261 lines
20 KiB
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261 lines
20 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1035.60843" ID-GBIF-Dataset="de9f0011-899b-4265-8330-33220b678a9b" ID-PMC="PMC8096804" ID-Pensoft-Pub="1313-2970-1035-1" ID-Pensoft-UUID="4C2258D4EE6754488B9280D3AB0447A1" ID-PubMed="33958931" ID-ZooBank="ED19022EA1704DB79587FEFE15D07854" ModsDocID="1313-2970-1035-1" checkinTime="1619617189537" checkinUser="pensoft" docAuthor="Marzuki, Mohammad Effendi bin, Liew, Thor-Seng & Mohd-Azlan, Jayasilan" docDate="2021" docId="1F165D15C0265803ADF5C630CE00282E" docLanguage="en" docName="ZooKeys 1035: 1-113" docOrigin="ZooKeys 1035" docPubDate="2021-04-27" docSource="http://dx.doi.org/10.3897/zookeys.1035.60843" docTitle="Everettia microrhytida Marzuki & Liew & Mohd-Azlan 2021, sp. nov." docType="treatment" docUuid="EDF2CDBE-F06C-423B-8442-F7AC1DC20BE7" docUuidSource="ZooBank" docVersion="4" id="4C2258D4EE6754488B9280D3AB0447A1" lastPageNumber="1" masterDocId="4C2258D4EE6754488B9280D3AB0447A1" masterDocTitle="Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species" masterLastPageNumber="113" masterPageNumber="1" pageNumber="1" updateTime="1668150216077" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Land snails and slugs of Bau limestone hills, Sarawak (Malaysia, Borneo), with the descriptions of 13 new species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Marzuki, Mohammad Effendi bin</mods:namePart>
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<mods:affiliation>Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia & Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88450, Kota Kinabalu, Sabah, Malaysia</mods:affiliation>
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<mods:nameIdentifier type="email">fendiemz@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liew, Thor-Seng</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-9437-5924</mods:nameIdentifier>
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<mods:affiliation>Institute for Tropical Biology and Conservation, Universiti Malaysia Sabah, Jalan UMS, 88450, Kota Kinabalu, Sabah, Malaysia</mods:affiliation>
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<mods:nameIdentifier type="email">thorsengliew@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mohd-Azlan, Jayasilan</mods:namePart>
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<mods:affiliation>Institute of Biodiversity and Environmental Conservation, Universiti Malaysia Sarawak, 94300, Kota Samarahan, Sarawak, Malaysia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-04-27</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1035</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>113</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1035.60843</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1035.60843</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1035-1</mods:identifier>
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<mods:identifier type="ZooBank">ED19022EA1704DB79587FEFE15D07854</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4C2258D4EE6754488B9280D3AB0447A1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="181753329" LSID="urn:lsid:zoobank.org:act:EDF2CDBE-F06C-423B-8442-F7AC1DC20BE7" httpUri="http://treatment.plazi.org/id/1F165D15C0265803ADF5C630CE00282E" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/EDF2CDBE-F06C-423B-8442-F7AC1DC20BE7" authority="Marzuki & Liew & Mohd-Azlan, 2021" authorityName="Marzuki & Liew & Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Dyakiidae" genus="Everettia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Everettia microrhytida" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="microrhytida" status="sp. nov.">Everettia microrhytida</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 41" captionStartId="F41" captionText="Figure 41. A Everettia cutteri (H. Adams, 1870) ME 8968 Gunung Doya Kapor B Everettia microrhytida, sp. nov., MZU. MOL. 20.25 Holotype Gunung Batu C Everettia minuta, sp. nov., MZU. MOL. 20.23 Holotype Gunung." figureDoi="10.3897/zookeys.1035.60843.figure41" httpUri="https://binary.pensoft.net/fig/536829" pageId="0" pageNumber="1">Figures 41B</figureCitation>
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<figureCitation captionStart="Figure 42" captionStartId="F42" captionText="Figure 42. Everettia spp. A-C Everettia minuta, sp. nov., MZU. MOL. 20.24 Paratype A Enlargement of the apical side showing the apex B Enlargement of the teleoconch showing the shell sculpture C Enlargement of the basal side of the shell D-F Everettia microrhytida, sp. nov., MZU. MOL. 20.26 Paratype D Enlargement of the apical side showing the apex E Enlargement of the teleoconch showing the shell sculpture F Enlargement of the basal side of the shell." figureDoi="10.3897/zookeys.1035.60843.figure42" httpUri="https://binary.pensoft.net/fig/536830" pageId="0" pageNumber="1">, 42D-F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<materialsCitation ID-GBIF-Occurrence="3093633321" collectingDate="2017-02-10" collectorName="M. E. Marzuki" country="Malaysia" county="Jambusan" latitude="1.3974028" location="Kuching Division" longLatPrecision="1" longitude="110.18889" municipality="Skio" specimenCount="1" typeStatus="Holotype">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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(SH
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<quantity metricMagnitude="-2" metricUnit="m" metricValue="1.214" unit="mm" value="12.14">12.14 mm</quantity>
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, SW 22.00 mm) (MZU.MOL.20.25),
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<collectingCountry name="Malaysia">Malaysia</collectingCountry>
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, Sarawak,
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<location LSID="urn:lsid:plazi:treatment:1F165D15C0265803ADF5C630CE00282E:77A8E3946EEF6130B70510C8CF8866F4" country="Malaysia" county="Jambusan" latitude="1.3974028" longLatPrecision="1" longitude="110.18889" municipality="Skio" name="Kuching Division">Kuching Division</location>
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,
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<location LSID="urn:lsid:plazi:treatment:1F165D15C0265803ADF5C630CE00282E:E783F7D57E451DA67CB21C204C82B07D" country="Malaysia" county="Jambusan" latitude="1.3974028" longLatPrecision="1" longitude="110.18889" municipality="Skio" name="Gunung Batu">Gunung Batu</location>
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, limestone outcrop along
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<collectingMunicipality>Skio</collectingMunicipality>
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road,
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<collectingCounty>Jambusan</collectingCounty>
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,
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<locationDeviation location="Bau">
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<quantity metricMagnitude="3" metricUnit="m" metricValue="3.8624256" unit="mi" value="2.4">2.4 miles</quantity>
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E Bau
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</locationDeviation>
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,
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<geoCoordinate degrees="1" direction="north" minutes="23" orientation="latitude" precision="1" seconds="50.65" value="1.3974028">1°23'50.65"N</geoCoordinate>
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,
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<geoCoordinate degrees="110" direction="east" minutes="11" orientation="longitude" precision="1" seconds="19.99" value="110.18889">110°11'19.99"E</geoCoordinate>
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, coll.
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<collectorName>M. E. Marzuki</collectorName>
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,
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<collectingDate value="2017-02-10">10.II.2017</collectingDate>
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</materialsCitation>
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.
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<materialsCitation ID-GBIF-Occurrence="3093633317" collectingDate="2011-07-10" collectorName="M. E. Marzuki" country="Malaysia" county="Jambusan" latitude="1.3974028" location="Kuching Division" longLatPrecision="1" longitude="110.18889" municipality="Skio" specimenCount="1" typeStatus="Paratypes">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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:
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<specimenCount type="generic">1 ex.</specimenCount>
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(MZU.MOL.20.26), the same locality as holotype, coll.
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<collectorName>M. E. Marzuki</collectorName>
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,
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<collectingDate value="2011-07-10">10.VII.2011</collectingDate>
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="3093633328" collectingDate="2015-09-27" collectorName="M. E. Marzuki" country="Malaysia" latitude="1.3233527" location="Kampung Padang Pan" longLatPrecision="1" longitude="110.062874" specimenCount="1" typeStatus="paratype">
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<specimenCount type="generic">1 ex.</specimenCount>
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(ME0006829), small limestone escarpment near
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<location LSID="urn:lsid:plazi:treatment:1F165D15C0265803ADF5C630CE00282E:3B75AB258F298CE5F2971B161D3BDC31" country="Malaysia" latitude="1.3233527" longLatPrecision="1" longitude="110.062874" name="Kampung Padang Pan">Kampung Padang Pan</location>
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,
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<locationDeviation location="Bau">
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<quantity metricMagnitude="4" metricUnit="m" metricValue="2.414016" unit="mi" value="15.0">15 miles</quantity>
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SW Bau
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</locationDeviation>
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,
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<geoCoordinate degrees="1" direction="north" minutes="19" orientation="latitude" precision="1" seconds="24.07" value="1.3233527">1°19'24.07"N</geoCoordinate>
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,
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<geoCoordinate degrees="110" direction="east" minutes="3" orientation="longitude" precision="1" seconds="46.34" value="110.062874">110°3'46.34"E</geoCoordinate>
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, coll.
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<collectorName>M. E. Marzuki</collectorName>
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,
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<collectingDate value="2015-09-27">27.IX.2015</collectingDate>
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="3093633381" collectingDate="2010-06-21" collectorName="M. E. Marzuki" country="Malaysia" latitude="1.3114027" location="Kampung Beratok" longLatPrecision="1" longitude="110.41032" municipality="Serian-Kuching" specimenCount="2" typeStatus="paratype">
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<specimenCount type="generic">2 ex.</specimenCount>
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(ME0003498), small limestone outcrop at
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<location LSID="urn:lsid:plazi:treatment:1F165D15C0265803ADF5C630CE00282E:0DFB0892624664399832B4582F7CD18E" country="Malaysia" latitude="1.3114027" longLatPrecision="1" longitude="110.41032" municipality="Serian-Kuching" name="Kampung Beratok">Kampung Beratok</location>
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,
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<collectingMunicipality>Serian-Kuching</collectingMunicipality>
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road,
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<locationDeviation location="Serian">
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<quantity metricMagnitude="4" metricUnit="m" metricValue="2.30136192" unit="mi" value="14.3">14.3 miles</quantity>
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NW Serian
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</locationDeviation>
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,
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<geoCoordinate degrees="1" direction="north" minutes="18" orientation="latitude" precision="1" seconds="41.05" value="1.3114027">1°18'41.05"N</geoCoordinate>
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,
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<geoCoordinate degrees="110" direction="east" minutes="24" orientation="longitude" precision="1" seconds="37.13" value="110.41032">110°24'37.13"E</geoCoordinate>
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, coll.
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<collectorName>M. E. Marzuki</collectorName>
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,
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<collectingDate value="2010-06-21">21.VI.2010</collectingDate>
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="3093633412" collectingDate="2017-04-23" collectorName="M. E. Marzuki" country="Sarawak" county="Kanan" latitude="1.4152666" location="Lobang Angin" longLatPrecision="1" longitude="110.137115" municipality="Sungai" specimenCount="6" typeStatus="paratype">
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<specimenCount type="generic">6 ex.</specimenCount>
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(ME0009145),
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<location LSID="urn:lsid:plazi:treatment:1F165D15C0265803ADF5C630CE00282E:249D3B85A3915908A5DDA44D1152ED1D" country="Sarawak" county="Kanan" latitude="1.4152666" longLatPrecision="1" longitude="110.137115" municipality="Sungai" name="Lobang Angin">Lobang Angin</location>
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(
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<location LSID="urn:lsid:plazi:treatment:1F165D15C0265803ADF5C630CE00282E:643A093412B9C5ABD7BA69221A5FC564" country="Sarawak" county="Kanan" latitude="1.4152666" longLatPrecision="1" longitude="110.137115" municipality="Sungai" name="Site">Site</location>
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3), limestone outcrop near
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<collectingMunicipality>Sungai</collectingMunicipality>
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<collectingCountry name="Malaysia">Sarawak</collectingCountry>
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<collectingCounty>Kanan</collectingCounty>
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,
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<locationDeviation location="Bau">
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<quantity metricMagnitude="3" metricUnit="m" metricValue="2.816352" unit="mi" value="1.75">1.75 miles</quantity>
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W of Bau
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</locationDeviation>
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,
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<geoCoordinate degrees="1" direction="north" minutes="24" orientation="latitude" precision="1" seconds="54.96" value="1.4152666">1°24'54.96"N</geoCoordinate>
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,
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<geoCoordinate degrees="110" direction="east" minutes="8" orientation="longitude" precision="1" seconds="13.62" value="110.137115">110°8'13.62"E</geoCoordinate>
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, coll.
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<collectorName>M. E. Marzuki</collectorName>
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,
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<collectingDate value="2017-04-23">23.IV.2017</collectingDate>
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</materialsCitation>
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;
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<materialsCitation ID-GBIF-Occurrence="3093633364" collectingDate="2018-05-12" collectorName="M. E. Marzuki" country="Malaysia" county="Kanan" latitude="1.4152666" location="Lobang Angin" longLatPrecision="1" longitude="110.137115" municipality="Sungai" specimenCount="1" typeStatus="paratype">
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<specimenCount type="generic">1 ex.</specimenCount>
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(ME0009845), the same locality, coll.
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<collectorName>M. E. Marzuki</collectorName>
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,
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<collectingDate value="2018-05-12">12.V.2018</collectingDate>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This species is similar to
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<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1854" class="Gastropoda" family="Dyakiidae" genus="Everettia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Everettia consul" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="consul">
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<emphasis italics="true" pageId="0" pageNumber="1">Everettia consul</emphasis>
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</taxonomicName>
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(Pfeiffer, 1854) in terms of general shape and size. However, it differs from
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<taxonomicName lsidName="E. consul" pageId="0" pageNumber="1" rank="species" species="consul">
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<emphasis italics="true" pageId="0" pageNumber="1">E. consul</emphasis>
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</taxonomicName>
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by lacking spiral sculpture and having only very fine (sometimes inconspicuous), somewhat wrinkled, puncture-like sculpture on both the apical and apertural sides.
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<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1854" class="Gastropoda" family="Dyakiidae" genus="Everettia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Everettia consul" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="consul">
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<emphasis italics="true" pageId="0" pageNumber="1">Everettia consul</emphasis>
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</taxonomicName>
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, on the other hand, has a shell with a more elevated spire and its shell surface has densely placed radial threads and somewhat cut by irregularly spaced spiral grooves.
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</paragraph>
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<caption doi="10.3897/zookeys.1035.60843.figure42" httpUri="https://binary.pensoft.net/fig/536830" pageId="0" pageNumber="1" start="Figure 42" startId="F42">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 42.</emphasis>
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<taxonomicName class="Gastropoda" family="Dyakiidae" genus="Everettia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Everettia" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Everettia</emphasis>
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</taxonomicName>
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spp.
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<emphasis bold="true" pageId="0" pageNumber="1">A-C</emphasis>
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<taxonomicName authorityName="Marzuki & Liew & Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Dyakiidae" genus="Everettia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Everettia minuta" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="minuta">
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<emphasis italics="true" pageId="0" pageNumber="1">Everettia minuta</emphasis>
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</taxonomicName>
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, sp. nov., MZU.MOL.20.24 Paratype
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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Enlargement of the apical side showing the apex
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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Enlargement of the teleoconch showing the shell sculpture
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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Enlargement of the basal side of the shell
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<emphasis bold="true" pageId="0" pageNumber="1">D-F</emphasis>
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<taxonomicName authorityName="Marzuki & Liew & Mohd-Azlan" authorityYear="2021" class="Gastropoda" family="Dyakiidae" genus="Everettia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Everettia microrhytida" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="microrhytida">
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<emphasis italics="true" pageId="0" pageNumber="1">Everettia microrhytida</emphasis>
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</taxonomicName>
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, sp. nov., MZU.MOL.20.26 Paratype
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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Enlargement of the apical side showing the apex
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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Enlargement of the teleoconch showing the shell sculpture
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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Enlargement of the basal side of the shell.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Shell moderately large, rather thin, translucent, pale to dark brown, spire slightly elevated. Surface with a glossy lustre. Whorls convex. Number of whorls <
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<normalizedToken originalValue="6¼">61/4</normalizedToken>
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. Protoconch: almost smooth, spiral striation absent with inconspicuous radial threads near the suture. Teleoconch with no: spiral sculpture but with very fine, inconspicuous, somewhat wrinkled, puncture-like shell sculptures on both apical and apertural sides. Radial sculpture of teleoconch very fine as well as inconspicuous growth lines, most conspicuous radial threads near the suture and below periphery. Periphery round; suture shallow. Aperture lunulate. Peristome simple; somewhat thickened and reflected on columellar side, not thickened nor reflected on basal and palatal sides. Umbilicus open, narrow; sometimes partly covered by reflected peristome; umbilical region moderately concave. Dimensions: shell height <12.58 mm; shell width <22.13 mm; diameters of the first three whorls 1.60 mm, 2.55 mm, and 4.90 mm, respectively; aperture height <8.94 mm; aperture width <11.66 mm.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Geographic distribution and habitat.</paragraph>
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<paragraph pageId="0" pageNumber="1">It is known from the Bau and Serian-Padawan limestone hill clusters. Living snails were observed foraging among leaf litter and plant debris near the cliff in a lowland limestone forest.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="etymology">
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<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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From the Greek
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<emphasis italics="true" pageId="0" pageNumber="1">
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mikro
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<normalizedToken originalValue="rytídes">rytides</normalizedToken>
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</emphasis>
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, meaning extremely small wrinkles, in reference to the shell sculpture.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |