187 lines
16 KiB
XML
187 lines
16 KiB
XML
<document ID-DOI="10.25221/fee.467.3" ID-ISSN="2713-2196" ID-Zenodo-Dep="7399592" approvalRequired="3" approvalRequired_for_taxonomicNames="2" approvalRequired_for_treatments="1" checkinTime="1670240547502" checkinUser="felipe" docAuthor="Aristov, D. S. & Rasnitsyn, A. P." docDate="2022" docId="307887CAFFEAA219FF40D7338CBFFBA5" docLanguage="en" docName="FarEastEntomol.467.17-22.pdf" docOrigin="Far Eastern Entomologist 467" docSource="http://dx.doi.org/10.25221/fee.467.3" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.2:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Permosialis ualentovae Novokshonov et Zhuzhgova 2004" docType="treatment" docVersion="3" lastPageNumber="19" masterDocId="CC41FFB2FFEBA21BFFA3D5078E3EFFA2" masterDocTitle="New and little known species of the genus Permosialis (Insecta: Palaeomanteida: Permosialidae) from the Middle Permian of Russia" masterLastPageNumber="22" masterPageNumber="17" pageNumber="18" updateTime="1672757110344" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New and little known species of the genus Permosialis (Insecta: Palaeomanteida: Permosialidae) from the Middle Permian of Russia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Aristov, D. S.</mods:namePart>
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<mods:affiliation>Borissak Paleontological Institute of the Russian Academy of Sciences, Profsoyuznaya str. 123, Moscow 117997, Russia. Cherepovets State University, Lunacharsky prospect 5, Cherepovets 162600, Vologda region, Russia.</mods:affiliation>
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<mods:nameIdentifier type="email">danil_aristov@mail.ru</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rasnitsyn, A. P.</mods:namePart>
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<mods:affiliation>Borissak Paleontological Institute of the Russian Academy of Sciences, Profsoyuznaya str. 123, Moscow 117997, Russia. Natural History Museum, London, UK.</mods:affiliation>
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<mods:nameIdentifier type="email">E-mail:</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Far Eastern Entomologist</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-12-02</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>467</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>17</mods:start>
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<mods:end>22</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.25221/fee.467.3</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.25221/fee.467.3</mods:identifier>
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<mods:identifier type="ISSN">2713-2196</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7399592</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:307887CAFFEAA219FF40D7338CBFFBA5" httpUri="http://treatment.plazi.org/id/307887CAFFEAA219FF40D7338CBFFBA5" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18">
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<subSubSection box="[227,807,564,586]" pageId="1" pageNumber="18" type="nomenclature">
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<paragraph blockId="1.[227,807,564,612]" box="[227,807,564,586]" pageId="1" pageNumber="18">
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<heading box="[227,807,564,586]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="18" reason="2">
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<taxonomicName authority="Novokshonov et Zhuzhgova, 2004" authorityName="Novokshonov et Zhuzhgova" authorityYear="2004" box="[227,807,564,586]" class="Insecta" family="Permosialidae" genus="Permosialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Miomoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="ualentovae">
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<emphasis bold="true" box="[227,807,564,586]" pageId="1" pageNumber="18">
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<emphasis bold="true" box="[227,457,564,586]" italics="true" pageId="1" pageNumber="18">Permosialis ualentovae</emphasis>
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Novokshonov et Zhuzhgova, 2004
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[227,310,590,612]" pageId="1" pageNumber="18" type="description">
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<paragraph blockId="1.[227,807,564,612]" box="[227,310,590,612]" pageId="1" pageNumber="18">
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<figureCitation box="[227,310,590,612]" captionStart="Figs 1, 2" captionStartId="1.[265,306,1585,1607]" captionTargetBox="[336,1017,669,1564]" captionTargetId="figure-102@1.[336,1018,665,1570]" captionTargetPageId="1" captionText="Figs 1, 2. Forewing of Permosialis ualentovae Novokshonov et Zhuzhgova, 2004, holotype PIN, No 1366/349. 1 – general appearance; 2 – reconstruction." figureDoi="http://doi.org/10.5281/zenodo.7399594" httpUri="https://zenodo.org/record/7399594/files/figure.png" pageId="1" pageNumber="18">Figs 1, 2</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection box="[265,1096,627,649]" pageId="1" pageNumber="18" type="reference_group">
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<paragraph blockId="1.[265,1096,627,649]" box="[265,1096,627,649]" pageId="1" pageNumber="18">
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<taxonomicName authority=": Novokshonov & Zhuzhgova, 2004: 183" authorityName=": Novokshonov & Zhuzhgova" authorityPageNumber="183" authorityYear="2004" box="[265,886,627,649]" class="Insecta" family="Permosialidae" genus="Permosialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Miomoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="ualentovae">
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<emphasis box="[265,491,627,648]" italics="true" pageId="1" pageNumber="18">Permosialis ualentovae</emphasis>
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: Novokshonov & Zhuzhgova, 2004: 183
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</taxonomicName>
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, fig. 6a, Pl. 10, fig. 2.
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</paragraph>
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</subSubSection>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7399594" ID-Zenodo-Dep="7399594" httpUri="https://zenodo.org/record/7399594/files/figure.png" pageId="1" pageNumber="18" startId="1.[265,306,1585,1607]" targetBox="[336,1017,669,1564]" targetPageId="1">
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<paragraph blockId="1.[227,1127,1585,1633]" pageId="1" pageNumber="18">
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Figs 1, 2. Forewing of
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<taxonomicName authority="Novokshonov et Zhuzhgova, 2004" authorityName="Novokshonov et Zhuzhgova" authorityYear="2004" box="[520,1123,1585,1607]" class="Insecta" family="Permosialidae" genus="Permosialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Miomoptera" pageId="1" pageNumber="18" phylum="Arthropoda" rank="species" species="ualentovae">
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<emphasis box="[520,753,1585,1606]" italics="true" pageId="1" pageNumber="18">Permosialis ualentovae</emphasis>
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Novokshonov et Zhuzhgova, 2004
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</taxonomicName>
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,
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<materialsCitation box="[227,916,1611,1633]" collectionCode="PIN" pageId="1" pageNumber="18" specimenCount="1" typeStatus="holotype">
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<typeStatus box="[227,311,1611,1633]" pageId="1" pageNumber="18">holotype</typeStatus>
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<collectionCode box="[317,358,1612,1633]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="1" pageNumber="18">PIN</collectionCode>
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, No 1366/349. 1 – general appearance; 2 – reconstruction
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</materialsCitation>
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.
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</paragraph>
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</caption>
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<subSubSection pageId="2" pageNumber="19" type="materials_examined">
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<paragraph blockId="2.[227,1127,262,1031]" pageId="2" pageNumber="19">
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<materialsCitation box="[265,1127,262,284]" collectionCode="PIN" country="Russia" location="No" pageId="2" pageNumber="19" specimenCount="1" typeStatus="holotype">
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MATERIAL.
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<typeStatus box="[405,494,262,284]" pageId="2" pageNumber="19">Holotype</typeStatus>
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<collectionCode box="[505,546,262,283]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="2" pageNumber="19">PIN</collectionCode>
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,
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<location LSID="urn:lsid:plazi:treatment:307887CAFFEAA219FF40D7338CBFFBA5:BD0E6007FFE9A219FD93D4018C73FEB9" box="[560,589,262,283]" country="Russia" name="No" pageId="2" pageNumber="19">No</location>
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1366/349, counterpart of forewing fragment;
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<collectingCountry box="[1057,1123,262,284]" name="Russia" pageId="2" pageNumber="19">Russia</collectingCountry>
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,
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</materialsCitation>
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<materialsCitation box="[227,1127,290,312]" location="Kityak River" municipality="Malmyzh district" pageId="2" pageNumber="19" specimenCount="1" stateProvince="Kirov">
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<collectingRegion box="[227,353,290,312]" country="Russia" name="Kirov" pageId="2" pageNumber="19">Kirov region</collectingRegion>
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,
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<collectingMunicipality box="[363,530,290,312]" pageId="2" pageNumber="19">Malmyzh district</collectingMunicipality>
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, left bank of the
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<location LSID="urn:lsid:plazi:treatment:307887CAFFEAA219FF40D7338CBFFBA5:BD0E6007FFE9A219FD17D4258D0FFE9A" box="[692,817,290,312]" municipality="Malmyzh district" name="Kityak River" pageId="2" pageNumber="19" stateProvince="Kirov">Kityak River</location>
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opposite the village of Bol’shoi
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</materialsCitation>
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Kityak; Upper Permian, Upper Kazanian, Belebei Formation; in Borissiak Paleontological Institute, Russian Academy of Sciences (
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<collectionCode box="[620,660,348,369]" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="2" pageNumber="19">PIN</collectionCode>
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RAS).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="19" type="diagnosis">
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<paragraph blockId="2.[227,1127,262,1031]" pageId="2" pageNumber="19">
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DIAGNOSIS. Differs from all other species of
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<taxonomicName authorityName="Martynov" authorityYear="1928" box="[720,835,377,398]" class="Insecta" family="Permosialidae" genus="Permosialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Miomoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="genus">
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<emphasis box="[720,835,377,398]" italics="true" pageId="2" pageNumber="19">Permosialis</emphasis>
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</taxonomicName>
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by color pattern (three pairs of margined pale spots on darkened background) and venationally from all but
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<taxonomicName authority="Aristov & Rasnitsyn, 2022" authorityName="Aristov & Rasnitsyn" authorityYear="2022" box="[987,1082,405,427]" class="Insecta" family="Permosialidae" genus="Permosialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Miomoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="belebei" status="sp.">
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<emphasis box="[987,1082,405,427]" italics="true" pageId="2" pageNumber="19">P. belebei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1089,1117,406,427]" pageId="2" pageNumber="19">sp.</taxonomicNameLabel>
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nov. by two RS forks placed close to each other and far from both R base and wing margin, and from
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<taxonomicName authority="Aristov & Rasnitsyn, 2022" authorityName="Aristov & Rasnitsyn" authorityYear="2022" box="[320,416,463,485]" class="Insecta" family="Permosialidae" genus="Permosialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Miomoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="belebei" status="sp. n.">
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<emphasis box="[320,416,463,485]" italics="true" pageId="2" pageNumber="19">P. belebei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[422,474,463,484]" pageId="2" pageNumber="19" rank="species">sp. n.</taxonomicNameLabel>
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(as well as from all or most of other congeners) by reduced (short and narrow) fork of CuA.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="19" type="description">
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<paragraph blockId="2.[227,1127,262,1031]" pageId="2" pageNumber="19">
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DESCRIPTION. Forewing comparatively narrow (not as wide oval as
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<taxonomicName authorityName="O. Martynova" authorityYear="1961" box="[960,1062,520,542]" class="Insecta" family="Permosialidae" genus="Permosialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Miomoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="sibirica">
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<emphasis box="[960,1062,520,542]" italics="true" pageId="2" pageNumber="19">P. sibirica</emphasis>
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</taxonomicName>
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), with characheristic color pattern of round dark margined pale spots on somewhat darkened background: two spots in subcostal area and one in each interradial, radiomedial, mediocubital and cubitoanal ones that form three vertical pairs (first subcostal spot with cubitoanal one, second subcostal with radiomedial, and interradial with mediocubital one). Costal and subcostal spaces both moderately wide (costal one wider), SC almost straight, with few almost regularly spaced short branchlets (6 visible). RS 3-branched, both forks placed close to each other but not to RS base. M stalk apparently free, fork leveled with first RS fork. M
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<subScript attach="right" box="[1119,1127,731,745]" fontSize="6" pageId="2" pageNumber="19">5</subScript>
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not preserved. CuA stalk and branches concave, fork short, narrow. CuP curved apical. 1A concave.
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</paragraph>
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<paragraph blockId="2.[227,1127,262,1031]" pageId="2" pageNumber="19">
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MEASUREMENTS. Forewing fragment length
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<quantity box="[736,825,808,830]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="2" pageNumber="19" unit="mm" value="12.6">12.6 mm</quantity>
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, supposed full length about 16 mm, width
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<quantity box="[337,413,836,858]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="19" unit="mm" value="7.5">7.5 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="19" type="discussion">
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<paragraph blockId="2.[227,1127,262,1031]" pageId="2" pageNumber="19">
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<materialsCitation collectionCode="NOTES" collectorName="Novokshonov & Zhuzhgova" pageId="2" pageNumber="19" specimenCount="1" typeStatus="paratype">
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<collectionCode box="[265,345,865,886]" pageId="2" pageNumber="19">NOTES</collectionCode>
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.
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<collectorName box="[359,494,865,887]" pageId="2" pageNumber="19">Novokshonov</collectorName>
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&
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<collectorName box="[535,643,865,887]" pageId="2" pageNumber="19">Zhuzhgova</collectorName>
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(2004: 183) list
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<specimenCount box="[823,975,865,887]" pageId="2" pageNumber="19" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
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of the present species. Of them, 1366/379 and 1366/463 are described as a different species
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<taxonomicName authority="Aristov & Rasnitsyn, 2022" authorityName="Aristov & Rasnitsyn" authorityYear="2022" box="[972,1068,894,916]" class="Insecta" family="Permosialidae" genus="Permosialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Miomoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="belebei" status="sp. n.">
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<emphasis box="[972,1068,894,916]" italics="true" pageId="2" pageNumber="19">P. belebei</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1075,1123,895,916]" pageId="2" pageNumber="19" rank="species">sp. n</taxonomicNameLabel>
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</materialsCitation>
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. herein (see below), and PIN, No. 1366/382 represents a fragmentary hind wing which can be hardly ascribed to
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<taxonomicName authorityName="Novokshonov et Zhuzhgova" authorityYear="2004" box="[418,554,951,973]" class="Insecta" family="Permosialidae" genus="Permosialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Miomoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="ualentovae">
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<emphasis box="[418,554,951,973]" italics="true" pageId="2" pageNumber="19">P. ualentovae</emphasis>
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</taxonomicName>
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because it shows a darker subapical transverse belt (vs. margined spots in fore wings of
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<taxonomicName authorityName="Novokshonov et Zhuzhgova" authorityYear="2004" box="[543,675,980,1002]" class="Insecta" family="Permosialidae" genus="Permosialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Miomoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="ualentovae">
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<emphasis box="[543,675,980,1002]" italics="true" pageId="2" pageNumber="19">P. ualentovae</emphasis>
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</taxonomicName>
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) and would be rather treated as
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<taxonomicName box="[982,1127,980,1002]" class="Insecta" family="Permosialidae" genus="Permosialis" kingdom="Animalia" order="Miomoptera" pageId="2" pageNumber="19" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis box="[982,1094,980,1001]" italics="true" pageId="2" pageNumber="19">Permosialis</emphasis>
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sp.
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</taxonomicName>
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until more conclusive information appears.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |