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<document id="86177B0B5B18B428F22152701AD92222" ID-CLB-Dataset="53267" ID-DOI="10.11646/zootaxa.5249.4.6" ID-GBIF-Dataset="2e9b0a37-bcc0-4a94-95b1-155a88faa86d" ID-ISSN="1175-5326" ID-Zenodo-Dep="7694629" ID-ZooBank="FEF1BDA2-8CF1-4D0A-ADAC-CD7F132C95FA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1677826210846" checkinUser="plazi" docAuthor="Da Silva, Irene M., Ferreira-Keppler, Ruth L. &amp; Cazorla, Carla G." docDate="2023" docId="03F987FCFF85FFDCDC9852CAFC33A28A" docLanguage="en" docName="zootaxa.5249.4.6.pdf" docOrigin="Zootaxa 5249 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5249.4.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Stilobezzia (Acanthohelea) medialuna Da Silva &amp; Ferreira-Keppler &amp; Cazorla 2023, n. sp." docType="treatment" docVersion="7" lastPageNumber="488" masterDocId="FFC0FF84FF84FFDFDC0F5715FFADA717" masterDocTitle="Stilobezzia Kieffer (Diptera: Ceratopogonidae) from the Brazilian Amazon: three new species, redescription of S. (Stilobezzia) maculata Lane and new records" masterLastPageNumber="499" masterPageNumber="485" pageNumber="486" updateTime="1699476158784" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="D9BE5F714FE61C5B8E73FF47514A48F8" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="CCD4EC2041090166F69FFDE9C52E6978">
<mods:title id="BBBB3CF94B23A83C5C8A969FA81022D6">Stilobezzia Kieffer (Diptera: Ceratopogonidae) from the Brazilian Amazon: three new species, redescription of S. (Stilobezzia) maculata Lane and new records</mods:title>
</mods:titleInfo>
<mods:name id="91E3CEF2C5D6233F8F00BFC45A5D6323" type="personal">
<mods:role id="9C03D8C542FFCB0179223B81AF3A4A43">
<mods:roleTerm id="E17107E4E5BBD0FA92D1A11378A1EF35">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8CC71744123C6B023CC607C246E3AD11">Da Silva, Irene M.</mods:namePart>
<mods:nameIdentifier id="E9224917B29557485FB0944C06A7C80B" type="ORCID">0000-0002-2528-6653</mods:nameIdentifier>
<mods:affiliation id="1F6615C6FCB803A06FF8C6BF9CF03A00">Instituto Nacional de Pesquisas da Amazônia (INPA), Coordenação de Pós-Graduação (COPOG), Divisão do Curso em Entomologia (DiEnt), Laboratório de Citotaxonomia e Insetos Aquáticos (LACIA), Manaus, AM, Brazil &amp; irenem. silva. bio @ gmail. com; https: // orcid. org / 0000 - 0002 - 2528 - 6653</mods:affiliation>
<mods:nameIdentifier id="0D6693D41E4C5665CA1F6BF55CD31D5C" type="email">irenem.silva.bio@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="46D43577731CF24535C1833336CD7C5F" type="personal">
<mods:role id="8E547A0AF340D1B23FAC3DA4CDE94211">
<mods:roleTerm id="DB221AAC564941D605AD7AA23336A965">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4E1E0AAA00FD983E00F4C916E42ECE5F">Ferreira-Keppler, Ruth L.</mods:namePart>
<mods:nameIdentifier id="4F2B5D8A338B799B56290358CD06F43F" type="ORCID">0000-0001-5485-0279</mods:nameIdentifier>
<mods:affiliation id="6A36C5D4162A2B4C349D10597C1FAB06">Instituto Nacional de Pesquisas da Amazônia (INPA), Coordenação de Pós-Graduação (COPOG), Divisão do Curso em Entomologia (DiEnt), Laboratório de Citotaxonomia e Insetos Aquáticos (LACIA), Manaus, AM, Brazil &amp; ruth @ inpa. gov. br; https: // orcid. org / 0000 - 0001 - 5485 - 0279</mods:affiliation>
<mods:nameIdentifier id="6960E8F73291349AD83F320068013DCE" type="email">ruth@inpa.gov.br</mods:nameIdentifier>
</mods:name>
<mods:name id="980A35D6A0C787C6D4208E09259BBAD1" type="personal">
<mods:role id="10C344EAAE26866665C24BC8B0AD751C">
<mods:roleTerm id="766C0A020DA477A6B385B6A1749F1741">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7EF74907B0AD726502D3521F98263D87">Cazorla, Carla G.</mods:namePart>
<mods:nameIdentifier id="449ED0574027505524FBB1E52A791571" type="ORCID">0000-0002-3644-8574</mods:nameIdentifier>
<mods:affiliation id="6883F493CD6BFA8393CD350434FAAF99">División Entomología, Museo de La Plata, Anexo Museo, 122 y 60 s / n La Plata, CP 1900, Argentina and Consejo Nacional de Investigaciones Científicas y técnicas (CONICET) &amp; carlacazorla @ fcnym. unlp. edu. ar; https: // orcid. org / 0000 - 0002 - 3644 - 8574</mods:affiliation>
<mods:nameIdentifier id="AFFFAAFDBB4E42F9F95C7956608B882C" type="email">carlacazorla@fcnym.unlp.edu.ar</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="EC2D68BA0F848A5E289878EDD209499A">text</mods:typeOfResource>
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<mods:titleInfo id="5A81422BF7CFC3EBFE615D37B46B64D7">
<mods:title id="72E456688C6ACD105CB88DEDFE45ABCB">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="2679CF05C25CF28098ABD03B6153C495">
<mods:date id="57C97B48DD3583C2F39BC436DAEA0653">2023</mods:date>
<mods:detail id="9048DE1B6CCDA96592D5BDE96CFE13C3" type="pubDate">
<mods:number id="BE78FA77A7300136186EE97291DC2946">2023-03-03</mods:number>
</mods:detail>
<mods:detail id="DB49248B366B52C80AF585FB1BA06656" type="volume">
<mods:number id="867DD93684E4087015EE78C260FC1749">5249</mods:number>
</mods:detail>
<mods:detail id="201676A892FDB42EC720A6BF5D5B1360" type="issue">
<mods:number id="E3087246291DAB1D0F2E18847684FA77">4</mods:number>
</mods:detail>
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<mods:start id="231B3A281CD1A395A653685BA25EDEEA">485</mods:start>
<mods:end id="1D20700EB71EB6A1AC389D2FA30D9A0B">499</mods:end>
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<mods:url id="31716EE37128E3B37E2AAD772CA959EA">http://dx.doi.org/10.11646/zootaxa.5249.4.6</mods:url>
</mods:location>
<mods:classification id="0D571AB7E26D61255DCB51DD38DD9E67">journal article</mods:classification>
<mods:identifier id="70D7B19CBC2F6FE1D978A142D9AE31CE" type="CLB-Dataset">53267</mods:identifier>
<mods:identifier id="7CFFE75D99A6D976E4C84C71632990FC" type="DOI">10.11646/zootaxa.5249.4.6</mods:identifier>
<mods:identifier id="EC3710D7941818CBFCE2CFB020A96E20" type="GBIF-Dataset">2e9b0a37-bcc0-4a94-95b1-155a88faa86d</mods:identifier>
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<mods:identifier id="7D2207CA3C7ECC87B0FE5746FB1E19A3" type="Zenodo-Dep">7694629</mods:identifier>
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</mods:mods>
<treatment id="03F987FCFF85FFDCDC9852CAFC33A28A" ID-DOI="http://doi.org/10.5281/zenodo.7694713" ID-GBIF-Taxon="209409377" ID-Zenodo-Dep="7694713" LSID="urn:lsid:plazi:treatment:03F987FCFF85FFDCDC9852CAFC33A28A" httpUri="http://treatment.plazi.org/id/03F987FCFF85FFDCDC9852CAFC33A28A" lastPageId="3" lastPageNumber="488" pageId="1" pageNumber="486">
<subSubSection id="C34A6561FF85FFDEDC9852CAFD00A2EE" box="[151,685,1503,1530]" pageId="1" pageNumber="486" type="nomenclature">
<paragraph id="8BEF36EAFF85FFDEDC9852CAFD00A2EE" blockId="1.[151,685,1503,1566]" box="[151,685,1503,1530]" pageId="1" pageNumber="486">
<heading id="D0A78186FF85FFDEDC9852CAFD00A2EE" bold="true" box="[151,685,1503,1530]" fontSize="11" level="1" pageId="1" pageNumber="486" reason="1">
<emphasis id="B924EAF8FF85FFDEDC9852CAFD00A2EE" bold="true" box="[151,685,1503,1530]" pageId="1" pageNumber="486">
<taxonomicName id="4C504D69FF85FFDEDC9852CAFDCEA2ED" ID-CoL="BDM9C" authority="Da Silva &amp; Ferreira-Keppler &amp; Cazorla, 2023" authorityName="Da Silva &amp; Ferreira-Keppler &amp; Cazorla" authorityYear="2023" box="[151,611,1503,1530]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="486" phylum="Arthropoda" rank="species" species="medialuna" status="sp. nov." subGenus="Acanthohelea">
<emphasis id="B924EAF8FF85FFDEDC9852CAFEB5A2ED" bold="true" box="[151,280,1503,1530]" italics="true" pageId="1" pageNumber="486">Stilobezzia</emphasis>
(
<emphasis id="B924EAF8FF85FFDEDD2652CAFE7CA2ED" bold="true" box="[297,465,1503,1530]" italics="true" pageId="1" pageNumber="486">Acanthohelea</emphasis>
)
<emphasis id="B924EAF8FF85FFDEDDED52CAFDCEA2ED" bold="true" box="[482,611,1503,1530]" italics="true" pageId="1" pageNumber="486">medialuna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2175783FF85FFDEDE6552CAFD00A2EE" box="[618,685,1503,1529]" pageId="1" pageNumber="486" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A6561FF85FFDEDC985116FEE7A109" box="[151,330,1539,1566]" pageId="1" pageNumber="486" type="description">
<paragraph id="8BEF36EAFF85FFDEDC985116FEE7A109" blockId="1.[151,685,1503,1566]" box="[151,330,1539,1566]" pageId="1" pageNumber="486">
(
<figureCitation id="136B2A6FFF85FFDEDC905116FEB5A10A" box="[159,280,1539,1565]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="1" pageNumber="486">Figs. 1–12</figureCitation>
,
<figureCitation id="136B2A6FFF85FFDEDD2A5116FEEFA109" box="[293,322,1539,1566]" captionStart="FIGURE 50" captionStartId="12.[151,250,1229,1253]" captionTargetBox="[160,1425,194,1194]" captionTargetId="figure-24@12.[153,1434,181,1205]" captionTargetPageId="12" captionText="FIGURE 50. Distribution of the new species of Stilobezzia Kieffer from Manaus, Amazonas, Brazil." figureDoi="http://doi.org/10.5281/zenodo.7694635" httpUri="https://zenodo.org/record/7694635/files/figure.png" pageId="1" pageNumber="486">50</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34A6561FF85FFDEDC98515EFC29A00E" pageId="1" pageNumber="486" type="materials_examined">
<paragraph id="8BEF36EAFF85FFDEDC98515EFB8EA1BA" blockId="1.[151,1437,1611,1998]" pageId="1" pageNumber="486">
<emphasis id="B924EAF8FF85FFDEDC98515EFEE8A172" bold="true" box="[151,325,1611,1637]" pageId="1" pageNumber="486">Type material:</emphasis>
<materialsCitation id="3B383CB7FF85FFDEDD45515EFDF0A1BA" ID-GBIF-Occurrence="4041566309" collectingDate="2017-12-01" collectingDateMax="2017-12-15" collectingDateMin="2017-12-01" collectingMethod="Malaise trap" collectorName="J. A. Rafael &amp; F. F. Xavier" country="Brazil" county="Base" latitude="-2.5891666" location="Manaus" longLatPrecision="21" longitude="-60.115276" municipality="Manaus" pageId="1" pageNumber="486" specimenCount="1" stateProvince="Amazonas" typeStatus="holotype">
<emphasis id="B924EAF8FF85FFDEDD45515EFDFDA172" bold="true" box="[330,592,1611,1637]" pageId="1" pageNumber="486">
<typeStatus id="54EB8848FF85FFDEDD45515EFE1AA172" box="[330,439,1611,1637]" pageId="1" pageNumber="486" type="holotype">Holotype</typeStatus>
<specimenCount id="9D56FD63FF85FFDEDDB3515EFE5BA172" box="[444,502,1611,1637]" pageId="1" pageNumber="486" type="male">male</specimenCount>
(
<taxonomicName id="4C504D69FF85FFDEDE0C515EFDE4A172" authorityName="Erwin" authorityYear="1978" box="[515,585,1611,1637]" class="Insecta" family="Carabidae" genus="Inpa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="485" phylum="Arthropoda" rank="genus">INPA</taxonomicName>
)
</emphasis>
labeled: “Holotype
<taxonomicName id="4C504D69FF85FFDEDF22515EFA22A172" authority="Silva &amp; Cazorla" authorityName="Da Silva &amp; Ferreira-Keppler &amp; Cazorla" authorityYear="2023" box="[813,1423,1611,1637]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="486" phylum="Arthropoda" rank="species" species="medialuna" subGenus="Acanthohelea">
<emphasis id="B924EAF8FF85FFDEDF22515EFC0BA172" box="[813,934,1611,1637]" italics="true" pageId="1" pageNumber="486">Stilobezzia</emphasis>
(
<emphasis id="B924EAF8FF85FFDEDFBB515EFBE3A172" box="[948,1102,1611,1637]" italics="true" pageId="1" pageNumber="486">Acanthohelea</emphasis>
)
<emphasis id="B924EAF8FF85FFDED852515EFB7EA172" box="[1117,1235,1611,1637]" italics="true" pageId="1" pageNumber="486">medialuna</emphasis>
Silva &amp; Cazorla
</taxonomicName>
”.
<collectingCountry id="F347767AFF85FFDEDC98517AFF52A19E" box="[151,255,1647,1673]" name="Brazil" pageId="1" pageNumber="486">BRAZIL</collectingCountry>
,
<collectingRegion id="4994F808FF85FFDEDD03517AFE28A19E" box="[268,389,1647,1673]" pageId="1" pageNumber="486">Amazonas</collectingRegion>
,
<collectingMunicipality id="6B8BAC90FF85FFDEDD9D5165FE40A19D" box="[402,493,1648,1674]" pageId="1" pageNumber="486">Manaus</collectingMunicipality>
, ZF-2,
<locationDeviation id="9A5D2FC8FF85FFDEDE4A5165FC8CA19E" box="[581,801,1647,1674]" pageId="1" pageNumber="486">Km 14, Base LBA</locationDeviation>
,
<geoCoordinate id="EE64502DFF85FFDEDF215164FC04A19B" box="[814,937,1649,1676]" degrees="02" direction="south" minutes="35" orientation="latitude" pageId="1" pageNumber="486" precision="15" seconds="21" value="-2.5891666">02º35′21″S</geoCoordinate>
–
<geoCoordinate id="EE64502DFF85FFDEDFB75164FBEEA19B" box="[952,1091,1649,1676]" degrees="60" direction="west" minutes="06" orientation="longitude" pageId="1" pageNumber="486" precision="15" seconds="55" value="-60.115276">60º06′55″W</geoCoordinate>
,
<collectingMethod id="52114EFDFF85FFDED85D517AFB4DA19D" box="[1106,1248,1647,1674]" pageId="1" pageNumber="486">Malaise trap</collectingMethod>
over stream,
<date id="FFEE102AFF85FFDED98F517AFEB4A1B9" pageId="1" pageNumber="486" value="2017-12-01" valueMax="2017-12-15" valueMin="2017-12-01">
<collectingDate id="EFAAE9C2FF85FFDED98F517AFEB4A1B9" pageId="1" pageNumber="486" value="2017-12-01" valueMax="2017-12-15" valueMin="2017-12-01">1– 15.xii.2017</collectingDate>
</date>
,
<collectorName id="26A5533CFF85FFDEDD2A5186FE0FA1BA" box="[293,418,1683,1709]" pageId="1" pageNumber="486">J.A. Rafael</collectorName>
and
<collectorName id="26A5533CFF85FFDEDDD75181FDF4A1BA" box="[472,601,1683,1710]" pageId="1" pageNumber="486">F.F. Xavier</collectorName>
:
</materialsCitation>
<materialsCitation id="3B383CB7FF85FFDEDE6E5186FB8EA1BA" ID-GBIF-Occurrence="4041566319" box="[609,1059,1683,1710]" collectingDate="2017-12-01" collectingDateMax="2017-12-15" collectingDateMin="2017-12-01" collectorName="J. A. Rafael &amp; F. F. Xavier" country="Brazil" county="Base" latitude="-2.5891666" location="Amazonas" longLatPrecision="21" longitude="-60.115276" municipality="Km" pageId="1" pageNumber="486" specimenCount="1" typeStatus="allotype">
<typeStatus id="54EB8848FF85FFDEDE6E5186FD6EA1BA" box="[609,707,1683,1709]" pageId="1" pageNumber="486" type="allotype">Allotype</typeStatus>
<specimenCount id="9D56FD63FF85FFDEDEC55186FCB4A1BA" box="[714,793,1683,1709]" pageId="1" pageNumber="486" type="female">female</specimenCount>
, same data as holotype.
</materialsCitation>
</paragraph>
<paragraph id="8BEF36EAFF85FFDEDCC851A2FC29A00E" blockId="1.[151,1437,1611,1998]" pageId="1" pageNumber="486">
<materialsCitation id="3B383CB7FF85FFDEDCC851A2FA31A1E2" ID-GBIF-Occurrence="4041566331" collectingDate="2018-09-18" collectingDateMax="2018-10-04" collectingDateMin="2018-09-18" collectingMethod="suspensa trap" collectionCode="R" collectorName="I. M. Da Silva &amp; R. L. Ferreira-Keppler" country="Brazil" county="Base" latitude="-2.5891666" location="Campina Biological Reserve" longLatPrecision="21" longitude="-60.051945" municipality="Campinarana" pageId="1" pageNumber="486" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
<emphasis id="B924EAF8FF85FFDEDCC851A2FE92A1C6" bold="true" box="[199,319,1719,1745]" pageId="1" pageNumber="486">
<typeStatus id="54EB8848FF85FFDEDCC851A2FE92A1C6" box="[199,319,1719,1745]" pageId="1" pageNumber="486" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="9D56FD63FF85FFDEDD4151A2FE05A1C6" box="[334,424,1719,1745]" pageId="1" pageNumber="486" type="male">3 males</specimenCount>
,
<specimenCount id="9D56FD63FF85FFDEDDBB51A2FD89A1C6" box="[436,548,1719,1745]" pageId="1" pageNumber="486" type="female">2 females</specimenCount>
, same data as holotype.
<location id="8E8F6031FF85FFDEDF3851A2FBD3A1C5" LSID="urn:lsid:plazi:treatment:03F987FCFF85FFDCDC9852CAFC33A28A:8E8F6031FF85FFDEDF3851A2FBD3A1C5" box="[823,1150,1719,1746]" country="Brazil" county="Base" latitude="-2.5891666" longLatPrecision="21" longitude="-60.051945" municipality="Campinarana" name="Campina Biological Reserve" pageId="1" pageNumber="486">Campina Biological Reserve</location>
,
<collectingMunicipality id="6B8BAC90FF85FFDED88551A2FA8FA1C6" box="[1162,1314,1719,1745]" pageId="1" pageNumber="486">Campinarana</collectingMunicipality>
,
<geoCoordinate id="EE64502DFF85FFDED92151ACFF08A1EF" degrees="02" direction="south" minutes="58" orientation="latitude" pageId="1" pageNumber="486" seconds="94" value="-2.58">02º58′94″ S</geoCoordinate>
–
<geoCoordinate id="EE64502DFF85FFDEDCBB51C8FEEAA1E1" box="[180,327,1756,1784]" degrees="60" direction="west" minutes="03" orientation="longitude" pageId="1" pageNumber="486" precision="15" seconds="07" value="-60.051945">60º03′07″ W</geoCoordinate>
,
<collectingMethod id="52114EFDFF85FFDEDD5B51C9FE40A1E1" box="[340,493,1756,1782]" pageId="1" pageNumber="486">suspensa trap</collectingMethod>
over stream,
<date id="FFEE102AFF85FFDEDE8451CEFCE9A1E2" box="[651,836,1755,1781]" pageId="1" pageNumber="486" value="2018-09-18" valueMax="2018-10-04" valueMin="2018-09-18">
<collectingDate id="EFAAE9C2FF85FFDEDE8451CEFCE9A1E2" box="[651,836,1755,1781]" pageId="1" pageNumber="486" value="2018-09-18" valueMax="2018-10-04" valueMin="2018-09-18">18.ix–04.x.2018</collectingDate>
</date>
,
<collectorName id="26A5533CFF85FFDEDF5D51C9FC42A1E2" box="[850,1007,1755,1782]" pageId="1" pageNumber="486">I.M. Da Silva</collectorName>
and
<collectorName id="26A5533CFF85FFDED82651C9FA88A1E2" box="[1065,1317,1755,1782]" pageId="1" pageNumber="486">R.L. Ferreira-Keppler</collectorName>
:
<specimenCount id="9D56FD63FF85FFDED93E51CEFA35A1E2" box="[1329,1432,1755,1781]" pageId="1" pageNumber="486" type="female">1 female</specimenCount>
;
</materialsCitation>
<materialsCitation id="3B383CB7FF85FFDEDC9851EAFE5BA00E" ID-GBIF-Occurrence="4041566312" box="[151,502,1791,1817]" collectingDate="2018-09-18" collectingDateMax="2018-10-04" collectingDateMin="2018-09-18" collectingMethod="suspensa trap" collectionCode="R" collectorName="I. M. Da Silva &amp; R. L. Ferreira-Keppler" country="Brazil" county="Base" latitude="-2.5891666" location="Campina Biological Reserve" longLatPrecision="21" longitude="-60.051945" municipality="Campinarana" pageId="1" pageNumber="486" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
<date id="FFEE102AFF85FFDEDC9851EAFE98A00E" box="[151,309,1791,1817]" pageId="1" pageNumber="486" value="2018-10-04" valueMax="2018-10-23" valueMin="2018-10-04">
<collectingDate id="EFAAE9C2FF85FFDEDC9851EAFE98A00E" box="[151,309,1791,1817]" pageId="1" pageNumber="486" value="2018-10-04" valueMax="2018-10-23" valueMin="2018-10-04">04–23.x.2018</collectingDate>
</date>
:
<specimenCount id="9D56FD63FF85FFDEDD4F51EAFE3BA00E" box="[320,406,1791,1817]" pageId="1" pageNumber="486" type="male">3 males</specimenCount>
(
<taxonomicName id="4C504D69FF85FFDEDDAA51EAFE46A00E" authorityName="Erwin" authorityYear="1978" box="[421,491,1791,1817]" class="Insecta" family="Carabidae" genus="Inpa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="485" phylum="Arthropoda" rank="genus">
<collectionCode id="ED41AE2FFF85FFDEDDAA51EAFE46A00E" box="[421,491,1791,1817]" collectionName="Brazil, Amazonas, Manaus, Instituto Nacional de Pesquisas da Amazoonia, Colecao Sistematica da Entomologia" pageId="1" pageNumber="486">INPA</collectionCode>
</taxonomicName>
);
</materialsCitation>
<materialsCitation id="3B383CB7FF85FFDEDDF251EAFC2CA00E" ID-GBIF-Occurrence="4041566328" box="[509,897,1791,1817]" collectingDate="2018-09-18" collectingDateMax="2018-10-04" collectingDateMin="2018-09-18" collectingMethod="suspensa trap" collectionCode="R" collectorName="I. M. Da Silva &amp; R. L. Ferreira-Keppler" country="Brazil" county="Base" latitude="-2.5891666" location="Campina Biological Reserve" longLatPrecision="21" longitude="-60.051945" municipality="Campinarana" pageId="1" pageNumber="486" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
<date id="FFEE102AFF85FFDEDDF251EAFD15A00E" box="[509,696,1791,1817]" pageId="1" pageNumber="486" value="2018-10-23" valueMax="2018-11-09" valueMin="2018-10-23">
<collectingDate id="EFAAE9C2FF85FFDEDDF251EAFD15A00E" box="[509,696,1791,1817]" pageId="1" pageNumber="486" value="2018-10-23" valueMax="2018-11-09" valueMin="2018-10-23">23.x–09.xi.2018</collectingDate>
</date>
:
<specimenCount id="9D56FD63FF85FFDEDECC51EAFC89A00E" box="[707,804,1791,1817]" pageId="1" pageNumber="486" type="female">1 female</specimenCount>
(
<taxonomicName id="4C504D69FF85FFDEDF3C51EAFCD4A00E" authorityName="Erwin" authorityYear="1978" box="[819,889,1791,1817]" class="Insecta" family="Carabidae" genus="Inpa" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="485" phylum="Arthropoda" rank="genus">
<collectionCode id="ED41AE2FFF85FFDEDF3C51EAFCD4A00E" box="[819,889,1791,1817]" collectionName="Brazil, Amazonas, Manaus, Instituto Nacional de Pesquisas da Amazoonia, Colecao Sistematica da Entomologia" pageId="1" pageNumber="486">INPA</collectionCode>
</taxonomicName>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6561FF85FFDEDCC85036FB66A0DA" pageId="1" pageNumber="486" type="diagnosis">
<paragraph id="8BEF36EAFF85FFDEDCC85036FB66A0DA" blockId="1.[151,1437,1611,1998]" pageId="1" pageNumber="486">
<emphasis id="B924EAF8FF85FFDEDCC85036FE97A02A" bold="true" box="[199,314,1827,1853]" pageId="1" pageNumber="486">Diagnosis</emphasis>
. Only extant species of
<taxonomicName id="4C504D69FF85FFDEDE415036FCD1A02A" box="[590,892,1827,1853]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="486" phylum="Arthropoda" rank="subGenus" subGenus="Acanthohelea">
<emphasis id="B924EAF8FF85FFDEDE415036FD6AA02A" box="[590,711,1827,1853]" italics="true" pageId="1" pageNumber="486">Stilobezzia</emphasis>
(
<emphasis id="B924EAF8FF85FFDEDED65036FCDEA02A" box="[729,883,1827,1853]" italics="true" pageId="1" pageNumber="486">Acanthohelea</emphasis>
)
</taxonomicName>
in the Neotropical Region with males that have elongated genitalia with stout crescent-shaped gonocoxites, and gonostylus curved 90° at mid-length with toothed apex and a lung-shaped hyaline membrane between the parameres and aedeagus. Females with tarsomere 5 of fore and mid legs with 3 pairs of batonnets and hind leg with 1 pair; abdomen dark brown except segment 10 and cerci paler, and sternite 8 with a large V-shaped posteromedian excavation and 2 ovoid spermathecae.
</paragraph>
</subSubSection>
<caption id="DF2F6662FF86FFDDDC985082FC64A0EF" ID-DOI="http://doi.org/10.5281/zenodo.7694633" ID-Zenodo-Dep="7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="2" pageNumber="487" startId="2.[151,264,1943,1967]" targetBox="[205,1378,179,1907]" targetPageId="2">
<paragraph id="8BEF36EAFF86FFDDDC985082FC64A0EF" blockId="2.[151,1436,1943,2040]" pageId="2" pageNumber="487">
<emphasis id="B924EAF8FF86FFDDDC985082FEE5A0B8" bold="true" box="[151,328,1943,1967]" pageId="2" pageNumber="487">FIGURES 1–12.</emphasis>
<taxonomicName id="4C504D69FF86FFDDDD40508DFD7FA0A7" authority="Da Silva &amp; Ferreira-Keppler &amp; Cazorla, 2023" authorityName="Da Silva &amp; Ferreira-Keppler &amp; Cazorla" authorityYear="2023" box="[335,722,1944,1968]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="487" phylum="Arthropoda" rank="species" species="medialuna" status="sp. nov." subGenus="Acanthohelea">
<emphasis id="B924EAF8FF86FFDDDD40508DFE10A0A7" box="[335,445,1944,1968]" italics="true" pageId="2" pageNumber="487">Stilobezzia</emphasis>
(
<emphasis id="B924EAF8FF86FFDDDDC3508DFDFAA0A7" box="[460,599,1944,1968]" italics="true" pageId="2" pageNumber="487">Acanthohelea</emphasis>
)
<emphasis id="B924EAF8FF86FFDDDE68508DFD7FA0A7" box="[615,722,1944,1968]" italics="true" pageId="2" pageNumber="487">medialuna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2175783FF86FFDDDED6508DFCBEA0A7" box="[729,787,1944,1968]" pageId="2" pageNumber="487" rank="species">
<emphasis id="B924EAF8FF86FFDDDED6508DFD4AA0A7" bold="true" box="[729,743,1944,1968]" pageId="2" pageNumber="487">n</emphasis>
.
<emphasis id="B924EAF8FF86FFDDDEFA508DFCBEA0A7" bold="true" box="[757,787,1944,1968]" pageId="2" pageNumber="487">sp.</emphasis>
</taxonomicNameLabel>
, male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view.
</paragraph>
</caption>
<subSubSection id="C34A6561FF87FFDCDCC85783FD45A306" pageId="3" pageNumber="488" type="description">
<paragraph id="8BEF36EAFF87FFDCDCC85783FB55A5DA" blockId="3.[151,1437,150,1437]" pageId="3" pageNumber="488">
<emphasis id="B924EAF8FF87FFDCDCC85783FEFCA7A7" bold="true" box="[199,337,150,176]" pageId="3" pageNumber="488">Description</emphasis>
.
<emphasis id="B924EAF8FF87FFDCDD525782FE3AA7A6" box="[349,407,151,177]" italics="true" pageId="3" pageNumber="488">Male</emphasis>
. Head (
<figureCitation id="136B2A6FFF87FFDCDDE45782FD87A7A6" box="[491,554,151,177]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="3" pageNumber="488">Fig. 1</figureCitation>
.) dark brown.Antenna with yellowish brown pedicel; light brown flagellomeres with pale brown basal ring; segments 11–12 with stout bristles at base; brown with dense plume; AR 0.90–1.02 (0.97; n=3). Palpus dark brown; segment 5 as long as 3; segment 3 with round deep sensory pit;
<collectingRegion id="4994F808FF87FFDCD8B357CAFB73A7EE" box="[1212,1246,223,249]" country="Brazil" name="Para" pageId="3" pageNumber="488">PR</collectingRegion>
1.81–2.50 (2.07; n=3). Thorax (
<figureCitation id="136B2A6FFF87FFDCDD315616FE04A60A" box="[318,425,259,285]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="3" pageNumber="488">Figs. 1–2</figureCitation>
) dark brown with stout setae, humeral pits slightly lighter; scutellum with 4 stout setae. Legs dark brown; basal 2/3 of fore and mid femora, paler; hind leg darkest, apex of tibia yellowish; tibial comb with 7–8 spines; tarsomeres pale brown; prothoracic TR 1.93–2.1 (1.99; n=4); mesothoracic TR 2.17–2.41 (2.24; n=4); metathoracic TR 1.77–1.93 (1.86; n=4). Wing (
<figureCitation id="136B2A6FFF87FFDCDEA0567AFD5FA69E" box="[687,754,366,393]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="3" pageNumber="488">Fig. 5</figureCitation>
) hyaline with dark radial cells; second radial cell 2X longer than first; cubital fork at level of base of r-m cross vein; wing length 1.30–1.36 (1.32; n=4) mm; width 0.40–0.42 (0.41; n=4) mm; costal ratio 0.75–0.76 (0.75; n=4). Halter stem pale brown, knob whitish. Abdomen (
<figureCitation id="136B2A6FFF87FFDCD8F156A2FAF6A6C6" box="[1278,1371,439,465]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="3" pageNumber="488">Fig. 1–2</figureCitation>
) dark brown; tergites 1–8 with brown lateral spots; pleural segments 7–8 yellowish brown. Genitalia (
<figureCitation id="136B2A6FFF87FFDCD8EF56CEFAE2A6E2" box="[1248,1359,475,501]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="3" pageNumber="488">Figs. 8–9</figureCitation>
). Dark brown: tergite 9 triangular distal portion slightly sclerotized, bilobed with apical seta; sternite 9 3X wider than long; cerci conspicuous, divergent. Gonocoxite greatly elongate, crescent-shaped 5.70X longer than wide. Gonostylus 1.80X shorter than gonocoxite, curved about 90º at mid-length; apical portion with toothed apex. Parameres (
<figureCitation id="136B2A6FFF87FFDCD9605552FF7CA592" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="3" pageNumber="488">Fig. 9–10</figureCitation>
) subparallel, slender, stem slightly wider at base, apical portions abruptly curved, apically directed. Aedeagus (
<figureCitation id="136B2A6FFF87FFDCDC90559AFF5CA5BE" box="[159,241,655,681]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="3" pageNumber="488">Fig. 11</figureCitation>
) composed of two convergent sclerites, each slender, slightly curved, with widened, blunt apices. Lungshaped hyaline membranes (
<figureCitation id="136B2A6FFF87FFDCDDD955A6FDB4A5DA" box="[470,537,691,717]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="3" pageNumber="488">Fig. 9</figureCitation>
) between parameres and aedeagus, apparently basally connected.
</paragraph>
<paragraph id="8BEF36EAFF87FFDCDCC855C2FD45A306" blockId="3.[151,1437,150,1437]" pageId="3" pageNumber="488">
<emphasis id="B924EAF8FF87FFDCDCC855C2FEB6A5E6" box="[199,283,727,753]" italics="true" pageId="3" pageNumber="488">Female</emphasis>
. Similar to male, with the following notable sexual differences: Head (
<figureCitation id="136B2A6FFF87FFDCD83155C2FB0FA5E6" box="[1086,1186,727,753]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="3" pageNumber="488">Fig. 3–4</figureCitation>
). AR 1.17–1.18 (1.17; n=2);
<collectingRegion id="4994F808FF87FFDCDCD455EEFF50A402" box="[219,253,763,789]" country="Brazil" name="Para" pageId="3" pageNumber="488">PR</collectingRegion>
2.44–3.00 (2.71; n=3). Mandible with 6–7 teeth. Thorax (
<figureCitation id="136B2A6FFF87FFDCDF8455EEFC77A402" box="[907,986,763,789]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="3" pageNumber="488">Figs. 3</figureCitation>
): claws 1.42X shorter than tarsomere 5; fore and mid tarsomeres with 3 pairs of batonnets, hind tarsomere with 1 pair; basal tooth smaller; prothoracic TR 2.03–2.17 (2.12; n=3); mesothoracic TR 2.21–2.52 (2.36; n=3); metathoracic TR 2.15–2.29 (2.22; n=3). Wing (
<figureCitation id="136B2A6FFF87FFDCD9605456FF0BA496" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="3" pageNumber="488">Fig. 7</figureCitation>
) with macrotrichiae on C, R
<subScript id="17D434AFFF87FFDCDDE7546CFE5DA490" attach="left" box="[488,496,889,903]" fontSize="6" pageId="3" pageNumber="488">1</subScript>
, R
<subScript id="17D434AFFF87FFDCDE1E546CFDB4A490" attach="left" box="[529,537,889,903]" fontSize="6" pageId="3" pageNumber="488">3</subScript>
, and end of M
<subScript id="17D434AFFF87FFDCDEB0546CFD6AA490" attach="left" box="[703,711,889,903]" fontSize="6" pageId="3" pageNumber="488">1</subScript>
; wing length 1.14–1.46 (1.33; n=3) mm; width 0.42–0.54 (0.47; n=3) mm; costal ratio 0.77–0.79 (0.78; n=3). Abdomen (
<figureCitation id="136B2A6FFF87FFDCDF2B549EFCD7A4B2" box="[804,890,907,933]" captionStart="FIGURES 1–12" captionStartId="2.[151,264,1943,1967]" captionTargetBox="[205,1378,179,1907]" captionTargetId="figure-24@2.[205,1382,179,1918]" captionTargetPageId="2" captionText="FIGURES 1–12. Stilobezzia (Acanthohelea) medialuna n. sp., male 1–2, 6, 8–11; female 3, 6–7, 12; 1. Habitus, lateral view; 2 Habitus, dorsal view; 3. Habitus, lateral view. 4. Head, dorsal view; 5. Wing; 6–7. Wing; 8–9. Genitalia, ventral view; 10. Parameres, ventral view; 11. Aedeagus, ventral view. 12. Genitalia, ventral view." figureDoi="http://doi.org/10.5281/zenodo.7694633" httpUri="https://zenodo.org/record/7694633/files/figure.png" pageId="3" pageNumber="488">Fig. 12</figureCitation>
). Sternite 8 with large V-shaped posteromedian excavation and 4 pairs of setae; sternite 9 with sinuous, sclerotized margins; sternite 10 with 2 pairs of setae; cerci triangular, short, paler. Two ovoid, sclerotized spermathecae, without necks, measuring 90–80 µm and 70–70 µm (n=3), plus, an ovoid rudimentary third spermatheca.
</paragraph>
</subSubSection>
<subSubSection id="C34A6561FF87FFDCDCC8530FFD06A322" box="[199,683,1050,1077]" pageId="3" pageNumber="488" type="distribution">
<paragraph id="8BEF36EAFF87FFDCDCC8530FFD06A322" blockId="3.[151,1437,150,1437]" box="[199,683,1050,1077]" pageId="3" pageNumber="488">
<emphasis id="B924EAF8FF87FFDCDCC8530FFEF4A323" bold="true" box="[199,345,1050,1076]" pageId="3" pageNumber="488">Distribution</emphasis>
.
<collectingCountry id="F347767AFF87FFDCDD68530EFE06A322" box="[359,427,1051,1077]" name="Brazil" pageId="3" pageNumber="488">Brazil</collectingCountry>
(Amazonas: Manaus).
</paragraph>
</subSubSection>
<subSubSection id="C34A6561FF87FFDCDCC8532AFF72A36A" pageId="3" pageNumber="488" type="etymology">
<paragraph id="8BEF36EAFF87FFDCDCC8532AFF72A36A" blockId="3.[151,1437,150,1437]" pageId="3" pageNumber="488">
<emphasis id="B924EAF8FF87FFDCDCC8532AFEE5A34E" bold="true" box="[199,328,1087,1113]" pageId="3" pageNumber="488">Etymology</emphasis>
. The specific name
<taxonomicName id="4C504D69FF87FFDCDE3D532AFD05A34E" authorityName="Da Silva &amp; Ferreira-Keppler &amp; Cazorla" authorityYear="2023" box="[562,680,1087,1113]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="medialuna">
<emphasis id="B924EAF8FF87FFDCDE3D532AFD05A34E" box="[562,680,1087,1113]" italics="true" pageId="3" pageNumber="488">medialuna</emphasis>
</taxonomicName>
refers to the shape of the gonocoxite, which resembles a crescent moon.
</paragraph>
</subSubSection>
<subSubSection id="C34A6561FF87FFDCDCC85392FC33A28A" pageId="3" pageNumber="488" type="discussion">
<paragraph id="8BEF36EAFF87FFDCDCC85392FC33A28A" blockId="3.[151,1437,150,1437]" pageId="3" pageNumber="488">
<emphasis id="B924EAF8FF87FFDCDCC85392FE99A3B6" bold="true" box="[199,308,1159,1185]" pageId="3" pageNumber="488">Remarks</emphasis>
. This species resembles the Patagonian species
<taxonomicName id="4C504D69FF87FFDCDF625392FE9FA3D2" authority="Cazorla &amp; Spinelli, 2007" authorityName="Cazorla &amp; Spinelli" authorityYear="2007" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="curvistyla" subGenus="Acanthohelea">
<emphasis id="B924EAF8FF87FFDCDF625392FC4BA3B6" box="[877,998,1159,1185]" italics="true" pageId="3" pageNumber="488">Stilobezzia</emphasis>
(
<emphasis id="B924EAF8FF87FFDCDFF45392FB38A3B6" box="[1019,1173,1159,1185]" italics="true" pageId="3" pageNumber="488">Acanthohelea</emphasis>
)
<emphasis id="B924EAF8FF87FFDCD8A55392FAB5A3B6" box="[1194,1304,1159,1185]" italics="true" pageId="3" pageNumber="488">curvistyla</emphasis>
<bibRefCitation id="EFC14B1BFF87FFDCD92B5392FE9FA3D2" author="Cazorla, C. G. &amp; Spinelli, G. R." pageId="3" pageNumber="488" pagination="181 - 187" refId="ref8488" refString="Cazorla, C. G. &amp; Spinelli, G. R. (2007) A new species of Patagonian Stilobezzia (Acanthohelea) and a redescription of S. (A.) nigerrima Ingram and Macfie (Diptera: Ceratopogonidae). The American Entomological Society, 133 (1), 181 - 187. https: // doi. org / 10.3157 / 0002 - 8320 (2007) 133 [181: ANSOPS] 2.0. CO; 2" type="journal article" year="2007">Cazorla &amp; Spinelli, 2007</bibRefCitation>
</taxonomicName>
by the shape of the aedeagus and the 90º curved gonostylus, however
<taxonomicName id="4C504D69FF87FFDCD82153BEFB1BA3D2" authorityName="Cazorla &amp; Spinelli" authorityYear="2007" box="[1070,1206,1195,1221]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="curvistyla">
<emphasis id="B924EAF8FF87FFDCD82153BEFB1BA3D2" box="[1070,1206,1195,1221]" italics="true" pageId="3" pageNumber="488">S. curvistyla</emphasis>
</taxonomicName>
can be distinguished by the pointed apex of the gonostylus, the sclerotized protuberance on the proximal inner margin of the gonocoxite and the conspicuous posterior pointed projection of tergite 9. Females of
<taxonomicName id="4C504D69FF87FFDCDFD853E6FBCFA21A" authorityName="Cazorla &amp; Spinelli" authorityYear="2007" box="[983,1122,1267,1293]" class="Insecta" family="Ceratopogonidae" genus="Stilobezzia" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="488" phylum="Arthropoda" rank="species" species="curvistyla">
<emphasis id="B924EAF8FF87FFDCDFD853E6FBCFA21A" box="[983,1122,1267,1293]" italics="true" pageId="3" pageNumber="488">S. curvistyla</emphasis>
</taxonomicName>
have much more elongated spermathecae and the posteromedian excavation of sternite 8 is U-shaped. The male genitalia of this new species also resemble
<emphasis id="B924EAF8FF87FFDCDD38522EFEE8A242" box="[311,325,1339,1365]" italics="true" pageId="3" pageNumber="488">S</emphasis>
. (
<emphasis id="B924EAF8FF87FFDCDD52522EFEC3A242" box="[349,366,1339,1365]" italics="true" pageId="3" pageNumber="488">A</emphasis>
.)
<emphasis id="B924EAF8FF87FFDCDD8A522EFE73A243" box="[389,478,1339,1364]" italics="true" pageId="3" pageNumber="488">guarani</emphasis>
<bibRefCitation id="EFC14B1BFF87FFDCDDEA522EFCADA242" author="Cazorla, C. G. &amp; Spinelli, G. R." box="[485,768,1339,1365]" pageId="3" pageNumber="488" pagination="45 - 56" refId="ref8573" refString="Cazorla, C. G. &amp; Spinelli, G. R. (2010) Four new Neotropical species of Stilobezzia (Acanthohelea) Kieffer (Diptera: Ceratopogonidae). Zootaxa, 2669 (1), 45 - 56. https: // doi. org / 10.11646 / zootaxa. 2669.1.3" type="journal article" year="2010">Cazorla &amp; Spinelli, 2010</bibRefCitation>
, from northeastern
<collectingCountry id="F347767AFF87FFDCDFD7522EFBEAA242" box="[984,1095,1339,1365]" name="Argentina" pageId="3" pageNumber="488">Argentina</collectingCountry>
and
<collectingCountry id="F347767AFF87FFDCD871522EFB4AA242" box="[1150,1255,1339,1365]" name="Paraguay" pageId="3" pageNumber="488">Paraguay</collectingCountry>
, by the shape of the aedeagus and the elongated parameres with curved apical portion, but differing in the shape of the gonocoxite, elongate with inner protuberance bearing two small sclerotized teeth.
</paragraph>
</subSubSection>
</treatment>
</document>
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