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<document id="6A8FDD5CDD0F4E032AF9DBFC1CD075A2" ID-CLB-Dataset="300009" ID-DOI="10.32225/bkpmnh.2021.50_55" ID-GBIF-Dataset="70224806-ca38-47ca-bd8a-63da3ac532f9" ID-ISSN="2189-6720" ID-Zenodo-Dep="12774639" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1721260925551" checkinUser="felipe" docAuthor="Watanabe, Kyohei" docDate="2021" docId="03F987931B64A736FF3C62BCFE9A76EA" docLanguage="en" docName="BullKanagawaPrefectMus.50.55-136.pdf" docOrigin="Bulletin of the Kanagawa Prefectural Museum (Natural Science) 50" docSource="https://doi.org/10.32225/bkpmnh.2021.50_55" docStyle="DocumentStyle:020B98DC1285EBBD02620724ECB44EDE.1:BullKanagawaPrefectMus.2013-.journal_article.english" docStyleId="020B98DC1285EBBD02620724ECB44EDE" docStyleName="BullKanagawaPrefectMus.2013-.journal_article.english" docStyleVersion="1" docTitle="Paraphylax yakushimensis Watanabe, 2021, sp. nov." docType="treatment" docUuid="98B819A5-A553-4E68-8A4F-4C442CB3A18F" docUuidSource="ZooBank" docVersion="9" lastPageNumber="89" masterDocId="FFC0FFEB1B45A714FFCA6633FFD77447" masterDocTitle="Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species" masterLastPageNumber="136" masterPageNumber="55" pageNumber="88" updateTime="1721329445643" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1AC0A9DD5CBAEE8FA9E2B25A0AC5BEC4">Taxonomic and Zoogeographic Study of the Japanese Phygadeuontinae (Hymenoptera, Ichneumonidae), with Descriptions of 17 New Species</mods:title>
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<mods:namePart id="1421893C7082A52D86B2CD0B8DE70277">Watanabe, Kyohei</mods:namePart>
<mods:affiliation id="0D57F7DF9118FB839E6603A62308EB30">Kanagawa Prefectural Museum of Natural History, 499 Iryuda, Odawara, Kanagawa 250 0031, Japan</mods:affiliation>
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<mods:title id="7DEF04AB89D175A2148C6C6455242BAE">Bulletin of the Kanagawa Prefectural Museum (Natural Science)</mods:title>
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<mods:date id="F05FF538256FB10F330412BDD238E1B3">2021</mods:date>
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<treatment id="03F987931B64A736FF3C62BCFE9A76EA" ID-DOI="http://doi.org/10.5281/zenodo.12773498" ID-GBIF-Taxon="235610851" ID-Zenodo-Dep="12773498" LSID="urn:lsid:zoobank.org:act:98B819A5-A553-4E68-8A4F-4C442CB3A18F" httpUri="http://treatment.plazi.org/id/03F987931B64A736FF3C62BCFE9A76EA" lastPageId="34" lastPageNumber="89" pageId="33" pageNumber="88">
<subSubSection id="C34A650E1B64A735FF3C62BCFD5570EF" box="[246,642,1167,1192]" pageId="33" pageNumber="88" type="nomenclature">
<paragraph id="8BEF36851B64A735FF3C62BCFD5570EF" blockId="33.[199,689,1167,1269]" box="[246,642,1167,1192]" pageId="33" pageNumber="88">
<heading id="D0A781E91B64A735FF3C62BCFD5570EF" box="[246,642,1167,1192]" centered="true" fontSize="10" level="2" pageId="33" pageNumber="88" reason="6">
<emphasis id="B924EA971B64A735FF3C62BCFD5570EF" bold="true" box="[246,642,1167,1192]" pageId="33" pageNumber="88">
<taxonomicName id="4C504D061B64A735FF3C62BCFDF370E0" authorityName="Watanabe" authorityYear="2021" box="[246,548,1167,1191]" class="Insecta" family="Ichneumonidae" genus="Paraphylax" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="88" phylum="Arthropoda" rank="species" species="yakushimensis" status="sp. nov.">
<emphasis id="B924EA971B64A735FF3C62BCFDF370E0" bold="true" box="[246,548,1167,1191]" italics="true" pageId="33" pageNumber="88">Paraphylax yakushimensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21757EC1B64A735FDE162A3FD5570EF" box="[555,642,1168,1192]" pageId="33" pageNumber="88" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B64A735FF0D6284FD667089" box="[199,689,1206,1231]" pageId="33" pageNumber="88" type="vernacular_names">
<paragraph id="8BEF36851B64A735FF0D6284FD667089" blockId="33.[199,689,1167,1269]" box="[199,689,1206,1231]" pageId="33" pageNumber="88">(New SJN: Yakushima-suji-togari-himebachi)</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B64A735FEF662EEFDEB70B2" box="[316,572,1245,1269]" pageId="33" pageNumber="88" type="description">
<paragraph id="8BEF36851B64A735FEF662EEFDEB70B2" blockId="33.[199,689,1167,1269]" box="[316,572,1245,1269]" pageId="33" pageNumber="88">
(
<figureCitation id="136B2A001B64A735FE8962EEFE1D70B2" box="[323,458,1245,1269]" captionStart="Fig" captionStartId="34.[190,224,1994,2016]" captionTargetBox="[189,1439,1237,1968]" captionTargetId="figure-402@34.[189,1439,1237,1968]" captionTargetPageId="34" captionText="Fig. 28. Paraphylax yakushimensis sp. nov., KPM-NK 55081, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesoscutum and scutellum, dorsal view; E, propodeum, dorsal view; F, fore wing; G, metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774699" httpUri="https://zenodo.org/record/12774699/files/figure.png" pageId="33" pageNumber="88">Figs 28AG</figureCitation>
,
<figureCitation id="136B2A001B64A735FE1F62EEFE2D70B2" box="[469,506,1245,1269]" captionStart="Fig" captionStartId="74.[191,224,1111,1133]" captionTargetBox="[193,1432,157,1080]" captionTargetId="figure-134@74.[189,1438,151,1085]" captionTargetPageId="74" captionText="Fig.65.Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Diaglyptidea conformis (Gmelin,1790) (B, KPM-NK 81650), Micraris ryukyuensis sp. nov. (C, KPM-NK 81873, holotype), Bathythrix kuwanae Viereck, 1912 (D, KPM-NK 81718), Bentyra ryukyuana sp. nov. (E, KPM-NK 81887, holotype), Chirotica matsukemushii (Matsumura, 1926) (F, KPM-NK 81906), Paraphylax politus sp. nov. (G, KPM-NK 55079, holotype), Pa. transstriatus sp. nov. (H, KPM-NK 55080, holotype), Pa. yakushimensis sp. nov. (I, KPM-NK 55081, holotype), Pa. yambarensis sp. nov. (J, KPM-NK 55085, holotype), Ethelurgus sodalis kumatai Kusigemati, 1983 (K, KPM- NK 81938), Aclastus etorofuensis (Uchida, 1936) (L, KPM-NK 81976), Hemiteles kuro sp. nov. (M, KPM-NK 55024, holotype), H. yamatonis sp. nov. (O, KPM-NK 55033, paratype), Isadelphus nigrus sp. nov. (P, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (Q, KPM-NK 81989, holotype), Mastrus molestae (Uchida, 1933) (R, KPM-NK 81987), Ma.takadai Momoi, 1970 (R, KPM-NK 81979), Gnotus chionops (Gravenhorst, 1829) (L, KPM-NK 82104), Gn. striatus (Uchida, 1930) (M, KPM-NK 82061), Orthizema semanotae Sheng &amp; Sun, 2014 (V, KPM-NK 82047), Nipponaetes haeussleri (Uchida, 1933) (W, KPM-NK 81832), and Surculus japonicus sp. nov. (X, KPM-NK 81861, holotype), females from Japan ― A-W, carinae around area superomedia of propodeum, dorsal view; X, T I, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774787" httpUri="https://zenodo.org/record/12774787/files/figure.png" pageId="33" pageNumber="88">65I</figureCitation>
,
<figureCitation id="136B2A001B64A735FDCC62EEFDE270B2" box="[518,565,1245,1269]" captionStart="Fig" captionStartId="75.[151,184,1580,1602]" captionTargetBox="[161,1381,163,1549]" captionTargetId="figure-5@75.[151,1398,151,1553]" captionTargetPageId="75" captionText="Fig. 66. Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Micraris ryukyuensis sp. nov. (B, KPM-NK 81873, holotype), Surculus japonicus sp. nov. (C, KPM-NK 81861, holotype), Bentyra ryukyuana sp. nov. (D, KPM-NK 81887, holotype), Paraphylax elegans sp. nov. (E, KPM-NK 55078, holotype), Pa. politus sp. nov. (F, KPM-NK 55079, holotype), Pa. transstriatus sp. nov. (G, KPM-NK 55080, holotype), Pa.yakushimensis sp. nov. (H, KPM-NK 55081, holotype), Pa. yambarensis sp. nov. (I, KPM-NK 55085, holotype), Hemiteles japonicus sp. nov. (J, KPM-NK 55021, holotype),H. kuro sp. nov. (K, KPM-NK 55024, holotype), H.maculipterus sp. nov. (L, KPM-NK 55025, holotype), H. yamatonis sp. nov. (M, KPM-NK 55032, holotype), Brachypimpla latipetiolator (Uchida, 1935) (N, O, KPM-NK 81973), Isadelphus nigrus sp. nov. (P, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (Q, KPM-NK 81989, holotype), Mastrus molestae (Uchida, 1933) (R, KPM-NK 81987), Megacara similis Sheng, 1999 (S, KPM-NK 81835), Tropistes shimizui sp. nov. (T, KPM-NK 81833, holotype), and Uchidella toichii sp. nov. (U, KPM-NK 82011, holotype), females from Japan ― A-N, P-U, apical part of ovipositor, lateral view (A, C-G, I-K, N, Q-U, left side; B, H, L, M, P, right side); O, apex of ovipositor sheath, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774789" httpUri="https://zenodo.org/record/12774789/files/figure.png" pageId="33" pageNumber="88">66H</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B64A735FF656318FE09712C" pageId="33" pageNumber="88" type="etymology">
<paragraph id="8BEF36851B64A735FF656318FE09712C" blockId="33.[151,738,1323,2087]" pageId="33" pageNumber="88">
<emphasis id="B924EA971B64A735FF656318FEFB7104" bold="true" box="[175,300,1323,1347]" pageId="33" pageNumber="88">Etymology</emphasis>
. The specific name is from Yakushima Island, where is locality of
<typeStatus id="54EB88271B64A735FE686360FE0D712C" box="[418,474,1363,1387]" pageId="33" pageNumber="88">types</typeStatus>
.
</paragraph>
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<subSubSection id="C34A650E1B64A735FF65634AFD35723C" pageId="33" pageNumber="88" type="materials_examined">
<paragraph id="8BEF36851B64A735FF65634AFD35723C" blockId="33.[151,738,1323,2087]" pageId="33" pageNumber="88">
<emphasis id="B924EA971B64A735FF65634AFEFA71D6" bold="true" box="[175,301,1401,1425]" pageId="33" pageNumber="88">Type series</emphasis>
.
<materialsCitation id="3B383CD81B64A735FEF7634AFDCB7198" ID-GBIF-Occurrence="4906826496" collectingDate="2007-11-02" collectingDateMax="2007-12-01" collectingDateMin="2007-11-02" collectorName="T. Yamauchi" country="Japan" location="Mt. Aiko-dake" pageId="33" pageNumber="88" specimenCode="KPM-NK 55081" specimenCount="1" stateProvince="Kagoshima" typeStatus="holotype">
<typeStatus id="54EB88271B64A735FEF7634AFE7471D6" box="[317,419,1401,1425]" pageId="33" pageNumber="88" type="holotype">Holotype</typeStatus>
:
<specimenCode id="DBF69EFE1B64A735FE646349FDB571D6" box="[430,610,1401,1425]" collectionCode="KPM-NK" pageId="33" pageNumber="88">KPM-NK 55081</specimenCode>
, F,
<collectingCountry id="F34776151B64A735FD41634AFD0A71D6" box="[651,733,1401,1425]" name="Japan" pageId="33" pageNumber="88">JAPAN</collectingCountry>
,
<collectingRegion id="4994F8671B64A735FF5D6393FE9171FF" box="[151,326,1440,1464]" country="Japan" name="Kagoshima" pageId="33" pageNumber="88">Kagoshima Pref.</collectingRegion>
, Yakushima Is.,
<location id="8E8F605E1B64A735FE276393FD5271FF" LSID="urn:lsid:plazi:treatment:03F987931B64A736FF3C62BCFE9A76EA:8E8F605E1B64A735FE276393FD5271FF" box="[493,645,1440,1464]" country="Japan" name="Mt. Aiko-dake" pageId="33" pageNumber="88" stateProvince="Kagoshima">Mt. Aiko-dake</location>
,
<date id="FFEE10451B64A735FD5A6393FECF7198" pageId="33" pageNumber="88" value="2007-11-02" valueMax="2007-12-01" valueMin="2007-11-02">
<collectingDate id="EFAAE9AD1B64A735FD5A6393FECF7198" pageId="33" pageNumber="88" value="2007-11-02" valueMax="2007-12-01" valueMin="2007-11-02">2. XI. 1. XII. 2007</collectingDate>
</date>
,
<collectorName id="26A553531B64A735FEE963F4FE737198" box="[291,420,1479,1503]" pageId="33" pageNumber="88">T. Yamauchi</collectorName>
leg. (MsT)
</materialsCitation>
.
<materialsCitation id="3B383CD81B64A735FDED63F4FF23726A" ID-GBIF-Occurrence="4906826506" collectingDate="2007-08-25" collectingDateMax="2007-09-22" collectingDateMin="2007-08-25" collectorName="Yakushima Is. &amp; T. Yamauchi" country="Japan" location="Mt. Aiko-dake" pageId="33" pageNumber="88" specimenCode="KPM-NK 55082" specimenCount="2" stateProvince="Kagoshima" typeStatus="paratype">
<typeStatus id="54EB88271B64A735FDED63F4FD547198" box="[551,643,1479,1503]" pageId="33" pageNumber="88" type="paratype">Paratype</typeStatus>
:
<collectingCountry id="F34776151B64A735FD4463F4FD097198" box="[654,734,1479,1503]" name="Japan" pageId="33" pageNumber="88">JAPAN</collectingCountry>
,
<specimenCode id="DBF69EFE1B64A735FF5D63DDFE927241" box="[151,325,1518,1542]" collectionCode="KPM-NK" pageId="33" pageNumber="88">KPM-NK 55082</specimenCode>
, F, same data of holotype,
<date id="FFEE10451B64A735FD9B63DDFF23726A" pageId="33" pageNumber="88" value="2007-08-25" valueMax="2007-09-22" valueMin="2007-08-25">
<collectingDate id="EFAAE9AD1B64A735FD9B63DDFF23726A" pageId="33" pageNumber="88" value="2007-08-25" valueMax="2007-09-22" valueMin="2007-08-25">25. VIII. 22. IX. 2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CD81B64A735FF346026FD09726A" ID-GBIF-Occurrence="4906826441" box="[254,734,1557,1581]" collectingDate="2007-09-28" collectingDateMax="2007-11-02" collectingDateMin="2007-09-28" collectorName="Yakushima Is. &amp; T. Yamauchi" country="Japan" location="Mt. Aiko-dake" pageId="33" pageNumber="88" specimenCode="KPM-NK 55083" specimenCount="1" stateProvince="Kagoshima" typeStatus="paratype">
<specimenCode id="DBF69EFE1B64A735FF346026FE7C726A" box="[254,427,1557,1581]" collectionCode="KPM-NK" pageId="33" pageNumber="88">KPM-NK 55083</specimenCode>
, F, ditto,
<date id="FFEE10451B64A735FDCD6026FD09726A" box="[519,734,1557,1581]" pageId="33" pageNumber="88" value="2007-09-28" valueMax="2007-11-02" valueMin="2007-09-28">
<collectingDate id="EFAAE9AD1B64A735FDCD6026FD09726A" box="[519,734,1557,1581]" pageId="33" pageNumber="88" value="2007-09-28" valueMax="2007-11-02" valueMin="2007-09-28">28. IX. 2. XI. 2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B383CD81B64A735FF5D600FFD35723C" ID-GBIF-Occurrence="4906826659" collectingDate="2006-08-25" collectingDateMax="2006-09-23" collectingDateMin="2006-08-25" collectorName="T. Yamauchi" country="Japan" location="Hanyama" pageId="33" pageNumber="88" specimenCode="KPM-NK 55084" specimenCount="1" stateProvince="Kagoshima" typeStatus="paratype">
<specimenCode id="DBF69EFE1B64A735FF5D600FFE877213" box="[151,336,1596,1620]" collectionCode="KPM-NK" pageId="33" pageNumber="88">KPM-NK 55084</specimenCode>
, F,
<collectingRegion id="4994F8671B64A735FEB0600FFDE37213" box="[378,564,1596,1620]" country="Japan" name="Kagoshima" pageId="33" pageNumber="88">Kagoshima Pref.</collectingRegion>
, Yakushima Is.,
<location id="8E8F605E1B64A735FF5D6050FF2E723C" LSID="urn:lsid:plazi:treatment:03F987931B64A736FF3C62BCFE9A76EA:8E8F605E1B64A735FF5D6050FF2E723C" box="[151,249,1635,1659]" country="Japan" name="Hanyama" pageId="33" pageNumber="88" stateProvince="Kagoshima">Hanyama</location>
,
<date id="FFEE10451B64A735FEC86051FE3D723C" box="[258,490,1634,1659]" pageId="33" pageNumber="88" value="2006-08-25" valueMax="2006-09-23" valueMin="2006-08-25">
<collectingDate id="EFAAE9AD1B64A735FEC86051FE3D723C" box="[258,490,1634,1659]" pageId="33" pageNumber="88" value="2006-08-25" valueMax="2006-09-23" valueMin="2006-08-25">25. VIII. 23. IX. 2006</collectingDate>
</date>
,
<collectorName id="26A553531B64A735FE386050FDB9723C" box="[498,622,1635,1659]" pageId="33" pageNumber="88">T. Yamauchi</collectorName>
leg. (MsT).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B64A736FF6560BAFE5775AC" lastPageId="34" lastPageNumber="89" pageId="33" pageNumber="88" type="description">
<paragraph id="8BEF36851B64A735FF6560BAFD47728F" blockId="33.[151,738,1323,2087]" pageId="33" pageNumber="88">
<emphasis id="B924EA971B64A735FF6560BAFEFB72E6" bold="true" box="[175,300,1673,1697]" pageId="33" pageNumber="88">Description</emphasis>
. Female (n=4). Body length 4.34.8 (HT: 4.8) mm. Body polished, covered with silver setae.
</paragraph>
<paragraph id="8BEF36851B64A735FF6560E4FADC7546" blockId="33.[151,738,1323,2087]" lastBlockId="33.[813,1400,155,2087]" pageId="33" pageNumber="88">
Head. Clypeus 1.92.0 (HT: 2.0) times as wide as maximum length, its anterior margin rounded. Face densely punctate, with a median weak convexity (
<figureCitation id="136B2A001B64A735FD7C6116FF127323" captionStart="Fig" captionStartId="34.[190,224,1994,2016]" captionTargetBox="[189,1439,1237,1968]" captionTargetId="figure-402@34.[189,1439,1237,1968]" captionTargetPageId="34" captionText="Fig. 28. Paraphylax yakushimensis sp. nov., KPM-NK 55081, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesoscutum and scutellum, dorsal view; E, propodeum, dorsal view; F, fore wing; G, metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774699" httpUri="https://zenodo.org/record/12774699/files/figure.png" pageId="33" pageNumber="88">Fig. 28B</figureCitation>
). MSL 1.0 times as long as BWM. Frons rugose, the rugae largely transverse. Gena punctate. Vertex punctate and largely transversely striate. OOL slightly shorter than POL. OOL and POL longer than OD. Occipital carina complete, its lower end joined with hypostomal carina distant from mandibular base. Upper tooth of mandible slightly longer than lower tooth. Base of mandible flat. Antenna with 2425 (HT: 24) flagellomeres. FL III 6.7 times as long as maximum depth in lateral view.
</paragraph>
<paragraph id="8BEF36851B64A735FC8F6723FC2070B2" blockId="33.[813,1400,155,2087]" pageId="33" pageNumber="88">
Mesosoma. Upper side of collar with a weak median carina. Lateral part of pronotum largely rugose except for a smooth area dorsally. Epomia short. Mesoscutum with a median longitudinal groove on the median lobe (
<figureCitation id="136B2A001B64A735FAC567B6FABB75DA" box="[1295,1388,389,413]" captionStart="Fig" captionStartId="34.[190,224,1994,2016]" captionTargetBox="[189,1439,1237,1968]" captionTargetId="figure-402@34.[189,1439,1237,1968]" captionTargetPageId="34" captionText="Fig. 28. Paraphylax yakushimensis sp. nov., KPM-NK 55081, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesoscutum and scutellum, dorsal view; E, propodeum, dorsal view; F, fore wing; G, metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774699" httpUri="https://zenodo.org/record/12774699/files/figure.png" pageId="33" pageNumber="88">Fig. 28D</figureCitation>
). Notaulus long, its posterior ends joined with each other posteriorly (
<figureCitation id="136B2A001B64A735FC6167E0FBDA75AC" box="[939,1037,467,491]" captionStart="Fig" captionStartId="34.[190,224,1994,2016]" captionTargetBox="[189,1439,1237,1968]" captionTargetId="figure-402@34.[189,1439,1237,1968]" captionTargetPageId="34" captionText="Fig. 28. Paraphylax yakushimensis sp. nov., KPM-NK 55081, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesoscutum and scutellum, dorsal view; E, propodeum, dorsal view; F, fore wing; G, metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774699" httpUri="https://zenodo.org/record/12774699/files/figure.png" pageId="33" pageNumber="88">Fig. 28D</figureCitation>
). Area along notaulus transversely striae (
<figureCitation id="136B2A001B64A735FCBB67C9FC187655" box="[881,975,506,530]" captionStart="Fig" captionStartId="34.[190,224,1994,2016]" captionTargetBox="[189,1439,1237,1968]" captionTargetId="figure-402@34.[189,1439,1237,1968]" captionTargetPageId="34" captionText="Fig. 28. Paraphylax yakushimensis sp. nov., KPM-NK 55081, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesoscutum and scutellum, dorsal view; E, propodeum, dorsal view; F, fore wing; G, metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774699" httpUri="https://zenodo.org/record/12774699/files/figure.png" pageId="33" pageNumber="88">Fig. 28D</figureCitation>
). Scutellum smooth except for lateral part partly rugulose, with a lateral longitudinal carina except for apex. Mesopleuron longitudinally striate except for smooth speculum and an anterior small smooth area. Posterior transverse carina of mesosternum complete. Metapleuron punctate, with a complete juxtacoxal carina. Anterior and posterior transverse carinae of propodeum complete (
<figureCitation id="136B2A001B64A735FA8464D0FC857765" captionStart="Fig" captionStartId="74.[191,224,1111,1133]" captionTargetBox="[193,1432,157,1080]" captionTargetId="figure-134@74.[189,1438,151,1085]" captionTargetPageId="74" captionText="Fig.65.Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Diaglyptidea conformis (Gmelin,1790) (B, KPM-NK 81650), Micraris ryukyuensis sp. nov. (C, KPM-NK 81873, holotype), Bathythrix kuwanae Viereck, 1912 (D, KPM-NK 81718), Bentyra ryukyuana sp. nov. (E, KPM-NK 81887, holotype), Chirotica matsukemushii (Matsumura, 1926) (F, KPM-NK 81906), Paraphylax politus sp. nov. (G, KPM-NK 55079, holotype), Pa. transstriatus sp. nov. (H, KPM-NK 55080, holotype), Pa. yakushimensis sp. nov. (I, KPM-NK 55081, holotype), Pa. yambarensis sp. nov. (J, KPM-NK 55085, holotype), Ethelurgus sodalis kumatai Kusigemati, 1983 (K, KPM- NK 81938), Aclastus etorofuensis (Uchida, 1936) (L, KPM-NK 81976), Hemiteles kuro sp. nov. (M, KPM-NK 55024, holotype), H. yamatonis sp. nov. (O, KPM-NK 55033, paratype), Isadelphus nigrus sp. nov. (P, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (Q, KPM-NK 81989, holotype), Mastrus molestae (Uchida, 1933) (R, KPM-NK 81987), Ma.takadai Momoi, 1970 (R, KPM-NK 81979), Gnotus chionops (Gravenhorst, 1829) (L, KPM-NK 82104), Gn. striatus (Uchida, 1930) (M, KPM-NK 82061), Orthizema semanotae Sheng &amp; Sun, 2014 (V, KPM-NK 82047), Nipponaetes haeussleri (Uchida, 1933) (W, KPM-NK 81832), and Surculus japonicus sp. nov. (X, KPM-NK 81861, holotype), females from Japan ― A-W, carinae around area superomedia of propodeum, dorsal view; X, T I, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774787" httpUri="https://zenodo.org/record/12774787/files/figure.png" pageId="33" pageNumber="88">Fig. 65I</figureCitation>
). Pleural carina complete. Lateral longitudinal carina of propodeum absent. Area basalis distinct (
<figureCitation id="136B2A001B64A735FB286502FAE2770E" box="[1250,1333,817,841]" captionStart="Fig" captionStartId="74.[191,224,1111,1133]" captionTargetBox="[193,1432,157,1080]" captionTargetId="figure-134@74.[189,1438,151,1085]" captionTargetPageId="74" captionText="Fig.65.Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Diaglyptidea conformis (Gmelin,1790) (B, KPM-NK 81650), Micraris ryukyuensis sp. nov. (C, KPM-NK 81873, holotype), Bathythrix kuwanae Viereck, 1912 (D, KPM-NK 81718), Bentyra ryukyuana sp. nov. (E, KPM-NK 81887, holotype), Chirotica matsukemushii (Matsumura, 1926) (F, KPM-NK 81906), Paraphylax politus sp. nov. (G, KPM-NK 55079, holotype), Pa. transstriatus sp. nov. (H, KPM-NK 55080, holotype), Pa. yakushimensis sp. nov. (I, KPM-NK 55081, holotype), Pa. yambarensis sp. nov. (J, KPM-NK 55085, holotype), Ethelurgus sodalis kumatai Kusigemati, 1983 (K, KPM- NK 81938), Aclastus etorofuensis (Uchida, 1936) (L, KPM-NK 81976), Hemiteles kuro sp. nov. (M, KPM-NK 55024, holotype), H. yamatonis sp. nov. (O, KPM-NK 55033, paratype), Isadelphus nigrus sp. nov. (P, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (Q, KPM-NK 81989, holotype), Mastrus molestae (Uchida, 1933) (R, KPM-NK 81987), Ma.takadai Momoi, 1970 (R, KPM-NK 81979), Gnotus chionops (Gravenhorst, 1829) (L, KPM-NK 82104), Gn. striatus (Uchida, 1930) (M, KPM-NK 82061), Orthizema semanotae Sheng &amp; Sun, 2014 (V, KPM-NK 82047), Nipponaetes haeussleri (Uchida, 1933) (W, KPM-NK 81832), and Surculus japonicus sp. nov. (X, KPM-NK 81861, holotype), females from Japan ― A-W, carinae around area superomedia of propodeum, dorsal view; X, T I, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774787" httpUri="https://zenodo.org/record/12774787/files/figure.png" pageId="33" pageNumber="88">Fig. 65I</figureCitation>
). Area superomedia indistinct laterally (
<figureCitation id="136B2A001B64A735FBB7656BFB097737" box="[1149,1246,856,880]" captionStart="Fig" captionStartId="34.[190,224,1994,2016]" captionTargetBox="[189,1439,1237,1968]" captionTargetId="figure-402@34.[189,1439,1237,1968]" captionTargetPageId="34" captionText="Fig. 28. Paraphylax yakushimensis sp. nov., KPM-NK 55081, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesoscutum and scutellum, dorsal view; E, propodeum, dorsal view; F, fore wing; G, metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774699" httpUri="https://zenodo.org/record/12774699/files/figure.png" pageId="33" pageNumber="88">Figs 28E</figureCitation>
,
<figureCitation id="136B2A001B64A735FB23656BFAD97737" box="[1257,1294,856,880]" captionStart="Fig" captionStartId="74.[191,224,1111,1133]" captionTargetBox="[193,1432,157,1080]" captionTargetId="figure-134@74.[189,1438,151,1085]" captionTargetPageId="74" captionText="Fig.65.Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Diaglyptidea conformis (Gmelin,1790) (B, KPM-NK 81650), Micraris ryukyuensis sp. nov. (C, KPM-NK 81873, holotype), Bathythrix kuwanae Viereck, 1912 (D, KPM-NK 81718), Bentyra ryukyuana sp. nov. (E, KPM-NK 81887, holotype), Chirotica matsukemushii (Matsumura, 1926) (F, KPM-NK 81906), Paraphylax politus sp. nov. (G, KPM-NK 55079, holotype), Pa. transstriatus sp. nov. (H, KPM-NK 55080, holotype), Pa. yakushimensis sp. nov. (I, KPM-NK 55081, holotype), Pa. yambarensis sp. nov. (J, KPM-NK 55085, holotype), Ethelurgus sodalis kumatai Kusigemati, 1983 (K, KPM- NK 81938), Aclastus etorofuensis (Uchida, 1936) (L, KPM-NK 81976), Hemiteles kuro sp. nov. (M, KPM-NK 55024, holotype), H. yamatonis sp. nov. (O, KPM-NK 55033, paratype), Isadelphus nigrus sp. nov. (P, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (Q, KPM-NK 81989, holotype), Mastrus molestae (Uchida, 1933) (R, KPM-NK 81987), Ma.takadai Momoi, 1970 (R, KPM-NK 81979), Gnotus chionops (Gravenhorst, 1829) (L, KPM-NK 82104), Gn. striatus (Uchida, 1930) (M, KPM-NK 82061), Orthizema semanotae Sheng &amp; Sun, 2014 (V, KPM-NK 82047), Nipponaetes haeussleri (Uchida, 1933) (W, KPM-NK 81832), and Surculus japonicus sp. nov. (X, KPM-NK 81861, holotype), females from Japan ― A-W, carinae around area superomedia of propodeum, dorsal view; X, T I, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774787" httpUri="https://zenodo.org/record/12774787/files/figure.png" pageId="33" pageNumber="88">65I</figureCitation>
). Anterior and posterior parts of propodeum transversely, obliquely, and irregularly rugulose. Median part of propodeum longitudinally rugulose (
<figureCitation id="136B2A001B64A735FBFB65FEFB4377A2" box="[1073,1172,973,997]" captionStart="Fig" captionStartId="34.[190,224,1994,2016]" captionTargetBox="[189,1439,1237,1968]" captionTargetId="figure-402@34.[189,1439,1237,1968]" captionTargetPageId="34" captionText="Fig. 28. Paraphylax yakushimensis sp. nov., KPM-NK 55081, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesoscutum and scutellum, dorsal view; E, propodeum, dorsal view; F, fore wing; G, metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774699" httpUri="https://zenodo.org/record/12774699/files/figure.png" pageId="33" pageNumber="88">Figs 28E</figureCitation>
,
<figureCitation id="136B2A001B64A735FB6B65FEFB1077A2" box="[1185,1223,973,997]" captionStart="Fig" captionStartId="74.[191,224,1111,1133]" captionTargetBox="[193,1432,157,1080]" captionTargetId="figure-134@74.[189,1438,151,1085]" captionTargetPageId="74" captionText="Fig.65.Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Diaglyptidea conformis (Gmelin,1790) (B, KPM-NK 81650), Micraris ryukyuensis sp. nov. (C, KPM-NK 81873, holotype), Bathythrix kuwanae Viereck, 1912 (D, KPM-NK 81718), Bentyra ryukyuana sp. nov. (E, KPM-NK 81887, holotype), Chirotica matsukemushii (Matsumura, 1926) (F, KPM-NK 81906), Paraphylax politus sp. nov. (G, KPM-NK 55079, holotype), Pa. transstriatus sp. nov. (H, KPM-NK 55080, holotype), Pa. yakushimensis sp. nov. (I, KPM-NK 55081, holotype), Pa. yambarensis sp. nov. (J, KPM-NK 55085, holotype), Ethelurgus sodalis kumatai Kusigemati, 1983 (K, KPM- NK 81938), Aclastus etorofuensis (Uchida, 1936) (L, KPM-NK 81976), Hemiteles kuro sp. nov. (M, KPM-NK 55024, holotype), H. yamatonis sp. nov. (O, KPM-NK 55033, paratype), Isadelphus nigrus sp. nov. (P, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (Q, KPM-NK 81989, holotype), Mastrus molestae (Uchida, 1933) (R, KPM-NK 81987), Ma.takadai Momoi, 1970 (R, KPM-NK 81979), Gnotus chionops (Gravenhorst, 1829) (L, KPM-NK 82104), Gn. striatus (Uchida, 1930) (M, KPM-NK 82061), Orthizema semanotae Sheng &amp; Sun, 2014 (V, KPM-NK 82047), Nipponaetes haeussleri (Uchida, 1933) (W, KPM-NK 81832), and Surculus japonicus sp. nov. (X, KPM-NK 81861, holotype), females from Japan ― A-W, carinae around area superomedia of propodeum, dorsal view; X, T I, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774787" httpUri="https://zenodo.org/record/12774787/files/figure.png" pageId="33" pageNumber="88">65I</figureCitation>
). Length of fore wing 3.63.9 (HT: 3.9) mm. Areolet absent (
<figureCitation id="136B2A001B64A735FAC665C7FABC704B" box="[1292,1387,1012,1036]" captionStart="Fig" captionStartId="34.[190,224,1994,2016]" captionTargetBox="[189,1439,1237,1968]" captionTargetId="figure-402@34.[189,1439,1237,1968]" captionTargetPageId="34" captionText="Fig. 28. Paraphylax yakushimensis sp. nov., KPM-NK 55081, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesoscutum and scutellum, dorsal view; E, propodeum, dorsal view; F, fore wing; G, metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774699" httpUri="https://zenodo.org/record/12774699/files/figure.png" pageId="33" pageNumber="88">Fig. 28F</figureCitation>
). Vein 2m-cu of fore wing with two bullae (
<figureCitation id="136B2A001B64A735FACD6228FABD7074" box="[1287,1386,1051,1075]" captionStart="Fig" captionStartId="34.[190,224,1994,2016]" captionTargetBox="[189,1439,1237,1968]" captionTargetId="figure-402@34.[189,1439,1237,1968]" captionTargetPageId="34" captionText="Fig. 28. Paraphylax yakushimensis sp. nov., KPM-NK 55081, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesoscutum and scutellum, dorsal view; E, propodeum, dorsal view; F, fore wing; G, metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774699" httpUri="https://zenodo.org/record/12774699/files/figure.png" pageId="33" pageNumber="88">Fig. 28F</figureCitation>
). Nervellus inclivous, intercepted slightly behind middle. Hind femur 5.15.2 (HT: 5.2) times as long as maximum depth in lateral view. Hind TS I: II: III: IV: V = 2.0: 0.90.95 (HT: 0.9): 0.650.7 (HT: 0.7): 0.30.4 (HT: 0.4): 0.4. Tarsal claws simple.
</paragraph>
<paragraph id="8BEF36851B64A735FC8F6336FAF7723C" blockId="33.[813,1400,155,2087]" pageId="33" pageNumber="88">
Metasoma. T I 1.7 times as long as maximum width, longitudinally striate except for smooth posterior margin (
<figureCitation id="136B2A001B64A735FCFE6361FC17712D" box="[820,960,1362,1386]" captionStart="Fig" captionStartId="34.[190,224,1994,2016]" captionTargetBox="[189,1439,1237,1968]" captionTargetId="figure-402@34.[189,1439,1237,1968]" captionTargetPageId="34" captionText="Fig. 28. Paraphylax yakushimensis sp. nov., KPM-NK 55081, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesoscutum and scutellum, dorsal view; E, propodeum, dorsal view; F, fore wing; G, metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774699" httpUri="https://zenodo.org/record/12774699/files/figure.png" pageId="33" pageNumber="88">Figs 28C, G</figureCitation>
). Median dorsal carina of T I indistinct. Dorsolateral carina of T I complete. T II and T III covered with longitudinal striae except for smooth posterior area on posterior 0.4 (
<figureCitation id="136B2A001B64A735FC2A63F4FBB27198" box="[992,1125,1479,1503]" captionStart="Fig" captionStartId="34.[190,224,1994,2016]" captionTargetBox="[189,1439,1237,1968]" captionTargetId="figure-402@34.[189,1439,1237,1968]" captionTargetPageId="34" captionText="Fig. 28. Paraphylax yakushimensis sp. nov., KPM-NK 55081, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesoscutum and scutellum, dorsal view; E, propodeum, dorsal view; F, fore wing; G, metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774699" httpUri="https://zenodo.org/record/12774699/files/figure.png" pageId="33" pageNumber="88">Figs 28C, G</figureCitation>
). T IV finely and sparsely punctate, with some weak longitudinal striae medially. T V and T VI largely smooth medially. Ovipositor sheath 0.70.75 (HT: 0.75) times as long as hind tibia. Ovipositor straight, with a nodus and ventral teeth (
<figureCitation id="136B2A001B64A735FB726050FAC2723C" box="[1208,1301,1635,1659]" captionStart="Fig" captionStartId="75.[151,184,1580,1602]" captionTargetBox="[161,1381,163,1549]" captionTargetId="figure-5@75.[151,1398,151,1553]" captionTargetPageId="75" captionText="Fig. 66. Acrolyta japonica sp. nov. (A, KPM-NK 54996, holotype), Micraris ryukyuensis sp. nov. (B, KPM-NK 81873, holotype), Surculus japonicus sp. nov. (C, KPM-NK 81861, holotype), Bentyra ryukyuana sp. nov. (D, KPM-NK 81887, holotype), Paraphylax elegans sp. nov. (E, KPM-NK 55078, holotype), Pa. politus sp. nov. (F, KPM-NK 55079, holotype), Pa. transstriatus sp. nov. (G, KPM-NK 55080, holotype), Pa.yakushimensis sp. nov. (H, KPM-NK 55081, holotype), Pa. yambarensis sp. nov. (I, KPM-NK 55085, holotype), Hemiteles japonicus sp. nov. (J, KPM-NK 55021, holotype),H. kuro sp. nov. (K, KPM-NK 55024, holotype), H.maculipterus sp. nov. (L, KPM-NK 55025, holotype), H. yamatonis sp. nov. (M, KPM-NK 55032, holotype), Brachypimpla latipetiolator (Uchida, 1935) (N, O, KPM-NK 81973), Isadelphus nigrus sp. nov. (P, KPM-NK 82003, holotype), Lochetica japonica sp. nov. (Q, KPM-NK 81989, holotype), Mastrus molestae (Uchida, 1933) (R, KPM-NK 81987), Megacara similis Sheng, 1999 (S, KPM-NK 81835), Tropistes shimizui sp. nov. (T, KPM-NK 81833, holotype), and Uchidella toichii sp. nov. (U, KPM-NK 82011, holotype), females from Japan ― A-N, P-U, apical part of ovipositor, lateral view (A, C-G, I-K, N, Q-U, left side; B, H, L, M, P, right side); O, apex of ovipositor sheath, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12774789" httpUri="https://zenodo.org/record/12774789/files/figure.png" pageId="33" pageNumber="88">Fig. 66H</figureCitation>
).
</paragraph>
<paragraph id="8BEF36851B64A736FC8F60B9FD637583" blockId="33.[813,1400,155,2087]" lastBlockId="34.[189,775,155,685]" lastPageId="34" lastPageNumber="89" pageId="33" pageNumber="88">
Coloration (
<figureCitation id="136B2A001B64A735FC0A60B9FB9272E5" box="[960,1093,1674,1698]" captionStart="Fig" captionStartId="34.[190,224,1994,2016]" captionTargetBox="[189,1439,1237,1968]" captionTargetId="figure-402@34.[189,1439,1237,1968]" captionTargetPageId="34" captionText="Fig. 28. Paraphylax yakushimensis sp. nov., KPM-NK 55081, holotype, female ― A, lateral habitus; B, head, frontal view; C, head, mesosoma, and metasoma, dorsal view; D, mesoscutum and scutellum, dorsal view; E, propodeum, dorsal view; F, fore wing; G, metasoma, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12774699" httpUri="https://zenodo.org/record/12774699/files/figure.png" pageId="33" pageNumber="88">Figs 28AG</figureCitation>
). Body (excluding wings and legs) black to blackish brown. Basal parts of antenna yellowish brown to reddish brown. Median parts of face and clypeus tinged with yellowish brown to reddish brown. Mandible yellow except for darkened apex. Palpi and tegula yellowish brown to yellow. Margins of pronotum, mesoscutum, and mesopleuron partly tinged with reddish brown. Scutellum and postscutellum dark reddish brown. T I reddish brown, its lateral sides more or less darkened. T II and T III reddish brown. T IV dark reddish brown anteriorly, reddish brown posteriorly. T V to T VII dark reddish brown except for reddish brown posterior margin. Membranous part of metasomal sternites whitish yellow. Ovipositor reddish brown. Wings hyaline, with a longitudinal clouded band just behind pterostigma and a small, clouded area subbasally. Veins yellowish brown to blackish brown. Pterostigma blackish brown. Legs reddish brown. Base of tibiae white. T I and T IV to T VII sometimes entirely tinged with reddish brown.
</paragraph>
<paragraph id="8BEF36851B67A736FF1F67E0FE5775AC" blockId="34.[189,775,155,685]" box="[213,384,467,491]" pageId="34" pageNumber="89">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B67A736FF1F67CAFD927655" box="[213,581,505,530]" pageId="34" pageNumber="89" type="distribution">
<paragraph id="8BEF36851B67A736FF1F67CAFD927655" blockId="34.[189,775,155,685]" box="[213,581,505,530]" pageId="34" pageNumber="89">
<emphasis id="B924EA971B67A736FF1F67CAFE807656" bold="true" box="[213,343,505,529]" pageId="34" pageNumber="89">Distribution</emphasis>
.
<collectingCountry id="F34776151B67A736FEA967C9FE4A7656" box="[355,413,506,529]" name="Japan" pageId="34" pageNumber="89">Japan</collectingCountry>
(Yakushima Is.).
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B67A736FF1F6413FE6C767F" box="[213,443,544,568]" pageId="34" pageNumber="89" type="biology_ecology">
<paragraph id="8BEF36851B67A736FF1F6413FE6C767F" blockId="34.[189,775,155,685]" box="[213,443,544,568]" pageId="34" pageNumber="89">
<emphasis id="B924EA971B67A736FF1F6413FE93767F" bold="true" box="[213,324,544,568]" pageId="34" pageNumber="89">Bionomics</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C34A650E1B67A736FF1F647BFE9A76EA" pageId="34" pageNumber="89" type="discussion">
<paragraph id="8BEF36851B67A736FF1F647BFE9A76EA" blockId="34.[189,775,155,685]" pageId="34" pageNumber="89">
<emphasis id="B924EA971B67A736FF1F647BFEE07627" bold="true" box="[213,311,584,608]" pageId="34" pageNumber="89">Remarks</emphasis>
. This species resembles
<taxonomicName id="4C504D061B67A736FDF8647BFF2776C1" authority="Momoi, 1966" authorityName="Momoi" authorityYear="1966" class="Insecta" family="Ichneumonidae" genus="Parallocota" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="89" phylum="Arthropoda" rank="species" species="saigusai">
<emphasis id="B924EA971B67A736FDF8647BFD787618" box="[562,687,584,607]" italics="true" pageId="34" pageNumber="89">Pa. saigusai</emphasis>
Momoi,
<quantity id="4CA89B601B67A736FF77645DFF2776C1" box="[189,240,622,646]" metricMagnitude="1" metricUnit="m" metricValue="4.99364" pageId="34" pageNumber="89" unit="in" value="1966.0">1966</quantity>
</taxonomicName>
in the body coloration but it can be distinguished by the above key.
</paragraph>
</subSubSection>
</treatment>
</document>