc1" ID-PMC="PMC7445195" ID-Pensoft-Pub="1313-2970-960-63" ID-Pensoft-UUID="52BF27E864625C32B58BD8C17C1AA574" ID-PubMed="32884398" ID-ZooBank="53E72CF9515F47E49350F123BC2D8442" ModsDocID="1313-2970-960-63" checkinTime="1597680765858" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Surbakti, Suriani, Sumoked, Bob & Balke, Michael" docDate="2020" docId="5C7CE25B131050B08E2490D22ACAABDE" docLanguage="en" docName="ZooKeys 960: 63-78" docOrigin="ZooKeys 960" docSource="http://dx.doi.org/10.3897/zookeys.960.55007" docTitle="Exocelina athesphatos Shaverdo & Surbakti & Sumoked & Balke 2020, sp. nov." docType="treatment" docUuid="B42F6969-FA5C-48A9-A542-ECDCC0A25CB4" docUuidSource="ZooBank" docVersion="4" id="52BF27E864625C32B58BD8C17C1AA574" lastPageNumber="63" masterDocId="52BF27E864625C32B58BD8C17C1AA574" masterDocTitle="Two new species of the Exocelina ekari group from New Guinea with strongly modified male antennae (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="78" masterPageNumber="63" pageNumber="63" updateTime="1668169273712" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species of the Exocelina ekari group from New Guinea with strongly modified male antennae (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Shaverdo, Helena</mods:namePart>
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<mods:affiliation>Naturhistorisches Museum Wien, Burgring 7, 1010 Vienna, Austria</mods:affiliation>
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<mods:nameIdentifier type="email">shaverdo@mail.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Surbakti, Suriani</mods:namePart>
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<mods:affiliation>Department of Biology, Universitas Cendrawasih, Waena, Papua, Indonesia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sumoked, Bob</mods:namePart>
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<mods:affiliation>Walian 2, Tomohon Selatan, N Sulawesi 95439, Indonesia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Balke, Michael</mods:namePart>
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<mods:affiliation>SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany and GeoBioCenter, Ludwig-Maximilians-University, Munich, Germany</mods:affiliation>
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<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>960</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>63</mods:start>
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<mods:end>78</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.960.55007</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.960.55007</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-960-63</mods:identifier>
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<mods:identifier type="ZooBank">53E72CF9515F47E49350F123BC2D8442</mods:identifier>
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<mods:identifier type="Pensoft-UUID">52BF27E864625C32B58BD8C17C1AA574</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="166240665" LSID="urn:lsid:zoobank.org:act:B42F6969-FA5C-48A9-A542-ECDCC0A25CB4" httpUri="http://treatment.plazi.org/id/5C7CE25B131050B08E2490D22ACAABDE" lastPageNumber="63" pageId="0" pageNumber="63">
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<subSubSection pageId="0" pageNumber="63" type="nomenclature">
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<paragraph pageId="0" pageNumber="63">
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<taxonomicName LSID="5C7CE25B-1310-50B0-8E24-90D22ACAABDE" authority="Shaverdo & Surbakti & Sumoked & Balke, 2020" authorityName="Shaverdo & Surbakti & Sumoked & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina athesphatos" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="athesphatos" status="sp. nov.">Exocelina athesphatos</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="63">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Exocelina athesphatos sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure1" httpUri="https://binary.pensoft.net/fig/442331" pageId="0" pageNumber="63">Figures 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Exocelina athesphatos sp. nov., paratype A median lobe in ventral view B median lobe in lateral view." figureDoi="10.3897/zookeys.960.55007.figure2" httpUri="https://binary.pensoft.net/fig/442332" pageId="0" pageNumber="63">, 2</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Exocelina athesphatos sp. nov., paratype A right paramere in external view B right male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.960.55007.figure3" httpUri="https://binary.pensoft.net/fig/442333" pageId="0" pageNumber="63">, 3</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dirt road from Ok Sibil to Ok Bap." figureDoi="10.3897/zookeys.960.55007.figure4" httpUri="https://binary.pensoft.net/fig/442334" pageId="0" pageNumber="63">, 4</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male antennae A Exocelina athesphatos sp. nov., paratype B E. tsinga sp. nov., paratype." figureDoi="10.3897/zookeys.960.55007.figure8" httpUri="https://binary.pensoft.net/fig/442338" pageId="0" pageNumber="63">, 8A</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Abdominal ventrite 6 A Exocelina athesphatos sp. nov., paratype B E. tsinga sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure9" httpUri="https://binary.pensoft.net/fig/442339" pageId="0" pageNumber="63">, 9A</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="type locality">
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<paragraph pageId="0" pageNumber="63">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="63">
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Indonesia: Papua Province: Pegunungan Bintang Regency, near Ok Bap,
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<geoCoordinate degrees="04" direction="south" minutes="49" orientation="latitude" precision="1" seconds="28.6" value="-4.824611">04°49'28.6"S</geoCoordinate>
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,
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<geoCoordinate degrees="140" direction="east" minutes="24" orientation="longitude" precision="1" seconds="47.0" value="140.41306">140°24'47.0"E</geoCoordinate>
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, 1,961 m a.s.l.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="type material">
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<paragraph pageId="0" pageNumber="63">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Holotype</emphasis>
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: male "Indonesia: Papua, nr Ok Bab [sic!], 1961 m, 8.vi.2015, -4.82460033148527, 140.413050251081, Sumoked" (MZB).
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Paratypes</emphasis>
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: 16 males, 9 females with the same label as the holotype (MZB, KSP).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="description">
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<paragraph pageId="0" pageNumber="63">Description.</paragraph>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Body size and form</emphasis>
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: Beetle medium-sized: TL-H 4.3-4.8 mm, TL 4.85-5.4 mm, MW 2.3-2.5 mm (holotype: TL-H 4.6 mm, TL 5 mm, MW 2.4 mm), with oblong-oval habitus.
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</paragraph>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Colouration</emphasis>
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: Dorsally piceous, sometimes with dark brown posterior part of head, middle and lateral parts of pronotum, and usually with dark brown elytral sutural lines; head appendages yellowish red, legs yellowish red to reddish brown (Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Exocelina athesphatos sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure1" httpUri="https://binary.pensoft.net/fig/442331" pageId="0" pageNumber="63">1</figureCitation>
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). Teneral specimens paler, reddish brown.
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</paragraph>
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<caption doi="10.3897/zookeys.960.55007.figure1" httpUri="https://binary.pensoft.net/fig/442331" pageId="0" pageNumber="63" start="Figure 1" startId="F1">
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" pageId="0" pageNumber="63">Figure 1.</emphasis>
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Habitus and colouration of
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<taxonomicName authorityName="Shaverdo & Surbakti & Sumoked & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina athesphatos" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="athesphatos">
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<emphasis italics="true" pageId="0" pageNumber="63">Exocelina athesphatos</emphasis>
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</taxonomicName>
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sp. nov., holotype.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Surface sculpture</emphasis>
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: Shiny dorsally, with inconspicuous to fine, distinct punctation and weakly impressed microreticulation. Head with dense punctation (spaces between punctures 1-2 times size of punctures), evidently finer and sparser anteriorly; diameter of punctures almost equal to diameter of cells of microreticulation. Pronotum with sparser and finer punctation than head. Elytra with very sparse and fine punctation, sometimes inconspicuous. Microreticulation of elytra weakly impressed, in some specimens slightly stronger. Pronotum and especially head with stronger microreticulation. Metaventrite and metacoxae distinctly microreticulate, metacoxal plates with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, strioles, and very fine and sparse punctation.
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</paragraph>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Structures</emphasis>
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: Pronotum with distinct, relatively narrow lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, slightly rounded anteriorly. Blade of prosternal process lanceolate, relatively broad, slightly convex, with distinct lateral bead and few setae. Abdominal ventrite 6 broadly rounded or almost truncate, with elongate medial impression.
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</paragraph>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Male</emphasis>
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: Antenna strongly modified (Figs
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus and colouration of Exocelina athesphatos sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure1" httpUri="https://binary.pensoft.net/fig/442331" pageId="0" pageNumber="63">1</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Male antennae A Exocelina athesphatos sp. nov., paratype B E. tsinga sp. nov., paratype." figureDoi="10.3897/zookeys.960.55007.figure8" httpUri="https://binary.pensoft.net/fig/442338" pageId="0" pageNumber="63">8A</figureCitation>
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): antennomere 2 strongly reduced, antennomeres 3 and 4 strongly enlarged (antennomere 3 the largest), antennomeres 5 and 6 distinctly enlarged, antennomeres 7-10 stout. Pro- and mesotarsomeres 1-3 dilated. Protarsomere 4 slightly dilated, with anterolateral angle shortly expanded (not visible in Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Exocelina athesphatos sp. nov., paratype A right paramere in external view B right male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.960.55007.figure3" httpUri="https://binary.pensoft.net/fig/442333" pageId="0" pageNumber="63">3B</figureCitation>
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due to a wrong angle, but evident for
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<taxonomicName lsidName="E. tsinga" pageId="0" pageNumber="63" rank="species" species="tsinga">
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<emphasis italics="true" pageId="0" pageNumber="63">E. tsinga</emphasis>
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</taxonomicName>
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sp. nov. in Fig.
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Exocelina tsinga sp. nov., paratype A right paramere in external view B right male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.960.55007.figure7" httpUri="https://binary.pensoft.net/fig/442337" pageId="0" pageNumber="63">7B</figureCitation>
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; the species are similar in this character) and with large, thick, slightly curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 18 and posterior row of 9 short, thick, pointed setae (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Exocelina athesphatos sp. nov., paratype A right paramere in external view B right male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.960.55007.figure3" httpUri="https://binary.pensoft.net/fig/442333" pageId="0" pageNumber="63">3B</figureCitation>
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). Median lobe long and slender, with slightly discontinuous outline (see in apical part), enlarged and thickened apex, and with extremely small, fine, sparse setae distally on lateral margins; apex distinctly curved downwards in lateral view and in ventral view, deeply concave, with divergent sides (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Exocelina athesphatos sp. nov., paratype A median lobe in ventral view B median lobe in lateral view." figureDoi="10.3897/zookeys.960.55007.figure2" httpUri="https://binary.pensoft.net/fig/442332" pageId="0" pageNumber="63">2</figureCitation>
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). Paramere with very deep dorsal notch, separating subdistal part; subdistal part is very large, broad, with fringe of six or seven very broad, flattened setae and more numerous thin, fine setae; proximal setae numerous, dense, thin, much more inconspicuous than subdistal (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Exocelina athesphatos sp. nov., paratype A right paramere in external view B right male protarsomeres 4 - 5 in ventral view." figureDoi="10.3897/zookeys.960.55007.figure3" httpUri="https://binary.pensoft.net/fig/442333" pageId="0" pageNumber="63">3A</figureCitation>
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). Abdominal ventrite 6 with relatively deep, elongate medial impression forming two small tubercles on both sides subapically and with 14-18 lateral striae on each side (Fig.
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Abdominal ventrite 6 A Exocelina athesphatos sp. nov., paratype B E. tsinga sp. nov., holotype." figureDoi="10.3897/zookeys.960.55007.figure9" httpUri="https://binary.pensoft.net/fig/442339" pageId="0" pageNumber="63">9A</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.960.55007.figure2" httpUri="https://binary.pensoft.net/fig/442332" pageId="0" pageNumber="63" start="Figure 2" startId="F2">
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" pageId="0" pageNumber="63">Figure 2.</emphasis>
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<taxonomicName authorityName="Shaverdo & Surbakti & Sumoked & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina athesphatos" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="athesphatos">
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<emphasis italics="true" pageId="0" pageNumber="63">Exocelina athesphatos</emphasis>
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</taxonomicName>
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sp. nov., paratype
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<emphasis bold="true" pageId="0" pageNumber="63">A</emphasis>
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median lobe in ventral view
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<emphasis bold="true" pageId="0" pageNumber="63">B</emphasis>
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median lobe in lateral view.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.960.55007.figure3" httpUri="https://binary.pensoft.net/fig/442333" pageId="0" pageNumber="63" start="Figure 3" startId="F3">
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" pageId="0" pageNumber="63">Figure 3.</emphasis>
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<taxonomicName authorityName="Shaverdo & Surbakti & Sumoked & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina athesphatos" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="athesphatos">
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<emphasis italics="true" pageId="0" pageNumber="63">Exocelina athesphatos</emphasis>
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</taxonomicName>
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sp. nov., paratype
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<emphasis bold="true" pageId="0" pageNumber="63">A</emphasis>
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right paramere in external view
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<emphasis bold="true" pageId="0" pageNumber="63">B</emphasis>
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right male protarsomeres 4-5 in ventral view.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="63">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="63">Female</emphasis>
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: Antennae and pro- and mesotarsi not modified. Abdominal ventrite 6 more rounded apically, with shallow elongate medial impression, without tubercles and lateral striae.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="affinities">
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<paragraph pageId="0" pageNumber="63">Affinities.</paragraph>
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<paragraph pageId="0" pageNumber="63">
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Based on shape of the modified male antennae and presence of pronotal bead, the new species can be placed close to the
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<taxonomicName lsidName="E. polita" pageId="0" pageNumber="63" rank="species" species="polita">
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<emphasis italics="true" pageId="0" pageNumber="63">E. polita</emphasis>
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</taxonomicName>
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species complex in the identification key. However, this is not a monophyletic group. According to a molecular phylogenetic analysis (Toussaint et al., in preparation), the two new species are sister species, on their own separate branch within the
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<taxonomicName lsidName="E. ekari" pageId="0" pageNumber="63" rank="species" species="ekari">
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<emphasis italics="true" pageId="0" pageNumber="63">E. ekari</emphasis>
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</taxonomicName>
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group.
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</paragraph>
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<paragraph pageId="0" pageNumber="63">
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<taxonomicName authorityName="Shaverdo & Surbakti & Sumoked & Balke" authorityYear="2020" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina athesphatos" order="Coleoptera" pageId="0" pageNumber="63" phylum="Arthropoda" rank="species" species="athesphatos">
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<emphasis italics="true" pageId="0" pageNumber="63">Exocelina athesphatos</emphasis>
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</taxonomicName>
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sp. nov. is similar to
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<taxonomicName lsidName="E. utowaensis" pageId="0" pageNumber="63" rank="species" species="utowaensis">
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<emphasis italics="true" pageId="0" pageNumber="63">E. utowaensis</emphasis>
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</taxonomicName>
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Shaverdo, Hendrich & Balke, 2012 in modifications of the abdominal ventrite 6 and general shape of the median lobe and paramere. But the new species distinctly differs from it in larger size (TL-H 3.4-3.8 mm in
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<taxonomicName lsidName="E. utowaensis" pageId="0" pageNumber="63" rank="species" species="utowaensis">
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<emphasis italics="true" pageId="0" pageNumber="63">E. utowaensis</emphasis>
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</taxonomicName>
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), having pronotal bead (absent in
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<taxonomicName lsidName="E. utowaensis" pageId="0" pageNumber="63" rank="species" species="utowaensis">
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<emphasis italics="true" pageId="0" pageNumber="63">E. utowaensis</emphasis>
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</taxonomicName>
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) and strongly modified male antennae (simple in
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<taxonomicName lsidName="E. utowaensis" pageId="0" pageNumber="63" rank="species" species="utowaensis">
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<emphasis italics="true" pageId="0" pageNumber="63">E. utowaensis</emphasis>
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</taxonomicName>
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). Additionally, in general shape of the median lobe and paramere as well as in the relatively narrow pronotal bead, the new species resembles
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<taxonomicName lsidName="E. oceai" pageId="0" pageNumber="63" rank="species" species="oceai">
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<emphasis italics="true" pageId="0" pageNumber="63">E. oceai</emphasis>
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</taxonomicName>
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Shaverdo, Hendrich & Balke, 2012, which is, however, much smaller (TL-H 3.35-3.8 mm) and has simple male antennae. For comparison with
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<taxonomicName lsidName="E. tsinga" pageId="0" pageNumber="63" rank="species" species="tsinga">
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<emphasis italics="true" pageId="0" pageNumber="63">E. tsinga</emphasis>
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</taxonomicName>
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sp. nov. see below.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="distribution">
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<paragraph pageId="0" pageNumber="63">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="63">Indonesia: Papua Province. The species is known only from the type locality.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="habitat">
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<paragraph pageId="0" pageNumber="63">Habitat.</paragraph>
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<paragraph pageId="0" pageNumber="63">
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The specimens were collected from small puddles, in roadside ditches besides a dirt road (in Fig.
|
|
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Dirt road from Ok Sibil to Ok Bap." figureDoi="10.3897/zookeys.960.55007.figure4" httpUri="https://binary.pensoft.net/fig/442334" pageId="0" pageNumber="63">4</figureCitation>
|
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at the left hand side). The beetles have been presumably been washed into these ditches from small forest creeks during heavy rainfalls.
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</paragraph>
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|
<caption doi="10.3897/zookeys.960.55007.figure4" httpUri="https://binary.pensoft.net/fig/442334" pageId="0" pageNumber="63" start="Figure 4" startId="F4">
|
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<paragraph pageId="0" pageNumber="63">
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|
<emphasis bold="true" pageId="0" pageNumber="63">Figure 4.</emphasis>
|
|
Dirt road from Ok Sibil to Ok Bap.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="63" type="etymology">
|
|
<paragraph pageId="0" pageNumber="63">Etymology.</paragraph>
|
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<paragraph pageId="0" pageNumber="63">
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|
The species name
|
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<emphasis italics="true" pageId="0" pageNumber="63">
|
|
<normalizedToken originalValue="αθέσφατος">αθέσφατος</normalizedToken>
|
|
</emphasis>
|
|
derives from Greek, meaning "unadulterated, pure". The name is an adjective in the nominative singular.
|
|
</paragraph>
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</subSubSection>
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</treatment>
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</document> |