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<document id="3F7EEAB793D982A8EEFFD0079800616F" ID-CLB-Dataset="116162" ID-DOI="10.35929/RSZ.0008" ID-GBIF-Dataset="093c4aba-a8f1-436b-a1ab-0995bf2f5225" ID-ISSN="0035-418" ID-Zenodo-Dep="5743571" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638275838110" checkinUser="carolina" docAuthor="Vagalinski, Boyan" docDate="2020" docId="450587FEFFEEF44FFC144E0AFBC7FBEF" docLanguage="en" docName="RevueSuisseZool.127.1.83-94.pdf" docOrigin="Revue suisse de Zoologie 127 (1)" docSource="http://dx.doi.org/10.35929/rsz.0008" docStyle="DocumentStyle:EA5207814960044D272DF5BE47DFF256.3:RevSuisseZool.2020-.journal_article.1cover" docStyleId="EA5207814960044D272DF5BE47DFF256" docStyleName="RevSuisseZool.2020-.journal_article.1cover" docStyleVersion="3" docTitle="Syrioiulus lohmanderi Vagalinski 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="88" masterDocId="B93CFF86FFECF449FFD14F0AFFADFFAE" masterDocTitle="A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae)" masterLastPageNumber="94" masterPageNumber="83" pageNumber="84" updateTime="1699471492402" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8CD192C23FFA624CAE9883F801D517F7">A new species of Syrioiulus Verhoeff, 1914 from Iran, with remarks on the taxonomy of the genus (Diplopoda: Julida: Julidae)</mods:title>
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<mods:namePart id="0556DDF0C4D9C447F9C9198A822DF3BF">Vagalinski, Boyan</mods:namePart>
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<mods:title id="F6B954412199744320FAE9B8B69315E7">Revue suisse de Zoologie</mods:title>
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<mods:date id="101F78DDFC07B4212F3C9E6A72653A06">2020</mods:date>
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<mods:number id="4320E7ECA0400C5D7E1AB09622D20184">2020-06-04</mods:number>
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<mods:identifier id="7F1193723A39360F70D095B23AAB1505" type="DOI">10.35929/RSZ.0008</mods:identifier>
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<treatment id="450587FEFFEEF44FFC144E0AFBC7FBEF" ID-DOI="http://doi.org/10.5281/zenodo.6974955" ID-GBIF-Taxon="198839709" ID-Zenodo-Dep="6974955" LSID="urn:lsid:plazi:treatment:450587FEFFEEF44FFC144E0AFBC7FBEF" httpUri="http://treatment.plazi.org/id/450587FEFFEEF44FFC144E0AFBC7FBEF" lastPageId="6" lastPageNumber="88" pageId="2" pageNumber="84">
<subSubSection id="85B66563FFEEF44BFC144E0AFABAFEB6" box="[965,1303,256,280]" pageId="2" pageNumber="84" type="nomenclature">
<paragraph id="CD1336E8FFEEF44BFC144E0AFABAFEB6" blockId="2.[965,1303,256,312]" box="[965,1303,256,280]" pageId="2" pageNumber="84">
<heading id="965B8184FFEEF44BFC144E0AFABAFEB6" bold="true" box="[965,1303,256,280]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="84" reason="2">
<emphasis id="FFD8EAFAFFEEF44BFC144E0AFABAFEB6" bold="true" box="[965,1303,256,280]" pageId="2" pageNumber="84">
<taxonomicName id="0AAC4D6BFFEEF44BFC144E0AFB16FEB6" ID-CoL="31ed5e90-30a8-413d-9108-6f6c61e4af18" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[965,1211,256,280]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFEEF44BFC144E0AFB16FEB6" bold="true" box="[965,1211,256,280]" italics="true" pageId="2" pageNumber="84">Syrioiulus lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFEEF44BFB134E0BFABAFEB6" box="[1218,1303,257,280]" pageId="2" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEEF44BFBEA4E2BFB0DFE96" box="[1083,1184,289,312]" pageId="2" pageNumber="84" type="description">
<paragraph id="CD1336E8FFEEF44BFBEA4E2BFB0DFE96" blockId="2.[965,1303,256,312]" box="[1083,1184,289,312]" pageId="2" pageNumber="84">
<figureCitation id="55972A6DFFEEF44BFBEA4E2BFB0DFE96" box="[1083,1184,289,312]" captionStart-0="Figs 1-4" captionStart-1="Figs 5-10" captionStart-2="Figs 11-14" captionStart-3="Figs 15-16" captionStartId-0="3.[151,192,1996,2017]" captionStartId-1="4.[151,192,1539,1560]" captionStartId-2="5.[151,192,1052,1073]" captionStartId-3="5.[151,192,2028,2049]" captionTargetBox-0="[155,1436,615,1953]" captionTargetBox-1="[198,1386,263,1488]" captionTargetBox-2="[179,1412,258,1005]" captionTargetBox-3="[151,1436,1266,1989]" captionTargetId-0="figure-159@3.[153,1438,615,1953]" captionTargetId-1="figure-212@4.[515,880,803,1077]" captionTargetId-2="figure-18@5.[518,886,510,772]" captionTargetId-3="figure-137@5.[151,1436,1266,1989]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionText-0="Figs 1-4. Syrioiulus lohmanderi sp. nov., overall appearance of male holotype (1, 2, 4) and female paratype (3), in MHNG. (1) Head and body rings 1-5. (2) Mid-body rings. (3) Head and collum. (4) Telson. Photos not to scale." captionText-1="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." captionText-2="Figs 11-14. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view.Abbreviations: cl = caudomesal lamella, ff = central filiform process, fp = fan-like process; m = mesomeral process; P = promere; s = solenomere; sp = shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm." captionText-3="Figs 15-16. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (15) Distal parts of left opisthomere, caudo-lateral view. (16) Solenomere, same aspect." figureDoi-0="http://doi.org/10.5281/zenodo.5743573" figureDoi-1="http://doi.org/10.5281/zenodo.5743577" figureDoi-2="http://doi.org/10.5281/zenodo.5743581" figureDoi-3="http://doi.org/10.5281/zenodo.5743585" httpUri-0="https://zenodo.org/record/5743573/files/figure.png" httpUri-1="https://zenodo.org/record/5743577/files/figure.png" httpUri-2="https://zenodo.org/record/5743581/files/figure.png" httpUri-3="https://zenodo.org/record/5743585/files/figure.png" pageId="2" pageNumber="84">Figs 1-16</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEEF44BFCEE4E6AFB0FFD79" pageId="2" pageNumber="84" type="materials_examined">
<paragraph id="CD1336E8FFEEF44BFCEE4E6AFB0FFD79" blockId="2.[831,1437,352,727]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4E6AFC00FED9" bold="true" box="[831,941,352,375]" pageId="2" pageNumber="84">Material:</emphasis>
<materialsCitation id="7DC43CB5FFEEF44BFC6F4E6AFC58FDD9" ID-GBIF-Occurrence="3881331305" collectingDate="1973-07-06" collectionCode="MNHG" collectorName="A. Senglet." country="Iran" isTailEnumeration="true" location="Gilan" pageId="2" pageNumber="84" specimenCount="1" specimenCount-adult="1" specimenCount-juvenile="1" stateProvince="Gilan" typeStatus="holotype">
<collectionCode id="ABBDAE2DFFEEF44BFC6F4E6AFBB9FED9" box="[958,1044,352,375]" collectionName="MNHG" pageId="2" pageNumber="84">MNHG</collectionCode>
;
<specimenCount id="DBAAFD61FFEEF44BFBF94E6AFBF1FED9" box="[1064,1116,352,375]" pageId="2" pageNumber="84" type="male">male</specimenCount>
<typeStatus id="1217884AFFEEF44BFBBF4E6AFB60FED9" box="[1134,1229,352,375]" pageId="2" pageNumber="84" type="holotype">holotype</typeStatus>
(fragmented into head, rings 1-6, left part of ring 7 and rest of body; gonopods, left antenna, leg-pair 1, leg-pair 2, penis, right leg 3, right edge of ring 7, and left mid-body leg dissected; left opisthomere prepared for
<collectionCode id="ABBDAE2DFFEEF44BFAD14EEAFA96FE59" box="[1280,1339,480,503]" pageId="2" pageNumber="84">SEM</collectionCode>
), sample field number 7314; label “
<collectingCountry id="B5BB7678FFEEF44BFBB04D0AFB23FDB9" box="[1121,1166,512,535]" name="Iran" pageId="2" pageNumber="84">Iran</collectingCountry>
(
<collectingRegion id="0F68F80AFFEEF44BFB4E4D0AFB72FDB9" box="[1183,1247,512,535]" country="Iran" name="Gilan" pageId="2" pageNumber="84">Gilan</collectingRegion>
), Siahkal, tamis., mousse, creux gros arbres” [Iran, Gilan Province, Siahkal, sifting, moss, old, hollow trees];
<date id="B9121028FFEEF44BFADB4D4AFA35FDF9" box="[1290,1432,576,599]" pageId="2" pageNumber="84" value="1973-07-06">
<collectingDate id="A956E9C0FFEEF44BFADB4D4AFA35FDF9" box="[1290,1432,576,599]" pageId="2" pageNumber="84" value="1973-07-06">06.JUL.1973</collectingDate>
</date>
; leg.
<collectorName id="6059533EFFEEF44BFCA54D6AFC58FDD9" box="[884,1013,608,631]" pageId="2" pageNumber="84">A. Senglet.</collectorName>
</materialsCitation>
<materialsCitation id="7DC43CB5FFEEF44BFBD24D6AFC59FD19" ID-GBIF-Occurrence="3881331302" collectionCode="MNHG" collectorName="A. Senglet." country="Iran" isTailEnumeration="true" location="Gilan" pageId="2" pageNumber="84" specimenCount-adult="1" specimenCount-juvenile="1" stateProvince="Gilan" typeStatus="paratype">
–
<collectionCode id="ABBDAE2DFFEEF44BFBCE4D6AFBD8FDD9" box="[1055,1141,608,631]" collectionName="MNHG" pageId="2" pageNumber="84">MNHG</collectionCode>
;
<specimenCount id="DBAAFD61FFEEF44BFB564D6AFB74FDD9" box="[1159,1241,608,631]" pageId="2" pageNumber="84" type="adult">1 adult</specimenCount>
<specimenCount id="DBAAFD61FFEEF44BFB394D6AFA9CFDD9" box="[1256,1329,608,631]" pageId="2" pageNumber="84" type="female">female</specimenCount>
<typeStatus id="1217884AFFEEF44BFAEE4D6AFA31FDD9" box="[1343,1436,608,631]" pageId="2" pageNumber="84" type="paratype">paratype</typeStatus>
(fragmented into head to ring 3 and rest of body; right vulva dissected);
</materialsCitation>
<materialsCitation id="7DC43CB5FFEEF44BFC2A4DAAFB33FD79" ID-GBIF-Occurrence="3881331311" collectingDate="1973-07-06" collectionCode="MNHG" collectorName="A. Senglet." country="Iran" isTailEnumeration="true" location="Gilan" pageId="2" pageNumber="84" specimenCount-juvenile="1" stateProvince="Gilan" typeStatus="paratype">
<specimenCount id="DBAAFD61FFEEF44BFC2A4DAAFBC8FD19" box="[1019,1125,672,695]" pageId="2" pageNumber="84" type="juvenile">1 juvenile</specimenCount>
female
<typeStatus id="1217884AFFEEF44BFB6D4DAAFAB5FD19" box="[1212,1304,672,695]" pageId="2" pageNumber="84" type="paratype">paratype</typeStatus>
(unbroken); same collecting data as
<typeStatus id="1217884AFFEEF44BFBEC4DCAFB33FD79" box="[1085,1182,704,727]" pageId="2" pageNumber="84" type="holotype">holotype</typeStatus>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEEF44BFCEE4DE5FAB3FCA8" box="[831,1310,751,774]" pageId="2" pageNumber="84" type="type_taxon">
<paragraph id="CD1336E8FFEEF44BFCEE4DE5FAB3FCA8" blockId="2.[831,1310,751,774]" box="[831,1310,751,774]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4DE5FC77FCA8" bold="true" box="[831,986,751,774]" pageId="2" pageNumber="84">
<typeStatus id="1217884AFFEEF44BFCEE4DE5FCDAFCA8" box="[831,887,751,774]" pageId="2" pageNumber="84">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="B5BB7678FFEEF44BFC314DE5FBA3FCA8" box="[992,1038,751,774]" name="Iran" pageId="2" pageNumber="84">Iran</collectingCountry>
,
<collectingRegion id="0F68F80AFFEEF44BFBC84DE5FB10FCA8" box="[1049,1213,751,774]" country="Iran" name="Gilan" pageId="2" pageNumber="84">Gilan Province</collectingRegion>
, Siahkal.
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEEF44BFCEE4C14FAD3FCDB" pageId="2" pageNumber="84" type="etymology">
<paragraph id="CD1336E8FFEEF44BFCEE4C14FAD3FCDB" blockId="2.[831,1436,798,885]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4C14FC68FC9B" bold="true" box="[831,965,798,821]" pageId="2" pageNumber="84">Etymology:</emphasis>
Named in memory of Hans Lohmander (1896-1961), a prominent millipede taxonomist who described six species currently assigned to
<taxonomicName id="0AAC4D6BFFEEF44BFADA4C54FADAFCDB" authorityName="Verhoeff" authorityYear="1914" box="[1291,1399,862,885]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEEF44BFADA4C54FADAFCDB" box="[1291,1399,862,885]" italics="true" pageId="2" pageNumber="84">Syrioiulus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEEF44BFCEE4C87FAA9FB6A" pageId="2" pageNumber="84" type="diagnosis">
<paragraph id="CD1336E8FFEEF44BFCEE4C87FAA9FB6A" blockId="2.[831,1437,909,1220]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4C87FC1AFC0A" bold="true" box="[831,951,909,932]" pageId="2" pageNumber="84">Diagnosis:</emphasis>
A blind species of
<taxonomicName id="0AAC4D6BFFEEF44BFB714C87FAA0FC0A" authorityName="Verhoeff" authorityYear="1914" box="[1184,1293,909,932]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEEF44BFB714C87FAA0FC0A" box="[1184,1293,909,932]" italics="true" pageId="2" pageNumber="84">Syrioiulus</emphasis>
</taxonomicName>
, particularly similar to
<taxonomicName id="0AAC4D6BFFEEF44BFC6D4CA7FACAFC6A" authority="(Lohmander, 1932 a)" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[956,1383,941,964]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="astrabadensis">
<emphasis id="FFD8EAFAFFEEF44BFC6D4CA7FBD8FC6A" box="[956,1141,941,964]" italics="true" pageId="2" pageNumber="84">S. astrabadensis</emphasis>
(
<bibRefCitation id="A93D4B19FFEEF44BFB5B4CA7FAF2FC6A" author="Lohmander H." box="[1162,1375,941,964]" pageId="2" pageNumber="84" pagination="1 - 44" refId="ref7038" refString="Lohmander H. 1932 a. Neue Diplopoden aus Persien. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles handlingar 5, Ser. B 3 (2): 1 - 44." type="journal article" year="1932">Lohmander, 1932a</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="0AAC4D6BFFEEF44BFCEE4CC7FB3EFC4A" authority="(Golovatch, 1983)" baseAuthorityName="Golovatch" baseAuthorityYear="1983" box="[831,1171,973,996]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="FFD8EAFAFFEEF44BFCEE4CC7FC16FC4A" box="[831,955,973,996]" italics="true" pageId="2" pageNumber="84">S. persicus</emphasis>
(
<bibRefCitation id="A93D4B19FFEEF44BFC1E4CC7FB26FC4A" author="Golovatch S. I." box="[975,1163,973,996]" pageId="2" pageNumber="84" pagination="157 - 169" refId="ref6753" refString="Golovatch S. I. 1983. A contribution to the millipede fauna of Iran (Diplopoda). Annalen des Naturhistorischen Museums in Wien (Serie B) 85: 157 - 169." type="journal article" year="1983">Golovatch, 1983</bibRefCitation>
)
</taxonomicName>
in gonopod and vulva structure. Different from
<taxonomicName id="0AAC4D6BFFEEF44BFB854CE7FAA8FBAA" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[1108,1285,1005,1028]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="astrabadensis">
<emphasis id="FFD8EAFAFFEEF44BFB854CE7FAA8FBAA" box="[1108,1285,1005,1028]" italics="true" pageId="2" pageNumber="84">S. astrabadensis</emphasis>
</taxonomicName>
mostly by its larger size and darker colouration, and by mesomeral process being subequal to rather than significantly shorter than solenomere. Different from
<taxonomicName id="0AAC4D6BFFEEF44BFAD04B47FAD5FBCA" baseAuthorityName="Golovatch" baseAuthorityYear="1983" box="[1281,1400,1101,1124]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="2" pageNumber="84" phylum="Arthropoda" rank="species" species="persicus">
<emphasis id="FFD8EAFAFFEEF44BFAD04B47FAD5FBCA" box="[1281,1400,1101,1124]" italics="true" pageId="2" pageNumber="84">S. persicus</emphasis>
</taxonomicName>
by absence of metazonal setae, and by presence of typical, pointed mesal apical denticle of promere (the same being a rounded lobe in the latter species).
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEEF44FFCEE4BD6FA3DFD5B" lastPageId="6" lastPageNumber="88" pageId="2" pageNumber="84" type="description">
<paragraph id="CD1336E8FFEEF44BFCEE4BD6FC6EFB5D" blockId="2.[831,1437,1244,2067]" box="[831,963,1244,1267]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4BD6FC6EFB5D" bold="true" box="[831,963,1244,1267]" pageId="2" pageNumber="84">Description</emphasis>
</paragraph>
<paragraph id="CD1336E8FFEEF44BFCEE4BF6FB50FA3D" blockId="2.[831,1437,1244,2067]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4BF6FC77FABD" box="[831,986,1276,1299]" italics="true" pageId="2" pageNumber="84">Measurements</emphasis>
:
<typeStatus id="1217884AFFEEF44BFC244BF6FBF5FABD" box="[1013,1112,1276,1299]" pageId="2" pageNumber="84" type="holotype">Holotype</typeStatus>
with 60+1+T body rings, length
<quantity id="0A549B0DFFEEF44BFC5E4A16FC4EFA9D" box="[911,995,1308,1331]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="84" unit="mm" value="29.0">29 mm</quantity>
, mid-body vertical diameter
<quantity id="0A549B0DFFEEF44BFAE14A16FA34FA9D" box="[1328,1433,1308,1331]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" pageId="2" pageNumber="84" unit="mm" value="1.65">1.65 mm</quantity>
. Adult female
<typeStatus id="1217884AFFEEF44BFC094A36FB99FAFD" box="[984,1076,1340,1363]" pageId="2" pageNumber="84" type="paratype">paratype</typeStatus>
with 59+1+T body rings, length
<quantity id="0A549B0DFFEEF44BFCEE4A56FC3FFADD" box="[831,914,1372,1395]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="2" pageNumber="84" unit="mm" value="28.0">28 mm</quantity>
, mid-body vertical diameter
<quantity id="0A549B0DFFEEF44BFB0B4A56FA99FADD" box="[1242,1332,1372,1395]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="84" unit="mm" value="1.9">1.9 mm</quantity>
. Juvenile female
<typeStatus id="1217884AFFEEF44BFC5E4A76FC46FA3D" box="[911,1003,1404,1427]" pageId="2" pageNumber="84" type="paratype">paratype</typeStatus>
with 49+4+T body rings.
</paragraph>
<paragraph id="CD1336E8FFEEF44BFCEE4A96FB6EF9DD" blockId="2.[831,1437,1244,2067]" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4A96FC6CFA1D" box="[831,961,1436,1459]" italics="true" pageId="2" pageNumber="84">Colouration</emphasis>
(after more than 40 years of ethanol preservation) (
<figureCitation id="55972A6DFFEEF44BFC0D4AB6FB97FA7D" box="[988,1082,1468,1491]" captionStart="Figs 1-4" captionStartId="3.[151,192,1996,2017]" captionTargetBox="[155,1436,615,1953]" captionTargetId="figure-159@3.[153,1438,615,1953]" captionTargetPageId="3" captionText="Figs 1-4. Syrioiulus lohmanderi sp. nov., overall appearance of male holotype (1, 2, 4) and female paratype (3), in MHNG. (1) Head and body rings 1-5. (2) Mid-body rings. (3) Head and collum. (4) Telson. Photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743573" httpUri="https://zenodo.org/record/5743573/files/figure.png" pageId="2" pageNumber="84">Figs 1-4</figureCitation>
): Mostly ochre brown, prozonae greyish with purple tinges; head with a dark, blurred, oval spot mesally to each antenna, and with a dark, narrow, axial line on vertex; body without axial line; legs and antennae light yellow to completely pallid; pre-anal ring and paraprocts darker ventrally.
</paragraph>
<paragraph id="CD1336E8FFEEF44AFCEE4976FE17FE97" blockId="2.[831,1437,1244,2067]" lastBlockId="3.[151,756,258,537]" lastPageId="3" lastPageNumber="85" pageId="2" pageNumber="84">
<emphasis id="FFD8EAFAFFEEF44BFCEE4976FB8EF93D" box="[831,1059,1660,1683]" italics="true" pageId="2" pageNumber="84">General morphology</emphasis>
(
<figureCitation id="55972A6DFFEEF44BFBE44976FB38F93D" box="[1077,1173,1660,1683]" captionStart="Figs 1-4" captionStartId="3.[151,192,1996,2017]" captionTargetBox="[155,1436,615,1953]" captionTargetId="figure-159@3.[153,1438,615,1953]" captionTargetPageId="3" captionText="Figs 1-4. Syrioiulus lohmanderi sp. nov., overall appearance of male holotype (1, 2, 4) and female paratype (3), in MHNG. (1) Head and body rings 1-5. (2) Mid-body rings. (3) Head and collum. (4) Telson. Photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743573" httpUri="https://zenodo.org/record/5743573/files/figure.png" pageId="2" pageNumber="84">Figs 1-4</figureCitation>
,
<figureCitation id="55972A6DFFEEF44BFB744976FB18F93D" box="[1189,1205,1660,1683]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="2" pageNumber="84">5</figureCitation>
): Ommatidia absent; vertigial setae absent; 5 supralabral setae in the adult specimens, and
<quantity id="0A549B0DFFEEF44BFC2749B6FB8BF97D" box="[1014,1062,1724,1747]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="2" pageNumber="84" unit="in" value="4.0">4 in</quantity>
the juvenile; 20-26 labral setae. Antennae (
<figureCitation id="55972A6DFFEEF44BFC6849D6FC53F95D" box="[953,1022,1756,1779]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="2" pageNumber="84">Fig. 5</figureCitation>
) about 1.2 times as long as head in both sexes; antennomeres 3, 4, and 5 subequal in length, about 0.8 times as long as 2, and 1.5 times as long as 6, 5 considerably thicker than 6; a whorl of sensilla basiconica at distal margins of antennomeres 5 and 6, those on 5 nearly as long as the four apical cones, somewhat longer than those on 6. Gnathochilarium of normal julid appearance; promentum relatively short: considerably less than half the length of lingual lamellae; each lingual lamella bearing 4-5 setae. Collum smooth, some specimens with one to several very shallow grooves near posterolateral corners.
</paragraph>
<paragraph id="CD1336E8FFEFF44AFF464E48FA2CFE17" blockId="3.[151,756,258,537]" lastBlockId="3.[831,1437,258,537]" pageId="3" pageNumber="85">Prozonae with very short and shallow, scattered, longitudinal striae. Metazonae rather shallowly and somewhat irregularly striated, with 5-8 striae in a square with sides equal to metazonal length just below ozopore level; hind margins slightly turned outwards, bearing no setae. Ozopores set in metazonae, at 1/4-1/3 of metazonal length behind pro-metazonal suture; the latter indistinct, especially on dorsal side, recognizable only as a barely visible concentric groove running through middle of ring. Mid-body legs 1.55 and 0.95 times as long as midbody diameter in the adult male and in the adult female, respectively. Tarsus of mid-body legs 1.2-1.3 times as long as tibia, and about 3 times as long as apical claw.</paragraph>
<paragraph id="CD1336E8FFEFF44DFCEE4EC8FDB0F899" blockId="3.[831,1437,258,537]" lastBlockId="4.[151,757,1728,2071]" lastPageId="4" lastPageNumber="86" pageId="3" pageNumber="85">
Telson (
<figureCitation id="55972A6DFFEFF44AFC464EC8FC71FE77" box="[919,988,450,473]" captionStart="Figs 1-4" captionStartId="3.[151,192,1996,2017]" captionTargetBox="[155,1436,615,1953]" captionTargetId="figure-159@3.[153,1438,615,1953]" captionTargetPageId="3" captionText="Figs 1-4. Syrioiulus lohmanderi sp. nov., overall appearance of male holotype (1, 2, 4) and female paratype (3), in MHNG. (1) Head and body rings 1-5. (2) Mid-body rings. (3) Head and collum. (4) Telson. Photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743573" httpUri="https://zenodo.org/record/5743573/files/figure.png" pageId="3" pageNumber="85">Fig. 4</figureCitation>
): Pre-anal ring dorsally with a few short setae; epiproct absent; hypoproct short and broadly rounded, tightly fitting under paraprocts in both sexes, with a pair of disto-median setae and a transverse row of around 10 setae proximally to the former. Paraprocts densely covered with long setae, without rows of short and stiff setae along caudal margins.
</paragraph>
<caption id="99D36660FFEFF44AFF4648C6FBD8F850" ID-DOI="http://doi.org/10.5281/zenodo.5743573" ID-Zenodo-Dep="5743573" httpUri="https://zenodo.org/record/5743573/files/figure.png" pageId="3" pageNumber="85" startId="3.[151,192,1996,2017]" targetBox="[155,1436,615,1953]" targetPageId="3">
<paragraph id="CD1336E8FFEFF44AFF4648C6FBD8F850" blockId="3.[151,1437,1996,2046]" pageId="3" pageNumber="85">
Figs 1-4.
<taxonomicName id="0AAC4D6BFFEFF44AFF2748C6FE60F84F" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[246,461,1996,2017]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="3" pageNumber="85" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFEFF44AFF2748C6FE60F84F" box="[246,461,1996,2017]" italics="true" pageId="3" pageNumber="85">Syrioiulus lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFEFF44AFE0248C6FD8CF84F" box="[467,545,1996,2017]" pageId="3" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
, overall appearance of male holotype (1, 2, 4) and female paratype (3), in MHNG. (1) Head and body rings 1-5. (2) Mid-body rings. (3) Head and collum. (4) Telson. Photos not to scale.
</paragraph>
</caption>
<caption id="99D36660FFE8F44DFF464909FBBAF9C0" ID-DOI="http://doi.org/10.5281/zenodo.5743577" ID-Zenodo-Dep="5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="4" pageNumber="86" startId="4.[151,192,1539,1560]" targetBox="[198,1386,263,1488]" targetPageId="4">
<paragraph id="CD1336E8FFE8F44DFF464909FBBAF9C0" blockId="4.[151,1437,1539,1646]" pageId="4" pageNumber="86">
Figs 5-10.
<taxonomicName id="0AAC4D6BFFE8F44DFED84909FE4CF9B6" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[265,481,1539,1560]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="4" pageNumber="86" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFE8F44DFED84909FE4CF9B6" box="[265,481,1539,1560]" italics="true" pageId="4" pageNumber="86">Syrioiulus lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFE8F44DFE394909FD9BF9B6" box="[488,566,1539,1560]" pageId="4" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
, structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations:
<emphasis id="FFD8EAFAFFE8F44DFD364937FD55F9FC" box="[743,760,1597,1618]" italics="true" pageId="4" pageNumber="86">ct</emphasis>
= central tube;
<emphasis id="FFD8EAFAFFE8F44DFC5A4937FC31F9FC" box="[907,924,1597,1618]" italics="true" pageId="4" pageNumber="86">lv</emphasis>
= lateral valve;
<emphasis id="FFD8EAFAFFE8F44DFBE44937FBFFF9FC" box="[1077,1106,1597,1618]" italics="true" pageId="4" pageNumber="86">ma</emphasis>
= mesal ampulla;
<emphasis id="FFD8EAFAFFE8F44DFAD14937FAB5F9FC" box="[1280,1304,1597,1618]" italics="true" pageId="4" pageNumber="86">mt</emphasis>
= mesal tube;
<emphasis id="FFD8EAFAFFE8F44DFED84953FE88F9C0" box="[265,293,1625,1646]" italics="true" pageId="4" pageNumber="86">mv</emphasis>
= mesal valve;
<emphasis id="FFD8EAFAFFE8F44DFE6C4953FE78F9C0" box="[445,469,1625,1646]" italics="true" pageId="4" pageNumber="86">op</emphasis>
= operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm.
</paragraph>
</caption>
<paragraph id="CD1336E8FFE8F44DFF46484AFA2BF839" blockId="4.[151,757,1728,2071]" lastBlockId="4.[831,1437,1728,2071]" pageId="4" pageNumber="86">
<emphasis id="FFD8EAFAFFE8F44DFF46484AFE3EF8F9" box="[151,403,1856,1879]" italics="true" pageId="4" pageNumber="86">Male sexual characters</emphasis>
: Mandibular stipites (see
<figureCitation id="55972A6DFFE8F44DFD7D484AFD47F8F9" box="[684,746,1856,1879]" captionStart="Figs 1-4" captionStartId="3.[151,192,1996,2017]" captionTargetBox="[155,1436,615,1953]" captionTargetId="figure-159@3.[153,1438,615,1953]" captionTargetPageId="3" captionText="Figs 1-4. Syrioiulus lohmanderi sp. nov., overall appearance of male holotype (1, 2, 4) and female paratype (3), in MHNG. (1) Head and body rings 1-5. (2) Mid-body rings. (3) Head and collum. (4) Telson. Photos not to scale." figureDoi="http://doi.org/10.5281/zenodo.5743573" httpUri="https://zenodo.org/record/5743573/files/figure.png" pageId="4" pageNumber="86">Fig. 1</figureCitation>
) forming rather small, rounded, frontoventral lobes. Legpair 1 (
<figureCitation id="55972A6DFFE8F44DFF0E488AFEB0F839" box="[223,285,1920,1943]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="4" pageNumber="86">Fig. 6</figureCitation>
) developed as compact hooks pointing mostly mesad, with a tapering tibial outgrowth and a minute, knob-like, tarsal remnant. Leg-pair 2 (
<figureCitation id="55972A6DFFE8F44DFDEF48CAFD2CF879" box="[574,641,1984,2007]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="4" pageNumber="86">Fig. 7</figureCitation>
) with two crested adhesive pads, one each on postfemur and tibia. Leg-pair 3 (
<figureCitation id="55972A6DFFE8F44DFEC3470AFEFDF7B9" box="[274,336,2048,2071]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="4" pageNumber="86">Fig. 8</figureCitation>
) and following pairs additionally with a vestigial pad on femur; all pads gradually diminishing in size towards telson, completely disappearing in posterior third of body. Pleurotergum 7 (
<figureCitation id="55972A6DFFE8F44DFB5E480AFB7DF8B9" box="[1167,1232,1792,1815]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="4" pageNumber="86">Fig. 9</figureCitation>
) ventrally forming large, rounded lobes originating from the border between pro- and metazona, protruding mostly ventrad. Penis soft, hyaline, with deeply divided, conoidal lobes ending up in fingertip-like terminal lamellae turned slightly caudad.
</paragraph>
<paragraph id="CD1336E8FFE8F44FFCEE48AAFC7AFED6" blockId="4.[831,1437,1728,2071]" lastBlockId="6.[831,1437,258,757]" lastPageId="6" lastPageNumber="88" pageId="4" pageNumber="86">
<emphasis id="FFD8EAFAFFE8F44DFCEE48AAFC00F819" box="[831,941,1952,1975]" italics="true" pageId="4" pageNumber="86">Gonopods</emphasis>
(
<figureCitation id="55972A6DFFE8F44DFC6C48AAFB9BF819" box="[957,1078,1952,1975]" captionStart-0="Figs 11-14" captionStart-1="Figs 15-16" captionStartId-0="5.[151,192,1052,1073]" captionStartId-1="5.[151,192,2028,2049]" captionTargetBox-0="[179,1412,258,1005]" captionTargetBox-1="[151,1436,1266,1989]" captionTargetId-0="figure-18@5.[518,886,510,772]" captionTargetId-1="figure-137@5.[151,1436,1266,1989]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="Figs 11-14. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view.Abbreviations: cl = caudomesal lamella, ff = central filiform process, fp = fan-like process; m = mesomeral process; P = promere; s = solenomere; sp = shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm." captionText-1="Figs 15-16. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (15) Distal parts of left opisthomere, caudo-lateral view. (16) Solenomere, same aspect." figureDoi-0="http://doi.org/10.5281/zenodo.5743581" figureDoi-1="http://doi.org/10.5281/zenodo.5743585" httpUri-0="https://zenodo.org/record/5743581/files/figure.png" httpUri-1="https://zenodo.org/record/5743585/files/figure.png" pageId="4" pageNumber="86">Figs 11-16</figureCitation>
): Promere (
<figureCitation id="55972A6DFFE8F44DFB6448AAFAAEF819" box="[1205,1283,1952,1975]" captionStart="Figs 11-14" captionStartId="5.[151,192,1052,1073]" captionTargetBox="[179,1412,258,1005]" captionTargetId="figure-18@5.[518,886,510,772]" captionTargetPageId="5" captionText="Figs 11-14. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view.Abbreviations: cl = caudomesal lamella, ff = central filiform process, fp = fan-like process; m = mesomeral process; P = promere; s = solenomere; sp = shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.5743581" httpUri="https://zenodo.org/record/5743581/files/figure.png" pageId="4" pageNumber="86">Fig. 11</figureCitation>
,
<emphasis id="FFD8EAFAFFE8F44DFAC048AAFA8CF819" box="[1297,1313,1952,1975]" italics="true" pageId="4" pageNumber="86">P</emphasis>
in
<figureCitation id="55972A6DFFE8F44DFA9748AAFA39F819" box="[1350,1428,1952,1975]" captionStart="Figs 11-14" captionStartId="5.[151,192,1052,1073]" captionTargetBox="[179,1412,258,1005]" captionTargetId="figure-18@5.[518,886,510,772]" captionTargetPageId="5" captionText="Figs 11-14. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view.Abbreviations: cl = caudomesal lamella, ff = central filiform process, fp = fan-like process; m = mesomeral process; P = promere; s = solenomere; sp = shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.5743581" httpUri="https://zenodo.org/record/5743581/files/figure.png" pageId="4" pageNumber="86">Fig. 12</figureCitation>
) relatively slender and spoon-like, tapering, as high as opisthomere; width of shaft about 0.7 times that of bowl; ridge (
<emphasis id="FFD8EAFAFFE8F44DFC09470AFC4FF7B9" box="[984,994,2048,2071]" italics="true" pageId="4" pageNumber="86">r</emphasis>
) massive and rounded, without freely protruding apical part; apical denticles well developed, clearly separated, mesal one running vertical, lateral one somewhat oblique, both with blunt tips pointing caudobasad. Opisthomere (
<figureCitation id="55972A6DFFEAF44FFEA8496BFE43F9D6" box="[377,494,1633,1656]" captionStart-0="Figs 11-14" captionStart-1="Figs 15-16" captionStartId-0="5.[151,192,1052,1073]" captionStartId-1="5.[151,192,2028,2049]" captionTargetBox-0="[179,1412,258,1005]" captionTargetBox-1="[151,1436,1266,1989]" captionTargetId-0="figure-18@5.[518,886,510,772]" captionTargetId-1="figure-137@5.[151,1436,1266,1989]" captionTargetPageId-0="5" captionTargetPageId-1="5" captionText-0="Figs 11-14. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view.Abbreviations: cl = caudomesal lamella, ff = central filiform process, fp = fan-like process; m = mesomeral process; P = promere; s = solenomere; sp = shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm." captionText-1="Figs 15-16. Syrioiulus lohmanderi sp. nov., gonopods of male holotype, in MHNG. (15) Distal parts of left opisthomere, caudo-lateral view. (16) Solenomere, same aspect." figureDoi-0="http://doi.org/10.5281/zenodo.5743581" figureDoi-1="http://doi.org/10.5281/zenodo.5743585" httpUri-0="https://zenodo.org/record/5743581/files/figure.png" httpUri-1="https://zenodo.org/record/5743585/files/figure.png" pageId="6" pageNumber="88">Figs 12-16</figureCitation>
) approximately halfway divided into mesomeral process (
<emphasis id="FFD8EAFAFFEAF44FFDC8498BFD81F936" box="[537,556,1665,1688]" italics="true" pageId="6" pageNumber="88">m</emphasis>
) and solenomere (
<emphasis id="FFD8EAFAFFEAF44FFF7149ABFF07F916" box="[160,170,1697,1720]" italics="true" pageId="6" pageNumber="88">s</emphasis>
), these being subequal in height, forming a broad sinus between them; mesomeral process running mostly straight, at considerable distance from solenomere, apically broadened and spade-like, with a strongly elongated mesal corner turned caudad, partly concealing solenomere, and with a nearly right-angled lateral corner bearing several minute spines/denticles; caudomesal lamella (
<emphasis id="FFD8EAFAFFEAF44FFF2F488EFEBCF835" box="[254,273,1924,1947]" italics="true" pageId="6" pageNumber="88">cl</emphasis>
) well-developed, forming a narrow ridge protruding completely mesad, thus being clearly visible in frontal and caudal views; solenomere slender, gradually bent anteriad, ending in two small processes [an anterior, fan-like (
<emphasis id="FFD8EAFAFFEAF44FFE55470CFE34F7B3" box="[388,409,2054,2077]" italics="true" pageId="6" pageNumber="88">fp</emphasis>
) and a posterior, shield-like one (
<emphasis id="FFD8EAFAFFEAF44FFC994E08FCCDFEB7" box="[840,864,258,281]" italics="true" pageId="6" pageNumber="88">sp</emphasis>
)] surrounding a deep central concavity; a minute central filiform process (
<emphasis id="FFD8EAFAFFEAF44FFB884E28FBC5FE97" box="[1113,1128,290,313]" italics="true" pageId="6" pageNumber="88">ff</emphasis>
) visible in certain aspects; anterior lamella vestigial, recognizable only by several apical fringes.
</paragraph>
<caption id="99D36660FFE9F44CFF464B16FED5FB0A" ID-DOI="http://doi.org/10.5281/zenodo.5743581" ID-Zenodo-Dep="5743581" httpUri="https://zenodo.org/record/5743581/files/figure.png" pageId="5" pageNumber="87" startId="5.[151,192,1052,1073]" targetBox="[179,1412,258,1005]" targetPageId="5">
<paragraph id="CD1336E8FFE9F44CFF464B16FED5FB0A" blockId="5.[151,1437,1052,1188]" pageId="5" pageNumber="87">
Figs 11-14.
<taxonomicName id="0AAC4D6BFFE9F44CFECD4B16FE5EFB9F" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[284,499,1052,1073]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="87" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFE9F44CFECD4B16FE5EFB9F" box="[284,499,1052,1073]" italics="true" pageId="5" pageNumber="87">Syrioiulus lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFE9F44CFE284B16FDEBFB9F" box="[505,582,1052,1073]" pageId="5" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
, gonopods of male holotype, in MHNG. (11) Left promere, caudal view. (12) Right pair of gonopods, mesal view. (13) Left opisthomere, caudal, slightly lateral view. (14) Distal parts of left opisthomere, lateral view. Abbreviations:
<emphasis id="FFD8EAFAFFE9F44CFE364B5FFE55FBC4" box="[487,504,1109,1130]" italics="true" pageId="5" pageNumber="87">cl</emphasis>
= caudomesal lamella,
<emphasis id="FFD8EAFAFFE9F44CFD084B5FFD4BFBC4" box="[729,742,1109,1130]" italics="true" pageId="5" pageNumber="87">ff</emphasis>
= central filiform process,
<emphasis id="FFD8EAFAFFE9F44CFC364B5FFC57FBC4" box="[999,1018,1109,1130]" italics="true" pageId="5" pageNumber="87">fp</emphasis>
= fan-like process;
<emphasis id="FFD8EAFAFFE9F44CFB644B5FFB6BFBC4" box="[1205,1222,1109,1130]" italics="true" pageId="5" pageNumber="87">m</emphasis>
= mesomeral process;
<emphasis id="FFD8EAFAFFE9F44CFECD4B78FE86FB29" box="[284,299,1138,1159]" italics="true" pageId="5" pageNumber="87">P</emphasis>
= promere;
<emphasis id="FFD8EAFAFFE9F44CFE734B78FE06FB29" box="[418,427,1138,1159]" italics="true" pageId="5" pageNumber="87">s</emphasis>
= solenomere;
<emphasis id="FFD8EAFAFFE9F44CFD934B78FDFAFB29" box="[578,599,1138,1159]" italics="true" pageId="5" pageNumber="87">sp</emphasis>
= shield-like process. Scale bar of Figs 11-13: 0.1 mm; Figs 14-15: 0.05 mm; Fig. 16: 0.01 mm.
</paragraph>
</caption>
<caption id="99D36660FFE9F44CFF4648E6FDD1F7B0" ID-DOI="http://doi.org/10.5281/zenodo.5743585" ID-Zenodo-Dep="5743585" httpUri="https://zenodo.org/record/5743585/files/figure.png" pageId="5" pageNumber="87" startId="5.[151,192,2028,2049]" targetBox="[151,1436,1266,1989]" targetPageId="5">
<paragraph id="CD1336E8FFE9F44CFF4648E6FDD1F7B0" blockId="5.[151,1436,2028,2078]" pageId="5" pageNumber="87">
Figs 15-16.
<taxonomicName id="0AAC4D6BFFE9F44CFECD48E6FE5CF7AF" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[284,497,2028,2049]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="5" pageNumber="87" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFE9F44CFECD48E6FE5CF7AF" box="[284,497,2028,2049]" italics="true" pageId="5" pageNumber="87">Syrioiulus lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFE9F44CFE2448E6FDECF7AF" box="[501,577,2028,2049]" pageId="5" pageNumber="87" rank="species">sp. nov.</taxonomicNameLabel>
, gonopods of male holotype, in MHNG. (15) Distal parts of left opisthomere, caudo-lateral view. (16) Solenomere, same aspect.
</paragraph>
</caption>
<paragraph id="CD1336E8FFEAF44FFCEE4E8BFA3DFD5B" blockId="6.[831,1437,258,757]" pageId="6" pageNumber="88">
<emphasis id="FFD8EAFAFFEAF44FFCEE4E8BFBF8FE36" box="[831,1109,385,408]" italics="true" pageId="6" pageNumber="88">Female sexual characters</emphasis>
: Leg-pair 1 considerably, legpair 2 slightly flattened antero-caudally. Vulva (
<figureCitation id="55972A6DFFEAF44FFA944EAAFA39FE19" box="[1349,1428,416,439]" captionStart="Figs 5-10" captionStartId="4.[151,192,1539,1560]" captionTargetBox="[198,1386,263,1488]" captionTargetId="figure-212@4.[515,880,803,1077]" captionTargetPageId="4" captionText="Figs 5-10. Syrioiulus lohmanderi sp. nov., structures of male holotype (5-9) and female paratype (10), in MHNG. (5) Antenna. (6) Leg-pair 1, frontal view. (7) Leg 2. (8) Leg 3. (9) Right ventral edge of pleurotergum 7, ventro-lateral view. (10) Right vulva, caudal, slightly mesal view.Abbreviations: ct = central tube; lv = lateral valve; ma = mesal ampulla; mt = mesal tube; mv = mesal valve; op = operculum. Scale bar of Fig. 8: 0.2 mm; the rest: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5743577" httpUri="https://zenodo.org/record/5743577/files/figure.png" pageId="6" pageNumber="88">Fig. 10</figureCitation>
) strongly compressed in the sagittal plane; bursa mostly symmetric: lateral valve (
<emphasis id="FFD8EAFAFFEAF44FFBB14EEAFBDEFE59" box="[1120,1139,480,503]" italics="true" pageId="6" pageNumber="88">lv</emphasis>
) slightly higher and more narrow than mesal one (
<emphasis id="FFD8EAFAFFEAF44FFBB14D0AFBD2FDB9" box="[1120,1151,512,535]" italics="true" pageId="6" pageNumber="88">mv</emphasis>
); median cleft relatively broad; operculum (
<emphasis id="FFD8EAFAFFEAF44FFBC64D15FB9FFD98" box="[1047,1074,543,566]" italics="true" pageId="6" pageNumber="88">op</emphasis>
) by far exceeding bursa, apical margin with a distinct mid-lateral incision and with several bumps along mesal section; setation moderately dense, randomly spread across whole surface of vulva. Receptaculum seminis consisting of a narrow, nearly straight central tube (
<emphasis id="FFD8EAFAFFEAF44FFBFC4DB4FBEDFD7B" box="[1069,1088,702,725]" italics="true" pageId="6" pageNumber="88">ct</emphasis>
) supporting a very short mesal tube (
<emphasis id="FFD8EAFAFFEAF44FFCAC4DD4FC35FD5B" box="[893,920,734,757]" italics="true" pageId="6" pageNumber="88">mt</emphasis>
) ending in a small, ovoid, mesal ampulla (
<emphasis id="FFD8EAFAFFEAF44FFAB14DD4FA2CFD5B" box="[1376,1409,734,757]" italics="true" pageId="6" pageNumber="88">ma</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="85B66563FFEAF44FFC914C06FBC7FBEF" pageId="6" pageNumber="88" type="discussion">
<paragraph id="CD1336E8FFEAF44FFC914C06FAC0FC6C" blockId="6.[832,1443,780,1089]" pageId="6" pageNumber="88">
<emphasis id="FFD8EAFAFFEAF44FFC914C06FC1DFC8D" bold="true" box="[832,944,780,803]" pageId="6" pageNumber="88">Remarks:</emphasis>
The
<typeStatus id="1217884AFFEAF44FFC3C4C06FBB6FC8D" box="[1005,1051,780,803]" pageId="6" pageNumber="88">type</typeStatus>
locality of the new species lies in the foothills of the
<collectingRegion id="0F68F80AFFEAF44FFBF64C26FB5BFCED" box="[1063,1270,812,835]" country="Iran" name="Alborz" pageId="6" pageNumber="88">Alborz Mountains</collectingRegion>
, the mountain range where most
<taxonomicName id="0AAC4D6BFFEAF44FFBC84C46FB2BFCCD" authorityName="Verhoeff" authorityYear="1914" box="[1049,1158,844,867]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="genus">
<emphasis id="FFD8EAFAFFEAF44FFBC84C46FB2BFCCD" box="[1049,1158,844,867]" italics="true" pageId="6" pageNumber="88">Syrioiulus</emphasis>
</taxonomicName>
species from
<collectingCountry id="B5BB7678FFEAF44FFAFE4C46FAF7FCCD" box="[1327,1370,844,867]" name="Iran" pageId="6" pageNumber="88">Iran</collectingCountry>
were exclusively recorded (
<figureCitation id="55972A6DFFEAF44FFBE94C66FB20FC2D" box="[1080,1165,876,899]" captionStart="Fig" captionStartId="8.[151,186,1806,1827]" captionTargetBox="[151,1435,257,1760]" captionTargetId="figure-4@8.[151,1436,257,1760]" captionTargetPageId="8" captionText="Fig. 18. Localities of Syrioiulus spp. in Iran. Ring: S. astrabadensis; cross: S. continentalis; square: S. discolor; pentagon: S. incarnatus; star: S. lohmanderi sp. nov.; filled circle: S. persicus; triangle: S. zarudnyi." figureDoi="http://doi.org/10.5281/zenodo.5743591" httpUri="https://zenodo.org/record/5743591/files/figure.png" pageId="6" pageNumber="88">Fig. 18</figureCitation>
). The only exception is
<taxonomicName id="0AAC4D6BFFEAF44FFC914C81FB05FC0C" authority="(Lohmander, 1932 a)" baseAuthorityName="Lohmander" baseAuthorityYear="1932" box="[832,1192,907,930]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="zarudnyi">
<emphasis id="FFD8EAFAFFEAF44FFC914C81FC10FC0C" box="[832,957,907,930]" italics="true" pageId="6" pageNumber="88">S. zarudnyi</emphasis>
(
<bibRefCitation id="A93D4B19FFEAF44FFC1E4C81FB0DFC0C" author="Lohmander H." box="[975,1184,907,930]" pageId="6" pageNumber="88" pagination="1 - 44" refId="ref7038" refString="Lohmander H. 1932 a. Neue Diplopoden aus Persien. Goteborgs Kungliga Vetenskaps- och Vitterhets-Samhalles handlingar 5, Ser. B 3 (2): 1 - 44." type="journal article" year="1932">Lohmander, 1932a</bibRefCitation>
)
</taxonomicName>
which is known also from two localities in the central part of the country.
</paragraph>
<paragraph id="CD1336E8FFEAF44FFC914CC1FBC7FBEF" blockId="6.[832,1443,780,1089]" pageId="6" pageNumber="88">
Considering the absence of ommatidia and the method (sifting dead organic matter) by which the studied specimens were collected,
<taxonomicName id="0AAC4D6BFFEAF44FFBA44B00FABEFB8F" authority="Vagalinski, 2020" authorityName="Vagalinski" authorityYear="2020" box="[1141,1299,1034,1057]" class="Diplopoda" family="Julidae" genus="Syrioiulus" kingdom="Animalia" order="Julida" pageId="6" pageNumber="88" phylum="Arthropoda" rank="species" species="lohmanderi" status="sp. nov.">
<emphasis id="FFD8EAFAFFEAF44FFBA44B00FABEFB8F" box="[1141,1299,1034,1057]" italics="true" pageId="6" pageNumber="88">S. lohmanderi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E4EB5781FFEAF44FFAF34B00FAD0FB8F" box="[1314,1405,1034,1057]" pageId="6" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
is likely an euedaphic species.
</paragraph>
</subSubSection>
</treatment>
</document>
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