treatments-xml/data/34/44/87/344487B2FFC5FFB25FE6FC8EFB0BFCA6.xml
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<document id="89DD0D09BDA7504853D815935E201121" ID-DOI="10.1080/00222933.2013.798699" ID-ISSN="1464-5262" ID-Zenodo-Dep="5197590" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628882222048" checkinUser="felipe" docAuthor="Kramer, Bernd &amp; Wink, Michael" docDate="2013" docId="344487B2FFC5FFB25FE6FC8EFB0BFCA6" docLanguage="en" docName="JNatHist.47.35-36.2327-2362.pdf" docOrigin="Journal of Natural History (J. Nat. Hist.) 47 (35 - 36)" docSource="http://dx.doi.org/10.1080/00222933.2013.798699" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Marcusenius multisquamatus Kramer &amp; Wink 2013, sp. nov." docType="treatment" docUuid="CE3474C9-C821-420C-9548-0D8F917BFCB9" docUuidSource="ZooBank" docVersion="5" lastPageNumber="2343" masterDocId="C87DFFCAFFCBFFA05E00FFCBFF9BFFB8" masterDocTitle="East-west differentiation in the Marcusenius macrolepidotus species complex in Southern Africa: the description of a new species for the lower Cunene River, Namibia (Teleostei: Mormyridae)" masterLastPageNumber="2362" masterPageNumber="2327" pageNumber="2339" updateTime="1699054295763" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E66E1ABD1C48EFC6E5F0C2B4806B4773">East-west differentiation in the Marcusenius macrolepidotus species complex in Southern Africa: the description of a new species for the lower Cunene River, Namibia (Teleostei: Mormyridae)</mods:title>
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<mods:name id="539E27BD6122F2EB4E60A04B69E0F7A2" type="personal">
<mods:role id="3A4D71E5E785D226170DF570E8FCF862">
<mods:roleTerm id="A99A95570CF6F7FE16529D6DE3DD61E7">Author</mods:roleTerm>
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<mods:namePart id="5BA80085911347ED8ED441C1D185F5D0">Kramer, Bernd</mods:namePart>
<mods:affiliation id="71BD4D37D85B03C1CF9917A0FAECEA1B">University of Regensburg, Zoological Institute, Universitätsstrasse 31, D- 93040 Regensburg, Germany;</mods:affiliation>
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<mods:roleTerm id="7E78D74BD6FC5E2E3FF25755D5F1A63D">Author</mods:roleTerm>
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<mods:namePart id="6B2E493DA4E832E03B0792A5CD69909D">Wink, Michael</mods:namePart>
<mods:affiliation id="36910D4B4C6D372B11E2B849DC399A8E">Heidelberg University, Institute of Pharmacy and Molecular Biotechnology, Im Neuenheimer Feld 364, D- 69120 Heidelberg, Germany</mods:affiliation>
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<mods:title id="6535EDCF57F6D51E0A997B93E38FFA8B">Journal of Natural History</mods:title>
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<mods:part id="26EDFA43E246F73FF0C7ED7F87FCF3C7">
<mods:date id="9F8DCC9616C5D41F9906F47E8F6EAA4C">2013</mods:date>
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<mods:title id="F9962DE6B19AD1F1A0CB40BA3C449B9D">J. Nat. Hist.</mods:title>
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<mods:number id="E909154551FF7A84050A710F57082423">2013-07-18</mods:number>
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<mods:number id="A96B1EF12536900AD09953B9ACFECE63">47</mods:number>
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<mods:number id="1B1EAE4629C01913DB38A56796FA3F5B">35 - 36</mods:number>
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<mods:url id="A1D66410F2149A0974CB6E229DFE9812">http://dx.doi.org/10.1080/00222933.2013.798699</mods:url>
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<subSubSection id="F4F7652FFFC5FFAE5FE6FC8EFC1AFCE5" box="[486,897,836,861]" pageId="14" pageNumber="2339" type="nomenclature">
<paragraph id="BC5236A4FFC5FFAE5FE6FC8EFC1AFCE5" blockId="14.[486,897,836,893]" box="[486,897,836,861]" pageId="14" pageNumber="2339">
<heading id="E71A81C8FFC5FFAE5FE6FC8EFC1AFCE5" box="[486,897,836,861]" centered="true" fontSize="10" level="2" pageId="14" pageNumber="2339" reason="2">
<taxonomicName id="7BED4D27FFC5FFAE5FE6FC8EFCBDFCE5" ID-CoL="3Y39B" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[486,806,836,861]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="14" pageNumber="2339" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFC5FFAE5FE6FC8EFCBDFCE5" bold="true" box="[486,806,836,861]" italics="true" pageId="14" pageNumber="2339">Marcusenius multisquamatus</emphasis>
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<taxonomicNameLabel id="95AA57CDFFC5FFAE5D2DFC8DFC1AFCE5" box="[813,897,838,861]" pageId="14" pageNumber="2339" rank="species">sp. nov.</taxonomicNameLabel>
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(
<figureCitation id="24D62A21FFC5FFAE5C7BFCAEFD70FCC5" box="[635,747,869,893]" captionStart="Figure 4" captionStartId="11.[143,210,1506,1528]" captionTargetBox="[168,1081,161,1473]" captionTargetId="figure-6@11.[168,1081,161,1473]" captionTargetPageId="11" captionText="Figure 4. Photographs of bulldog fish from various origins. (A) Marcusenius macrolepidotus (Peters, 1852), SAIAB 60847, coll. R. Bills 1 August 1999, Lower Zambezi; (B) Marcusenius angolensis (Boulenger, 1905), BMNH 1905.5.29.64 (holotype); (C) Marcusenius altisambesi, coll. F. H. van der Bank and B. Kramer, 21 August 1999, Upper Zambezi, Kalimbeza, live specimen of 16.5 cm SL photographed 3 July 2003 in aquarium; (D) M. altisambesi, coll. F.H. van der Bank and B. Kramer, 11/12 August 2004, Okavango River, live fish of SL 13 cm photographed 20 April 2006 in aquarium; (E) Marcusenius multisquamatus sp. nov., coll. B. Kramer and Ernst Swartz, coll. and photographed on 19 August 2006, Cunene River, below Ruacana Falls, specimen RUAC01, SL 15.4 cm." figureDoi="http://doi.org/10.5281/zenodo.5197598" httpUri="https://zenodo.org/record/5197598/files/figure.png" pageId="14" pageNumber="2339">Figure 4E</figureCitation>
)
</paragraph>
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<emphasis id="8E99EAB6FFC5FFAE5ED3FC09FEE5FC61" box="[211,382,962,985]" italics="true" pageId="14" pageNumber="2339">Type specimens</emphasis>
</heading>
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<paragraph id="BC5236A4FFC5FFAE5ED3FC22FDDDFA98" blockId="14.[211,1172,962,1726]" pageId="14" pageNumber="2339">
<materialsCitation id="0C853CF9FFC5FFAE5ED3FC22FB0BFBD9" collectingDate="2006-08-15" collectionCode="SAIAB, KUNE" collectorName="E. Swartz &amp; Swartz, B &amp; L. da Costa" country="Namibia" county="Falls" elevation="300" latitude="-17.001944" location="Cunene River" longLatPrecision="21" longitude="13.2491665" municipality="Hot Springs Campsite" pageId="14" pageNumber="2339" specimenCode="SAIAB 78781, KUNE24" specimenCount="1" typeStatus="holotype">
<typeStatus id="63568806FFC5FFAE5ED3FC22FED9FBB9" box="[211,322,1001,1025]" pageId="14" pageNumber="2339" type="holotype">Holotype</typeStatus>
:
<specimenCode id="EC4B9EDFFFC5FFAE5F4CFC22FE73FBB9" box="[332,488,1001,1025]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="14" pageNumber="2339" type="Other type of research institution/biorepository">SAIAB 78781</specimenCode>
(field no.
<specimenCode id="EC4B9EDFFFC5FFAE5C58FC22FD51FBB9" box="[600,714,1001,1025]" collectionCode="KUNE" country="China" lsid="urn:lsid:biocol.org:col:13450" name="Kunming Institute of Ecology, Academia Sinica" pageId="14" pageNumber="2339" type="Herbarium">KUNE24</specimenCode>
), live SL
<quantity id="7B159B41FFC5FFAE5D31FC22FC11FBB9" box="[817,906,1001,1025]" metricMagnitude="-1" metricUnit="m" metricValue="2.09" pageId="14" pageNumber="2339" unit="cm" value="20.9">20.9 cm</quantity>
, fixed SL
<quantity id="7B159B41FFC5FFAE5DF9FC22FBC9FBB9" box="[1017,1106,1001,1025]" metricMagnitude="-1" metricUnit="m" metricValue="2.02" pageId="14" pageNumber="2339" unit="cm" value="20.2">20.2 cm</quantity>
, fixed TL
<quantity id="7B159B41FFC5FFAE5EFEFBC2FEC2FB99" box="[254,345,1033,1057]" metricMagnitude="-1" metricUnit="m" metricValue="2.29" pageId="14" pageNumber="2339" unit="cm" value="22.9">22.9 cm</quantity>
, male,
<collectingCountry id="C4FA7634FFC5FFAE5FA8FBC2FD94FB99" box="[424,527,1033,1057]" name="Namibia" pageId="14" pageNumber="2339">Namibia</collectingCountry>
:
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:
<location id="B932607FFFC5FFAE5CC5FBC1FCCBFB99" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC5FFAE5CC5FBC1FCCBFB99" box="[709,848,1033,1057]" country="Namibia" county="Falls" latitude="-17.001944" longLatPrecision="21" longitude="13.2491665" municipality="Hot Springs Campsite" name="Epupa Falls" pageId="14" pageNumber="2339">Epupa Falls</location>
,
<collectingMunicipality id="5C36ACDEFFC5FFAE5D5BFBC1FBC2FB99" box="[859,1113,1033,1057]" pageId="14" pageNumber="2339">Hot Springs Campsite</collectingMunicipality>
, estimated
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</quantity>
upstream from the
<collectingCounty id="55334E28FFC5FFAE5C40FBE2FDE3FBF9" box="[576,632,1065,1089]" pageId="14" pageNumber="2339">Falls</collectingCounty>
,
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17
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00
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07
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S
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,
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13
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14
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57
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E
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, about
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altitude,
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<collectingDate id="D817E98CFFC5FFAE5ED3FB82FE1DFBD9" box="[211,390,1097,1121]" pageId="14" pageNumber="2339" value="2006-08-15">15 August 2006</collectingDate>
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, coll.
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, B. Kramer and
<collectorName id="11185372FFC5FFAE5CF8FB81FCE4FBD9" box="[760,895,1097,1121]" pageId="14" pageNumber="2339">L. da Costa</collectorName>
at ≤
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<elevation id="37C0D197FFC5FFAE5DBFFB82FC64FBD9" box="[959,1023,1097,1121]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="2339" unit="m" value="1.5">1.5 m</elevation>
</quantity>
water depth
</materialsCitation>
.
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<typeStatus id="63568806FFC5FFAE5ED3FBA2FEDEFB39" box="[211,325,1129,1153]" pageId="14" pageNumber="2339" type="paratype">Paratypes</typeStatus>
:
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(2)
</materialsCitation>
,
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</materialsCitation>
;
<materialsCitation id="0C853CF9FFC5FFAE5CBEFBA2FCF3FB39" accessionNumber="ZSM 38526" box="[702,872,1129,1153]" collectionCode="ZSM" country="Namibia" pageId="14" pageNumber="2339" specimenCount="1" typeStatus="paratype">
<collectionCode id="DAFCAE61FFC5FFAE5CBEFBA2FD62FB39" box="[702,761,1129,1153]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
38526 (2)
</materialsCitation>
,
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<collectionCode id="DAFCAE61FFC5FFAE5D72FBA2FC36FB39" box="[882,941,1129,1153]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
38527 (2)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC5FFAE5A26FBA2FDD9FA98" collectingDate="2006-08-12" collectingDateMax="2006-08-17" collectingDateMin="2006-08-12" collectorName="E. Swartz &amp; B. Kramer &amp; L. da Costa" country="Namibia" county="Cunene River" elevation="300" latitude="-17.001944" location="Cunene River" longLatPrecision="21" longitude="13.2491665" municipality="Falls" pageId="14" pageNumber="2339" specimenCount="1" typeStatus="paratype">
size range
<quantity id="7B159B41FFC5FFAE5ED3FB43FEF2FB18" box="[211,361,1160,1184]" metricMagnitude="-1" metricUnit="m" metricValue="1.515" metricValueMax="2.02" metricValueMin="1.01" pageId="14" pageNumber="2339" unit="cm" value="15.149999999999999" valueMax="20.2" valueMin="10.1">10.120.2 cm</quantity>
SL,
<collectingCountry id="C4FA7634FFC5FFAE5FA1FB43FD93FB18" box="[417,520,1160,1184]" name="Namibia" pageId="14" pageNumber="2339">Namibia</collectingCountry>
:
<location id="B932607FFFC5FFAE5C14FB43FD2EFB18" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC5FFAE5C14FB43FD2EFB18" box="[532,693,1160,1184]" country="Namibia" county="Cunene River" latitude="-17.001944" longLatPrecision="21" longitude="13.2491665" municipality="Falls" name="Cunene River" pageId="14" pageNumber="2339">Cunene River</location>
:
<location id="B932607FFFC5FFAE5CC0FB42FCD7FB18" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC5FFAE5CC0FB42FCD7FB18" box="[704,844,1160,1184]" country="Namibia" county="Cunene River" latitude="-17.001944" longLatPrecision="21" longitude="13.2491665" municipality="Falls" name="Epupa Falls" pageId="14" pageNumber="2339">Epupa Falls</location>
, Hot Springs Campsite, estimated
<quantity id="7B159B41FFC5FFAE5F1FFB63FEFFFB78" box="[287,356,1192,1216]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="2339" unit="m" value="300.0">
<elevation id="37C0D197FFC5FFAE5F1FFB63FEFFFB78" box="[287,356,1192,1216]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="2339" unit="m" value="300.0">300 m</elevation>
</quantity>
upstream from the
<collectingMunicipality id="5C36ACDEFFC5FFAE5C40FB63FDE3FB78" box="[576,632,1192,1216]" pageId="14" pageNumber="2339">Falls</collectingMunicipality>
,
<geoCoordinate id="D9D95063FFC5FFAE5C83FB63FC9CFB78" box="[643,775,1188,1216]" degrees="17" direction="south" minutes="00" orientation="latitude" pageId="14" pageNumber="2339" precision="15" seconds="07" value="-17.001944">
17
<superScript id="4B989BECFFC5FFAE5C9DFB6FFD3DFB0D" attach="left" box="[669,678,1188,1205]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
00
<superScript id="4B989BECFFC5FFAE5CC2FB6FFD5CFB0D" attach="left" box="[706,711,1188,1205]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
07
<superScript id="4B989BECFFC5FFAE5CE3FB6FFD75FB0D" attach="left" box="[739,750,1188,1205]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC5FFAE5D12FB63FC01FB78" box="[786,922,1188,1216]" degrees="13" direction="east" minutes="14" orientation="longitude" pageId="14" pageNumber="2339" precision="15" seconds="57" value="13.2491665">
13
<superScript id="4B989BECFFC5FFAE5D2CFB6FFCAEFB0D" attach="left" box="[812,821,1188,1205]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
14
<superScript id="4B989BECFFC5FFAE5D51FB6FFCCDFB0D" attach="left" box="[849,854,1188,1205]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
57
<superScript id="4B989BECFFC5FFAE5D72FB6FFCE6FB0D" attach="left" box="[882,893,1188,1205]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
E
</geoCoordinate>
, about
<quantity id="7B159B41FFC5FFAE5DECFB63FBAAFB78" box="[1004,1073,1192,1216]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="2339" unit="m" value="600.0">
<elevation id="37C0D197FFC5FFAE5DECFB63FBAAFB78" box="[1004,1073,1192,1216]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="14" pageNumber="2339" unit="m" value="600.0">600 m</elevation>
</quantity>
altitude, coll.
<collectorName id="11185372FFC5FFAE5F0CFB02FEE5FB58" box="[268,382,1224,1248]" pageId="14" pageNumber="2339">E. Swartz</collectorName>
,
<collectorName id="11185372FFC5FFAE5F8CFB02FD9CFB58" box="[396,519,1225,1248]" pageId="14" pageNumber="2339">B. Kramer</collectorName>
, and
<collectorName id="11185372FFC5FFAE5C47FB02FD4BFB58" box="[583,720,1224,1248]" pageId="14" pageNumber="2339">L. da Costa</collectorName>
at ≤
<quantity id="7B159B41FFC5FFAE5D14FB03FCCEFB58" box="[788,853,1224,1248]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="2339" unit="m" value="1.5">
<elevation id="37C0D197FFC5FFAE5D14FB03FCCEFB58" box="[788,853,1224,1248]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="2339" unit="m" value="1.5">1.5 m</elevation>
</quantity>
water depth,
<collectingCounty id="55334E28FFC5FFAE5DF6FB03FB08FB58" box="[1014,1171,1224,1248]" pageId="14" pageNumber="2339">Cunene River</collectingCounty>
water at Hot Springs: Saturday,
<date id="C8531064FFC5FFAE5C42FB23FD6EFAB8" box="[578,757,1256,1280]" pageId="14" pageNumber="2339" value="2006-08-12">
<collectingDate id="D817E98CFFC5FFAE5C42FB23FD6EFAB8" box="[578,757,1256,1280]" pageId="14" pageNumber="2339" value="2006-08-12">12 August 2006</collectingDate>
</date>
, 12.50 h: 19.9
<superScript id="4B989BECFFC5FFAE5D9BFB2FFC3FFB4D" attach="right" box="[923,932,1252,1269]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
C, 48 µS
<superScript id="4B989BECFFC5FFAE5A13FB22FBD7FB4D" attach="left" box="[1043,1100,1252,1280]" fontSize="7" pageId="14" pageNumber="2339">cm1</superScript>
, from 11 Augure 2006
<date id="C8531064FFC5FFAE5F93FAC3FDD9FA98" box="[403,578,1288,1312]" pageId="14" pageNumber="2339" value="2006-08-17">
<collectingDate id="D817E98CFFC5FFAE5F93FAC3FDD9FA98" box="[403,578,1288,1312]" pageId="14" pageNumber="2339" value="2006-08-17">17 August 2006</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFC5FFAE5EFEFAE2FBE5FA67" blockId="14.[211,1172,962,1726]" pageId="14" pageNumber="2339">
<emphasis id="8E99EAB6FFC5FFAE5EFEFAE2FEF4FAF8" box="[254,367,1321,1344]" italics="true" pageId="14" pageNumber="2339">Non-types</emphasis>
.
<materialsCitation id="0C853CF9FFC5FFAE5F7DFAE3FDD8FAF8" box="[381,579,1320,1344]" collectionCode="SAIAB" pageId="14" pageNumber="2339" specimenCode="SAIAB 78785" specimenCount="1">
<specimenCode id="EC4B9EDFFFC5FFAE5F7DFAE3FD81FAF8" box="[381,538,1320,1344]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="14" pageNumber="2339" type="Other type of research institution/biorepository">SAIAB 78785</specimenCode>
(2)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC5FFAE5C4EFAE3FC8FFAF8" box="[590,788,1320,1344]" collectionCode="SAIAB" pageId="14" pageNumber="2339" specimenCode="SAIAB 78789" specimenCount="1">
<specimenCode id="EC4B9EDFFFC5FFAE5C4EFAE3FD70FAF8" box="[590,747,1320,1344]" collectionCode="SAIAB" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34984" lsid="urn:lsid:biocol.org:col:34984" name="South African Institute for Aquatic Biodiversity" pageId="14" pageNumber="2339" type="Other type of research institution/biorepository">SAIAB 78789</specimenCode>
(2)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC5FFAE5D20FAE3FC30FAF8" accessionNumber="ZSM 38528" box="[800,939,1320,1344]" collectionCode="ZSM" pageId="14" pageNumber="2339" specimenCount="1">
<collectionCode id="DAFCAE61FFC5FFAE5D20FAE3FCC0FAF8" box="[800,859,1320,1344]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
38528,
</materialsCitation>
<materialsCitation id="0C853CF9FFC5FFAE5DB2FAE3FBC5FAF8" accessionNumber="ZSM 38529" box="[946,1118,1320,1344]" collectionCode="ZSM" pageId="14" pageNumber="2339" specimenCount="1">
<collectionCode id="DAFCAE61FFC5FFAE5DB2FAE3FC76FAF8" box="[946,1005,1320,1344]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
38529 (2)
</materialsCitation>
,
<materialsCitation id="0C853CF9FFC5FFAE5A6AFAE3FBE1FA67" collectingDate="2004-08-19" collectingDateMax="2006-08-23" collectingDateMin="2004-08-19" collectorName="E. Swartz &amp; B. Kramer" elevation="800" latitude="-17.406666" location="Falls" longLatPrecision="21" longitude="14.216944" municipality="Ruacana Falls" pageId="14" pageNumber="2339" specimenCount="1" stateProvince="Cunene">
size range
<quantity id="7B159B41FFC5FFAE5F1CFA83FE2FFAD8" box="[284,436,1352,1376]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" metricValueMax="1.54" metricValueMin="1.16" pageId="14" pageNumber="2339" unit="cm" value="13.5" valueMax="15.4" valueMin="11.6">11.615.4 cm</quantity>
SL (live),
<collectingMunicipality id="5C36ACDEFFC5FFAE5C31FA82FD4CFAD8" box="[561,727,1352,1376]" pageId="14" pageNumber="2339">Ruacana Falls</collectingMunicipality>
, Hippo Pool Campsite, just below the
<location id="B932607FFFC5FFAE5ED3FAA3FE90FA38" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC5FFAE5ED3FAA3FE90FA38" box="[211,267,1384,1408]" latitude="-17.406666" longLatPrecision="21" longitude="14.216944" municipality="Ruacana Falls" name="Falls" pageId="14" pageNumber="2339" stateProvince="Cunene">Falls</location>
,
<geoCoordinate id="D9D95063FFC5FFAE5F17FAA3FE07FA38" box="[279,412,1380,1408]" degrees="17" direction="south" minutes="24" orientation="latitude" pageId="14" pageNumber="2339" precision="15" seconds="24" value="-17.406666">
17
<superScript id="4B989BECFFC5FFAE5F31FAAFFEA1FACD" attach="left" box="[305,314,1380,1397]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
24
<superScript id="4B989BECFFC5FFAE5F56FAAFFEC0FACD" attach="left" box="[342,347,1380,1397]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
24
<superScript id="4B989BECFFC5FFAE5F77FAAFFE19FACD" attach="left" box="[375,386,1380,1397]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC5FFAE5FA8FAA3FDA9FAC7" box="[424,562,1380,1408]" degrees="14" direction="east" minutes="13" orientation="longitude" pageId="14" pageNumber="2339" precision="15" seconds="01" value="14.216944">
14
<superScript id="4B989BECFFC5FFAE5FC2FAAFFE50FACD" attach="left" box="[450,459,1380,1397]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
1301” E
</geoCoordinate>
, about
<quantity id="7B159B41FFC5FFAE5C87FAA3FD56FAC7" box="[647,717,1384,1408]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="2339" unit="m" value="800.0">
<elevation id="37C0D197FFC5FFAE5C87FAA3FD56FAC7" box="[647,717,1384,1408]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="2339" unit="m" value="800.0">800 m</elevation>
</quantity>
altitude, coll.
<collectorName id="11185372FFC5FFAE5D6FFAA3FC47FA38" box="[879,988,1384,1408]" pageId="14" pageNumber="2339">E. Swartz</collectorName>
and
<collectorName id="11185372FFC5FFAE5A16FAA3FB0BFAC7" box="[1046,1168,1384,1407]" pageId="14" pageNumber="2339">B. Kramer</collectorName>
, at ≤
<quantity id="7B159B41FFC5FFAE5F09FA4CFEDCFA27" box="[265,327,1415,1439]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="2339" unit="m" value="1.5">
<elevation id="37C0D197FFC5FFAE5F09FA4CFEDCFA27" box="[265,327,1415,1439]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="2339" unit="m" value="1.5">1.5 m</elevation>
</quantity>
water depth,
<collectingRegion id="7E29F846FFC5FFAE5FDFFA4CFDAEFA27" box="[479,565,1415,1439]" country="Angola" name="Cunene" pageId="14" pageNumber="2339">Cunene</collectingRegion>
River water at Hippo Pool: Saturday,
<date id="C8531064FFC5FFAE5DDFFA4CFB14FA27" box="[991,1167,1415,1439]" pageId="14" pageNumber="2339" value="2004-08-19">
<collectingDate id="D817E98CFFC5FFAE5DDFFA4CFB14FA27" box="[991,1167,1415,1439]" pageId="14" pageNumber="2339" value="2004-08-19">19 August 2004</collectingDate>
</date>
, 10.00 h: 21.1
<superScript id="4B989BECFFC5FFAE5F74FA6FFEE6FA0D" attach="right" box="[372,381,1444,1461]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
C, 45.8 µS
<superScript id="4B989BECFFC5FFAE5C08FA63FDDAFA0C" attach="left" box="[520,577,1443,1471]" fontSize="7" pageId="14" pageNumber="2339">cm1</superScript>
;
<date id="C8531064FFC5FFAE5C55FA6CFC96FA07" box="[597,781,1447,1471]" pageId="14" pageNumber="2339" value="2006-08-20">
<collectingDate id="D817E98CFFC5FFAE5C55FA6CFC96FA07" box="[597,781,1447,1471]" pageId="14" pageNumber="2339" value="2006-08-20">20 August 2006</collectingDate>
</date>
, 10.00 h, 19.8
<superScript id="4B989BECFFC5FFAE5DBEFA6FFC5CFA0D" attach="right" box="[958,967,1444,1461]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
C, 45.4 µS
<superScript id="4B989BECFFC5FFAE5A51FA63FB11FA0C" attach="left" box="[1105,1162,1443,1471]" fontSize="7" pageId="14" pageNumber="2339">cm1</superScript>
; 21 August, 10.18 h, 19.4
<superScript id="4B989BECFFC5FFAE5FE8FA08FE6AFA6C" attach="right" box="[488,497,1475,1492]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
C, 44.2 µS
<superScript id="4B989BECFFC5FFAE5C70FA03FD32FA6C" attach="left" box="[624,681,1475,1503]" fontSize="7" pageId="14" pageNumber="2339">cm1</superScript>
; from
<date id="C8531064FFC5FFAE5CF6FA0CFC38FA67" box="[758,931,1479,1503]" pageId="14" pageNumber="2339" value="2006-08-18">
<collectingDate id="D817E98CFFC5FFAE5CF6FA0CFC38FA67" box="[758,931,1479,1503]" pageId="14" pageNumber="2339" value="2006-08-18">18 August 2006</collectingDate>
</date>
to
<date id="C8531064FFC5FFAE5DC8FA0CFBE1FA67" box="[968,1146,1479,1503]" pageId="14" pageNumber="2339" value="2006-08-23">
<collectingDate id="D817E98CFFC5FFAE5DC8FA0CFBE1FA67" box="[968,1146,1479,1503]" pageId="14" pageNumber="2339" value="2006-08-23">23 August 2006</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFC5FFAE5EFEFA2CFD30F906" blockId="14.[211,1172,962,1726]" pageId="14" pageNumber="2339">
<materialsCitation id="0C853CF9FFC5FFAE5EFEFA2CFDA8FA47" accessionNumber="ZSM 41761" box="[254,563,1511,1535]" collectionCode="ZSM" pageId="14" pageNumber="2339" specimenCount="1">
<collectionCode id="DAFCAE61FFC5FFAE5EFEFA2CFEA2FA47" box="[254,313,1511,1535]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
41761 (11), specimens
</materialsCitation>
<materialsCitation id="0C853CF9FFC5FFAE5C39FA2CFDC3FA47" box="[569,600,1511,1535]" collectionCode="R" pageId="14" pageNumber="2339" specimenCode="R1" specimenCount="1">
<specimenCode id="EC4B9EDFFFC5FFAE5C39FA2CFDC3FA47" box="[569,600,1511,1535]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="2339">R1</specimenCode>
</materialsCitation>
<materialsCitation id="0C853CF9FFC5FFAE5C69FA2CFB9AF9A7" collectingDate="2009-12-15" collectionCode="R" collectorName="F. H. van der Bank" elevation="2" latitude="-17.2601" location="River" longLatPrecision="1" longitude="11.764867" pageId="14" pageNumber="2339" specimenCode="R11" specimenCount="1" stateProvince="Cunene">
<specimenCode id="EC4B9EDFFFC5FFAE5C69FA2CFD01FA47" box="[617,666,1511,1535]" collectionCode="R" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="2339">R11</specimenCode>
, from the
<collectingRegion id="7E29F846FFC5FFAE5D0AFA2CFCFBFA47" box="[778,864,1511,1535]" country="Angola" name="Cunene" pageId="14" pageNumber="2339">Cunene</collectingRegion>
<location id="B932607FFFC5FFAE5D67FA2CFC3EFA47" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC5FFAE5D67FA2CFC3EFA47" box="[871,933,1511,1535]" latitude="-17.2601" longLatPrecision="1" longitude="11.764867" name="River" pageId="14" pageNumber="2339" stateProvince="Cunene">River</location>
mouth,
<geoCoordinate id="D9D95063FFC5FFAE5A03FA2CFB14FA47" box="[1027,1167,1507,1535]" degrees="17" direction="south" minutes="15.606" orientation="latitude" pageId="14" pageNumber="2339" precision="1" value="-17.2601">
17
<superScript id="4B989BECFFC5FFAE5A1EFA28FBBCFA4C" attach="none" box="[1054,1063,1507,1524]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
15.606
<superScript id="4B989BECFFC5FFAE5A72FA28FBECFA4C" attach="left" box="[1138,1143,1507,1524]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC5FFAE5ED3F9CCFEFEF9A7" box="[211,357,1539,1567]" degrees="11" direction="east" minutes="45.892" orientation="longitude" pageId="14" pageNumber="2339" precision="1" value="11.764867">
11
<superScript id="4B989BECFFC5FFAE5EEEF9C8FF6CF9AC" attach="none" box="[238,247,1539,1556]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
45.892
<superScript id="4B989BECFFC5FFAE5F42F9C8FEDCF9AC" attach="left" box="[322,327,1539,1556]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
E
</geoCoordinate>
, altitude
<quantity id="7B159B41FFC5FFAE5FD1F9CCFD9BF9A7" box="[465,512,1543,1567]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="2339" unit="m" value="2.0">
<elevation id="37C0D197FFC5FFAE5FD1F9CCFD9BF9A7" box="[465,512,1543,1567]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="2339" unit="m" value="2.0">2 m</elevation>
</quantity>
,
<date id="C8531064FFC5FFAE5C0CF9CCFD78F9A7" box="[524,739,1543,1567]" pageId="14" pageNumber="2339" value="2009-12-15">
<collectingDate id="D817E98CFFC5FFAE5C0CF9CCFD78F9A7" box="[524,739,1543,1567]" pageId="14" pageNumber="2339" value="2009-12-15">15 December 2009</collectingDate>
</date>
, coll.
<collectorName id="11185372FFC5FFAE5D28F9C3FB9AF9A7" box="[808,1025,1543,1567]" pageId="14" pageNumber="2339">F.H. van der Bank</collectorName>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC5FFAE5A0EF9CCFB0BF9F8" accessionNumber="ZSM 41762" collectingDate="2010-11-08" collectionCode="ZSM" collectorName="S. Voges" latitude="-17.272083" longLatPrecision="1" longitude="11.7862835" pageId="14" pageNumber="2339" specimenCount="1" stateProvince="Cunene">
<collectionCode id="DAFCAE61FFC5FFAE5A0EF9CCFBD2F9A7" box="[1038,1097,1543,1567]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
41762 (2), specimens 49 and 49,
<geoCoordinate id="D9D95063FFC5FFAE5FF3F9ECFDE4F987" box="[499,639,1571,1599]" degrees="17" direction="south" minutes="16.325" orientation="latitude" pageId="14" pageNumber="2339" precision="1" value="-17.272083">
17
<superScript id="4B989BECFFC5FFAE5C0DF9E8FD8DF98C" attach="left" box="[525,534,1571,1588]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
16.325
<superScript id="4B989BECFFC5FFAE5C62F9E8FDFCF98C" attach="left" box="[610,615,1571,1588]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC5FFAE5C8AF9ECFC81F987" box="[650,794,1571,1599]" degrees="11" direction="east" minutes="47.177" orientation="longitude" pageId="14" pageNumber="2339" precision="1" value="11.7862835">
11
<superScript id="4B989BECFFC5FFAE5CA4F9E8FD36F98C" attach="left" box="[676,685,1571,1588]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
47.177
<superScript id="4B989BECFFC5FFAE5CF9F9E8FD65F98C" attach="left" box="[761,766,1571,1588]" fontSize="7" pageId="14" pageNumber="2339"></superScript>
E
</geoCoordinate>
,
<date id="C8531064FFC5FFAE5D25F9ECFC75F987" box="[805,1006,1575,1599]" pageId="14" pageNumber="2339" value="2010-11-08">
<collectingDate id="D817E98CFFC5FFAE5D25F9ECFC75F987" box="[805,1006,1575,1599]" pageId="14" pageNumber="2339" value="2010-11-08">8 November 2010</collectingDate>
</date>
, coll.
<collectorName id="11185372FFC5FFAE5A2FF9ECFB0BF9F8" box="[1071,1168,1575,1600]" pageId="14" pageNumber="2339">S. Voges</collectorName>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC5FFAE5ED3F98CFC65F9D8" accessionNumber="ZSM 41765" box="[211,1022,1607,1632]" collectingDate="2011-01-17" collectionCode="ZSM" collectorName="S. Voges" pageId="14" pageNumber="2339" specimenCount="1" stateProvince="Cunene">
<collectionCode id="DAFCAE61FFC5FFAE5ED3F98CFE95F9E7" box="[211,270,1607,1631]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
41765, specimen C113, same place,
<date id="C8531064FFC5FFAE5CA3F98CFCC7F9E7" box="[675,860,1607,1632]" pageId="14" pageNumber="2339" value="2011-01-17">
<collectingDate id="D817E98CFFC5FFAE5CA3F98CFCC7F9E7" box="[675,860,1607,1632]" pageId="14" pageNumber="2339" value="2011-01-17">17 January 2011</collectingDate>
</date>
, coll.
<collectorName id="11185372FFC5FFAE5D9DF98CFC65F9D8" box="[925,1022,1607,1632]" pageId="14" pageNumber="2339">S. Voges</collectorName>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC5FFAE5A08F98CFCD4F938" accessionNumber="ZSM 41763" collectingDate="2011-07-13" collectionCode="ZSM" collectorName="S. Voges" pageId="14" pageNumber="2339" specimenCount="1" stateProvince="Cunene">
<collectionCode id="DAFCAE61FFC5FFAE5A08F98CFBD8F9E7" box="[1032,1091,1607,1631]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
41763, specimen Ü7, same place,
<date id="C8531064FFC5FFAE5C0AF9ACFD3BF9C7" box="[522,672,1639,1663]" pageId="14" pageNumber="2339" value="2011-07-13">
<collectingDate id="D817E98CFFC5FFAE5C0AF9ACFD3BF9C7" box="[522,672,1639,1663]" pageId="14" pageNumber="2339" value="2011-07-13">13 July 2011</collectingDate>
</date>
, coll.
<collectorName id="11185372FFC5FFAE5CEAF9ACFCD4F938" box="[746,847,1639,1664]" pageId="14" pageNumber="2339">S. Voges</collectorName>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC5FFAE5D5DF9ACFC35F927" accessionNumber="ZSM 41764" collectingDate="2011-11-22" collectionCode="ZSM" collectorName="S. Voges. The" pageId="14" pageNumber="2339" specimenCount="1" stateProvince="Cunene">
<collectionCode id="DAFCAE61FFC5FFAE5D5DF9ACFC03F9C7" box="[861,920,1639,1663]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="14" pageNumber="2339" type="Museum">ZSM</collectionCode>
41764 (9), specimens Ä110Ä118, same place,
<date id="C8531064FFC5FFAE5FF4F94CFD54F927" box="[500,719,1671,1695]" pageId="14" pageNumber="2339" value="2011-11-22">
<collectingDate id="D817E98CFFC5FFAE5FF4F94CFD54F927" box="[500,719,1671,1695]" pageId="14" pageNumber="2339" value="2011-11-22">22 November 2011</collectingDate>
</date>
, coll.
<collectorName id="11185372FFC5FFAE5D14F94CFC35F927" box="[788,942,1671,1695]" pageId="14" pageNumber="2339">S. Voges. The</collectorName>
</materialsCitation>
specimens from the
<collectingRegion id="7E29F846FFC5FFAE5ED3F96DFEB2F906" box="[211,297,1702,1726]" country="Angola" name="Cunene" pageId="14" pageNumber="2339">Cunene</collectingRegion>
mouth were not studied for
<collectionCode id="DAFCAE61FFC5FFAE5C69F96DFD3CF906" box="[617,679,1702,1726]" pageId="14" pageNumber="2339">EOD</collectionCode>
.
</paragraph>
<paragraph id="BC5236A4FFC4FFAF5EBAFF6CFD90FEA7" blockId="15.[143,1104,167,542]" pageId="15" pageNumber="2340">
<materialsCitation id="0C853CF9FFC4FFAF5EBAFF6CFD9CFEA7" collectingDate="2009-12-15" collectionCode="IPMB" collectorName="F. H. van der Bank" country="Namibia" county="River Mouth" latitude="-17.2601" location="Samples" longLatPrecision="1" longitude="11.764867" municipality="Cunene River" pageId="15" pageNumber="2340" specimenCode="IPMB 57459-57469" specimenCount="1">
<location id="B932607FFFC4FFAF5EBAFF6CFE83FF07" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5EBAFF6CFE83FF07" box="[186,280,167,191]" country="Namibia" county="River Mouth" latitude="-17.2601" longLatPrecision="1" longitude="11.764867" municipality="Cunene River" name="Samples" pageId="15" pageNumber="2340">Samples</location>
examined for genetics. DNA samples are stored at the
<location id="B932607FFFC4FFAF5DCAFF6CFE99FF67" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5DCAFF6CFE99FF67" country="Namibia" county="River Mouth" latitude="-17.2601" longLatPrecision="1" longitude="11.764867" municipality="Cunene River" name="Institute of Pharmacy" pageId="15" pageNumber="2340">Institute of Pharmacy</location>
and
<location id="B932607FFFC4FFAF5F4CFF0CFDEEFF67" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5F4CFF0CFDEEFF67" box="[332,629,199,223]" country="Namibia" county="River Mouth" latitude="-17.2601" longLatPrecision="1" longitude="11.764867" municipality="Cunene River" name="Molecular Biotechnology" pageId="15" pageNumber="2340">Molecular Biotechnology</location>
,
<location id="B932607FFFC4FFAF5C89FF0CFC11FF67" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5C89FF0CFC11FF67" box="[649,906,199,223]" country="Namibia" county="River Mouth" latitude="-17.2601" longLatPrecision="1" longitude="11.764867" municipality="Cunene River" name="Heidelberg University" pageId="15" pageNumber="2340">Heidelberg University</location>
(
<collectionCode id="DAFCAE61FFC4FFAF5DA2FF0CFC76FF67" box="[930,1005,199,223]" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB</collectionCode>
).
<specimenCode id="EC4B9EDFFFC4FFAF5A09FF03FEBEFF47" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 5745957469</specimenCode>
,
<collectingCountry id="C4FA7634FFC4FFAF5F31FF2CFE03FF47" box="[305,408,231,255]" name="Namibia" pageId="15" pageNumber="2340">Namibia</collectingCountry>
:
<location id="B932607FFFC4FFAF5FA4FF2CFD07FF47" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5FA4FF2CFD07FF47" box="[420,668,231,255]" country="Namibia" county="River Mouth" latitude="-17.2601" longLatPrecision="1" longitude="11.764867" municipality="Cunene River" name="Cunene River Mouth" pageId="15" pageNumber="2340">
<collectingMunicipality id="5C36ACDEFFC4FFAF5FA4FF2CFDDAFF47" box="[420,577,231,255]" pageId="15" pageNumber="2340">
Cunene
<collectingCounty id="55334E28FFC4FFAF5C03FF2CFDDAFF47" box="[515,577,231,255]" pageId="15" pageNumber="2340">River</collectingCounty>
</collectingMunicipality>
Mouth
</location>
,
<geoCoordinate id="D9D95063FFC4FFAF5CA9FF2CFCACFF47" box="[681,823,227,255]" degrees="17" direction="south" minutes="15.606" orientation="latitude" pageId="15" pageNumber="2340" precision="1" value="-17.2601">
17
<superScript id="4B989BECFFC4FFAF5CC3FF28FD57FF4C" attach="left" box="[707,716,227,244]" fontSize="7" pageId="15" pageNumber="2340"></superScript>
15.606 S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC4FFAF5D44FF2CFC4CFF47" box="[836,983,227,255]" degrees="11" direction="east" minutes="45.892" orientation="longitude" pageId="15" pageNumber="2340" precision="1" value="11.764867">
11
<superScript id="4B989BECFFC4FFAF5D5FFF28FCF3FF4C" attach="none" box="[863,872,227,244]" fontSize="7" pageId="15" pageNumber="2340"></superScript>
45.892 E
</geoCoordinate>
, coll.
<collectorName id="11185372FFC4FFAF5A1DFF23FEB2FEA7" pageId="15" pageNumber="2340">F.H. van der Bank</collectorName>
,
<date id="C8531064FFC4FFAF5F34FECCFD9CFEA7" box="[308,519,263,287]" pageId="15" pageNumber="2340" value="2009-12-15">
<collectingDate id="D817E98CFFC4FFAF5F34FECCFD9CFEA7" box="[308,519,263,287]" pageId="15" pageNumber="2340" value="2009-12-15">15 December 2009</collectingDate>
</date>
</materialsCitation>
;
</paragraph>
<paragraph id="BC5236A4FFC4FFAF5EBAFEE3FE58FE27" blockId="15.[143,1104,167,542]" pageId="15" pageNumber="2340">
<materialsCitation id="0C853CF9FFC4FFAF5EBAFEE3FC38FEE7" collectingDate="2006-08-14" collectionCode="IPMB" collectorName="E. Swartz &amp; B. Kramer" country="Namibia" latitude="-17.001944" location="Cunene River" longLatPrecision="21" longitude="13.2491665" municipality="Epupa Falls" pageId="15" pageNumber="2340" specimenCode="IPMB 43971-43974" specimenCount="1">
<specimenCode id="EC4B9EDFFFC4FFAF5EBAFEE3FE31FE87" box="[186,426,295,319]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 4397143974</specimenCode>
,
<collectingCountry id="C4FA7634FFC4FFAF5FC3FEECFDB1FE87" box="[451,554,295,319]" name="Namibia" pageId="15" pageNumber="2340">Namibia</collectingCountry>
:
<location id="B932607FFFC4FFAF5C42FEECFD74FE87" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5C42FEECFD74FE87" box="[578,751,295,319]" country="Namibia" latitude="-17.001944" longLatPrecision="21" longitude="13.2491665" municipality="Epupa Falls" name="Cunene River" pageId="15" pageNumber="2340">Cunene River</location>
:
<collectingMunicipality id="5C36ACDEFFC4FFAF5D07FEE3FC05FE87" box="[775,926,295,319]" pageId="15" pageNumber="2340">Epupa Falls</collectingMunicipality>
,
<geoCoordinate id="D9D95063FFC4FFAF5DB7FEECFBD0FE87" box="[951,1099,291,319]" degrees="17" direction="south" minutes="00" orientation="latitude" pageId="15" pageNumber="2340" precision="15" seconds="07" value="-17.001944">
17
<superScript id="4B989BECFFC4FFAF5DD2FEE8FC40FE8C" attach="none" box="[978,987,291,308]" fontSize="7" pageId="15" pageNumber="2340"></superScript>
0007” S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC4FFAF5E8FFE8CFEBFFEE7" box="[143,292,323,351]" degrees="13" direction="east" minutes="14" orientation="longitude" pageId="15" pageNumber="2340" precision="15" seconds="57" value="13.2491665">
13
<superScript id="4B989BECFFC4FFAF5EAAFE88FF28FEEC" attach="none" box="[170,179,323,340]" fontSize="7" pageId="15" pageNumber="2340"></superScript>
1457” E
</geoCoordinate>
, coll.
<collectorName id="11185372FFC4FFAF5F7DFE83FE6FFEE7" box="[381,500,327,351]" pageId="15" pageNumber="2340">E. Swartz</collectorName>
and
<collectorName id="11185372FFC4FFAF5C43FE83FD5DFEE7" box="[579,710,328,351]" pageId="15" pageNumber="2340">B. Kramer</collectorName>
,
<date id="C8531064FFC4FFAF5CDCFE8CFC38FEE7" box="[732,931,327,351]" pageId="15" pageNumber="2340" value="2006-08-14">
<collectingDate id="D817E98CFFC4FFAF5CDCFE8CFC38FEE7" box="[732,931,327,351]" pageId="15" pageNumber="2340" value="2006-08-14">14 August 2006</collectingDate>
</date>
</materialsCitation>
,
<geoCoordinate id="D9D95063FFC4FFAF5DBAFE8CFBD0FEE7" box="[954,1099,323,351]" degrees="17" direction="south" minutes="00" orientation="latitude" pageId="15" pageNumber="2340" precision="15" seconds="07" value="-17.001944">
17
<superScript id="4B989BECFFC4FFAF5DD4FE88FC46FEEC" attach="left" box="[980,989,323,340]" fontSize="7" pageId="15" pageNumber="2340"></superScript>
0007” S
</geoCoordinate>
, E
<geoCoordinate id="D9D95063FFC4FFAF5EA0FEACFEA2FEC7" box="[160,313,355,383]" degrees="013" direction="east" minutes="14" orientation="longitude" pageId="15" pageNumber="2340" precision="15" seconds="57" value="13.2491665">
013
<superScript id="4B989BECFFC4FFAF5EC9FEA8FF49FECC" attach="none" box="[201,210,355,372]" fontSize="7" pageId="15" pageNumber="2340"></superScript>
1457” E
</geoCoordinate>
;
<materialsCitation id="0C853CF9FFC4FFAF5F46FEA3FC10FEC7" box="[326,907,359,383]" collectingDate="2006-08-15" collectionCode="IPMB" country="Namibia" pageId="15" pageNumber="2340" specimenCode="IPMB 43975-43978" specimenCount="1">
<specimenCode id="EC4B9EDFFFC4FFAF5F46FEA3FDB1FEC7" box="[326,554,359,383]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 4397543978</specimenCode>
, as before, but
<date id="C8531064FFC4FFAF5CD8FEACFC10FEC7" box="[728,907,359,383]" pageId="15" pageNumber="2340" value="2006-08-15">
<collectingDate id="D817E98CFFC4FFAF5CD8FEACFC10FEC7" box="[728,907,359,383]" pageId="15" pageNumber="2340" value="2006-08-15">15 August 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC4FFAF5D98FEA3FE24FE27" collectingDate="2006-08-17" collectionCode="IPMB" country="Namibia" pageId="15" pageNumber="2340" specimenCode="IPMB 43993" specimenCount="1">
<specimenCode id="EC4B9EDFFFC4FFAF5D98FEA3FBB7FEC7" box="[920,1068,359,383]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 43993</specimenCode>
, as before, but
<date id="C8531064FFC4FFAF5F0EFE4CFE24FE27" box="[270,447,391,415]" pageId="15" pageNumber="2340" value="2006-08-17">
<collectingDate id="D817E98CFFC4FFAF5F0EFE4CFE24FE27" box="[270,447,391,415]" pageId="15" pageNumber="2340" value="2006-08-17">17 August 2006</collectingDate>
</date>
</materialsCitation>
;
</paragraph>
<paragraph id="BC5236A4FFC4FFAF5EBAFE6CFC8AFDA6" blockId="15.[143,1104,167,542]" pageId="15" pageNumber="2340">
<materialsCitation id="0C853CF9FFC4FFAF5EBAFE6CFD2AFE66" collectingDate="2006-08-19" collectionCode="IPMB" collectorName="E. Swartz &amp; B. Kramer" country="Namibia" latitude="-17.406666" location="Cunene River" longLatPrecision="21" longitude="14.216944" municipality="Ruacana Falls" pageId="15" pageNumber="2340" specimenCode="IPMB 43980" specimenCount="1">
<specimenCode id="EC4B9EDFFFC4FFAF5EBAFE6CFED6FE06" box="[186,333,422,446]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 43980</specimenCode>
,
<collectingCountry id="C4FA7634FFC4FFAF5F58FE6DFE24FE06" box="[344,447,422,446]" name="Namibia" pageId="15" pageNumber="2340">Namibia</collectingCountry>
:
<location id="B932607FFFC4FFAF5FCAFE6DFDF2FE06" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5FCAFE6DFDF2FE06" box="[458,617,422,446]" country="Namibia" latitude="-17.406666" longLatPrecision="21" longitude="14.216944" municipality="Ruacana Falls" name="Cunene River" pageId="15" pageNumber="2340">Cunene River</location>
:
<collectingMunicipality id="5C36ACDEFFC4FFAF5C74FE6CFC8DFE06" box="[628,790,422,446]" pageId="15" pageNumber="2340">Ruacana Falls</collectingMunicipality>
,
<geoCoordinate id="D9D95063FFC4FFAF5D21FE6DFC3CFE06" box="[801,935,419,446]" degrees="17" direction="south" minutes="24" orientation="latitude" pageId="15" pageNumber="2340" precision="15" seconds="24" value="-17.406666">
17
<superScript id="4B989BECFFC4FFAF5D3CFE68FCDEFE0C" attach="none" box="[828,837,419,436]" fontSize="7" pageId="15" pageNumber="2340"></superScript>
2424” S
</geoCoordinate>
,
<geoCoordinate id="D9D95063FFC4FFAF5DB3FE6DFBD1FE06" box="[947,1098,419,446]" degrees="014" direction="east" minutes="13" orientation="longitude" pageId="15" pageNumber="2340" precision="15" seconds="01" value="14.216944">
014
<superScript id="4B989BECFFC4FFAF5DDBFE68FC7FFE0C" attach="none" box="[987,996,419,436]" fontSize="7" pageId="15" pageNumber="2340"></superScript>
1301” E
</geoCoordinate>
, coll.
<collectorName id="11185372FFC4FFAF5EC8FE0CFEACFE66" box="[200,311,454,478]" pageId="15" pageNumber="2340">E. Swartz</collectorName>
and
<collectorName id="11185372FFC4FFAF5F74FE0CFE75FE66" box="[372,494,455,478]" pageId="15" pageNumber="2340">B. Kramer</collectorName>
,
<date id="C8531064FFC4FFAF5FFCFE0DFD2AFE66" box="[508,689,454,478]" pageId="15" pageNumber="2340" value="2006-08-19">
<collectingDate id="D817E98CFFC4FFAF5FFCFE0DFD2AFE66" box="[508,689,454,478]" pageId="15" pageNumber="2340" value="2006-08-19">19 August 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC4FFAF5CBFFE0CFED3FE46" collectingDate="2006-08-21" collectionCode="IPMB" country="Namibia" pageId="15" pageNumber="2340" specimenCode="IPMB 43986, 43988" specimenCount="1">
<specimenCode id="EC4B9EDFFFC4FFAF5CBFFE0CFCCFFE66" box="[703,852,454,478]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 43986</specimenCode>
,
<specimenCode id="EC4B9EDFFFC4FFAF5D61FE0DFC3CFE66" box="[865,935,454,478]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">43988</specimenCode>
, as before, but
<date id="C8531064FFC4FFAF5E8FFE2DFED3FE46" box="[143,328,486,510]" pageId="15" pageNumber="2340" value="2006-08-21">
<collectingDate id="D817E98CFFC4FFAF5E8FFE2DFED3FE46" box="[143,328,486,510]" pageId="15" pageNumber="2340" value="2006-08-21">21 August 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC4FFAF5F57FE2CFCFBFE46" box="[343,864,486,510]" collectingDate="2006-08-22" collectionCode="IPMB" country="Namibia" pageId="15" pageNumber="2340" specimenCode="IPMB 43990" specimenCount="1">
<specimenCode id="EC4B9EDFFFC4FFAF5F57FE2CFE75FE46" box="[343,494,486,510]" collectionCode="IPMB" country="Germany" httpUri="http://grbio.org/cool/yfri-k4dg" name="Institut fur Pharmazie und Molekulare Biotechnologie" pageId="15" pageNumber="2340">IPMB 43990</specimenCode>
, as before, but
<date id="C8531064FFC4FFAF5CA7FE2DFCFBFE46" box="[679,864,486,510]" pageId="15" pageNumber="2340" value="2006-08-22">
<collectingDate id="D817E98CFFC4FFAF5CA7FE2DFCFBFE46" box="[679,864,486,510]" pageId="15" pageNumber="2340" value="2006-08-22">22 August 2006</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="0C853CF9FFC4FFAF5D6FFE2DFC96FDA6" accessionNumber="KC202227, KC202230, KC202238, KC202258" collectionCode="KC" country="Namibia" location="GenBank" pageId="15" pageNumber="2340" specimenCount="1">
<location id="B932607FFFC4FFAF5D6FFE2DFC40FE46" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFC4FFAF5D6FFE2DFC40FE46" box="[879,987,486,510]" country="Namibia" name="GenBank" pageId="15" pageNumber="2340">GenBank</location>
accession numbers: (
<accessionNumber id="A3BEAB47FFC4FFAF5F05FDCDFEE5FDA6" box="[261,382,518,542]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC202227" pageId="15" pageNumber="2340">
<collectionCode id="DAFCAE61FFC4FFAF5F05FDCDFEB0FDA6" box="[261,299,518,542]" pageId="15" pageNumber="2340">KC</collectionCode>
202227
</accessionNumber>
-
<accessionNumber id="A3BEAB47FFC4FFAF5F86FDCDFD9BFDA6" box="[390,512,518,542]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC202230" pageId="15" pageNumber="2340">KC202230</accessionNumber>
;
<accessionNumber id="A3BEAB47FFC4FFAF5C0BFDCDFD1EFDA6" box="[523,645,518,542]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC202238" pageId="15" pageNumber="2340">KC202238</accessionNumber>
-
<accessionNumber id="A3BEAB47FFC4FFAF5C8CFDCDFC9DFDA6" box="[652,774,518,542]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KC202258" pageId="15" pageNumber="2340">
<collectionCode id="DAFCAE61FFC4FFAF5C8CFDCDFD29FDA6" box="[652,690,518,542]" pageId="15" pageNumber="2340">KC</collectionCode>
202258
</accessionNumber>
)
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFC4FFAF5E8FFDAFFE85FDC3" blockId="15.[143,1103,611,706]" box="[143,286,611,635]" pageId="15" pageNumber="2340">
<heading id="E71A81C8FFC4FFAF5E8FFDAFFE85FDC3" box="[143,286,611,635]" fontSize="10" level="3" pageId="15" pageNumber="2340" reason="5">
<emphasis id="8E99EAB6FFC4FFAF5E8FFDAFFE85FDC3" box="[143,286,611,635]" italics="true" pageId="15" pageNumber="2340">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFC4FFAF5E8FFD41FF6EFD7A" blockId="15.[143,1103,611,706]" pageId="15" pageNumber="2340">
<collectingRegion id="7E29F846FFC4FFAF5E8FFD41FF7EFD1A" box="[143,229,650,674]" country="Angola" name="Cunene" pageId="15" pageNumber="2340">Cunene</collectingRegion>
River just above the Epupa Falls (Angolan/Namibian border, locality no. 9 on
<figureCitation id="24D62A21FFC4FFAF5E8FFD61FF76FD7A" box="[143,237,682,706]" captionStart="Figure 1" captionStartId="4.[211,278,932,954]" captionTargetBox="[213,1170,167,899]" captionTargetId="figure-5@4.[213,1170,167,899]" captionTargetPageId="4" captionText="Figure 1. Map of sampling locations in southern Africa. (A) Type locality for Marcusenius macrolepidotus (Peters, 1852), Tete on the Lower Zambezi River; (B) origin of M. macrolepidotus, SAIAB 60847, from Marromeu, Lower Zambezi delta (Figure 4A); (C) origin of M. macrolepidotus from the Buzi System, same location as SAIAB 67369, presently alive in aquarium; (D) likely type region for Marcusenius angolensis (Boulenger, 1905), the Lower Quanza; (E) Marcusenius altisambesi Kramer et al., 2007, type locality on the Upper Zambezi River; (F) M. altisambesi from the Kwando River; (G) M. altisambesi from the Okavango delta; (H, I) Marcusenius multisquamatus sp. nov., stretching from below the Ruacana Falls to above the Epupa Falls (I), the type locality; (J) specimens from the Cunene River Mouth." figureDoi="http://doi.org/10.5281/zenodo.5197592" httpUri="https://zenodo.org/record/5197592/files/figure.png" pageId="15" pageNumber="2340">Figure 1</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFC4FFAF5E8FFCCBFC89FBA6" pageId="15" pageNumber="2340" type="diagnosis">
<paragraph id="BC5236A4FFC4FFAF5E8FFCCBFF60FCAF" blockId="15.[143,1104,768,1054]" box="[143,251,768,791]" pageId="15" pageNumber="2340">
<heading id="E71A81C8FFC4FFAF5E8FFCCBFF60FCAF" box="[143,251,768,791]" fontSize="10" level="3" pageId="15" pageNumber="2340" reason="5">
<emphasis id="8E99EAB6FFC4FFAF5E8FFCCBFF60FCAF" box="[143,251,768,791]" italics="true" pageId="15" pageNumber="2340">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFC4FFAF5E8FFCECFC89FBA6" blockId="15.[143,1104,768,1054]" pageId="15" pageNumber="2340">Body moderately long, prominent mobile and forward-extending mental lobe on lower jaw, median fins set well back with dorsal fin shorter than and originating behind anal fin, depth of caudal peduncle 38% (3443%) of its length, 24 (2325) dorsal fin rays, 30 (2831) anal fin rays, 59 (5664) scales in lateral series, 13 (1216) scales around caudal peduncle, HL (head length) 20% (1921%) of SL, BD (body depth) 29% (2732%) of SL, LD (dorsal fin length) 19.4% (18.122.3%) of SL, LSo (length of snout) 48% (4550%) HL, LA (anal fin length) 24.2% (22.725.3%) of SL, CPL (length of caudal peduncle) 18.3% (16.319.9%) of SL. (See also Remarks.)</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFC4FFB05E8FFB90FD72FA47" lastPageId="16" lastPageNumber="2341" pageId="15" pageNumber="2340" type="description">
<paragraph id="BC5236A4FFC4FFAF5E8FFB90FE95FBCA" blockId="15.[143,1104,1115,1689]" box="[143,270,1115,1138]" pageId="15" pageNumber="2340">
<heading id="E71A81C8FFC4FFAF5E8FFB90FE95FBCA" box="[143,270,1115,1138]" fontSize="10" level="3" pageId="15" pageNumber="2340" reason="5">
<emphasis id="8E99EAB6FFC4FFAF5E8FFB90FE95FBCA" box="[143,270,1115,1138]" italics="true" pageId="15" pageNumber="2340">Description</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFC4FFB05E8FFB49FD6FFA32" blockId="15.[143,1104,1115,1689]" lastBlockId="16.[211,1172,1362,1418]" lastPageId="16" lastPageNumber="2341" pageId="15" pageNumber="2340">
Head with terminal mouth well in front of eye, mental lobe on lower jaw protruding beyond upper jaw. Head and body dorsolaterally compressed. Dorsal fin situated about two-thirds of standard length from snout, obliquely oriented, anteriorly higher and posteriorly lower, distal margin sometimes only slightly crescent-shaped with anterior two or three rays longer than posterior rays, number of rays 23 (
<emphasis id="8E99EAB6FFC4FFAF5D88FAC8FC0EFAA2" box="[904,917,1283,1306]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 5), 24 (
<emphasis id="8E99EAB6FFC4FFAF5A04FAC8FB8AFAA2" box="[1028,1041,1283,1306]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 4), 25 (
<emphasis id="8E99EAB6FFC4FFAF5EBAFAE8FF5CFA82" box="[186,199,1315,1338]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 6); anal fin opposite dorsal fin with distinctly more anterior origin, obliquely oriented, anteriorly lower and posteriorly higher, anterior rays longer than posterior ones, especially in males where they also appear stronger and often darkened, distal margin crescent-shaped (in males only posterior to rounded, elongated anterior part of fin), number of rays 28 (
<emphasis id="8E99EAB6FFC4FFAF5FB9FA69FE5DFA01" box="[441,454,1442,1465]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 1), 29 (
<emphasis id="8E99EAB6FFC4FFAF5C34FA69FDDAFA01" box="[564,577,1442,1465]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 4), 30 (
<emphasis id="8E99EAB6FFC4FFAF5CAFFA69FD27FA01" box="[687,700,1442,1465]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 7), 31 (
<emphasis id="8E99EAB6FFC4FFAF5D2BFA69FCA3FA01" box="[811,824,1442,1465]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 3). Scales cycloid with reticulate striae, scales extending anteriorly to operculum and pectoral fins (beyond pelvic fins). Scales on caudal peduncle circumference, 12 (
<emphasis id="8E99EAB6FFC4FFAF5D0FFA29FC87FA41" box="[783,796,1506,1529]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 5), 13 (
<emphasis id="8E99EAB6FFC4FFAF5D8AFA29FC0CFA41" box="[906,919,1506,1529]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 4), 14 (
<emphasis id="8E99EAB6FFC4FFAF5A04FA29FB8AFA41" box="[1028,1041,1506,1529]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 5), 16 (
<emphasis id="8E99EAB6FFC4FFAF5EB9F9C9FF5DF9A1" box="[185,198,1538,1561]" italics="true" pageId="15" pageNumber="2340">n</emphasis>
= 1) Caudal peduncle relatively deep, subcylindrical entire length, usually 18.3% (16.319.9%) in SL (
<tableCitation id="F16F031FFFC4FFAF5F73F9EAFE47F981" box="[371,476,1569,1593]" captionStart="Table 1" captionStartId="21.[58,113,594,616]" captionText="Table 1. Comparison of anatomical characters in three Marcusenius species from different southern African origins (Cunene MS = multisquamatus sp. nov., L Zambezi ML = macrolepidotus, Okavango AS and U Zambezi AS = altisambesi). Multivariate analysis of variance (MANOVA). p values in the body of the table not shown when&gt;0.05." pageId="15" pageNumber="2340">Table A1</tableCitation>
, in Appendix A). Electric organ discharge biphasic with weak pre-potential (
<figureCitation id="24D62A21FFC4FFAF5F7BF98AFE45F9E1" box="[379,478,1601,1625]" captionStart="Figure 5" captionStartId="16.[211,278,1076,1098]" captionTargetBox="[213,1170,168,1043]" captionTargetId="figure-5@16.[213,1170,168,1043]" captionTargetPageId="16" captionText="Figure 5. Electric organ discharges (EODs) recorded from fish from various origins. (A) Five specimens of Marcusenius macrolepidotus originating from the Buzi River. (B) Marcusenius altisambesi originating from the the Upper Zambezi System. Note P and N phases of exaggerated duration in male individual. (C) M. altisambesi originating from the Okavango delta. (D) Marcusenius multisquamatus sp. nov. originating from the Cunene River. M, male, F, female, J, juvenile individuals below size where sexual maturity is possible. Size given is SL. Note that in Buzi individuals an initial miniature potential is lacking which is present in the other forms of bulldog fish shown here (see EODs with clipped peaks, amplified × 3)." figureDoi="http://doi.org/10.5281/zenodo.5197600" httpUri="https://zenodo.org/record/5197600/files/figure.png" pageId="15" pageNumber="2340">Figure 5</figureCitation>
). Males approaching sexual maturity develop a kink in the base of the anal fin (e.g.
<figureCitation id="24D62A21FFC4FFAF5C08F9ABFDE6F9C0" box="[520,637,1632,1656]" captionStart="Figure 4" captionStartId="11.[143,210,1506,1528]" captionTargetBox="[168,1081,161,1473]" captionTargetId="figure-6@11.[168,1081,161,1473]" captionTargetPageId="11" captionText="Figure 4. Photographs of bulldog fish from various origins. (A) Marcusenius macrolepidotus (Peters, 1852), SAIAB 60847, coll. R. Bills 1 August 1999, Lower Zambezi; (B) Marcusenius angolensis (Boulenger, 1905), BMNH 1905.5.29.64 (holotype); (C) Marcusenius altisambesi, coll. F. H. van der Bank and B. Kramer, 21 August 1999, Upper Zambezi, Kalimbeza, live specimen of 16.5 cm SL photographed 3 July 2003 in aquarium; (D) M. altisambesi, coll. F.H. van der Bank and B. Kramer, 11/12 August 2004, Okavango River, live fish of SL 13 cm photographed 20 April 2006 in aquarium; (E) Marcusenius multisquamatus sp. nov., coll. B. Kramer and Ernst Swartz, coll. and photographed on 19 August 2006, Cunene River, below Ruacana Falls, specimen RUAC01, SL 15.4 cm." figureDoi="http://doi.org/10.5281/zenodo.5197598" httpUri="https://zenodo.org/record/5197598/files/figure.png" pageId="15" pageNumber="2340">Figure 4C</figureCitation>
) that is absent in juveniles and females where the anal fin base is straight. Colour in life: brownish grey with many distinct dark-brown blotches, except on head and belly, purple hue depending on the angle of light incidence, paired fins light and transparent.
</paragraph>
<caption id="E892662CFFDBFFB05ED3FBFFFC02FAAE" ID-DOI="http://doi.org/10.5281/zenodo.5197600" ID-Zenodo-Dep="5197600" httpUri="https://zenodo.org/record/5197600/files/figure.png" pageId="16" pageNumber="2341" startId="16.[211,278,1076,1098]" targetBox="[213,1170,168,1043]" targetPageId="16">
<paragraph id="BC5236A4FFDBFFB05ED3FBFFFC02FAAE" blockId="16.[211,1172,1076,1302]" pageId="16" pageNumber="2341">
Figure 5. Electric organ discharges (EODs) recorded from fish from various origins. (A) Five specimens of
<taxonomicName id="7BED4D27FFDBFFB05F64FB99FD19FBDF" baseAuthorityName="Peters" baseAuthorityYear="1852" box="[356,642,1105,1127]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="16" pageNumber="2341" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFDBFFB05F64FB99FD19FBDF" box="[356,642,1105,1127]" italics="true" pageId="16" pageNumber="2341">Marcusenius macrolepidotus</emphasis>
</taxonomicName>
originating from the Buzi River. (B)
<taxonomicName id="7BED4D27FFDBFFB05A16FB99FED9FB3C" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="16" pageNumber="2341" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFDBFFB05A16FB99FED9FB3C" italics="true" pageId="16" pageNumber="2341">Marcusenius altisambesi</emphasis>
</taxonomicName>
originating from the the Upper Zambezi System. Note
<emphasis id="8E99EAB6FFDBFFB05D7AFBA4FC11FB3C" box="[890,906,1135,1156]" italics="true" pageId="16" pageNumber="2341">P</emphasis>
and
<emphasis id="8E99EAB6FFDBFFB05DBEFBA4FC4BFB3C" box="[958,976,1135,1156]" italics="true" pageId="16" pageNumber="2341">N</emphasis>
phases of exaggerated duration in male individual. (C)
<taxonomicName id="7BED4D27FFDBFFB05C52FB47FD7DFB1A" box="[594,742,1164,1186]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="16" pageNumber="2341" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFDBFFB05C52FB47FD7DFB1A" box="[594,742,1164,1186]" italics="true" pageId="16" pageNumber="2341">M. altisambesi</emphasis>
</taxonomicName>
originating from the Okavango delta. (D)
<taxonomicName id="7BED4D27FFDBFFB05ED3FB61FE68FB07" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[211,499,1193,1215]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="16" pageNumber="2341" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDBFFB05ED3FB61FE68FB07" box="[211,499,1193,1215]" italics="true" pageId="16" pageNumber="2341">Marcusenius multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFDBFFB05FFAFB62FDDCFB06" box="[506,583,1193,1214]" pageId="16" pageNumber="2341" rank="species">sp. nov.</taxonomicNameLabel>
originating from the Cunene River. M, male, F, female, J, juvenile individuals below size where sexual maturity is possible. Size given is SL. Note that in Buzi individuals an initial miniature potential is lacking which is present in the other forms of bulldog fish shown here (see EODs with clipped peaks, amplified × 3).
</paragraph>
</caption>
<paragraph id="BC5236A4FFDBFFB05ED3FA74FE5DFA6F" blockId="16.[211,745,1471,1535]" box="[211,454,1471,1495]" pageId="16" pageNumber="2341">
<heading id="E71A81C8FFDBFFB05ED3FA74FE5DFA6F" box="[211,454,1471,1495]" fontSize="10" level="3" pageId="16" pageNumber="2341" reason="5">
<emphasis id="8E99EAB6FFDBFFB05ED3FA74FE5DFA6F" box="[211,454,1471,1495]" italics="true" pageId="16" pageNumber="2341">Colour in preservation</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFDBFFB05ED3FA2CFD72FA47" blockId="16.[211,745,1471,1535]" box="[211,745,1511,1535]" pageId="16" pageNumber="2341">Medium brown, with darker, irregular blotches.</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFDBFFB15ED3F9FFFCC4FD86" lastPageId="17" lastPageNumber="2342" pageId="16" pageNumber="2341" type="biology_ecology">
<paragraph id="BC5236A4FFDBFFB05ED3F9FFFEB7F9F4" blockId="16.[211,1172,1588,1716]" box="[211,300,1588,1612]" pageId="16" pageNumber="2341">
<heading id="E71A81C8FFDBFFB05ED3F9FFFEB7F9F4" box="[211,300,1588,1612]" fontSize="10" level="3" pageId="16" pageNumber="2341" reason="5">
<emphasis id="8E99EAB6FFDBFFB05ED3F9FFFEB7F9F4" box="[211,300,1588,1612]" italics="true" pageId="16" pageNumber="2341">Ecology</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFDBFFB15ED3F997FCC4FD86" blockId="16.[211,1172,1588,1716]" lastBlockId="17.[143,1104,167,574]" lastPageId="17" lastPageNumber="2342" pageId="16" pageNumber="2341">
The
<collectingRegion id="7E29F846FFDBFFB05F06F997FEC7F9CC" box="[262,348,1628,1652]" country="Angola" name="Cunene" pageId="16" pageNumber="2341">Cunene</collectingRegion>
is a major, perennial and independent river that arises from the Angolan central highlands of
<collectingRegion id="7E29F846FFDBFFB05FC0F9B0FE7EF92B" box="[448,485,1659,1683]" country="Angola" name="Bie" pageId="16" pageNumber="2341">Bié</collectingRegion>
and flows southward towards the Namibian border, shortly before it turns west and breaches the coastal mountain ranges (Zebra and Baynes Mountains) to drain into the Atlantic. In the section between Ruacana Falls and Epupa Falls, water level was regulated by a hydroelectric company (NamPower) at Ruacana Falls. The Ruacana Falls were bare rock and completely dry, apparently because the water dammed above Ruacana Falls (Calueque Dam) was all fed into the hydroelectric power turbines. When the water level below the dam was kept high, fishing with gill nets and other methods generally proved unproductive. The Epupa Falls consist of a main fall with many lesser falls beside this over a wide front, and mormyrid EODs were demonstrated with an electro-acoustic, custom-built “fish detector” also below the Falls although the fish were not caught. Although we were warned of a high incidence of crocodiles we saw only a few, and no hippopotami. River borders were covered mainly by dense semi-aquatic shrubs at Epupa Falls, and dense reed beds also with shrubs at Ruacana Falls. Palm trees (Makalani palms) were common at Epupa, much less so at Ruacana where dicotyledonous trees dominated.
</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFDAFFB15E8FFDB8FC08FCDF" pageId="17" pageNumber="2342" type="distribution">
<paragraph id="BC5236A4FFDAFFB15E8FFDB8FE8FFD33" blockId="17.[143,1103,627,723]" box="[143,276,627,651]" pageId="17" pageNumber="2342">
<heading id="E71A81C8FFDAFFB15E8FFDB8FE8FFD33" box="[143,276,627,651]" fontSize="10" level="3" pageId="17" pageNumber="2342" reason="5">
<emphasis id="8E99EAB6FFDAFFB15E8FFDB8FE8FFD33" box="[143,276,627,651]" italics="true" pageId="17" pageNumber="2342">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFDAFFB15E8FFD51FB9BFD6A" blockId="17.[143,1103,627,723]" pageId="17" pageNumber="2342">
<materialsCitation id="0C853CF9FFDAFFB15E8FFD51FC67FD6A" location="This river" municipality="Ruacana Falls" pageId="17" pageNumber="2342" specimenCount="1" stateProvince="Cunene">
At present known only from the lower
<collectingRegion id="7E29F846FFDAFFB15C4BFD51FD3AFD0A" box="[587,673,666,690]" country="Angola" name="Cunene" pageId="17" pageNumber="2342">Cunene</collectingRegion>
River, from just below
<collectingMunicipality id="5C36ACDEFFDAFFB15DAEFD50FBD4FD0A" box="[942,1103,666,690]" pageId="17" pageNumber="2342">Ruacana Falls</collectingMunicipality>
to the
<collectingRegion id="7E29F846FFDAFFB15ED7FD71FEB6FD6A" box="[215,301,698,722]" country="Angola" name="Cunene" pageId="17" pageNumber="2342">Cunene</collectingRegion>
mouth.
<location id="B932607FFFDAFFB15F8DFD71FE62FD6A" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFDAFFB15F8DFD71FE62FD6A" box="[397,505,698,722]" municipality="Ruacana Falls" name="This river" pageId="17" pageNumber="2342" stateProvince="Cunene">This river</location>
section forms the Angolan/Namibian border
</materialsCitation>
.
</paragraph>
<paragraph id="BC5236A4FFDAFFB15E8FFCCCFEB8FCA7" blockId="17.[143,1103,775,871]" box="[143,291,775,799]" pageId="17" pageNumber="2342">
<heading id="E71A81C8FFDAFFB15E8FFCCCFEB8FCA7" box="[143,291,775,799]" fontSize="10" level="3" pageId="17" pageNumber="2342" reason="5">
<emphasis id="8E99EAB6FFDAFFB15E8FFCCCFEB8FCA7" box="[143,291,775,799]" italics="true" pageId="17" pageNumber="2342">Relationships</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFDAFFB15E8FFCE4FC08FCDF" blockId="17.[143,1103,775,871]" pageId="17" pageNumber="2342">
Closest relationships are assumed with
<taxonomicName id="7BED4D27FFDAFFB15C40FCFBFD79FCFF" box="[576,738,815,839]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFDAFFB15C40FCFBFD79FCFF" box="[576,738,815,839]" italics="true" pageId="17" pageNumber="2342">M. altisambesi</emphasis>
</taxonomicName>
to the east of
<taxonomicName id="7BED4D27FFDAFFB15D7BFCFBFBD4FCFF" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[891,1103,815,839]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDAFFB15D7BFCFBFBD4FCFF" box="[891,1103,815,839]" italics="true" pageId="17" pageNumber="2342">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFDAFFB15E8FFC84FF7FFCDE" box="[143,228,847,870]" pageId="17" pageNumber="2342" rank="species">sp. nov.</taxonomicNameLabel>
on the basis of morphological similarity, EODs and genetics.
</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFDAFFB15E8FFC50FC18FB83" pageId="17" pageNumber="2342" type="etymology">
<paragraph id="BC5236A4FFDAFFB15E8FFC50FE9CFC0B" blockId="17.[143,1104,923,1083]" box="[143,263,923,947]" pageId="17" pageNumber="2342">
<heading id="E71A81C8FFDAFFB15E8FFC50FE9CFC0B" box="[143,263,923,947]" fontSize="10" level="3" pageId="17" pageNumber="2342" reason="5">
<emphasis id="8E99EAB6FFDAFFB15E8FFC50FE9CFC0B" box="[143,263,923,947]" italics="true" pageId="17" pageNumber="2342">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFDAFFB15E8FFC0FFC18FB83" blockId="17.[143,1104,923,1083]" pageId="17" pageNumber="2342">
<taxonomicName id="7BED4D27FFDAFFB15E8FFC0FFE54FC63" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[143,463,963,987]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDAFFB15E8FFC0FFE54FC63" box="[143,463,963,987]" italics="true" pageId="17" pageNumber="2342">Marcusenius multisquamatus</emphasis>
</taxonomicName>
<materialsCitation id="0C853CF9FFDAFFB15FD6FC0FFCE4FB83" collectionCode="BMNH" location="Mossamedes" municipality="southern African" pageId="17" pageNumber="2342" specimenCount="1" stateProvince="Cunene">
<taxonomicNameLabel id="95AA57CDFFDAFFB15FD6FC0FFDB0FC63" box="[470,555,964,987]" pageId="17" pageNumber="2342" rank="species">sp. nov.</taxonomicNameLabel>
refers to the highest number of lateral line scales among the different forms of
<collectingMunicipality id="5C36ACDEFFDAFFB15FFAFC28FD5AFC43" box="[506,705,995,1019]" pageId="17" pageNumber="2342">southern African</collectingMunicipality>
bulldog fish (excluding the three
<location id="B932607FFFDAFFB15E8FFBC8FEBBFBA3" LSID="urn:lsid:plazi:treatment:344487B2FFC5FFB25FE6FC8EFB0BFCA6:B932607FFFDAFFB15E8FFBC8FEBBFBA3" box="[143,288,1027,1051]" municipality="southern African" name="Mossamedes" pageId="17" pageNumber="2342" stateProvince="Cunene">Mossamedes</location>
/
<collectingRegion id="7E29F846FFDAFFB15F2CFBC8FE19FBA3" box="[300,386,1027,1051]" country="Angola" name="Cunene" pageId="17" pageNumber="2342">Cunene</collectingRegion>
specimens (
<collectionCode id="DAFCAE61FFDAFFB15C12FBC8FDF0FBA3" box="[530,619,1027,1051]" pageId="17" pageNumber="2342">BMNH</collectionCode>
1907.6.29.231233) from any location on the
<collectingRegion id="7E29F846FFDAFFB15EB9FBE8FE94FB83" box="[185,271,1059,1083]" country="Angola" name="Cunene" pageId="17" pageNumber="2342">Cunene</collectingRegion>
up to
<quantity id="7B159B41FFDAFFB15F58FBE8FE37FB83" box="[344,428,1059,1083]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="2342" unit="km" value="300.0">300 km</quantity>
north of the Angolan/Namibian border)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F4F7652FFFDAFFB25E8FFBA4FB0BFCA6" lastPageId="18" lastPageNumber="2343" pageId="17" pageNumber="2342" type="discussion">
<paragraph id="BC5236A4FFDAFFB15E8FFBA4FF6AFB3F" blockId="17.[143,1104,1135,1709]" box="[143,241,1135,1159]" pageId="17" pageNumber="2342">
<heading id="E71A81C8FFDAFFB15E8FFBA4FF6AFB3F" box="[143,241,1135,1159]" fontSize="10" level="3" pageId="17" pageNumber="2342" reason="5">
<emphasis id="8E99EAB6FFDAFFB15E8FFBA4FF6AFB3F" box="[143,241,1135,1159]" italics="true" pageId="17" pageNumber="2342">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="BC5236A4FFDAFFB15E8FFB5CFDEFFAD6" blockId="17.[143,1104,1135,1709]" pageId="17" pageNumber="2342">
Compared with the
<taxonomicName id="7BED4D27FFDAFFB15F6DFB53FD9EFB17" authorityName="Boulenger" authorityYear="1905" box="[365,517,1175,1199]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="8E99EAB6FFDAFFB15F6DFB53FD9EFB17" box="[365,517,1175,1199]" italics="true" pageId="17" pageNumber="2342">M. angolensis</emphasis>
</taxonomicName>
<typeStatus id="63568806FFDAFFB15C0AFB5CFDF4FB17" box="[522,623,1175,1199]" pageId="17" pageNumber="2342" type="holotype">holotype</typeStatus>
,
<taxonomicName id="7BED4D27FFDAFFB15C7AFB53FCD5FB17" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[634,846,1175,1199]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDAFFB15C7AFB53FCD5FB17" box="[634,846,1175,1199]" italics="true" pageId="17" pageNumber="2342">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFDAFFB15D54FB53FC33FB17" box="[852,936,1176,1199]" pageId="17" pageNumber="2342" rank="species">sp. nov.</taxonomicNameLabel>
specimens had lower counts in nD, no. of dorsal fin rays (maximum,
<quantity id="7B159B41FFDAFFB15D0CFB7CFCDCFB77" box="[780,839,1207,1231]" metricMagnitude="-1" metricUnit="m" metricValue="6.35" pageId="17" pageNumber="2342" unit="in" value="25.0">25 in</quantity>
the latter versus
<quantity id="7B159B41FFDAFFB15A15FB7CFBD4FB77" box="[1045,1103,1207,1231]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="17" pageNumber="2342" unit="in" value="26.0">26 in</quantity>
the former, that is, no overlap) and nA, number of anal fin rays (maximum, 31 versus 33), shorter LA, anal fin length (maximum, 0.253 versus 0.258 of SL) and PDL, predorsal length (maximum, 0.665 versus 0.674 of SL), smaller ratio HL/Na, head length/separation of nares (maximum, 15.52 versus 15.96), but a greater BD, body depth (minimum 0.271 versus 0.266 of SL).
</paragraph>
<paragraph id="BC5236A4FFDAFFB25EBAFABDFD63FE27" blockId="17.[143,1104,1135,1709]" lastBlockId="18.[211,1172,167,798]" lastPageId="18" lastPageNumber="2343" pageId="17" pageNumber="2342">
When compared with the other
<taxonomicName id="7BED4D27FFDAFFB15C4CFABCFCAEFA36" box="[588,821,1398,1422]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="8E99EAB6FFDAFFB15C4CFABCFD4CFA36" box="[588,727,1399,1422]" italics="true" pageId="17" pageNumber="2342">Marcusenius</emphasis>
species
</taxonomicName>
within the
<collectingRegion id="7E29F846FFDAFFB15DCFFABDFBDAFA36" box="[975,1089,1398,1422]" country="Namibia" name="Kavango West" pageId="17" pageNumber="2342">Okavango</collectingRegion>
Kwando
<collectingRegion id="7E29F846FFDAFFB15EFBFA5DFEC6FA16" box="[251,349,1430,1454]" country="Namibia" name="Zambezi" pageId="17" pageNumber="2342">Zambezi</collectingRegion>
System,
<taxonomicName id="7BED4D27FFDAFFB15FC8FA5CFD04FA16" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[456,671,1430,1454]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDAFFB15FC8FA5CFD04FA16" box="[456,671,1430,1454]" italics="true" pageId="17" pageNumber="2342">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFDAFFB15CA9FA5CFC9BFA16" box="[681,768,1431,1454]" pageId="17" pageNumber="2342" rank="species">sp. nov.</taxonomicNameLabel>
is characterized by a specific morphology and EOD in multivariate analysis, specific bands in genomic ISSR fingerprinting, and as a monophyletic taxon in mitochondrial DNA (mtDNA) cytochrome
<emphasis id="8E99EAB6FFDAFFB15A42FA1DFBD4FA56" box="[1090,1103,1494,1518]" italics="true" pageId="17" pageNumber="2342">b</emphasis>
analysis. To identify a specimen in hand it is best to rely on several characters in combination to exclude mistakes due to outliers. The 90th percentile of the distribution of HL (measured as HL/SL, head length to standard length) of
<taxonomicName id="7BED4D27FFDAFFB15D77F9FCFBCBF9F6" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[887,1104,1590,1614]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDAFFB15D77F9FCFBCBF9F6" box="[887,1104,1590,1614]" italics="true" pageId="17" pageNumber="2342">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFDAFFB15E8FF99DFF7FF9D5" box="[143,228,1622,1645]" pageId="17" pageNumber="2342" rank="species">sp. nov.</taxonomicNameLabel>
specimens, is shorter than the 10th percentile for
<taxonomicName id="7BED4D27FFDAFFB15D13F99CFC79F9D6" baseAuthorityName="Peters" baseAuthorityYear="1852" box="[787,994,1622,1646]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFDAFFB15D13F99CFC79F9D6" box="[787,994,1622,1646]" italics="true" pageId="17" pageNumber="2342">M. macrolepidotus</emphasis>
</taxonomicName>
(together with
<taxonomicName id="7BED4D27FFDAFFB15ECBF9BDFEFDF936" authorityName="Boulenger" authorityYear="1905" box="[203,358,1654,1678]" class="Actinopterygii" family="Mormyridae" genus="Gnathonemus" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="angolensis">
<emphasis id="8E99EAB6FFDAFFB15ECBF9BDFEFDF936" box="[203,358,1654,1678]" italics="true" pageId="17" pageNumber="2342">M. angolensis</emphasis>
</taxonomicName>
, shortest HL of all). The BD/SL ratio (body depth to standard length) of
<taxonomicName id="7BED4D27FFDAFFB15F0CF95DFE7DF915" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[268,486,1685,1709]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFDAFFB15F0CF95DFE7DF915" box="[268,486,1685,1709]" italics="true" pageId="17" pageNumber="2342">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFDAFFB15FF2F95DFDD0F915" box="[498,587,1686,1709]" pageId="17" pageNumber="2342" rank="species">sp. nov.</taxonomicNameLabel>
overlaps with that of
<taxonomicName id="7BED4D27FFDAFFB15D56F95DFBB3F915" baseAuthorityName="Peters" baseAuthorityYear="1852" box="[854,1064,1685,1709]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="17" pageNumber="2342" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFDAFFB15D56F95DFBB3F915" box="[854,1064,1685,1709]" italics="true" pageId="17" pageNumber="2342">M. macrolepidotus</emphasis>
</taxonomicName>
by less than one quartile; the same holds true for the distributions of nA (no. of anal fin rays), nD (no. of dorsal fin rays), LD/SL (ratio of dorsal fin length to standard length) in which the means or medians are greater for
<taxonomicName id="7BED4D27FFD9FFB25D52FF23FBB1FF47" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[850,1066,231,255]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25D52FF23FBB1FF47" box="[850,1066,231,255]" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25A35FF23FB14FF47" box="[1077,1167,232,255]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
, and SPc, number of scales around caudal peduncle in which the median for
<taxonomicName id="7BED4D27FFD9FFB25A38FEC3FECCFE87" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25A38FEC3FECCFE87" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25F5FFEE3FE2FFE87" box="[351,436,296,319]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
is smaller. The EOD of
<taxonomicName id="7BED4D27FFD9FFB25CC6FEE3FC07FE87" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[710,924,295,319]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25CC6FEE3FC07FE87" box="[710,924,295,319]" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25DA4FEE3FC62FE87" box="[932,1017,296,319]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
has a leading head-negativity of miniature amplitude that is usually not present in the
<taxonomicName id="7BED4D27FFD9FFB25DF8FE83FEC1FEC7" authority="EOD." authorityName="EOD." class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD9FFB25DF8FE83FE95FEC7" italics="true" pageId="18" pageNumber="2343">M. macrolepidotus</emphasis>
EOD.
</taxonomicName>
<taxonomicName id="7BED4D27FFD9FFB25F63FEA3FD3EFEC7" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[355,677,359,383]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25F63FEA3FD3EFEC7" box="[355,677,359,383]" italics="true" pageId="18" pageNumber="2343">Marcusenius multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25CAFFEA3FC9DFEC7" box="[687,774,360,383]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="7BED4D27FFD9FFB25D43FEA3FB8FFEC7" box="[835,1044,359,383]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="macrolepidotus">
<emphasis id="8E99EAB6FFD9FFB25D43FEA3FB8FFEC7" box="[835,1044,359,383]" italics="true" pageId="18" pageNumber="2343">M. macrolepidotus</emphasis>
</taxonomicName>
are clearly differentiated in ISSR bands 2, 6 and 8 (
<tableCitation id="F16F031FFFD9FFB25C97FE4CFD7CFE27" box="[663,743,391,415]" captionStart="Table 4" captionStartId="30.[213,268,169,191]" captionText="Table 4. Inter-simple-sequence-repeat profile of Marcusenius altisambesi, Marcusenius multisquamatus sp. nov. and Marcusenius macrolepidotus (selection of informative bands, numbers 1 to 13); x = band is present; formatting [bold/italic/bold-italic/clean]: highlighting bands characteristic of species." pageId="18" pageNumber="2343">Table 4</tableCitation>
).
</paragraph>
<paragraph id="BC5236A4FFD9FFB25EFEFE6DFB0BFCA6" blockId="18.[211,1172,167,798]" pageId="18" pageNumber="2343">
There is less than a 10% overlap of distributions between the greater LSo/HL (length of snout to head length) of
<taxonomicName id="7BED4D27FFD9FFB25C6AFE0CFCDAFE67" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[618,833,455,479]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25C6AFE0CFCDAFE67" box="[618,833,455,479]" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25D4AFE0CFC3AFE66" box="[842,929,455,478]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
specimens compared with that of
<taxonomicName id="7BED4D27FFD9FFB25F63FE2CFD93FE47" box="[355,520,487,511]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFD9FFB25F63FE2CFD93FE47" box="[355,520,487,511]" italics="true" pageId="18" pageNumber="2343">M. altisambesi</emphasis>
</taxonomicName>
from the Upper
<collectingRegion id="7E29F846FFD9FFB25CCDFE2DFCA9FE46" box="[717,818,486,510]" country="Namibia" name="Zambezi" pageId="18" pageNumber="2343">Zambezi</collectingRegion>
, and less than 25% overlap for
<taxonomicName id="7BED4D27FFD9FFB25ED3FDCCFEE2FDA6" box="[211,377,518,542]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFD9FFB25ED3FDCCFEE2FDA6" box="[211,377,518,542]" italics="true" pageId="18" pageNumber="2343">M. altisambesi</emphasis>
</taxonomicName>
from the
<collectingRegion id="7E29F846FFD9FFB25FF0FDCDFDFDFDA6" box="[496,614,518,542]" country="Namibia" name="Kavango West" pageId="18" pageNumber="2343">Okavango</collectingRegion>
. Also, there is less than one quartile overlap for LA/SL (anal fin length to SL), LSc/HL (length of snout to head length) and CL/HL (chin length to head length) of
<taxonomicName id="7BED4D27FFD9FFB25C28FD8CFD51FDE6" box="[552,714,582,606]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFD9FFB25C28FD8CFD51FDE6" box="[552,714,582,606]" italics="true" pageId="18" pageNumber="2343">M. altisambesi</emphasis>
</taxonomicName>
from the Upper
<collectingRegion id="7E29F846FFD9FFB25D84FD8DFC7DFDE6" box="[900,998,582,606]" country="Namibia" name="Zambezi" pageId="18" pageNumber="2343">Zambezi</collectingRegion>
compared with
<taxonomicName id="7BED4D27FFD9FFB25ED3FDACFE30FDC6" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[211,427,614,638]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25ED3FDACFE30FDC6" box="[211,427,614,638]" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25FB4FDACFD91FDC6" box="[436,522,615,638]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
and less than one quartile overlap for lower SLS (no. of lateral line scales) of
<taxonomicName id="7BED4D27FFD9FFB25FBDFD4CFDFBFD26" box="[445,608,646,670]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="altisambesi">
<emphasis id="8E99EAB6FFD9FFB25FBDFD4CFDFBFD26" box="[445,608,646,670]" italics="true" pageId="18" pageNumber="2343">M. altisambesi</emphasis>
</taxonomicName>
from the
<collectingRegion id="7E29F846FFD9FFB25CCEFD4DFCDAFD26" box="[718,833,646,670]" country="Namibia" name="Kavango West" pageId="18" pageNumber="2343">Okavango</collectingRegion>
compared with
<taxonomicName id="7BED4D27FFD9FFB25DF7FD4CFE8EFD06" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25DF7FD4CFE8EFD06" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25F1BFD6DFEEBFD06" box="[283,368,678,702]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
<collectingRegion id="7E29F846FFD9FFB25F76FD6DFE72FD06" box="[374,489,678,702]" country="Namibia" name="Kavango West" pageId="18" pageNumber="2343">Okavango</collectingRegion>
bulldog fish are distinguished by an SPc (no. of scales around caudal peduncle) of exclusively 12 (median 12, same median for Upper
<collectingRegion id="7E29F846FFD9FFB25DF6FD0EFBC3FD65" box="[1014,1112,709,733]" country="Namibia" name="Zambezi" pageId="18" pageNumber="2343">Zambezi</collectingRegion>
bulldog fish) whereas SPc ranges from
<quantity id="7B159B41FFD9FFB25C5EFD2EFD26FD45" box="[606,701,741,765]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" metricValueMax="4.064" metricValueMin="3.048" pageId="18" pageNumber="2343" unit="in" value="14.0" valueMax="16.0" valueMin="12.0">1216 in</quantity>
<taxonomicName id="7BED4D27FFD9FFB25CC5FD2DFC01FD46" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[709,922,742,766]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25CC5FD2DFC01FD46" box="[709,922,742,766]" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25DA2FD2DFC6CFD45" box="[930,1015,742,765]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
(median, 13). ISSR band 2 is specific for
<taxonomicName id="7BED4D27FFD9FFB25C02FCCDFD4CFCA6" authority="Kramer &amp; Wink, 2013" authorityName="Kramer &amp; Wink" authorityYear="2013" box="[514,727,774,798]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="multisquamatus" status="sp. nov.">
<emphasis id="8E99EAB6FFD9FFB25C02FCCDFD4CFCA6" box="[514,727,774,798]" italics="true" pageId="18" pageNumber="2343">M. multisquamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="95AA57CDFFD9FFB25CDEFCCDFCA9FCA5" box="[734,818,774,797]" pageId="18" pageNumber="2343" rank="species">sp. nov.</taxonomicNameLabel>
and band 6 for
<emphasis id="8E99EAB6FFD9FFB25DE5FCCDFB0BFCA6" box="[997,1168,774,798]" italics="true" pageId="18" pageNumber="2343">
<taxonomicName id="7BED4D27FFD9FFB25DE5FCCDFB17FCA6" box="[997,1164,774,798]" class="Actinopterygii" family="Mormyridae" genus="Marcusenius" kingdom="Animalia" order="Osteoglossiformes" pageId="18" pageNumber="2343" phylum="Chordata" rank="species" species="altisambesi">M. altisambesi</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>