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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1002.58017" ID-GBIF-Dataset="8bedb86f-0c38-4738-b1b5-1dab270c95d2" ID-PMC="PMC7746671" ID-Pensoft-Pub="1313-2970-1002-1" ID-Pensoft-UUID="92722606C9CB50B6A63A1345E68167F9" ID-PubMed="33363429" ID-ZooBank="C9EAF9DCD87D402FAC35CC4983D2E142" ModsDocID="1313-2970-1002-1" checkinTime="1607628106955" checkinUser="pensoft" docAuthor="Kaltenbach, Thomas, Garces, Jhoana M. &amp; Gattolliat, Jean-Luc" docDate="2020" docId="45859324A12F529FB9AB75DE737FE918" docLanguage="en" docName="ZooKeys 1002: 1-114" docOrigin="ZooKeys 1002" docPubDate="2020-12-10" docSource="http://dx.doi.org/10.3897/zookeys.1002.58017" docTitle="Labiobaetis pangantihoni Kaltenbach &amp; Garces &amp; Gattolliat 2020, sp. nov." docType="treatment" docUuid="AAA2FE3C-DA89-41AA-A20A-4AF457BB0848" docUuidSource="ZooBank" docVersion="4" id="92722606C9CB50B6A63A1345E68167F9" lastPageNumber="1" masterDocId="92722606C9CB50B6A63A1345E68167F9" masterDocTitle="The success story of Labiobaetis Novikova &amp; Kluge in the Philippines (Ephemeroptera, Baetidae), with description of 18 new species" masterLastPageNumber="114" masterPageNumber="1" pageNumber="1" updateTime="1668146493680" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>The success story of Labiobaetis Novikova &amp; Kluge in the Philippines (Ephemeroptera, Baetidae), with description of 18 new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kaltenbach, Thomas</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8052-0388</mods:nameIdentifier>
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
<mods:nameIdentifier type="email">thomas.kaltenbach@bluewin.ch</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Garces, Jhoana M.</mods:namePart>
<mods:affiliation>Department of Biology, School of Science and Engineering, Ateneo de Manila University, Quezon City, 1108 Metro Manila, Philippines</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gattolliat, Jean-Luc</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5873-5083</mods:nameIdentifier>
<mods:affiliation>Museum of Zoology, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="pubDate">
<mods:number>2020-12-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1002</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>114</mods:end>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1002.58017</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1002.58017</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1002-1</mods:identifier>
<mods:identifier type="ZooBank">C9EAF9DCD87D402FAC35CC4983D2E142</mods:identifier>
<mods:identifier type="Pensoft-UUID">92722606C9CB50B6A63A1345E68167F9</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="170725680" LSID="urn:lsid:zoobank.org:act:AAA2FE3C-DA89-41AA-A20A-4AF457BB0848" httpUri="http://treatment.plazi.org/id/45859324A12F529FB9AB75DE737FE918" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="http://zoobank.org/AAA2FE3C-DA89-41AA-A20A-4AF457BB0848" authority="Kaltenbach &amp; Garces &amp; Gattolliat, 2020" authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis pangantihoni" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pangantihoni" status="sp. nov.">Labiobaetis pangantihoni</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Labiobaetis pangantihoni sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure19" httpUri="https://binary.pensoft.net/fig/484684" pageId="0" pageNumber="1">Figures 19</figureCitation>
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pangantihoni sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure20" httpUri="https://binary.pensoft.net/fig/484685" pageId="0" pageNumber="1">, 20</figureCitation>
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Habitus, larvae, dorsal view a Labiobaetis gamay sp. nov. b Labiobaetis pangantihoni sp. nov. c Labiobaetis tagbanwa sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure43" httpUri="https://binary.pensoft.net/fig/484708" pageId="0" pageNumber="1">, 43b</figureCitation>
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Distribution of Labiobaetis in the Philippines a Group Labiobaetis operosus b Group Labiobaetis sumigarensis c Group Labiobaetis vallus." figureDoi="10.3897/zookeys.1002.58017.figure49" httpUri="https://binary.pensoft.net/fig/484714" pageId="0" pageNumber="1">, 49a</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Larva.</emphasis>
Following combination of characters: A) dorsal surface of labrum with submarginal arc of one plus nine or ten long, feathered setae; B) labial palp segment II with a rather slender, thumb-like distomedial protuberance, segment III slightly pentagonal; C) fore femur rather broad, length 3.4
<normalizedToken originalValue="×">x</normalizedToken>
maximum width, dorsal margin with eleven or 12 curved, spine-like setae; D) claw with 10-13 denticles; E) paraproct distally not expanded, with ca. 16 stout marginal spines.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Larva</emphasis>
(Figs
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Labiobaetis pangantihoni sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure19" httpUri="https://binary.pensoft.net/fig/484684" pageId="0" pageNumber="1">19</figureCitation>
,
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pangantihoni sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure20" httpUri="https://binary.pensoft.net/fig/484685" pageId="0" pageNumber="1">20</figureCitation>
,
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Habitus, larvae, dorsal view a Labiobaetis gamay sp. nov. b Labiobaetis pangantihoni sp. nov. c Labiobaetis tagbanwa sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure43" httpUri="https://binary.pensoft.net/fig/484708" pageId="0" pageNumber="1">43b</figureCitation>
). Body length 3.5-4 mm. Cerci ca. 2/3 of body length. Paracercus ca. 2/3 of cerci length. Antenna approx. 2.5
<normalizedToken originalValue="×">x</normalizedToken>
as long as head length.
</paragraph>
<caption doi="10.3897/zookeys.1002.58017.figure19" httpUri="https://binary.pensoft.net/fig/484684" pageId="0" pageNumber="1" start="Figure 19" startId="F19">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 19.</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis pangantihoni" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pangantihoni">
<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis pangantihoni</emphasis>
</taxonomicName>
sp. nov., larva morphology
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
labrum
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
right mandible
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
right prostheca
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
left mandible
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
left prostheca
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
hypopharynx and superlinguae
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
maxilla
<emphasis bold="true" pageId="0" pageNumber="1">h</emphasis>
labium
<emphasis bold="true" pageId="0" pageNumber="1">i</emphasis>
apex of paraglossa.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Colouration</emphasis>
.
</emphasis>
Head, thorax, and abdomen dorsally brown, with bright pattern as in Fig.
<figureCitation captionStart="Figure 43" captionStartId="F43" captionText="Figure 43. Habitus, larvae, dorsal view a Labiobaetis gamay sp. nov. b Labiobaetis pangantihoni sp. nov. c Labiobaetis tagbanwa sp. nov." figureDoi="10.3897/zookeys.1002.58017.figure43" httpUri="https://binary.pensoft.net/fig/484708" pageId="0" pageNumber="1">43b</figureCitation>
, fore protoptera brown with bright striation. Head, thorax, and abdomen ventrally light brown, genae brown, abdominal segments VIII and IX darker brown. Legs light brown, femur with distomedial brown spot, apex brown, tibia distally brown. Caudalii light brown with a brown band at ca. 1/3 of cerci length.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Antenna</emphasis>
(Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pangantihoni sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure20" httpUri="https://binary.pensoft.net/fig/484685" pageId="0" pageNumber="1">20f</figureCitation>
) with scape and pedicel subcylindrical, with well-developed distolateral process at scape.
</paragraph>
<caption doi="10.3897/zookeys.1002.58017.figure20" httpUri="https://binary.pensoft.net/fig/484685" pageId="0" pageNumber="1" start="Figure 20" startId="F20">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 20.</emphasis>
<taxonomicName authorityName="Kaltenbach &amp; Garces &amp; Gattolliat" authorityYear="2020" class="Insecta" family="Baetidae" genus="Labiobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Labiobaetis pangantihoni" order="Ephemeroptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="pangantihoni">
<emphasis italics="true" pageId="0" pageNumber="1">Labiobaetis pangantihoni</emphasis>
</taxonomicName>
sp. nov., larva morphology
<emphasis bold="true" pageId="0" pageNumber="1">a</emphasis>
foreleg
<emphasis bold="true" pageId="0" pageNumber="1">b</emphasis>
fore claw
<emphasis bold="true" pageId="0" pageNumber="1">c</emphasis>
tergum IV
<emphasis bold="true" pageId="0" pageNumber="1">d</emphasis>
gill IV
<emphasis bold="true" pageId="0" pageNumber="1">e</emphasis>
paraproct
<emphasis bold="true" pageId="0" pageNumber="1">f</emphasis>
antennal scape
<emphasis bold="true" pageId="0" pageNumber="1">g</emphasis>
metanotum.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Labrum</emphasis>
(Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Labiobaetis pangantihoni sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure19" httpUri="https://binary.pensoft.net/fig/484684" pageId="0" pageNumber="1">19a</figureCitation>
). Rectangular, length 0.7
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Distal margin with medial emargination and a small process. Dorsally with medium, fine, simple setae scattered over surface; submarginal arc of setae composed of one plus nine or ten long, feathered setae. Ventrally with marginal row of setae composed of lateral and anterolateral long, feathered setae and medial long, bifid setae; ventral surface with ca. four short, spine-like setae near lateral and anterolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Right mandible</emphasis>
(Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Labiobaetis pangantihoni sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure19" httpUri="https://binary.pensoft.net/fig/484684" pageId="0" pageNumber="1">19b, c</figureCitation>
). Incisor and kinetodontium fused. Incisor with five denticles; kinetodontium with three denticles, inner margin of innermost denticle without a row of thin setae. Prostheca robust, apically denticulate. Margin between prostheca and mola slightly convex, with minute denticles. Tuft of setae at apex of mola present.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Left mandible</emphasis>
(Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Labiobaetis pangantihoni sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure19" httpUri="https://binary.pensoft.net/fig/484684" pageId="0" pageNumber="1">19d, e</figureCitation>
). Incisor and kinetodontium fused. Incisor with four denticles; kinetodontium with three denticles. Prostheca robust, apically with small denticles and comb-shaped structure. Margin between prostheca and mola straight. Subtriangular process long and slender, above level of area between prostheca and mola. Denticles of mola apically constricted. Tuft of setae at apex of mola present.
</paragraph>
<paragraph pageId="0" pageNumber="1">Both mandibles with lateral margins almost straight. Basal half with fine, simple setae scattered over dorsal surface.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Hypopharynx and superlinguae</emphasis>
(Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Labiobaetis pangantihoni sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure19" httpUri="https://binary.pensoft.net/fig/484684" pageId="0" pageNumber="1">19f</figureCitation>
). Lingua approx. as long as superlinguae. Lingua longer than broad; medial tuft of stout setae well developed, short; distal half laterally expanded. Superlinguae distally rounded; lateral margin rounded; fine, long, simple setae along distal margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Maxilla</emphasis>
(Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Labiobaetis pangantihoni sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure19" httpUri="https://binary.pensoft.net/fig/484684" pageId="0" pageNumber="1">19g</figureCitation>
). Galea-lacinia ventrally with two simple, apical setae under canines. Inner dorsal row of setae with three denti-setae, distal denti-seta tooth-like, middle and proximal denti-setae slender, bifid and pectinate. Medially with one bipectinate, spine-like seta and four or five long, simple setae. Maxillary palp 1.3
<normalizedToken originalValue="×">x</normalizedToken>
as long as length of galea-lacinia; 2-segmented; palp segment II 1.3
<normalizedToken originalValue="×">x</normalizedToken>
length of segment I; setae on maxillary palp fine, simple, scattered over surface of segments I and II; apex of last segment rounded, with excavation at inner distolateral margin.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Labium</emphasis>
(Fig.
<figureCitation captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Labiobaetis pangantihoni sp. nov., larva morphology a labrum b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium i apex of paraglossa." figureDoi="10.3897/zookeys.1002.58017.figure19" httpUri="https://binary.pensoft.net/fig/484684" pageId="0" pageNumber="1">19h, i</figureCitation>
). Glossa basally broad, narrowing toward apex; shorter than paraglossa; inner margin with ca. eight spine-like setae, distalmost seta much longer than other setae; apex with two long and one medium, robust, pectinate setae; outer margin with six medium to long, spine-like setae; ventral surface with short, fine, simple, scattered setae. Paraglossa sub-rectangular, curved inward; apex rounded; with three rows of long, robust, distally pectinate setae in apical area and two medium, simple setae in anteromedial area; dorsally with a row of four long, spine-like, simple setae near inner margin. Labial palp with segment I 0.9
<normalizedToken originalValue="×">x</normalizedToken>
length of segments II and III combined. Segment I ventrally with short, fine, simple setae. Segment II with rather slender, thumb-like distomedial protuberance; distomedial protuberance 0.8
<normalizedToken originalValue="×">x</normalizedToken>
width of base of segment III; ventral surface with short, fine, simple setae; dorsally with a row of four spine-like, simple setae near outer margin. Segment III slightly pentagonal; apex slightly truncate; length 1.1
<normalizedToken originalValue="×">x</normalizedToken>
width; ventrally covered with short, spine-like, simple setae and short, fine, simple setae.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Hind protoptera</emphasis>
(Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pangantihoni sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure20" httpUri="https://binary.pensoft.net/fig/484685" pageId="0" pageNumber="1">20g</figureCitation>
) well developed.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Foreleg</emphasis>
(Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pangantihoni sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure20" httpUri="https://binary.pensoft.net/fig/484685" pageId="0" pageNumber="1">20a, b</figureCitation>
). Ratio of foreleg segments 1.2:1.0:0.6:0.2.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Femur</emphasis>
.
</emphasis>
Length ca. 3
<normalizedToken originalValue="×">x</normalizedToken>
maximum width. Dorsal margin with eleven or twelve curved, spine-like setae; length of setae 0.16
<normalizedToken originalValue="×">x</normalizedToken>
maximum width of femur. Apex rounded, with a pair of spine-like setae and some short, stout setae. Many stout, lanceolate setae scattered along ventral margin; femoral patch absent.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Tibia</emphasis>
.
</emphasis>
Dorsal margin with a row of short, spine-like setae, near margin another row of short, spine-like setae. Ventral margin with row of short to medium, curved, spine-like setae, on apex some longer and partly bipectinate setae and a tuft of fine, simple setae. Anterior surface scattered with stout, lanceolate setae. Patellotibial suture present on basal half.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Tarsus</emphasis>
.
</emphasis>
Dorsal margin with a row of short stout setae and fine, simple setae. Ventral margin with a row of curved, spine-like setae. Claw with one row of 10-13 denticles; distally pointed; with ca. four stripes; subapical setae absent.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Terga</emphasis>
(Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pangantihoni sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure20" httpUri="https://binary.pensoft.net/fig/484685" pageId="0" pageNumber="1">20c</figureCitation>
). Surface with irregular rows of U-shaped scale bases and scattered fine, simple setae. Posterior margin of tergum IV with triangular spines, wider than long.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gills</emphasis>
(Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pangantihoni sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure20" httpUri="https://binary.pensoft.net/fig/484685" pageId="0" pageNumber="1">20d</figureCitation>
). Present on segments I-VII. Margin with small denticles intercalating fine simple setae. Tracheae extending from main trunk to inner and outer margins. Gill I ca. 1/2 length of segment II. Gill IV as long as length of segments V and 1/2 VI combined. Gill VII as long as length of segments VIII and 1/4 IX combined.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paraproct</emphasis>
(Fig.
<figureCitation captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Labiobaetis pangantihoni sp. nov., larva morphology a foreleg b fore claw c tergum IV d gill IV e paraproct f antennal scape g metanotum." figureDoi="10.3897/zookeys.1002.58017.figure20" httpUri="https://binary.pensoft.net/fig/484685" pageId="0" pageNumber="1">20e</figureCitation>
). Distally not expanded, with ca. 16 stout, marginal spines. Surface scattered with U-shaped scale bases, fine, simple setae and micropores. Cercotractor with numerous small, marginal spines.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">Dedicated to Mr. Clister V. Pangantihon, outstanding collector, entomologist and project assistant of the AdMU Biodiversity Laboratory.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Philippines: Palawan (Fig.
<figureCitation captionStart="Figure 49" captionStartId="F49" captionText="Figure 49. Distribution of Labiobaetis in the Philippines a Group Labiobaetis operosus b Group Labiobaetis sumigarensis c Group Labiobaetis vallus." figureDoi="10.3897/zookeys.1002.58017.figure49" httpUri="https://binary.pensoft.net/fig/484714" pageId="0" pageNumber="1">49a</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="biological aspects">
<paragraph pageId="0" pageNumber="1">Biological aspects.</paragraph>
<paragraph pageId="0" pageNumber="1">The specimens were collected at altitudes from sea level to 180 m, partly on submerged wood in runs or riffles.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
.
</emphasis>
Philippines • larva; Palawan, Balabac, Danglis River near the road;
<geoCoordinate degrees="07" direction="north" minutes="57" orientation="latitude" precision="15" seconds="39" value="7.960833">07°57'39&quot;N</geoCoordinate>
,
<geoCoordinate degrees="117" direction="east" minutes="02" orientation="longitude" precision="15" seconds="59" value="117.04972">117°02'59&quot;E</geoCoordinate>
; 170 m; 2019; leg. Pelingen and Pangantihon; on slide; GBIFCH 00592336; PNM.
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
.
</emphasis>
Philippines • 9 larvae; same data as holotype; 3 on slides; GenBank: MT830959; GBIFCH 00592321, GBIFCH 00763684, GBIFCH 00592335; ZSM, temporarily stored in AdMU; 5 in alcohol; GBIFCH 00515426; PCSD; 1 in alcohol; GBIFCH 00515400; AdMU • 1 larva; Palawan, Balabac, Danglis Falls/Busay;
<geoCoordinate degrees="07" direction="north" minutes="57" orientation="latitude" precision="15" seconds="39" value="7.960833">07°57'39&quot;N</geoCoordinate>
,
<geoCoordinate degrees="117" direction="east" minutes="02" orientation="longitude" precision="15" seconds="59" value="117.04972">117°02'59&quot;E</geoCoordinate>
; 180 m; 2019; leg. Pelingen and Pangantihon; in alcohol; GBIFCH 00515427; AdMU • 5 larvae; Palawan, Quezon, Isugod, Aramaywan River;
<geoCoordinate degrees="09" direction="north" minutes="21" orientation="latitude" precision="15" seconds="07" value="9.351945">09°21'07&quot;N</geoCoordinate>
,
<geoCoordinate degrees="118" direction="east" minutes="08" orientation="longitude" precision="15" seconds="26" value="118.14056">118°08'26&quot;E</geoCoordinate>
; 14 m; 2019; leg. Pangantihon and Pelingen; 1 on slide; GBIFCH 00592331; AdMU; 2 in alcohol; GBIFCH 00515429; PNM; 2 in alcohol; GBIFCH 00515401, GBIFCH 00515402; AdMU • 2 larvae; Palawan, Quezon, Aramaywan River;
<geoCoordinate degrees="09" direction="north" minutes="18" orientation="latitude" precision="15" seconds="25" value="9.306945">09°18'25&quot;N</geoCoordinate>
,
<geoCoordinate degrees="118" direction="east" minutes="07" orientation="longitude" precision="15" seconds="42" value="118.128334">118°07'42&quot;E</geoCoordinate>
; 20 m; 2019; leg. Pangantihon and Pelingen; in alcohol; GBIFCH 00515428; AdMU.
</paragraph>
</subSubSection>
</treatment>
</document>