347 lines
47 KiB
XML
347 lines
47 KiB
XML
<document id="7FB299D37D7FE8483E0ABAC1686AE44F" ID="10.11646/zootaxa.4216.6.3" ID-CLB-Dataset="37326" ID-DOI="10.5281/zenodo.242442" ID-GBIF-Dataset="cd7b958e-d906-4cac-8582-f412619f550d" ID-ISSN="1175-5326" ID-Zenodo-Dep="242442" ID-ZooBank="099626D3-839B-438D-94B8-C89A5227A574" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484320646478" checkinUser="plazi" docAuthor="Ojeda-Escoto, Juan Carlos, Villalobos, José Luis & Álvarez, Fernando" docDate="2017" docId="2F0387BA9B1FFFEC86F7FEADFCA71090" docLanguage="en" docName="zootaxa.4216.6.3.pdf" docOrigin="Zootaxa 4216 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudothelphusa ixtapan Ojeda-Escoto, Villalobos & Álvarez, 2017, n. sp." docType="treatment" docVersion="7" lastPageNumber="566" masterDocId="D33AFFC29B1AFFEB8660FFE1FF8C1728" masterDocTitle="Three new species of freshwater crabs of the genus Pseudothelphusa (De Saussure, 1857) (Crustacea: Decapoda: Pseudothelphusidae) from Mexico" masterLastPageNumber="571" masterPageNumber="559" pageNumber="564" updateTime="1698435521958" updateUser="ExternalLinkService">
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<mods:title id="B1F628A7AED21E4EB47A2E37E07D37F6">Three new species of freshwater crabs of the genus Pseudothelphusa (De Saussure, 1857) (Crustacea: Decapoda: Pseudothelphusidae) from Mexico</mods:title>
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<mods:namePart id="8D457AE36BF7896F00E56771E832FA61">Ojeda-Escoto, Juan Carlos</mods:namePart>
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<mods:namePart id="6633F26868D62A1F5367A93FF40D87AE">Villalobos, José Luis</mods:namePart>
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<mods:namePart id="CAE40DE9459A2E83D68722134136909A">Álvarez, Fernando</mods:namePart>
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<taxonomicName id="60AA4D2F9B1FFFEE86F7FEADFE44164E" ID-CoL="4PD9D" box="[151,456,332,358]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="ixtapan" status="sp. nov.">
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<emphasis id="95DEEABE9B1FFFEE86F7FEADFE44164E" bold="true" box="[151,456,332,358]" italics="true" pageId="5" pageNumber="564">Pseudothelphusa ixtapan</emphasis>
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<taxonomicNameLabel id="8EED57C59B1FFFEE87AFFEADFD9D164E" box="[463,529,332,358]" pageId="5" pageNumber="564" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<paragraph id="A71536AC9B1FFFEE86F7FE8FFE9E16AF" blockId="5.[151,529,332,391]" box="[151,274,366,391]" pageId="5" pageNumber="564">
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(
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<figureCitation id="3F912A299B1FFFEE86FFFE8FFF6216AF" box="[159,238,366,391]" captionStart="FIGURE 4" captionStartId="6.[151,250,1958,1980]" captionTargetBox="[206,1388,635,1933]" captionTargetId="figure@6.[198,1388,625,1937]" captionTargetPageId="6" captionText="FIGURE 4. Pseudothelphusa ixtapan n. sp., male holotype, CNCR 343: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242446/files/figure.png" pageId="5" pageNumber="564">Figs. 4</figureCitation>
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,
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<figureCitation id="3F912A299B1FFFEE869AFE8FFE8416AF" box="[250,264,366,391]" captionStart="FIGURE 5" captionStartId="7.[151,250,1752,1774]" captionTargetBox="[154,1436,193,1726]" captionTargetId="figure@7.[151,1436,193,1730]" captionTargetPageId="7" captionText="FIGURE 5. Pseudothelphusa ixtapan n. sp., male holotype CNCR 343, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudo-marginal projection; mp, mesial process; ms, marginal suture; osc, opening of sperm channel." httpUri="https://zenodo.org/record/242447/files/figure.png" pageId="5" pageNumber="564">5</figureCitation>
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)
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</paragraph>
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<subSubSection id="EFB065279B1FFFEE86F7FE57FB271522" pageId="5" pageNumber="564" type="reference_group">
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<treatmentCitationGroup id="87BA11829B1FFFEE86F7FE57FD5916C3" pageId="5" pageNumber="564">
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<taxonomicName id="60AA4D2F9B1FFFEE86F7FE57FC7F16E4" ID-CoL="6WHWG" authority="Rodriguez & Smalley, 1969: 81" authorityName="Rodriguez & Smalley" authorityPageNumber="81" authorityYear="1969" box="[151,1011,438,460]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="564" phylum="Arthropoda" rank="subSpecies" species="dugesi" subGenus="Pseudothelphusa" subSpecies="belliana">
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<emphasis id="95DEEABE9B1FFFEE86F7FE57FEC816E4" box="[151,324,438,460]" italics="true" pageId="5" pageNumber="564">Pseudothelphusa</emphasis>
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(
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<emphasis id="95DEEABE9B1FFFEE8733FE57FD8C16E4" box="[339,512,438,460]" italics="true" pageId="5" pageNumber="564">Pseudothelphusa</emphasis>
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)
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<emphasis id="95DEEABE9B1FFFEE846EFE57FD2516E4" box="[526,681,438,460]" italics="true" pageId="5" pageNumber="564">dugesi belliana</emphasis>
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<treatmentCitation id="260B10BD9B1FFFEE84D0FE57FC7F16E4" author="Rodriguez" box="[688,1011,438,460]" page="81" pageId="5" pageNumber="564" year="1969">
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<bibRefCitation id="C33B4B5D9B1FFFEE84D0FE57FC4216E4" author="Rodriguez" box="[688,974,438,460]" pageId="5" pageNumber="564" refString="Rodriguez, G. & Smalley, A. E. (1969) Los cangrejos de agua dulce de Mexico de la familia Pseudothelphusidae (Crustacea, Brachyura). Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Serie Ciencias del Mar y Limnologia, 1, 69 - 112." type="journal article" year="1969">Rodríguez & Smalley, 1969</bibRefCitation>
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: 81
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</treatmentCitation>
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</taxonomicName>
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, fig. 13, pl. 9.
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<taxonomicName id="60AA4D2F9B1FFFEE86F7FE34FD0D16C3" ID-CoL="6WHWG" authority="Rodriguez, 1982: 135" authorityName="Rodriguez" authorityPageNumber="135" authorityYear="1982" box="[151,641,469,491]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="belliana">
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<emphasis id="95DEEABE9B1FFFEE86F7FE34FE1116C3" box="[151,413,469,491]" italics="true" pageId="5" pageNumber="564">Pseudothelphusa belliana</emphasis>
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<treatmentCitation id="260B10BD9B1FFFEE87C3FE34FD0D16C3" author="Rodriguez" box="[419,641,469,491]" page="135" pageId="5" pageNumber="564" year="1982">
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<bibRefCitation id="C33B4B5D9B1FFFEE87C3FE34FDDC16C3" author="Rodriguez" box="[419,592,469,491]" pageId="5" pageNumber="564" refString="Rodriguez, G. (1982) Les crabes d'eau douce d'Amerique. Famille des Pseudothelphusidae. Faune Tropicale, 22, 1 - 223." type="journal article" year="1982">Rodríguez, 1982</bibRefCitation>
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: 135
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</treatmentCitation>
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</taxonomicName>
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, fig. 88.
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</paragraph>
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<paragraph id="A71536AC9B1FFFEE86F7FE15FB271522" blockId="5.[151,1195,438,522]" box="[151,1195,500,522]" pageId="5" pageNumber="564">
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<treatmentCitationGroup id="87BA11829B1FFFEE86F7FE15FB271522" box="[151,1195,500,522]" pageId="5" pageNumber="564">
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<taxonomicName id="60AA4D2F9B1FFFEE86F7FE15FDED1522" ID-CoL="6WHWG" box="[151,609,500,522]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="564" phylum="Arthropoda" rank="species" species="belliana" subGenus="Pseudothelphusa">
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<emphasis id="95DEEABE9B1FFFEE86F7FE15FEC81522" box="[151,324,500,522]" italics="true" pageId="5" pageNumber="564">Pseudothelphusa</emphasis>
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(
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<emphasis id="95DEEABE9B1FFFEE8733FE15FD8C1522" box="[339,512,500,522]" italics="true" pageId="5" pageNumber="564">Pseudothelphusa</emphasis>
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)
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<emphasis id="95DEEABE9B1FFFEE846EFE15FDED1522" box="[526,609,500,522]" italics="true" pageId="5" pageNumber="564">belliana</emphasis>
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</taxonomicName>
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form
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<taxonomicName id="60AA4D2F9B1FFFEE84FFFE14FD671522" ID-CoL="4PD9D" box="[671,747,501,522]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="ixtapan" status="sp. nov.">Ixtapan</taxonomicName>
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de la Panocha,
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<treatmentCitation id="260B10BD9B1FFFEE85EAFE15FBEC1522" author="Villalobos" box="[906,1120,500,522]" page="248" pageId="5" pageNumber="564" year="2005">
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<bibRefCitation id="C33B4B5D9B1FFFEE85EAFE15FBBA1522" author="Villalobos" box="[906,1078,500,522]" pageId="5" pageNumber="564" refString="Villalobos, J. L. (2005) Sistematica de los cangrejos de agua dulce de Mexico, tribu Pseudothelphusini Ortmann, 1897 (Crustacea: Decapoda: Brachyura: Pseudothelphusidae). Analisis filogenetico, biogeografico y descripcion de especies nuevas. Ph. D. Dissertation, Facultad de Ciencias, Universidad Nacional Autonoma de Mexico, 394 pp." type="book" year="2005">Villalobos, 2005</bibRefCitation>
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:248
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</treatmentCitation>
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, pl. 18.
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material.
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, cl
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, cb
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;
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near
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<taxonomicName id="60AA4D2F9B1FFFEE8244FDD6FBF41566" box="[1060,1144,567,590]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="ixtapan" status="sp. nov.">Ixtapan</taxonomicName>
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<collectorName id="0A5F537A9B1FFFEE82E1FDD7FAAC1567" box="[1153,1312,566,591]" pageId="5" pageNumber="564">de la Panocha</collectorName>
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(
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,
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;
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</quantity>
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asl),
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<location id="A27560779B1FFFEE87D6FDBAFDB5155B" LSID="urn:lsid:plazi:treatment:2F0387BA9B1FFFEC86F7FEADFCA71090:A27560779B1FFFEE87D6FDBAFDB5155B" box="[438,569,602,627]" country="Mexico" county="Tejupilco" latitude="18.9" longLatPrecision="1273" longitude="-100.15" municipality="Municipality" name="San Miguel" pageId="5" pageNumber="564">San Miguel</location>
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<taxonomicName id="60AA4D2F9B1FFFEE8421FDBAFD14155A" box="[577,664,603,626]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="566" phylum="Arthropoda" rank="species" species="ixtapan" status="sp. nov.">Ixtapan</taxonomicName>
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,
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of
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, state of
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,
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<collectingCountry id="DFBD763C9B1FFFEE82F1FDBBFB66155B" box="[1169,1258,602,627]" name="Mexico" pageId="5" pageNumber="564">Mexico</collectingCountry>
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;
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<collectingDate id="C350E9849B1FFFEE8296FDBBFA1B155B" box="[1270,1431,602,627]" pageId="5" pageNumber="564" value="1961-04-25">25 April 1961</collectingDate>
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</date>
|
|
</materialsCitation>
|
|
; coll. anonymous; CNCR 343. This individual was considered as
|
|
<taxonomicName id="60AA4D2F9B1FFFEE8512FD9FFB6915BE" box="[882,1253,638,662]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="564" phylum="Arthropoda" rank="subSpecies" species="dugesi" subSpecies="belliana">
|
|
<emphasis id="95DEEABE9B1FFFEE8512FD9FFB6915BE" box="[882,1253,638,662]" italics="true" pageId="5" pageNumber="564">Pseudothelphusa dugesi belliana</emphasis>
|
|
</taxonomicName>
|
|
and included as part of the material examined by
|
|
<bibRefCitation id="C33B4B5D9B1FFFEE846BFD43FCCB1593" author="Rodriguez" box="[523,839,674,699]" pageId="5" pageNumber="564" refString="Rodriguez, G. & Smalley, A. E. (1969) Los cangrejos de agua dulce de Mexico de la familia Pseudothelphusidae (Crustacea, Brachyura). Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, Serie Ciencias del Mar y Limnologia, 1, 69 - 112." type="journal article" year="1969">Rodríguez & Smalley, 1969</bibRefCitation>
|
|
: 81, pl. 9 catalog number UNAM
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|
<date id="D314106C9B1FFFEE82A5FD42FA901593" box="[1221,1308,674,699]" pageId="5" pageNumber="564" value="1961-04-25">4 25 61</date>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="EFB065279B1FFFED86A7FD27FC191635" lastPageId="6" lastPageNumber="565" pageId="5" pageNumber="564" type="description">
|
|
<paragraph id="A71536AC9B1FFFEE86A7FD27FB85136F" blockId="5.[151,1437,566,2031]" pageId="5" pageNumber="564">
|
|
<emphasis id="95DEEABE9B1FFFEE86A7FD27FEDD15F7" bold="true" box="[199,337,710,735]" pageId="5" pageNumber="564">Description</emphasis>
|
|
. Dorsal surface of carapace smooth, central portion flat, puntacted (
|
|
<figureCitation id="3F912A299B1FFFEE8218FD27FB4C15F7" box="[1144,1216,710,735]" captionStart="FIGURE 4" captionStartId="6.[151,250,1958,1980]" captionTargetBox="[206,1388,635,1933]" captionTargetId="figure@6.[198,1388,625,1937]" captionTargetPageId="6" captionText="FIGURE 4. Pseudothelphusa ixtapan n. sp., male holotype, CNCR 343: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242446/files/figure.png" pageId="5" pageNumber="564">Fig. 4</figureCitation>
|
|
A). Frontal portion lacking superior frontal border, curved downward reaching inferior frontal border, partially visible dorsally (
|
|
<figureCitation id="3F912A299B1FFFEE830EFD0BFF2A140F" captionStart="FIGURE 4" captionStartId="6.[151,250,1958,1980]" captionTargetBox="[206,1388,635,1933]" captionTargetId="figure@6.[198,1388,625,1937]" captionTargetPageId="6" captionText="FIGURE 4. Pseudothelphusa ixtapan n. sp., male holotype, CNCR 343: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242446/files/figure.png" pageId="5" pageNumber="564">Fig. 4</figureCitation>
|
|
A, D). Front smooth, moderately bilobed, inferior frontal border slightly projected, sinuous, with evident middle concavity, slight convexity at level of antennular fossae (
|
|
<figureCitation id="3F912A299B1FFFEE8551FCD3FCF51463" box="[817,889,818,843]" captionStart="FIGURE 4" captionStartId="6.[151,250,1958,1980]" captionTargetBox="[206,1388,635,1933]" captionTargetId="figure@6.[198,1388,625,1937]" captionTargetPageId="6" captionText="FIGURE 4. Pseudothelphusa ixtapan n. sp., male holotype, CNCR 343: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242446/files/figure.png" pageId="5" pageNumber="564">Fig. 4</figureCitation>
|
|
D). Postfrontal lobes well formed with shallow anterior depression. Frontal portion of carapace anterior to postfrontal lobes convex. Median frontal groove shallow, well-marked, dividing front anteriorly reaching inferior frontal border; posteriorly separating postfrontal lobes, becoming obsolete little after. Cervical grooves well marked, wide, straight, not reaching anterolateral margin; posterior end curved. Regions moderately indicated, gastric, branchial swollen; cardiac, intestinal depressed (
|
|
<figureCitation id="3F912A299B1FFFEE8776FC07FED714D7" box="[278,347,998,1023]" captionStart="FIGURE 4" captionStartId="6.[151,250,1958,1980]" captionTargetBox="[206,1388,635,1933]" captionTargetId="figure@6.[198,1388,625,1937]" captionTargetPageId="6" captionText="FIGURE 4. Pseudothelphusa ixtapan n. sp., male holotype, CNCR 343: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242446/files/figure.png" pageId="5" pageNumber="564">Fig. 4</figureCitation>
|
|
A). Anterolateral margin moderately prominent in lateral view, serrated, with 19 blunt tubercles on left side, 23 on right side, from cervical groove to mid branchial region; area between orbit, cervical groove depressed with fused granules. Posterior margin of carapace straight (
|
|
<figureCitation id="3F912A299B1FFFEE85C2FBCFFC69136F" box="[930,997,1070,1095]" captionStart="FIGURE 4" captionStartId="6.[151,250,1958,1980]" captionTargetBox="[206,1388,635,1933]" captionTargetId="figure@6.[198,1388,625,1937]" captionTargetPageId="6" captionText="FIGURE 4. Pseudothelphusa ixtapan n. sp., male holotype, CNCR 343: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242446/files/figure.png" pageId="5" pageNumber="564">Fig. 4</figureCitation>
|
|
A).
|
|
</paragraph>
|
|
<paragraph id="A71536AC9B1FFFEE86A7FBB3FA911287" blockId="5.[151,1437,566,2031]" pageId="5" pageNumber="564">
|
|
External angle of orbit faintly developed, with shallow basal notch; internal angle extending into interior of orbital cavity as moderate keel (
|
|
<figureCitation id="3F912A299B1FFFEE879FFB97FDC813A7" box="[511,580,1142,1167]" captionStart="FIGURE 4" captionStartId="6.[151,250,1958,1980]" captionTargetBox="[206,1388,635,1933]" captionTargetId="figure@6.[198,1388,625,1937]" captionTargetPageId="6" captionText="FIGURE 4. Pseudothelphusa ixtapan n. sp., male holotype, CNCR 343: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242446/files/figure.png" pageId="5" pageNumber="564">Fig. 4</figureCitation>
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|
D). Basal article of antennal peduncle separated from inferior frontal margin by narrow hiatus. In frontal view, antennules, antennular fossae, interantennular septum concealed by front (
|
|
<figureCitation id="3F912A299B1FFFEE830FFB7BFF2B13FF" captionStart="FIGURE 4" captionStartId="6.[151,250,1958,1980]" captionTargetBox="[206,1388,635,1933]" captionTargetId="figure@6.[198,1388,625,1937]" captionTargetPageId="6" captionText="FIGURE 4. Pseudothelphusa ixtapan n. sp., male holotype, CNCR 343: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242446/files/figure.png" pageId="5" pageNumber="564">Fig. 4</figureCitation>
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|
D); internal margin of second antennular article with small distal projection; ventrally antennular fossae wider in middle, inferior frontal margin slightly sinuous. Opercular plate of antennal gland ovoidal, with small central constriction, tuft of setae on external third. Epistome, portions surrounding buccal cavity with dense patch of setae; lateral portions with rounded granules; epistomal tooth triangular, apex directed downward at same level of interantennular septum (
|
|
<figureCitation id="3F912A299B1FFFEE87CAFAAFFE7C124F" box="[426,496,1358,1383]" captionStart="FIGURE 4" captionStartId="6.[151,250,1958,1980]" captionTargetBox="[206,1388,635,1933]" captionTargetId="figure@6.[198,1388,625,1937]" captionTargetPageId="6" captionText="FIGURE 4. Pseudothelphusa ixtapan n. sp., male holotype, CNCR 343: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242446/files/figure.png" pageId="5" pageNumber="564">Fig. 4</figureCitation>
|
|
D). Opening of efferent branchial channel subquadrangular, ratio wide/length 1.36. Ischium of third maxilliped trapezoidal, slightly longer than wide; merus narrower than ischium, superior margin rounded, with shallow, rounded notch before the insertion of palp; ratio exopod/ischium 0.67 (
|
|
<figureCitation id="3F912A299B1FFFEE82D7FA77FB701287" box="[1207,1276,1430,1455]" captionStart="FIGURE 4" captionStartId="6.[151,250,1958,1980]" captionTargetBox="[206,1388,635,1933]" captionTargetId="figure@6.[198,1388,625,1937]" captionTargetPageId="6" captionText="FIGURE 4. Pseudothelphusa ixtapan n. sp., male holotype, CNCR 343: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242446/files/figure.png" pageId="5" pageNumber="564">Fig. 4</figureCitation>
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|
C).
|
|
</paragraph>
|
|
<paragraph id="A71536AC9B1FFFEE86A7FA5BFBAD1117" blockId="5.[151,1437,566,2031]" pageId="5" pageNumber="564">
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|
Chelipeds asymmetrical. Merus of larger cheliped with row of blunt tubercles along internal margin, carpus with triangular spine on internal margin. Chela with internal surface smooth, globose; fingers gaping, curved inward distally, with low, triangular teeth on cutting edges (
|
|
<figureCitation id="3F912A299B1FFFEE853EF9E3FC2B1133" box="[862,935,1538,1563]" captionStart="FIGURE 4" captionStartId="6.[151,250,1958,1980]" captionTargetBox="[206,1388,635,1933]" captionTargetId="figure@6.[198,1388,625,1937]" captionTargetPageId="6" captionText="FIGURE 4. Pseudothelphusa ixtapan n. sp., male holotype, CNCR 343: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace." httpUri="https://zenodo.org/record/242446/files/figure.png" pageId="5" pageNumber="564">Fig. 4</figureCitation>
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|
B). Dactylus rather narrow, widely curved, dorsally ornamented with several longitudinal rows of small pits, black granules.
|
|
</paragraph>
|
|
<paragraph id="A71536AC9B1FFFED86A7F9ABFE8F17D0" blockId="5.[151,1437,566,2031]" lastBlockId="6.[151,1437,151,573]" lastPageId="6" lastPageNumber="565" pageId="5" pageNumber="564">
|
|
Male gonopod moderately strong. In mesial view, distal crest of caudomarginal projection curved; distal lobe as triangular subacute tooth, separated from proximal lobe by U-shaped notch, with apex not reaching beyond cephalic margin (
|
|
<figureCitation id="3F912A299B1FFFEE8701F973FE2B1183" box="[353,423,1682,1707]" captionStart="FIGURE 5" captionStartId="7.[151,250,1752,1774]" captionTargetBox="[154,1436,193,1726]" captionTargetId="figure@7.[151,1436,193,1730]" captionTargetPageId="7" captionText="FIGURE 5. Pseudothelphusa ixtapan n. sp., male holotype CNCR 343, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudo-marginal projection; mp, mesial process; ms, marginal suture; osc, opening of sperm channel." httpUri="https://zenodo.org/record/242447/files/figure.png" pageId="5" pageNumber="564">Fig. 5</figureCitation>
|
|
A). Proximal lobe longer than wide, oval-shaped, proximally directed; cephalic margin curved (
|
|
<figureCitation id="3F912A299B1FFFEE8696F957FEB711E7" box="[246,315,1718,1743]" captionStart="FIGURE 5" captionStartId="7.[151,250,1752,1774]" captionTargetBox="[154,1436,193,1726]" captionTargetId="figure@7.[151,1436,193,1730]" captionTargetPageId="7" captionText="FIGURE 5. Pseudothelphusa ixtapan n. sp., male holotype CNCR 343, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudo-marginal projection; mp, mesial process; ms, marginal suture; osc, opening of sperm channel." httpUri="https://zenodo.org/record/242447/files/figure.png" pageId="5" pageNumber="564">Fig. 5</figureCitation>
|
|
A). In cephalic view, distal lobe of caudo-marginal projection directed cephalically, aligned with proximal lobe; internal border close to inner expansion of mesial process. Proximal lobe slightly oblique with respect to longitudinal axis of gonopod; distal half of internal surface swollen with ill-defined border; cephalic margin straight (
|
|
<figureCitation id="3F912A299B1FFFEE873FF8C3FE261013" box="[351,426,1826,1851]" captionStart="FIGURE 5" captionStartId="7.[151,250,1752,1774]" captionTargetBox="[154,1436,193,1726]" captionTargetId="figure@7.[151,1436,193,1730]" captionTargetPageId="7" captionText="FIGURE 5. Pseudothelphusa ixtapan n. sp., male holotype CNCR 343, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudo-marginal projection; mp, mesial process; ms, marginal suture; osc, opening of sperm channel." httpUri="https://zenodo.org/record/242447/files/figure.png" pageId="5" pageNumber="564">Fig. 5</figureCitation>
|
|
B). Mesial process as rectangular laminar expansion, distal margin perpendicular to longitudinal axis of gonopod, lateral margin with acute median tooth, series of small acute teeth on proximal angle (
|
|
<figureCitation id="3F912A299B1FFFEE86FFF88BFF6B10AB" box="[159,231,1898,1923]" captionStart="FIGURE 5" captionStartId="7.[151,250,1752,1774]" captionTargetBox="[154,1436,193,1726]" captionTargetId="figure@7.[151,1436,193,1730]" captionTargetPageId="7" captionText="FIGURE 5. Pseudothelphusa ixtapan n. sp., male holotype CNCR 343, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudo-marginal projection; mp, mesial process; ms, marginal suture; osc, opening of sperm channel." httpUri="https://zenodo.org/record/242447/files/figure.png" pageId="5" pageNumber="564">Fig. 5</figureCitation>
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|
B). Superior margin almost straight, ending internally in rectangular inner expansion, rounded, closing apical cavity; same margin developing into central crest in apical cavity, delimiting field of spines. Inferior margin widely rounded. In lateral, mesial views, apical cavity oriented caudo-cephalically (
|
|
<figureCitation id="3F912A299B1FFFEE8238F853FB1110E3" box="[1112,1181,1970,1995]" captionStart="FIGURE 5" captionStartId="7.[151,250,1752,1774]" captionTargetBox="[154,1436,193,1726]" captionTargetId="figure@7.[151,1436,193,1730]" captionTargetPageId="7" captionText="FIGURE 5. Pseudothelphusa ixtapan n. sp., male holotype CNCR 343, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudo-marginal projection; mp, mesial process; ms, marginal suture; osc, opening of sperm channel." httpUri="https://zenodo.org/record/242447/files/figure.png" pageId="5" pageNumber="564">Fig. 5</figureCitation>
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|
A, C). In caudal view, distal third straight, tilted mesially, distal crest concave (
|
|
<figureCitation id="3F912A299B1FFFEE854AF837FCFC10C7" box="[810,880,2006,2031]" captionStart="FIGURE 5" captionStartId="7.[151,250,1752,1774]" captionTargetBox="[154,1436,193,1726]" captionTargetId="figure@7.[151,1436,193,1730]" captionTargetPageId="7" captionText="FIGURE 5. Pseudothelphusa ixtapan n. sp., male holotype CNCR 343, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudo-marginal projection; mp, mesial process; ms, marginal suture; osc, opening of sperm channel." httpUri="https://zenodo.org/record/242447/files/figure.png" pageId="5" pageNumber="564">Fig. 5</figureCitation>
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|
D). In apical view, cavity U-shaped, opening of sperm channel in caudal position; field of spines wide, close to lateral crest, more than 60 terminal pore setae, some organized in caudo-cephalic rows; internally cavity armed with hook-shaped spinules along mesial, caudal surfaces (
|
|
<figureCitation id="3F912A299B1CFFED86FFFF3EFF6E17D0" box="[159,226,223,248]" captionStart="FIGURE 5" captionStartId="7.[151,250,1752,1774]" captionTargetBox="[154,1436,193,1726]" captionTargetId="figure@7.[151,1436,193,1730]" captionTargetPageId="7" captionText="FIGURE 5. Pseudothelphusa ixtapan n. sp., male holotype CNCR 343, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudo-marginal projection; mp, mesial process; ms, marginal suture; osc, opening of sperm channel." httpUri="https://zenodo.org/record/242447/files/figure.png" pageId="6" pageNumber="565">Fig. 5</figureCitation>
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E).
|
|
</paragraph>
|
|
<paragraph id="A71536AC9B1CFFED86A7FEE5FC191635" blockId="6.[151,1437,151,573]" box="[199,917,260,285]" pageId="6" pageNumber="565">
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|
<emphasis id="95DEEABE9B1CFFED86A7FEE5FD881635" bold="true" box="[199,516,260,285]" pageId="6" pageNumber="565">Geographical distribution.</emphasis>
|
|
Only known from the type locality.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="EFB065279B1CFFED86A7FEC6FEE3164D" pageId="6" pageNumber="565" type="etymology">
|
|
<paragraph id="A71536AC9B1CFFED86A7FEC6FEE3164D" blockId="6.[151,1437,151,573]" pageId="6" pageNumber="565">
|
|
<emphasis id="95DEEABE9B1CFFED86A7FEC6FEC21668" bold="true" box="[199,334,295,320]" pageId="6" pageNumber="565">Etymology.</emphasis>
|
|
The specific epithet is the name of the
|
|
<typeStatus id="7811880E9B1CFFED8568FEC9FCB41668" box="[776,824,296,320]" pageId="6" pageNumber="565">type</typeStatus>
|
|
locality,
|
|
<taxonomicName id="60AA4D2F9B1CFFED85C0FEC9FC7B1617" box="[928,1015,296,319]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="566" phylum="Arthropoda" rank="species" species="ixtapan" status="sp. nov.">Ixtapan</taxonomicName>
|
|
, in the state of
|
|
<collectingCountry id="DFBD763C9B1CFFED82C2FEC6FB771668" box="[1186,1275,295,320]" name="Mexico" pageId="6" pageNumber="565">
|
|
<collectingRegion id="656EF84E9B1CFFED82C2FEC6FB771668" box="[1186,1275,295,320]" country="Mexico" name="Mexico" pageId="6" pageNumber="565">Mexico</collectingRegion>
|
|
</collectingCountry>
|
|
; it is used as a noun in apposition.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="EFB065279B1CFFEC86A7FE8EFCA71090" lastPageId="7" lastPageNumber="566" pageId="6" pageNumber="565" type="discussion">
|
|
<paragraph id="A71536AC9B1CFFED86A7FE8EFAD91514" blockId="6.[151,1437,151,573]" pageId="6" pageNumber="565">
|
|
<emphasis id="95DEEABE9B1CFFED86A7FE8EFEB716A0" bold="true" box="[199,315,367,392]" pageId="6" pageNumber="565">Remarks.</emphasis>
|
|
Based on the morphology of the first gonopod,
|
|
<taxonomicName id="60AA4D2F9B1CFFED850EFE91FB0016AF" box="[878,1164,368,392]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="ixtapan" status="sp. nov.">
|
|
<emphasis id="95DEEABE9B1CFFED850EFE91FB0016AF" box="[878,1164,368,392]" italics="true" pageId="6" pageNumber="565">Pseudothelphusa ixtapan</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="95DEEABE9B1CFFED82F5FE8EFB5B16A0" bold="true" box="[1173,1239,367,392]" pageId="6" pageNumber="565">
|
|
<taxonomicNameLabel id="8EED57C59B1CFFED82F5FE8EFB5B16A0" box="[1173,1239,367,392]" pageId="6" pageNumber="565" rank="species">n. sp.</taxonomicNameLabel>
|
|
</emphasis>
|
|
is close to those species of
|
|
<taxonomicName id="60AA4D2F9B1CFFED877AFE75FE551684" box="[282,473,404,428]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="95DEEABE9B1CFFED877AFE75FE551684" box="[282,473,404,428]" italics="true" pageId="6" pageNumber="565">Pseudothelphusa</emphasis>
|
|
</taxonomicName>
|
|
with a subrectangular mesial process, like
|
|
<taxonomicName id="60AA4D2F9B1CFFED8587FE74FA971684" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" box="[999,1307,404,429]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="dilatata">
|
|
<emphasis id="95DEEABE9B1CFFED8587FE74FBEC1684" box="[999,1120,404,428]" italics="true" pageId="6" pageNumber="565">P. dilatata</emphasis>
|
|
<bibRefCitation id="C33B4B5D9B1CFFED820DFE75FA971684" author="Rathbun" box="[1133,1307,404,429]" pageId="6" pageNumber="565" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537." type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="60AA4D2F9B1CFFED834CFE74FEB216F8" authority="Rathbun, 1898" authorityName="Rathbun" authorityYear="1898" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="digueti">
|
|
<emphasis id="95DEEABE9B1CFFED834CFE74FA101684" box="[1324,1436,404,428]" italics="true" pageId="6" pageNumber="565">P. digueti</emphasis>
|
|
<bibRefCitation id="C33B4B5D9B1CFFED86F7FE56FEB216F8" author="Rathbun" box="[151,318,439,464]" pageId="6" pageNumber="565" refString="Rathbun, M. J. (1898) A contribution to a knowledge of the fresh-water crabs of America. The Pseudothelphusinae. Proceedings of the United States National Museum, 21 (1158), 507 - 537." type="journal article" year="1898">Rathbun, 1898</bibRefCitation>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="60AA4D2F9B1CFFED8718FE58FD4916F8" authority="Pretzmann, 1968" authorityName="Pretzmann" authorityYear="1968" box="[376,709,439,464]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="morelosis">
|
|
<emphasis id="95DEEABE9B1CFFED8718FE58FE7716F8" box="[376,507,440,464]" italics="true" pageId="6" pageNumber="565">P. morelosis</emphasis>
|
|
Pretzmann, 1968
|
|
</taxonomicName>
|
|
. The differences among these species are in the ornamentation of the mesial process,
|
|
<taxonomicName id="60AA4D2F9B1CFFED8712FE3CFE6D16DC" box="[370,481,477,500]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="ixtapan" status="sp. nov.">
|
|
<emphasis id="95DEEABE9B1CFFED8712FE3CFE6D16DC" box="[370,481,477,500]" italics="true" pageId="6" pageNumber="565">P. ixtapan</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="95DEEABE9B1CFFED8788FE3DFDAB16DD" bold="true" box="[488,551,476,501]" pageId="6" pageNumber="565">
|
|
<taxonomicNameLabel id="8EED57C59B1CFFED8788FE3DFDAB16DD" box="[488,551,476,501]" pageId="6" pageNumber="565" rank="species">n. sp.</taxonomicNameLabel>
|
|
</emphasis>
|
|
has only one lateral strong tooth and the inferior border is armed with a row of five acute spinules. In
|
|
<taxonomicName id="60AA4D2F9B1CFFED87F3FDE0FD891530" box="[403,517,512,536]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="dilatata">
|
|
<emphasis id="95DEEABE9B1CFFED87F3FDE0FD891530" box="[403,517,512,536]" italics="true" pageId="6" pageNumber="565">P. dilatata</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="60AA4D2F9B1CFFED8474FDE0FDF01530" box="[532,636,512,536]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="digueti">
|
|
<emphasis id="95DEEABE9B1CFFED8474FDE0FDF01530" box="[532,636,512,536]" italics="true" pageId="6" pageNumber="565">P. digueti</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="60AA4D2F9B1CFFED84D3FDE0FCB61530" box="[691,826,512,536]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="morelosis">
|
|
<emphasis id="95DEEABE9B1CFFED84D3FDE0FCB61530" box="[691,826,512,536]" italics="true" pageId="6" pageNumber="565">P. morelosis</emphasis>
|
|
</taxonomicName>
|
|
the lateral margin of the mesial process has more than one spiniform tooth; the inferior border is smooth in
|
|
<taxonomicName id="60AA4D2F9B1CFFED8486FDC4FCE61514" box="[742,874,548,572]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="morelosis">
|
|
<emphasis id="95DEEABE9B1CFFED8486FDC4FCE61514" box="[742,874,548,572]" italics="true" pageId="6" pageNumber="565">P. morelosis</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="60AA4D2F9B1CFFED85C4FDC4FB811514" box="[932,1037,548,572]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="digueti">
|
|
<emphasis id="95DEEABE9B1CFFED85C4FDC4FB811514" box="[932,1037,548,572]" italics="true" pageId="6" pageNumber="565">P. digueti</emphasis>
|
|
</taxonomicName>
|
|
, with denticles in
|
|
<taxonomicName id="60AA4D2F9B1CFFED82BCFDC4FAC21514" box="[1244,1358,548,572]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="dilatata">
|
|
<emphasis id="95DEEABE9B1CFFED82BCFDC4FAC21514" box="[1244,1358,548,572]" italics="true" pageId="6" pageNumber="565">P. dilatata</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<caption id="F3D566249B1CFFED86F7F847FE3E10F3" httpUri="https://zenodo.org/record/242446/files/figure.png" pageId="6" pageNumber="565" targetBox="[206,1388,635,1933]" targetPageId="6">
|
|
<paragraph id="A71536AC9B1CFFED86F7F847FE3E10F3" blockId="6.[151,1436,1958,2011]" pageId="6" pageNumber="565">
|
|
<emphasis id="95DEEABE9B1CFFED86F7F847FE9F1094" bold="true" box="[151,275,1958,1980]" pageId="6" pageNumber="565">FIGURE 4.</emphasis>
|
|
<taxonomicName id="60AA4D2F9B1CFFED877AF847FD951095" box="[282,537,1958,1981]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="565" phylum="Arthropoda" rank="species" species="ixtapan" status="sp. nov.">
|
|
<emphasis id="95DEEABE9B1CFFED877AF847FD951095" box="[282,537,1958,1981]" italics="true" pageId="6" pageNumber="565">Pseudothelphusa ixtapan</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="95DEEABE9B1CFFED8440F846FDD61094" bold="true" box="[544,602,1959,1980]" pageId="6" pageNumber="565">
|
|
<taxonomicNameLabel id="8EED57C59B1CFFED8440F846FDD61094" box="[544,602,1959,1980]" pageId="6" pageNumber="565" rank="species">n. sp.</taxonomicNameLabel>
|
|
</emphasis>
|
|
, male holotype, CNCR 343: A, dorsal view; B, major chela; C, third maxillipeds; D, frontal view of carapace.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="F3D566249B1DFFEC86F7F939FC661003" httpUri="https://zenodo.org/record/242447/files/figure.png" pageId="7" pageNumber="566" targetBox="[154,1436,193,1726]" targetPageId="7">
|
|
<paragraph id="A71536AC9B1DFFEC86F7F939FC661003" blockId="7.[151,1436,1752,1835]" pageId="7" pageNumber="566">
|
|
<emphasis id="95DEEABE9B1DFFEC86F7F939FE9F11C5" bold="true" box="[151,275,1752,1774]" pageId="7" pageNumber="566">FIGURE 5.</emphasis>
|
|
<taxonomicName id="60AA4D2F9B1DFFEC877BF939FD9711C6" box="[283,539,1752,1774]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="ixtapan" status="sp. nov.">
|
|
<emphasis id="95DEEABE9B1DFFEC877BF939FD9711C6" box="[283,539,1752,1774]" italics="true" pageId="7" pageNumber="566">Pseudothelphusa ixtapan</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="95DEEABE9B1DFFEC8442F939FDD011C5" bold="true" box="[546,604,1752,1773]" pageId="7" pageNumber="566">
|
|
<taxonomicNameLabel id="8EED57C59B1DFFEC8442F939FDD011C5" box="[546,604,1752,1773]" pageId="7" pageNumber="566" rank="species">n. sp.</taxonomicNameLabel>
|
|
</emphasis>
|
|
, male holotype CNCR 343, left first gonopod: A, mesial view; B, cephalic view; C, lateral view; D, caudal view; E, apical view; F, total cephalic view. Abbreviations: ac, apex cavity; cmp, caudo-marginal projection; mp, mesial process; ms, marginal suture; osc, opening of sperm channel.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="A71536AC9B1DFFEC86A7F8B6FCA71090" blockId="7.[151,1436,1879,1976]" pageId="7" pageNumber="566">
|
|
<materialsCitation id="17C23CF19B1DFFEC86A7F8B6FCA71090" ID-GBIF-Occurrence="1424565289" country="Mexico" location="Balsas River" pageId="7" pageNumber="566" specimenCount="1" stateProvince="Guerrero">
|
|
The new species is distributed in the same area where
|
|
<taxonomicName id="60AA4D2F9B1DFFEC8526F8B8FC5C1047" box="[838,976,1880,1904]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="mexicana">
|
|
<emphasis id="95DEEABE9B1DFFEC8526F8B8FC5C1047" box="[838,976,1880,1904]" italics="true" pageId="7" pageNumber="566">P. mexicana</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="60AA4D2F9B1DFFEC826DF8B8FB231047" box="[1037,1199,1880,1904]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="granatensis">
|
|
<emphasis id="95DEEABE9B1DFFEC826DF8B8FB231047" box="[1037,1199,1880,1904]" italics="true" pageId="7" pageNumber="566">P. granatensis</emphasis>
|
|
</taxonomicName>
|
|
occur, in the border region between the states of
|
|
<collectingRegion id="656EF84E9B1DFFEC8782F89DFDCB10BC" box="[482,583,1916,1940]" country="Mexico" name="Guerrero" pageId="7" pageNumber="566">Guerrero</collectingRegion>
|
|
and
|
|
<collectingCountry id="DFBD763C9B1DFFEC84E2F89DFD5710BD" box="[642,731,1916,1941]" name="Mexico" pageId="7" pageNumber="566">Mexico</collectingCountry>
|
|
.
|
|
<taxonomicName id="60AA4D2F9B1DFFEC8487F89DFB9710BC" box="[743,1051,1916,1940]" class="Malacostraca" family="Pseudothelphusidae" genus="Pseudothelphusa" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="566" phylum="Arthropoda" rank="species" species="morelosis">
|
|
<emphasis id="95DEEABE9B1DFFEC8487F89DFB9710BC" box="[743,1051,1916,1940]" italics="true" pageId="7" pageNumber="566">Pseudothelphusa morelosis</emphasis>
|
|
</taxonomicName>
|
|
, is another related species with a geographically close distribution in the
|
|
<location id="A27560779B1DFFEC8433F87EFD6D1090" LSID="urn:lsid:plazi:treatment:2F0387BA9B1FFFEC86F7FEADFCA71090:A27560779B1DFFEC8433F87EFD6D1090" box="[595,737,1951,1976]" country="Mexico" name="Balsas River" pageId="7" pageNumber="566" stateProvince="Guerrero">Balsas River</location>
|
|
basin.
|
|
</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |