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<document id="5714A772DC40EAF445710BB6D8266029" ID-CLB-Dataset="31714" ID-DOI="10.11646/zootaxa.4339.1.1" ID-GBIF-Dataset="e30650b4-4b5d-4031-8f07-e57668e07101" ID-ISSN="1175-5326" ID-Zenodo-Dep="1037681" ID-ZooBank="F5885D17-F256-4F6B-9D5D-1555637C116C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1509092316730" checkinUser="plazi" docAuthor="He, Liang &amp; Zhou, Hong-Zhang" docDate="2017" docId="03F987B68829FFBB1CCAFF3ED92EF856" docLanguage="en" docName="zootaxa.4339.1.1.pdf" docOrigin="Zootaxa 4339 (1)" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Ocypus (Pseudocypus) anguliculminis He &amp; Zhou 2017, sp. nov." docType="treatment" docVersion="7" lastPageNumber="17" masterDocId="FFC0FFCE8839FFAB1C5DFFA6DA56FFC5" masterDocTitle="Taxonomy of the subgenus Pseudocypus Mulsant &amp; Rey (Coleoptera: Staphylinidae: Staphylinini) and eight new species from China" masterLastPageNumber="77" masterPageNumber="1" pageNumber="17" updateTime="1698470325630" updateUser="ExternalLinkService">
<mods:mods id="D5FB7DBF950AB2EAFA0D56F28DF3376C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9E2862D33246083E460EC3EF5FD153B3">
<mods:title id="6E8557D9AB976A434F71CB36E7D15E18">Taxonomy of the subgenus Pseudocypus Mulsant &amp; Rey (Coleoptera: Staphylinidae: Staphylinini) and eight new species from China</mods:title>
</mods:titleInfo>
<mods:name id="D77B64452A9087540B53DA462D75EBAC" type="personal">
<mods:role id="7A7249439254FB3E61C817C26CC3C5E6">
<mods:roleTerm id="C6F80DC54B6B449A440DB5A1E82A6F22">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1903A6BD1188089A1DAB838B5D109A62">He, Liang</mods:namePart>
</mods:name>
<mods:name id="58092652B74E563EC95DCEC00D323AD6" type="personal">
<mods:role id="BAD6AC1292441AFBBAC6204CB13624D4">
<mods:roleTerm id="2057C76D878864295E4C15511310F74C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="81CE1D0F800159E8C8AC60B3811B29EB">Zhou, Hong-Zhang</mods:namePart>
</mods:name>
<mods:typeOfResource id="5F16E6C96108193EB0170075C509A963">text</mods:typeOfResource>
<mods:relatedItem id="A9FF553184DCB49450EAD482CC156093" type="host">
<mods:titleInfo id="A1D3F12F144FAAF2504E0FC1B8026A91">
<mods:title id="449EDB05917B177FE7606FA7A2E56EC9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="77C32B9F5983B439A1A5C1A9F9C6D5A7">
<mods:date id="CDF3CC3AE18F8BEAC1DE0140F4871992">2017</mods:date>
<mods:detail id="DC9CCF9AF9A6E0C2F09944635BAE752A" type="pubDate">
<mods:number id="6754B76F6E015E243003BF9E7EF29500">2017-10-27</mods:number>
</mods:detail>
<mods:detail id="43EC17A4F5F8258D30D36640F9B3C04A" type="volume">
<mods:number id="BE1BB8A77DBEA7A20C640E32A4B6214A">4339</mods:number>
</mods:detail>
<mods:detail id="6DF7B0F97E6D83CD18498A78DED6735F" type="issue">
<mods:number id="E8AAED81462BD9F3FFF2E276287B61FD">1</mods:number>
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<mods:start id="C26D9619CC392781B0A76CC10F6F1C3A">1</mods:start>
<mods:end id="92AD20D9163C5044BB9E9344463098C5">77</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="8FE3779D7724E730347BEA1B2B8A490F">journal article</mods:classification>
<mods:identifier id="0274FBC3819CECA6F81124414121C058" type="CLB-Dataset">31714</mods:identifier>
<mods:identifier id="F3A655A205C84FC66191C3F90EC4A8DE" type="DOI">10.11646/zootaxa.4339.1.1</mods:identifier>
<mods:identifier id="5B82A89358289CEFB391E92F1B463C76" type="GBIF-Dataset">e30650b4-4b5d-4031-8f07-e57668e07101</mods:identifier>
<mods:identifier id="7D80971BD7AC282BD4B462F6682344DE" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="99B50FF51CE5A264EEB87F8938316643" type="Zenodo-Dep">1037681</mods:identifier>
<mods:identifier id="5C786C13BAA44983C71559790BB28BAB" type="ZooBank">F5885D17-F256-4F6B-9D5D-1555637C116C</mods:identifier>
</mods:mods>
<treatment id="03F987B68829FFBB1CCAFF3ED92EF856" ID-DOI="http://doi.org/10.5281/zenodo.6014877" ID-GBIF-Taxon="135476898" ID-Zenodo-Dep="6014877" LSID="urn:lsid:plazi:treatment:03F987B68829FFBB1CCAFF3ED92EF856" httpUri="http://treatment.plazi.org/id/03F987B68829FFBB1CCAFF3ED92EF856" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="C34A652B8829FFBB1CCAFF3ED8BEFF77" box="[151,744,151,178]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BEF36A08829FFBB1CCAFF3ED8BEFF77" blockId="16.[151,744,151,211]" box="[151,744,151,178]" pageId="16" pageNumber="17">
<heading id="D0A781CC8829FFBB1CCAFF3ED8BEFF77" bold="true" box="[151,744,151,178]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B924EAB28829FFBB1CCAFF3ED8BEFF77" bold="true" box="[151,744,151,178]" pageId="16" pageNumber="17">
3.
<taxonomicName id="4C504D238829FFBB1CE9FF31D8D5FF74" ID-CoL="48MDP" authority="He &amp; Zhou, 2017" authorityName="He &amp; Zhou" authorityYear="2017" box="[180,643,151,178]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="anguliculminis" status="sp. nov." subGenus="Pseudocypus">
<emphasis id="B924EAB28829FFBB1CE9FF31DB5BFF74" bold="true" box="[180,269,151,177]" italics="true" pageId="16" pageNumber="17">Ocypus</emphasis>
(
<emphasis id="B924EAB28829FFBB1D42FF31DBEDFF74" bold="true" box="[287,443,151,177]" italics="true" pageId="16" pageNumber="17">Pseudocypus</emphasis>
)
<emphasis id="B924EAB28829FFBB1D91FF31D8D5FF74" bold="true" box="[460,643,151,177]" italics="true" pageId="16" pageNumber="17">anguliculminis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757C98829FFBB1ED7FF3ED8BEFF77" box="[650,744,152,178]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A652B8829FFBB1CCAFF1CDB16FF16" box="[151,320,186,211]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BEF36A08829FFBB1CCAFF1CDB16FF16" blockId="16.[151,744,151,211]" box="[151,320,186,211]" pageId="16" pageNumber="17">
(
<figureCitation id="136B2A258829FFBB1CC2FF1CDB6EFF16" box="[159,312,186,211]" captionStart="FIGURE 2 - 3" captionStartId="15.[151,250,1310,1332]" captionTargetBox="[349,1244,211,1268]" captionTargetId="figure@15.[343,1250,203,1272]" captionTargetPageId="15" captionText="FIGURE 2 - 3. Ocypus (Pseudocypus) denticulminis sp. nov.: A, B and C: variations of teeth on mandibles (1.0 mm); D. habitus of one of the special large specimens collected together with Ocypus (Pseudocypus) persiciculminis sp. nov. (2.0 mm). Scale in parentheses." httpUri="https://zenodo.org/record/1037691/files/figure.png" pageId="16" pageNumber="17">Figs. 3-1, 3-2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34A652B8829FFBB1CCAFEA4DF3BFE36" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BEF36A08829FFBB1CCAFEA4D99EFEDE" blockId="16.[151,1437,258,1939]" box="[151,968,258,283]" pageId="16" pageNumber="17">
<materialsCitation id="3B383CFD8829FFBB1CCAFEA4D99EFEDE" ID-GBIF-Occurrence="1676080661" box="[151,968,258,283]" country="China" county="Danba County" location="Danba County" pageId="16" pageNumber="17" specimenCount="1" stateProvince="Sichuan Province" typeStatus="holotype">
<emphasis id="B924EAB28829FFBB1CCAFEA4DB6FFEDE" bold="true" box="[151,313,258,283]" pageId="16" pageNumber="17">
<typeStatus id="54EB88028829FFBB1CCAFEA4DA84FEDE" box="[151,210,258,283]" pageId="16" pageNumber="17">Type</typeStatus>
locality:
</emphasis>
<collectingCountry id="F34776308829FFBB1D1DFEA5DBCAFEDE" box="[320,412,259,283]" name="China" pageId="16" pageNumber="17">CHINA</collectingCountry>
,
<collectingRegion id="4994F8428829FFBB1DFBFEA4D839FEDE" box="[422,623,258,283]" country="China" name="Sichuan" pageId="16" pageNumber="17">Sichuan Province</collectingRegion>
,
<collectingCounty id="628E4E2C8829FFBB1E24FEA4D94BFEDE" box="[633,797,258,283]" pageId="16" pageNumber="17">Danba County</collectingCounty>
, Geshizaxiang.
</materialsCitation>
</paragraph>
<paragraph id="8BEF36A08829FFBB1C9AFE80DF3BFE36" blockId="16.[151,1437,258,1939]" pageId="16" pageNumber="17">
<materialsCitation id="3B383CFD8829FFBB1C9AFE80DFCFFEA6" ID-GBIF-Occurrence="1676080669" collectingDate="2001-07-20" collectingDateMax="2001-07-22" collectingDateMin="2001-07-20" collectorName="Xiaodong Yu &amp; Hongzhang Zhou" country="China" county="Danba County" elevation="2640" location="Geshizaxiang" pageId="16" pageNumber="17" specimenCount="2" stateProvince="Sichuan Province" typeStatus="holotype">
<emphasis id="B924EAB28829FFBB1C9AFE80DBBAFEFA" bold="true" box="[199,492,294,319]" pageId="16" pageNumber="17">
<typeStatus id="54EB88028829FFBB1C9AFE80DB54FEFA" box="[199,258,294,319]" pageId="16" pageNumber="17">Type</typeStatus>
material.
<typeStatus id="54EB88028829FFBB1D22FE80DBBAFEFA" box="[383,492,294,319]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
</emphasis>
male,
<emphasis id="B924EAB28829FFBB1E66FE80D95DFEFA" bold="true" box="[571,779,294,319]" pageId="16" pageNumber="17">
<collectingCountry id="F34776308829FFBB1E66FE80D8CBFEFA" box="[571,669,294,319]" name="China" pageId="16" pageNumber="17">CHINA</collectingCountry>
:
<collectingRegion id="4994F8428829FFBB1EF7FE80D95DFEFA" box="[682,779,294,319]" country="China" name="Sichuan" pageId="16" pageNumber="17">Sichuan</collectingRegion>
</emphasis>
Province: west of
<collectingCounty id="628E4E2C8829FFBB1FB9FE80DEDCFEFA" box="[996,1162,294,319]" pageId="16" pageNumber="17">Danba County</collectingCounty>
,
<location id="8E8F607B8829FFBB18CBFE80DF67FEFA" LSID="urn:lsid:plazi:treatment:03F987B68829FFBB1CCAFF3ED92EF856:8E8F607B8829FFBB18CBFE80DF67FEFA" box="[1174,1329,294,319]" country="China" county="Danba County" name="Geshizaxiang" pageId="16" pageNumber="17" stateProvince="Sichuan Province">Geshizaxiang</location>
,
<elevation id="007DD1938829FFBB1962FE80DFC1FEFB" box="[1343,1431,294,319]" metricMagnitude="3" metricUnit="m" metricValue="2.6399999999999997" pageId="16" pageNumber="17" unit="m" value="2640.0">
<quantity id="4CA89B458829FFBB1962FE80DFC1FEFB" box="[1343,1431,294,319]" metricMagnitude="3" metricUnit="m" metricValue="2.6399999999999997" pageId="16" pageNumber="17" unit="m" value="2640.0">2640 m</quantity>
</elevation>
, mixed broad-leaved forest,
<collectingMethod id="52114EB78829FFBB1DBDFEECD83FFEA7" box="[480,617,330,355]" pageId="16" pageNumber="17">pitfall traps</collectingMethod>
,
<collectingDate id="EFAAE9888829FFBB1E21FEEDD967FEA6" box="[636,817,331,355]" pageId="16" pageNumber="17" value="2001-07-20" valueMax="2001-07-22" valueMin="2001-07-20">
<date id="FFEE10608829FFBB1E21FEEDD967FEA6" box="[636,817,331,355]" pageId="16" pageNumber="17" value="2001-07-20" valueMax="2001-07-22" valueMin="2001-07-20">20–22.VII.2001</date>
</collectingDate>
, coll.
<collectorName id="26A553768829FFBB1FDEFEECDE48FEA7" box="[899,1054,330,355]" pageId="16" pageNumber="17">Xiaodong Yu</collectorName>
&amp;
<collectorName id="26A553768829FFBB180DFEECDF4CFEA6" box="[1104,1306,330,355]" pageId="16" pageNumber="17">Hongzhang Zhou</collectorName>
(IZ-CAS)
</materialsCitation>
.
<materialsCitation id="3B383CFD8829FFBB1CCAFEC8DEE1FE42" ID-GBIF-Occurrence="1676080674" box="[151,1207,366,391]" collectingDate="2001-07-20" collectingDateMax="2001-07-22" collectingDateMin="2001-07-20" collectorName="Xiaodong Yu &amp; Hongzhang Zhou" country="China" county="Danba County" elevation="2640" location="Geshizaxiang" pageId="16" pageNumber="17" specimenCount="15" specimenCount-female="6" specimenCount-male="9" stateProvince="Sichuan Province" typeStatus="paratype">
<typeStatus id="54EB88028829FFBB1CCAFEC8DB59FE42" box="[151,271,366,391]" pageId="16" pageNumber="17" type="paratype">
<emphasis id="B924EAB28829FFBB1CCAFEC8DB59FE42" bold="true" box="[151,271,366,391]" pageId="16" pageNumber="17">Paratypes</emphasis>
</typeStatus>
:
<emphasis id="B924EAB28829FFBB1D43FEC8DBBBFE42" bold="true" box="[286,493,366,391]" pageId="16" pageNumber="17">
<collectingCountry id="F34776308829FFBB1D43FEC8DBD6FE42" box="[286,384,366,391]" name="China" pageId="16" pageNumber="17">CHINA</collectingCountry>
:
<collectingRegion id="4994F8428829FFBB1DD1FEC8DBBBFE42" box="[396,493,366,391]" country="China" name="Sichuan" pageId="16" pageNumber="17">Sichuan</collectingRegion>
</emphasis>
Province:
<specimenCount id="9D56FD298829FFBB1E3AFEC9D896FE42" box="[615,704,366,391]" pageId="16" pageNumber="17" type="male">9 males</specimenCount>
,
<specimenCount id="9D56FD298829FFBB1E91FEC8D96AFE42" box="[716,828,366,391]" pageId="16" pageNumber="17" type="female">6 females</specimenCount>
, same data as holotype (IZ-CAS)
</materialsCitation>
;
<materialsCitation id="3B383CFD8829FFBB189EFEC8D8DFFE0A" ID-GBIF-Occurrence="1676080685" collectingDate="2001-07-20" collectingDateMax="2001-07-22" collectingDateMin="2001-07-20" collectorName="Geshizaxiang &amp; Xiaodong Yu &amp; Hongzhang Zhou" country="China" county="Danba County" elevation="2610" location="Danba County" pageId="16" pageNumber="17" specimenCount="8" specimenCount-female="3" specimenCount-male="5" stateProvince="Sichuan Province" typeStatus="paratype">
<specimenCount id="9D56FD298829FFBB189EFEC8DF4AFE42" box="[1219,1308,366,391]" pageId="16" pageNumber="17" type="male">5 males</specimenCount>
,
<specimenCount id="9D56FD298829FFBB1975FEC9DFCEFE42" box="[1320,1432,366,391]" pageId="16" pageNumber="17" type="female">3 females</specimenCount>
, west of
<collectingCounty id="628E4E2C8829FFBB1CAFFE34DBC1FE6E" box="[242,407,402,427]" pageId="16" pageNumber="17">Danba County</collectingCounty>
,
<collectorName id="26A553768829FFBB1DFEFE34D868FE6E" box="[419,574,402,427]" pageId="16" pageNumber="17">Geshizaxiang</collectorName>
,
<elevation id="007DD1938829FFBB1E16FE34D8F5FE6F" box="[587,675,402,427]" metricMagnitude="3" metricUnit="m" metricValue="2.61" pageId="16" pageNumber="17" unit="m" value="2610.0">
<quantity id="4CA89B458829FFBB1E16FE34D8F5FE6F" box="[587,675,402,427]" metricMagnitude="3" metricUnit="m" metricValue="2.61" pageId="16" pageNumber="17" unit="m" value="2610.0">2610 m</quantity>
</elevation>
, mixed broadleaf-conifer forest,
<collectingMethod id="52114EB78829FFBB1845FE34DECAFE6F" box="[1048,1180,402,427]" pageId="16" pageNumber="17">pitfall traps</collectingMethod>
,
<collectingDate id="EFAAE9888829FFBB18F4FE35DF08FE6E" box="[1193,1374,403,427]" pageId="16" pageNumber="17" value="2001-07-20" valueMax="2001-07-22" valueMin="2001-07-20">
<date id="FFEE10608829FFBB18F4FE35DF08FE6E" box="[1193,1374,403,427]" pageId="16" pageNumber="17" value="2001-07-20" valueMax="2001-07-22" valueMin="2001-07-20">20–22.VII.2001</date>
</collectingDate>
, coll.
<collectorName id="26A553768829FFBB1CCAFE10DB7DFE0B" box="[151,299,438,463]" pageId="16" pageNumber="17">Xiaodong Yu</collectorName>
&amp;
<collectorName id="26A553768829FFBB1D13FE10D844FE0A" box="[334,530,438,463]" pageId="16" pageNumber="17">Hongzhang Zhou</collectorName>
(IZ-CAS)
</materialsCitation>
;
<materialsCitation id="3B383CFD8829FFBB1EC9FE11DF3BFE36" ID-GBIF-Occurrence="1676080688" collectingDate="2001-07-20" collectingDateMax="2001-07-22" collectingDateMin="2001-07-20" collectorName="Geshizaxiang &amp; Xiaodong Yu &amp; Hongzhang Zhou" country="China" county="Danba County" elevation="2650" location="Danba County" pageId="16" pageNumber="17" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Sichuan Province" typeStatus="paratype">
<specimenCount id="9D56FD298829FFBB1EC9FE11D8BAFE0A" box="[660,748,438,463]" pageId="16" pageNumber="17" type="male">2 males</specimenCount>
,
<specimenCount id="9D56FD298829FFBB1EA5FE11D931FE0A" box="[760,871,438,463]" pageId="16" pageNumber="17" type="female">3 females</specimenCount>
, west of
<collectingCounty id="628E4E2C8829FFBB1F94FE10DE38FE0A" box="[969,1134,438,463]" pageId="16" pageNumber="17">Danba County</collectingCounty>
,
<collectorName id="26A553768829FFBB1825FE10DF45FE0A" box="[1144,1299,438,463]" pageId="16" pageNumber="17">Geshizaxiang</collectorName>
,
<elevation id="007DD1938829FFBB1943FE10DAF9FE37" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="2.68" metricValueMin="2.62" pageId="16" pageNumber="17" unit="m" value="2650.0" valueMax="2680.0" valueMin="2620.0">
<quantity id="4CA89B458829FFBB1943FE10DAF9FE37" metricMagnitude="3" metricUnit="m" metricValue="2.65" metricValueMax="2.68" metricValueMin="2.62" pageId="16" pageNumber="17" unit="m" value="2650.0" valueMax="2680.0" valueMin="2620.0">2620–2680 m</quantity>
</elevation>
, mixed broad-leaved forest,
<collectingMethod id="52114EB78829FFBB1DB2FE7CD827FE37" box="[495,625,474,499]" pageId="16" pageNumber="17">pitfall traps</collectingMethod>
,
<collectingDate id="EFAAE9888829FFBB1E20FE7DD964FE36" box="[637,818,475,499]" pageId="16" pageNumber="17" value="2001-07-20" valueMax="2001-07-22" valueMin="2001-07-20">
<date id="FFEE10608829FFBB1E20FE7DD964FE36" box="[637,818,475,499]" pageId="16" pageNumber="17" value="2001-07-20" valueMax="2001-07-22" valueMin="2001-07-20">20–22.VII.2001</date>
</collectingDate>
, coll.
<collectorName id="26A553768829FFBB1F2BFE7CDE5CFE37" box="[886,1034,474,499]" pageId="16" pageNumber="17">Xiaodong Yu</collectorName>
&amp;
<collectorName id="26A553768829FFBB1873FE7CDEA7FE36" box="[1070,1265,474,499]" pageId="16" pageNumber="17">Hongzhang Zhou</collectorName>
(IZ-CAS).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A652B8829FFBB1C9AFE58D9E4F97E" pageId="16" pageNumber="17" type="description">
<paragraph id="8BEF36A08829FFBB1C9AFE58DBBFFDFE" blockId="16.[151,1437,258,1939]" pageId="16" pageNumber="17">
<emphasis id="B924EAB28829FFBB1C9AFE58DB2BFDD2" bold="true" box="[199,381,510,535]" pageId="16" pageNumber="17">Measurements.</emphasis>
Body length:
<quantity id="4CA89B458829FFBB1E43FE58D893FDD3" box="[542,709,510,535]" metricMagnitude="-2" metricUnit="m" metricValue="2.245" metricValueMax="2.43" metricValueMin="2.06" pageId="16" pageNumber="17" unit="mm" value="22.45" valueMax="24.3" valueMin="20.6">20.6–24.3 mm</quantity>
. CL: 0.76; EL: 3.22; ELS: 1.48; EW: 3.85; HL: 2.99; HW: 3.89; PO: 1.66; PL: 4.09; PW: 3.68.
</paragraph>
<paragraph id="8BEF36A08829FFBB1C9AFDE0DBFAFD0F" blockId="16.[151,1437,258,1939]" pageId="16" pageNumber="17">
<emphasis id="B924EAB28829FFBB1C9AFDE0DB0FFD9A" bold="true" box="[199,345,582,607]" pageId="16" pageNumber="17">Description.</emphasis>
Large sized species. Body robust, entirely black; maxillary and labial palpi dark brunneous; antennae black, with apical segments gradually becoming paler; legs black; protibia and protarsus covered with dense brown pubescence; pubescence of dorsal side of head and pronotum black, and that of elytra and abdominal tergites almost the same.
</paragraph>
<paragraph id="8BEF36A08829FFBB1C9AFD70D941FA96" blockId="16.[151,1437,258,1939]" pageId="16" pageNumber="17">Head of rounded quadrangular shape, moderately large, with rounded lateral sides and posterior angles, distinctly wider than long (ratio 1.30), posterior half slightly becoming narrower towards basal portion; tempora moderately long. Eyes relatively small and slightly protruding, considerably shorter than tempora (ratio 0.46) in dorsal view. Punctation and pubescence on dorsal surface of head relatively dense and fine, puncture intervals about as long as or smaller than diameters of punctures; punctation gradually becoming sparser toward clypeus; no impunctate midline present. Gular sutures distinctly separated. Antenna moderately long, segment 3 distinctly longer than segment 2 (ratio 1.38), segments 4 to 7 longer than wide, segment 8 to 10 about as long as wide, segment 11 longer than segment 10 (ratio 1.18), all segments except segment 11 gradually becoming shorter toward apex. Pronotum vaguely longer than wide (ratio 1.11), with almost parallel lateral sides; narrow marginal groove disappearing downwards at anterior third of pronotal length; impunctate midline distinct; punctation and pubescence on disc sparser than that on dorsal surface of head, puncture intervals larger than diameters of punctures. Pronotal hypomeron lacking microsetae. Scutellum finely punctate and setose on entire surface. Elytra short, moderately depressed at base, slightly dilated posteriad; elytral length shorter than pronotum along midline, with a ratio of 0.36 along suture and a ratio of 0.79 along sides; punctation much finer and denser than that on head disc, combined with very fine and dense microsculpture. Wings markedly reduced and budlike. Abdomen tergite 2 densely and finely punctate and pubescent on entire surface; all visible abdominal tergites very densely and finely punctate and pubescent, punctation becoming slightly sparser and coarser toward apex of abdomen; no yellowish tomentose pubescence present on visible tergites 4 and 5.</paragraph>
<paragraph id="8BEF36A08829FFBB1C9AFAF8DBC1F98A" blockId="16.[151,1437,258,1939]" pageId="16" pageNumber="17">
<emphasis id="B924EAB28829FFBB1C9AFAF8DB5DFAB2" bold="true" box="[199,267,1374,1399]" pageId="16" pageNumber="17">Male.</emphasis>
Sternite 8 with wide and markedly deep, acutely triangular medioapical emargination (
<figureCitation id="136B2A258829FFBB194BFAF8DFD9FAB2" box="[1302,1423,1374,1399]" captionStart="FIGURE 2 - 3" captionStartId="15.[151,250,1310,1332]" captionTargetBox="[349,1244,211,1268]" captionTargetId="figure@15.[343,1250,203,1272]" captionTargetPageId="15" captionText="FIGURE 2 - 3. Ocypus (Pseudocypus) denticulminis sp. nov.: A, B and C: variations of teeth on mandibles (1.0 mm); D. habitus of one of the special large specimens collected together with Ocypus (Pseudocypus) persiciculminis sp. nov. (2.0 mm). Scale in parentheses." httpUri="https://zenodo.org/record/1037691/files/figure.png" pageId="16" pageNumber="17">Fig. 3-2 J</figureCitation>
). Sternite 9 with narrow and shallow, acute apical emarginate, tapered basal portion moderately long (
<figureCitation id="136B2A258829FFBB194EFA24DFC6FA5E" box="[1299,1424,1410,1435]" captionStart="FIGURE 2 - 3" captionStartId="15.[151,250,1310,1332]" captionTargetBox="[349,1244,211,1268]" captionTargetId="figure@15.[343,1250,203,1272]" captionTargetPageId="15" captionText="FIGURE 2 - 3. Ocypus (Pseudocypus) denticulminis sp. nov.: A, B and C: variations of teeth on mandibles (1.0 mm); D. habitus of one of the special large specimens collected together with Ocypus (Pseudocypus) persiciculminis sp. nov. (2.0 mm). Scale in parentheses." httpUri="https://zenodo.org/record/1037691/files/figure.png" pageId="16" pageNumber="17">Fig. 3-2 H</figureCitation>
). Tergite 10 wide, shaped as in
<figureCitation id="136B2A258829FFBB1DB8FA00D80BFA7A" box="[485,605,1446,1471]" captionStart="FIGURE 2 - 3" captionStartId="15.[151,250,1310,1332]" captionTargetBox="[349,1244,211,1268]" captionTargetId="figure@15.[343,1250,203,1272]" captionTargetPageId="15" captionText="FIGURE 2 - 3. Ocypus (Pseudocypus) denticulminis sp. nov.: A, B and C: variations of teeth on mandibles (1.0 mm); D. habitus of one of the special large specimens collected together with Ocypus (Pseudocypus) persiciculminis sp. nov. (2.0 mm). Scale in parentheses." httpUri="https://zenodo.org/record/1037691/files/figure.png" pageId="16" pageNumber="17">Fig. 3-2 A</figureCitation>
. Aedeagus large and long, almost totally black, strongly sclerotized (
<figureCitation id="136B2A258829FFBB1939FA00DB24FA26" captionStart="FIGURE 2 - 3" captionStartId="15.[151,250,1310,1332]" captionTargetBox="[349,1244,211,1268]" captionTargetId="figure@15.[343,1250,203,1272]" captionTargetPageId="15" captionText="FIGURE 2 - 3. Ocypus (Pseudocypus) denticulminis sp. nov.: A, B and C: variations of teeth on mandibles (1.0 mm); D. habitus of one of the special large specimens collected together with Ocypus (Pseudocypus) persiciculminis sp. nov. (2.0 mm). Scale in parentheses." httpUri="https://zenodo.org/record/1037691/files/figure.png" pageId="16" pageNumber="17">Figs. 3-1 B, C, 3-2 B–D</figureCitation>
); apical portion of median lobe elongate and narrow (
<figureCitation id="136B2A258829FFBB1F82FA6CDE01FA26" box="[991,1111,1482,1507]" captionStart="FIGURE 2 - 3" captionStartId="15.[151,250,1310,1332]" captionTargetBox="[349,1244,211,1268]" captionTargetId="figure@15.[343,1250,203,1272]" captionTargetPageId="15" captionText="FIGURE 2 - 3. Ocypus (Pseudocypus) denticulminis sp. nov.: A, B and C: variations of teeth on mandibles (1.0 mm); D. habitus of one of the special large specimens collected together with Ocypus (Pseudocypus) persiciculminis sp. nov. (2.0 mm). Scale in parentheses." httpUri="https://zenodo.org/record/1037691/files/figure.png" pageId="16" pageNumber="17">Fig. 3-2 F</figureCitation>
); apical portion of paramere slightly asymmetrical, almost square-shaped (
<figureCitation id="136B2A258829FFBB1EC6FA48D926F9C2" box="[667,880,1518,1543]" captionStart="FIGURE 2 - 3" captionStartId="15.[151,250,1310,1332]" captionTargetBox="[349,1244,211,1268]" captionTargetId="figure@15.[343,1250,203,1272]" captionTargetPageId="15" captionText="FIGURE 2 - 3. Ocypus (Pseudocypus) denticulminis sp. nov.: A, B and C: variations of teeth on mandibles (1.0 mm); D. habitus of one of the special large specimens collected together with Ocypus (Pseudocypus) persiciculminis sp. nov. (2.0 mm). Scale in parentheses." httpUri="https://zenodo.org/record/1037691/files/figure.png" pageId="16" pageNumber="17">Figs. 3-1 D, 3-2 G</figureCitation>
); all sensory peg setae arranged in two lines near lateral sides and apical ends of paramere; several sensory peg setae joined together, forming two hornlike structure near apex (
<figureCitation id="136B2A258829FFBB1D4FF990DBDDF98A" box="[274,395,1590,1615]" captionStart="FIGURE 2 - 3" captionStartId="15.[151,250,1310,1332]" captionTargetBox="[349,1244,211,1268]" captionTargetId="figure@15.[343,1250,203,1272]" captionTargetPageId="15" captionText="FIGURE 2 - 3. Ocypus (Pseudocypus) denticulminis sp. nov.: A, B and C: variations of teeth on mandibles (1.0 mm); D. habitus of one of the special large specimens collected together with Ocypus (Pseudocypus) persiciculminis sp. nov. (2.0 mm). Scale in parentheses." httpUri="https://zenodo.org/record/1037691/files/figure.png" pageId="16" pageNumber="17">Fig. 3-1 D</figureCitation>
).
</paragraph>
<paragraph id="8BEF36A08829FFBB1C9AF9FCD9E4F97E" blockId="16.[151,1437,258,1939]" pageId="16" pageNumber="17">
<emphasis id="B924EAB28829FFBB1C9AF9FCDB73F9B6" bold="true" box="[199,293,1626,1651]" pageId="16" pageNumber="17">Female.</emphasis>
Second gonocoxite robust, moderately long (
<figureCitation id="136B2A258829FFBB1F7CF9FCD98DF9B6" box="[801,987,1626,1651]" captionStart="FIGURE 2 - 3" captionStartId="15.[151,250,1310,1332]" captionTargetBox="[349,1244,211,1268]" captionTargetId="figure@15.[343,1250,203,1272]" captionTargetPageId="15" captionText="FIGURE 2 - 3. Ocypus (Pseudocypus) denticulminis sp. nov.: A, B and C: variations of teeth on mandibles (1.0 mm); D. habitus of one of the special large specimens collected together with Ocypus (Pseudocypus) persiciculminis sp. nov. (2.0 mm). Scale in parentheses." httpUri="https://zenodo.org/record/1037691/files/figure.png" pageId="16" pageNumber="17">
Figs. 3-1 E 3-
<date id="FFEE10608829FFBB1FE7F9FDD98DF9B6" box="[954,987,1627,1651]" pageId="16" pageNumber="17">2 I</date>
</figureCitation>
). Tergite 10 of female moderately wide, with apical portion strongly sclerotized; lateral sides each with a large semi-circular indentation; apical portion gradually narrowed toward subacute apex, sparsely setose (
<figureCitation id="136B2A258829FFBB1F72F904D9F2F97E" box="[815,932,1698,1723]" captionStart="FIGURE 2 - 3" captionStartId="15.[151,250,1310,1332]" captionTargetBox="[349,1244,211,1268]" captionTargetId="figure@15.[343,1250,203,1272]" captionTargetPageId="15" captionText="FIGURE 2 - 3. Ocypus (Pseudocypus) denticulminis sp. nov.: A, B and C: variations of teeth on mandibles (1.0 mm); D. habitus of one of the special large specimens collected together with Ocypus (Pseudocypus) persiciculminis sp. nov. (2.0 mm). Scale in parentheses." httpUri="https://zenodo.org/record/1037691/files/figure.png" pageId="16" pageNumber="17">Fig. 3-2 E</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34A652B8829FFBB1C9AF960D886F88E" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BEF36A08829FFBB1C9AF960DB54F8C6" blockId="16.[151,1437,258,1939]" pageId="16" pageNumber="17">
<emphasis id="B924EAB28829FFBB1C9AF960DB36F91A" bold="true" box="[199,352,1734,1759]" pageId="16" pageNumber="17">Distribution.</emphasis>
<taxonomicName id="4C504D238829FFBB1D3BF961D831F91B" authority="He &amp; Zhou, 2017" authorityName="He &amp; Zhou" authorityYear="2017" box="[358,615,1734,1758]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="anguliculminis" status="sp. nov.">
<emphasis id="B924EAB28829FFBB1D3BF961D831F91B" box="[358,615,1734,1758]" italics="true" pageId="16" pageNumber="17">Ocypus anguliculminis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757C98829FFBB1E33F960D890F91A" box="[622,710,1734,1759]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B924EAB28829FFBB1E33F960D890F91A" bold="true" box="[622,710,1734,1759]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
is at present known only from Danba County in western
<collectingRegion id="4994F8428829FFBB191CF960DAA8F8C6" country="China" name="Sichuan" pageId="16" pageNumber="17">Sichuan Province</collectingRegion>
.
</paragraph>
<paragraph id="8BEF36A08829FFBB1C9AF8A8D886F88E" blockId="16.[151,1437,258,1939]" pageId="16" pageNumber="17">
<emphasis id="B924EAB28829FFBB1C9AF8A8DB47F8E2" bold="true" box="[199,273,1806,1831]" pageId="16" pageNumber="17">Notes.</emphasis>
<taxonomicName id="4C504D238829FFBB1D46F8A9D876F8E3" authority="He &amp; Zhou, 2017" authorityName="He &amp; Zhou" authorityYear="2017" box="[283,544,1806,1830]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="anguliculminis" status="sp. nov.">
<emphasis id="B924EAB28829FFBB1D46F8A9D876F8E3" box="[283,544,1806,1830]" italics="true" pageId="16" pageNumber="17">Ocypus anguliculminis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757C98829FFBB1E76F8A8D8DEF8E2" box="[555,648,1806,1831]" pageId="16" pageNumber="17" rank="species">
<emphasis id="B924EAB28829FFBB1E76F8A8D8DEF8E2" bold="true" box="[555,648,1806,1831]" pageId="16" pageNumber="17">sp. nov.</emphasis>
</taxonomicNameLabel>
can be easily distinguished from congeners by the aedeagus of the males and the special shaped tergite 10 of females.
</paragraph>
</subSubSection>
<subSubSection id="C34A652B8829FFBB1C9AF8F0D92EF856" pageId="16" pageNumber="17" type="etymology">
<paragraph id="8BEF36A08829FFBB1C9AF8F0D92EF856" blockId="16.[151,1437,258,1939]" pageId="16" pageNumber="17">
<emphasis id="B924EAB28829FFBB1C9AF8F0DB10F8AA" bold="true" box="[199,326,1878,1903]" pageId="16" pageNumber="17">Etymology</emphasis>
. The specific epithet,
<taxonomicName id="4C504D238829FFBB1E13F8F0D8A2F8AB" authorityName="He &amp; Zhou" authorityYear="2017" box="[590,756,1878,1902]" class="Insecta" family="Staphylinidae" genus="Ocypus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="anguliculminis">
<emphasis id="B924EAB28829FFBB1E13F8F0D8A2F8AB" box="[590,756,1878,1902]" italics="true" pageId="16" pageNumber="17">anguliculminis</emphasis>
</taxonomicName>
, is from Latin words &quot;
<emphasis id="B924EAB28829FFBB1858F8F0DE08F8AB" box="[1029,1118,1878,1902]" italics="true" pageId="16" pageNumber="17">angulus</emphasis>
&quot;(angle like, to indicate the special form of parameral apex) and &quot;
<emphasis id="B924EAB28829FFBB1E63F8DDD8D9F856" box="[574,655,1915,1939]" italics="true" pageId="16" pageNumber="17">culmen</emphasis>
&quot; (
<emphasis id="B924EAB28829FFBB1EF6F8DDD95CF856" box="[683,778,1915,1939]" italics="true" pageId="16" pageNumber="17">culminis</emphasis>
, tip, top).
</paragraph>
</subSubSection>
</treatment>
</document>
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