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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.130.1418" ID-GBIF-Dataset="1282b9f1-72a1-4673-84ed-972248edf7fc" ID-PMC="PMC3260760" ID-Pensoft-Pub="1313-2970-130-199" ID-PubMed="22259277" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-130-199" ModsDocOrigin="ZooKeys 130" ModsDocTitle="Two new species of Sinosmylites Hong (Neuroptera, Berothidae) from the Middle Jurassic of China, with notes on Mesoberothidae" checkinTime="1451249876786" checkinUser="pensoft" docAuthor="Makarkin, Vladimir N., Yang, Qiang &amp; Ren, Dong" docDate="2011" docId="4A85CC1BDB23D094ED9A73733A30C8A4" docLanguage="en" docName="ZooKeys 130: 199-215" docOrigin="ZooKeys 130" docSource="http://dx.doi.org/10.3897/zookeys.130.1418" docTitle="Sinosmylites Hong 1983, sit. n." docType="treatment" docVersion="3" lastPageNumber="201" masterDocId="FF9AFFACB9355618FF8C496EFF96130C" masterDocTitle="Two new species of Sinosmylites Hong (Neuroptera, Berothidae) from the Middle Jurassic of China, with notes on Mesoberothidae" masterLastPageNumber="215" masterPageNumber="199" pageNumber="201" updateTime="1668152189463" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Two new species of Sinosmylites Hong (Neuroptera, Berothidae) from the Middle Jurassic of China, with notes on Mesoberothidae</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Makarkin, Vladimir N.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Yang, Qiang</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ren, Dong</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
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<mods:part>
<mods:date>2011</mods:date>
<mods:detail type="volume">
<mods:number>130</mods:number>
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<mods:extent unit="page">
<mods:start>199</mods:start>
<mods:end>215</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.130.1418</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.130.1418</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-130-199</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152030855" LSID="urn:lsid:plazi:treatment:4A85CC1BDB23D094ED9A73733A30C8A4" httpUri="http://treatment.plazi.org/id/4A85CC1BDB23D094ED9A73733A30C8A4" lastPageNumber="201" pageId="2" pageNumber="201">
<subSubSection pageId="2" pageNumber="201" type="nomenclature">
<paragraph pageId="2" pageNumber="201">
Genus
<taxonomicName LSID="http://species-id.net/wiki/Sinosmylites" authority="Hong, 1983" authorityName="Hong" authorityYear="1983" class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="genus">Sinosmylites Hong, 1983</taxonomicName>
<taxonomicNameLabel pageId="2" pageNumber="201">sit. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="201" type="reference_group">
<paragraph pageId="2" pageNumber="201">
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="genus">Sinosmylites</taxonomicName>
Hong, 1983: 94, 198 [
<taxonomicName family="Osmylitidae" lsidName="" pageId="2" pageNumber="201" rank="family">Osmylitidae</taxonomicName>
];
<bibRefCitation author="Ren, D" journalOrPublisher="Seismic Publishing House, Beijing" pageId="9" pageNumber="208" title="Faunae and stratigraphy of Jurassic-Cretaceous in Beijing and the adjacent areas." year="1995">Ren et al. 1995</bibRefCitation>
: 101 [?
<taxonomicName family="Osmylidae" lsidName="" pageId="2" pageNumber="201" rank="family">Osmylidae</taxonomicName>
];
<bibRefCitation author="Ren, D" journalOrPublisher="Acta Zootaxonomica Sinica" pageId="9" pageNumber="208" pagination="461 - 479" title="On the new fossil genera and species of Neuroptera (Insecta) from the Late Jurassic of northeast China." volume="21" year="1996">Ren and Guo 1996</bibRefCitation>
: 466 [
<normalizedToken originalValue="osmylid-like">'osmylid-like'</normalizedToken>
<taxonomicName class="Magnoliopsida" family="Malvaceae" genus="Neuroptera" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Neuroptera" order="Malvales" pageId="2" pageNumber="201" phylum="Tracheophyta" rank="genus">Neuroptera</taxonomicName>
];
<bibRefCitation author="Makarkin, VN" journalOrPublisher="Zootaxa" pageId="8" pageNumber="207" pagination="15 - 23" title="Substitute names for three genera of fossil Neuroptera, with taxonomic notes." volume="1054" year="2005">Makarkin and Archibald 2005</bibRefCitation>
: 15, 16, 18, 19 [probably
<taxonomicName family="Prohemerobiidae" lsidName="" pageId="2" pageNumber="201" rank="family">Prohemerobiidae</taxonomicName>
];
<bibRefCitation pageId="2" pageNumber="201">Yang et al. 2010</bibRefCitation>
: 177 [
<taxonomicName family="Osmylidae" lsidName="" pageId="2" pageNumber="201" rank="family">Osmylidae</taxonomicName>
].
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="201" type="type species">
<paragraph pageId="2" pageNumber="201">Type species.</paragraph>
<paragraph pageId="2" pageNumber="201">
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites pectinatus" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="species" species="pectinatus">Sinosmylites pectinatus</taxonomicName>
Hong, 1983, by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="201" type="diagnosis">
<paragraph pageId="2" pageNumber="201">Diagnosis.</paragraph>
<paragraph pageId="2" pageNumber="201">
Forewing: costal space strongly narrowed basally; humeral veinlet not recurrent and branched; Sc, R1 fused distally; Sc+R1 with 9-11 veinlets, mostly simple; all subcostal veinlets simple; M forked far distal to origin of Rs; CuA pectinate, with seven branches; few crossveins in radial space arranged mainly in 1-2
<normalizedToken originalValue="inner">'inner'</normalizedToken>
gradate series.
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="201" type="species included">
<paragraph pageId="2" pageNumber="201">Species included.</paragraph>
<paragraph pageId="2" pageNumber="201">
Three species from the Middle Jurassic of China (Jiulongshan Formation):
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites pectinatus" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="species" species="pectinatus">Sinosmylites pectinatus</taxonomicName>
(Liaoning Province),
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites fumosus" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="species" species="fumosus">Sinosmylites fumosus</taxonomicName>
sp. n. and
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites rasnitsyni" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="species" species="rasnitsyni">Sinosmylites rasnitsyni</taxonomicName>
sp. n. (Inner Mongolia).
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="201" type="comments">
<paragraph pageId="2" pageNumber="201">Comments.</paragraph>
<paragraph pageId="2" pageNumber="201">
The venation of these two new species is very similar to that of
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites pectinatus" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="species" species="pectinatus">Sinosmylites pectinatus</taxonomicName>
. The latter species is represented by a nearly complete forewing (
<bibRefCitation author="Hong, Y" journalOrPublisher="Geological Publishing House, Beijing" pageId="8" pageNumber="207" title="Middle Jurassic fossil insects in North China." year="1983">Hong 1983</bibRefCitation>
). Unfortunately, however, it is quite poorly figured, and its type is now lost. Nevertheless, all main features of
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites fumosus" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="species" species="fumosus">Sinosmylites fumosus</taxonomicName>
sp. n. and
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites rasnitsyni" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="species" species="rasnitsyni">Sinosmylites rasnitsyni</taxonomicName>
sp. n. forewings agree well with those confirmed of
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites pectinatus" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="species" species="pectinatus">Sinosmylites pectinatus</taxonomicName>
, i.e., similar size (length 5.5 mm in
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites pectinatus" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="species" species="pectinatus">Sinosmylites pectinatus</taxonomicName>
; about 6.5 mm in
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites fumosus" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="species" species="fumosus">Sinosmylites fumosus</taxonomicName>
sp. n.; 6.7 mm in
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites rasnitsyni" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="species" species="rasnitsyni">Sinosmylites rasnitsyni</taxonomicName>
sp. n.), coloration (a single, more or less fuscous color), and the venational character states are as provided in the generic diagnosis. The few differences between the type species and the two new species (e.g., the presence of two
<normalizedToken originalValue="inner">'inner'</normalizedToken>
gradate series, and the CuP twice forked in
<taxonomicName class="Insecta" family="Berothidae" genus="Sinosmylites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sinosmylites pectinatus" order="Neuroptera" pageId="2" pageNumber="201" phylum="Arthropoda" rank="species" species="pectinatus">Sinosmylites pectinatus</taxonomicName>
) are at most specific.
</paragraph>
</subSubSection>
</treatment>
</document>