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<document id="85B0C80734F0C1F65F463BEF1CFB0DED" ID-DOI="10.1111/j.1096-3642.2009.00526.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443023" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630454731098" checkinUser="felipe" docAuthor="Liew, Thor-Seng, Schilthuizen, Menno &amp; Vermeulen, Jaap Jan" docDate="2009" docId="03F9878AFF99E8140CC988FA22A1FEFD" docLanguage="en" docName="j.1096-3642.2009.00526.x.pdf" docOrigin="Zoological Journal of the Linnean Society 157 (3)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Everettia paulbasintali Liew &amp; Schilthuizen &amp; Vermeulen 2009, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="537" masterDocId="FFC0FFF2FF8FE8030C238C132365FFCA" masterDocTitle="Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo" masterLastPageNumber="550" masterPageNumber="515" pageNumber="537" updateTime="1699276879878" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="5451E4273FB3E3985E4A1FF4B131BEAA">Systematic revision of the genus Everettia Godwin-Austen, 1891 (Mollusca: Gastropoda: Dyakiidae) in Sabah, northern Borneo</mods:title>
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<mods:namePart id="073297C406D832255CD3F983C6E5CD8F">Liew, Thor-Seng</mods:namePart>
</mods:name>
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<mods:namePart id="98DDD0E02000B08EEB8CA228A423A2C3">Schilthuizen, Menno</mods:namePart>
</mods:name>
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<mods:namePart id="B7FF5D9F91EBF623931F3396C158C2BB">Vermeulen, Jaap Jan</mods:namePart>
</mods:name>
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<mods:title id="86DFEAAC3B8E872816E801AF1F6DFD6E">Zoological Journal of the Linnean Society</mods:title>
</mods:titleInfo>
<mods:part id="34553BF03AD0F10036F9E0F3157502B3">
<mods:date id="B01BA51DA278F7D4E308F642160B0A5D">2009</mods:date>
<mods:detail id="93C46578FAD3492B64A31B66A04C3675" type="pubDate">
<mods:number id="591D1ACD3CA186FE0CD951C36771D923">2009-10-26</mods:number>
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<mods:detail id="D02DAE943FBC834CD8F2FE4F3A793CA6" type="volume">
<mods:number id="A64AC27B60EF9C8B51A435D79284EC0B">157</mods:number>
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<mods:number id="7A7573820F38C74F0E5736C5CB522A35">3</mods:number>
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<mods:url id="1C0E0B3142B68E0529126CF45ED8F16E">https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2009.00526.x</mods:url>
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<mods:classification id="0C287F6F6C8CEAF4DF49312CCB1C8728">journal article</mods:classification>
<mods:identifier id="63129B8F3F013D1560F72C4D237950E6" type="DOI">10.1111/j.1096-3642.2009.00526.x</mods:identifier>
<mods:identifier id="997880D745649D73FF333BFAE5E1CE19" type="ISSN">0024-4082</mods:identifier>
<mods:identifier id="55DC5F1ECB6662CDE48F3CB004AD2141" type="Zenodo-Dep">5443023</mods:identifier>
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<treatment id="03F9878AFF99E8140CC988FA22A1FEFD" LSID="urn:lsid:plazi:treatment:03F9878AFF99E8140CC988FA22A1FEFD" httpUri="http://treatment.plazi.org/id/03F9878AFF99E8140CC988FA22A1FEFD" lastPageId="23" lastPageNumber="537" pageId="22" pageNumber="537">
<subSubSection id="C34A6517FF99E8150CC988FA21E5FA6A" pageId="22" pageNumber="537" type="nomenclature">
<paragraph id="8BEF369CFF99E8150CC988FA21F9FB34" blockId="22.[234,668,1257,1278]" box="[234,668,1257,1278]" pageId="22" pageNumber="537">
<heading id="D0A781F0FF99E8150CC988FA21F9FB34" box="[234,668,1257,1278]" centered="true" fontSize="9" level="2" pageId="22" pageNumber="537" reason="2">
<taxonomicName id="4C504D1FFF99E8150CC988FA2148FB34" ID-CoL="6HM7S" authority="Liew &amp; Schilthuizen &amp; Vermeulen, 2009" authorityName="Liew &amp; Schilthuizen &amp; Vermeulen" authorityYear="2009" box="[234,557,1257,1278]" class="Gastropoda" family="Dyakiidae" genus="Everettia" kingdom="Animalia" order="Stylommatophora" pageId="22" pageNumber="537" phylum="Mollusca" rank="species" species="paulbasintali" status="sp. nov.">EVERETTIA PAULBASINTALI</taxonomicName>
<emphasis id="B924EA8EFF99E8150E1488FE21F9FB34" bold="true" box="[567,668,1257,1278]" pageId="22" pageNumber="537">
<taxonomicNameLabel id="A21757F5FF99E8150E1488FE21F9FB34" box="[567,668,1257,1278]" pageId="22" pageNumber="537" rank="species">SP. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BEF369CFF99E8150CB3890321E5FA6A" blockId="22.[144,759,1296,1440]" pageId="22" pageNumber="537">
<emphasis id="B924EA8EFF99E8150CB3890323B0FAEF" box="[144,213,1296,1317]" italics="true" pageId="22" pageNumber="537">Types:</emphasis>
<materialsCitation id="3B383CC1FF99E8150CC189032119FA6A" collectingDate="2007-08" collectionCode="T" collectorName="S. Liew &amp; Johny Lapidin. Date" country="Malaysia" county="State" latitude="4.45" location="Tawau Hills Park" longLatPrecision="1306" longitude="117.916664" municipality="Tawau District" pageId="22" pageNumber="537" specimenCode="BMNH 20080628" specimenCount="1" stateProvince="Sabah" typeStatus="holotype">
<collectingCountry id="F347760CFF99E8150CC18903220EFAEF" box="[226,363,1296,1317]" name="Malaysia" pageId="22" pageNumber="537">MALAYSIA</collectingCountry>
.
<collectingCounty id="628E4E10FF99E8150D58890322DDFAEF" box="[379,440,1296,1317]" pageId="22" pageNumber="537">State</collectingCounty>
of
<collectingRegion id="4994F87EFF99E8150DC689032155FAEC" box="[485,560,1296,1318]" country="Malaysia" name="Sabah" pageId="22" pageNumber="537">Sabah</collectingRegion>
.
<collectingMunicipality id="6B8BACE6FF99E8150E6389032196FAEC" box="[576,755,1296,1318]" pageId="22" pageNumber="537">Tawau District</collectingMunicipality>
,
<location id="8E8F6047FF99E8150CB3893C2201FA8E" LSID="urn:lsid:plazi:treatment:03F9878AFF99E8140CC988FA22A1FEFD:8E8F6047FF99E8150CB3893C2201FA8E" box="[144,356,1326,1348]" country="Malaysia" county="State" latitude="4.45" longLatPrecision="1306" longitude="117.916664" municipality="Tawau District" name="Tawau Hills Park" pageId="22" pageNumber="537" stateProvince="Sabah">Tawau Hills Park</location>
(
<geoCoordinate id="EE64505BFF99E8150D5B893D22B0FA8E" box="[376,469,1325,1348]" degrees="04" direction="north" minutes="27" orientation="latitude" pageId="22" pageNumber="537" precision="925" value="4.45">
04°27
<emphasis id="B924EA8EFF99E8150D9A893E22DAFA8E" box="[441,447,1325,1348]" italics="true" pageId="22" pageNumber="537">′</emphasis>
N
</geoCoordinate>
,
<geoCoordinate id="EE64505BFF99E8150DC5893D2128FA8E" box="[486,589,1325,1348]" degrees="117" direction="east" minutes="55" orientation="longitude" pageId="22" pageNumber="537" precision="925" value="117.916664">
117°55
<emphasis id="B924EA8EFF99E8150E10893E215CFA8E" box="[563,569,1325,1348]" italics="true" pageId="22" pageNumber="537">′</emphasis>
E
</geoCoordinate>
). Collected by T.-
<collectorName id="26A5534AFF99E8150C93895E2263FAA9" box="[176,262,1357,1379]" pageId="22" pageNumber="537">S. Liew</collectorName>
and
<collectorName id="26A5534AFF99E8150D64895E2159FAA8" box="[327,572,1357,1379]" pageId="22" pageNumber="537">Johny Lapidin. Date</collectorName>
:
<date id="FFEE105CFF99E8150E68895E21D5FAA9" box="[587,688,1357,1379]" pageId="22" pageNumber="537" value="2007-08">
<collectingDate id="EFAAE9B4FF99E8150E68895E21D5FAA9" box="[587,688,1357,1379]" pageId="22" pageNumber="537" value="2007-08">viii.2007</collectingDate>
</date>
,
<typeStatus id="54EB883EFF99E8150E9C895E23A1FA4B" pageId="22" pageNumber="537" type="holotype">holotype</typeStatus>
, BOR/MOL 5498,
<figureCitation id="136B2A19FF99E8150DB68978216FFA4B" box="[405,522,1387,1409]" captionStart="Figure 6" captionStartId="12.[144,223,1616,1635]" captionTargetBox="[306,1260,198,1584]" captionTargetId="figure-74@12.[303,1263,195,1586]" captionTargetPageId="12" captionText="Figure 6. Shells. A, Everettia lapidini sp. nov.; B, Everettia klemmantanica; C, Everettia subconsul; D, Everettia paulbasintali sp. nov.; E, Everettia themis; F, Everettia layanglayang sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443035" httpUri="https://zenodo.org/record/5443035/files/figure.png" pageId="22" pageNumber="537">Figure 6D</figureCitation>
;
<specimenCount id="9D56FD15FF99E8150E34897821A7FA4B" box="[535,706,1387,1409]" pageId="22" pageNumber="537" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
, two in
<specimenCode id="DBF69EE7FF99E8150C938998221CFA6A" box="[176,377,1418,1440]" collectionCode="BMNH" pageId="22" pageNumber="537">BMNH 20080628</specimenCode>
and two in SP 13060
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34A6517FF99E8140CB389DA22A1FEFD" lastPageId="23" lastPageNumber="538" pageId="22" pageNumber="537" type="etymology">
<paragraph id="8BEF369CFF99E8150CB389DA22F6F9D6" blockId="22.[144,758,1481,1564]" pageId="22" pageNumber="537">
<emphasis id="B924EA8EFF99E8150CB389DA2268FA14" box="[144,269,1481,1502]" italics="true" pageId="22" pageNumber="537">Etymology:</emphasis>
This species is named after Paul Basintal, director of
<collectingRegion id="4994F87EFF99E8150D3289FB223CFA34" box="[273,345,1512,1534]" country="Malaysia" name="Sabah" pageId="22" pageNumber="537">Sabah</collectingRegion>
Parks, who has served for over 33 years in
<collectingRegion id="4994F87EFF99E8150CDB8A152225F9D6" box="[248,320,1542,1564]" country="Malaysia" name="Sabah" pageId="22" pageNumber="537">Sabah</collectingRegion>
Parks.
</paragraph>
<paragraph id="8BEF369CFF99E8150CB38A562193F8A4" blockId="22.[144,759,1605,1903]" pageId="22" pageNumber="537">
<emphasis id="B924EA8EFF99E8150CB38A562208F990" box="[144,365,1605,1626]" italics="true" pageId="22" pageNumber="537">Material examined:</emphasis>
<collectingRegion id="4994F87EFF99E8150D568A5522ABF991" box="[373,462,1606,1627]" country="Malaysia" name="Sabah" pageId="22" pageNumber="537">SABAH</collectingRegion>
– Tabin Wildlife Reserve: BOR/MOL 924, Head Quarters,
<date id="FFEE105CFF99E8150E368A772108F9B0" box="[533,621,1636,1658]" pageId="22" pageNumber="537" value="2000-03">iii.2000</date>
; BOR/MOL 926, Base Camp
<date id="FFEE105CFF99E8150D728A9022FAF952" box="[337,415,1667,1688]" pageId="22" pageNumber="537" value="2000-10">x.2000</date>
; BOR/MOL 962,
<date id="FFEE105CFF99E8150E418A9021C8F952" box="[610,685,1667,1688]" pageId="22" pageNumber="537" value="2001-05">v.2001</date>
; BOR/ MOL 964,
<date id="FFEE105CFF99E8150D318AB22239F97C" box="[274,348,1697,1718]" pageId="22" pageNumber="537" value="2001-05">v.2001</date>
; BOR/MOL 928, Tabin Limestone,
<date id="FFEE105CFF99E8150CB38AD323B8F91F" box="[144,221,1728,1749]" pageId="22" pageNumber="537" value="2000-10">x.2000</date>
. Danum Valley: BOR/MOL 5271,
<date id="FFEE105CFF99E8150E418AD321C8F91F" box="[610,685,1728,1749]" pageId="22" pageNumber="537" value="2000-05">v.2000</date>
; BOR/ MOL 977,
<date id="FFEE105CFF99E8150D3C8ACC220FF93E" box="[287,362,1759,1780]" pageId="22" pageNumber="537" value="2000-05">v.2000</date>
. Imbak Valley: BOR/MOL 927, Transit Camp,
<date id="FFEE105CFF99E8150D758AEE22C9F8D9" box="[342,428,1789,1811]" pageId="22" pageNumber="537" value="2000-06">vi.2000</date>
; BOR/MOL 5264, Transit Camp,
<date id="FFEE105CFF99E8150CCA8B0F225AF8F8" box="[233,319,1820,1842]" pageId="22" pageNumber="537" value="2000-06">vi.2000</date>
; SP 12102,
<date id="FFEE105CFF99E8150DFF8B0F215FF8F8" box="[476,570,1820,1842]" pageId="22" pageNumber="537" value="2000-07">vii.2000</date>
. Kinabatangan: BOR/MOL 1294, Batu Materis,
<date id="FFEE105CFF99E8150E3C8B29210FF885" box="[543,618,1850,1871]" pageId="22" pageNumber="537" value="2002-05">v.2002</date>
; BOR/MOL 2310, Sukau, 2003; BOR/MOL 3597, Gomantong,
</paragraph>
<paragraph id="8BEF369CFF99E8150F058CD5274BFDA3" blockId="22.[806,1422,198,618]" pageId="22" pageNumber="537">
<date id="FFEE105CFF99E8150F058CD5200BFF16" box="[806,878,198,220]" pageId="22" pageNumber="537" value="2004-01">i.2004</date>
; BOR/MOL 5267, Bod Tai,
<date id="FFEE105CFF99E81508838CD5278EFF11" box="[1184,1259,198,219]" pageId="22" pageNumber="537" value="2002-05">v.2002</date>
. Tawau: BOR/ MOL 1297, Tawau Hills Park,
<date id="FFEE105CFF99E81508928CF72665FF30" box="[1201,1280,228,250]" pageId="22" pageNumber="537" value="2002-02">ii.2002</date>
; BOR/MOL 4268, Tawau Hills Park,
<date id="FFEE105CFF99E815087E8D1027CDFED3" box="[1117,1192,259,281]" pageId="22" pageNumber="537" value="2005-05">v.2005</date>
; SP 12704, Tawau Hills Park,
<date id="FFEE105CFF99E8150F9D8D312763FEF2" box="[958,1030,290,312]" pageId="22" pageNumber="537" value="2006-01">i.2006</date>
; SP 12902, Tawau Hills Park, Mount Magdelena,
<date id="FFEE105CFF99E81508318D532707FE9C" box="[1042,1122,320,342]" pageId="22" pageNumber="537" value="2007-02">ii.2007</date>
; SP 12193, Tawau Hills Park,
<date id="FFEE105CFF99E8150F4C8D4C20A1FEBF" box="[879,964,351,373]" pageId="22" pageNumber="537" value="2002-09">ix.2002</date>
; SP 12195, Tawau Hills Park,
<date id="FFEE105CFF99E81509178D4C26EDFEBF" box="[1332,1416,351,373]" pageId="22" pageNumber="537" value="2002-09">ix.2002</date>
; SP 12196, Tawau Hills Park,
<date id="FFEE105CFF99E81508848D6E2799FE59" box="[1191,1276,381,403]" pageId="22" pageNumber="537" value="2002-09">ix.2002</date>
; SP 12226, Tawau Hills Park,
<date id="FFEE105CFF99E81508258D8F2738FE78" box="[1030,1117,412,434]" pageId="22" pageNumber="537" value="2003-03">iii.2003</date>
; JJ 1876, Batu Baturong,
<date id="FFEE105CFF99E8150F058DA8200BFE1B" box="[806,878,443,465]" pageId="22" pageNumber="537" value="1987-01">i.1987</date>
; JJ 7592, Batu Baturong,
<date id="FFEE105CFF99E815088D8DA8266EFE1B" box="[1198,1291,443,465]" pageId="22" pageNumber="537" value="2000-07">vii.2000</date>
; JJ 13165, Tawau Hills Park,
<date id="FFEE105CFF99E81508358DCA2707FE25" box="[1046,1122,473,495]" pageId="22" pageNumber="537" value="2005-05">v.2005</date>
; JJ 13195, Tawau Hills Park,
<date id="FFEE105CFF99E8150F508DEB20DBFDC4" box="[883,958,504,526]" pageId="22" pageNumber="537" value="2005-05">v.2005</date>
; JJ 13208, Tawau Hills Park,
<date id="FFEE105CFF99E815091D8DEB26ECFDC4" box="[1342,1417,504,526]" pageId="22" pageNumber="537" value="2005-05">v.2005</date>
. Lahad Datu: JJ 1255, Segama Valley, Kirk’s Cave,
<date id="FFEE105CFF99E8150F058E26201EFD81" box="[806,891,565,587]" pageId="22" pageNumber="537" value="1986-09">ix.1986</date>
; JJ 7499, Segama valley,
<date id="FFEE105CFF99E81508998E262673FD81" box="[1210,1302,565,587]" pageId="22" pageNumber="537" value="2000-07">vii.2000</date>
; JJ 7789, Segama valley,
<date id="FFEE105CFF99E8150FFF8E47274FFDA3" box="[988,1066,596,617]" pageId="22" pageNumber="537" value="2000-10">x.2000</date>
.
</paragraph>
<paragraph id="8BEF369CFF99E8150F058E822740FD0F" blockId="22.[806,1421,657,709]" pageId="22" pageNumber="537">
<emphasis id="B924EA8EFF99E8150F058E822736FD6C" box="[806,1107,657,678]" italics="true" pageId="22" pageNumber="537">Diagnostic characteristics:</emphasis>
Shell surface with densely placed radial threads.
</paragraph>
<paragraph id="8BEF369CFF99E8150F058EFE2712F991" blockId="22.[806,1422,749,1627]" pageId="22" pageNumber="537">
<emphasis id="B924EA8EFF99E8150F058EFE20CBFCC8" box="[806,942,749,770]" italics="true" pageId="22" pageNumber="537">Description:</emphasis>
Shell (
<figureCitation id="136B2A19FF99E81508338EFE270EFCC9" box="[1040,1131,749,771]" captionStart="Figure 6" captionStartId="12.[144,223,1616,1635]" captionTargetBox="[306,1260,198,1584]" captionTargetId="figure-74@12.[303,1263,195,1586]" captionTargetPageId="12" captionText="Figure 6. Shells. A, Everettia lapidini sp. nov.; B, Everettia klemmantanica; C, Everettia subconsul; D, Everettia paulbasintali sp. nov.; E, Everettia themis; F, Everettia layanglayang sp. nov. Scale bar = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.5443035" httpUri="https://zenodo.org/record/5443035/files/figure.png" pageId="22" pageNumber="537">Figs 6D</figureCitation>
,
<figureCitation id="136B2A19FF99E815085C8EFE27BDFCC8" box="[1151,1240,749,770]" captionStart="Figure 10" captionStartId="17.[164,243,1552,1571]" captionTargetBox="[325,1283,195,1522]" captionTargetId="figure-92@17.[323,1283,195,1522]" captionTargetPageId="17" captionText="Figure 10. Shell surface sculptures (scanning electron micrographs). A, upper surface of Everettia jasilini sp. nov.; B, underside of E. jasilini sp. nov.; C, upper surface of Everettia safriei sp. nov.; D; underside of Everettia paulbasintali sp. nov.; E, upper surface of E. paulbasintali sp. nov.; F, underside of Everettia klemmantanica; G, upper surface on the fourth whorl of E. klemmantanica; H, upper surface on the second whorl of E. klemmantanica. Scale bar = 100 Mm." figureDoi="http://doi.org/10.5281/zenodo.5443043" httpUri="https://zenodo.org/record/5443043/files/figure.png" pageId="22" pageNumber="537">10D, E</figureCitation>
): large, rather thin, yellowish brown. Spire moderately elevated, outer whorls slightly shouldered below the suture. Periphery slightly shouldered to almost rounded, more distinctly angular in juveniles. Above the periphery, shell silky with very densely placed radial threads and somewhat cut by irregularly spaced spiral grooves. Below the periphery, shell has fine, densely placed spiral grooves. Height up to
<quantity id="4CA89B79FF99E81509078FF226EDFC3D" box="[1316,1416,993,1015]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="22" pageNumber="537" unit="mm" value="19.5">19.5 mm</quantity>
; width up to
<quantity id="4CA89B79FF99E8150FE98813274AFBDF" box="[970,1071,1024,1046]" metricMagnitude="-2" metricUnit="m" metricValue="2.55" pageId="22" pageNumber="537" unit="mm" value="25.5">25.5 mm</quantity>
; diameter of the first three whorls 0.8–0.9, 0.7–0.8, and
<quantity id="4CA89B79FF99E815084F880D2796FBFE" box="[1132,1267,1054,1076]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="22" pageNumber="537" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
, respectively; number of whorls up to five and a half; height aperture up to
<quantity id="4CA89B79FF99E8150F91884F2766FBBB" box="[946,1027,1116,1137]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="22" pageNumber="537" unit="mm" value="13.0">13 mm</quantity>
; width aperture up to
<quantity id="4CA89B79FF99E8150907884F26ECFBBB" box="[1316,1417,1116,1138]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="22" pageNumber="537" unit="mm" value="13.5">13.5 mm</quantity>
. Genitalia (
<figureCitation id="136B2A19FF99E8150F8A8869209AFB5B" box="[937,1023,1146,1169]" captionStart="Figure 7" captionStartId="13.[164,243,1755,1774]" captionTargetBox="[167,1442,198,1722]" captionTargetId="figure-0@13.[164,1444,194,1725]" captionTargetPageId="13" captionText="Figure 7. Genitalia. A, Everettia klemmantanica; B, Everettia lapidini sp. nov.; C, Everettia paulbasintali sp. nov.; D, Everettia subconsul; E, Everettia themis; F, Everettia layanglayang sp. nov.; G, Everettia jucundior sp. nov.; H, Everettia interior sp. nov.; I, Everettia corrugata corrugata." figureDoi="http://doi.org/10.5281/zenodo.5443037" httpUri="https://zenodo.org/record/5443037/files/figure.png" pageId="22" pageNumber="537">Fig. 7C</figureCitation>
): maximum length from genital opening to the end of dart-sac (before the visible gland tubules) up to
<quantity id="4CA89B79FF99E8150FE888AB2779FB07" box="[971,1052,1208,1229]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="537" unit="mm" value="14.0">14 mm</quantity>
. The penis, dart-sac, and vagina with thick muscular walls. P, V, GO, BC, and DS arranged around the short genital atrium, limited space between the openings of each of the genital structures. BC about one-quarter to one-third of total DS length. Animal: black bands on either side of the tentacles separated by a white band that occupies the space between the eye tentacles, and extends backward to the end of the mantle and downward to just below the eye tentacles. Other parts of the animal are covered by regularly spaced oblique dotted belts. The mantle is covered by black irregular markings and followed by regularly spaced radial darker brown bands at the last half whorl.
</paragraph>
<paragraph id="8BEF369CFF99E8150F058A9020F5F93E" blockId="22.[806,1421,1667,1780]" pageId="22" pageNumber="537">
<emphasis id="B924EA8EFF99E8150F058A90270FF952" box="[806,1130,1667,1688]" italics="true" pageId="22" pageNumber="537">Distribution and habitat:</emphasis>
Primary forest until
<quantity id="4CA89B79FF99E8150F058AB22018F97D" box="[806,893,1697,1719]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="22" pageNumber="537" unit="m" value="1400.0">1400 m</quantity>
.
<collectingRegion id="4994F87EFF99E8150FA88AB220B3F97D" box="[907,982,1697,1719]" country="Malaysia" name="Sabah" pageId="22" pageNumber="537">Sabah</collectingRegion>
: Danum Valley, Kinabatangan, Tabin Wildlife Reserve, Tawau Hills Park, Imbak Valley (
<figureCitation id="136B2A19FF99E8150F0D8ACD20E1F93E" box="[814,900,1758,1780]" captionStart="Figure 5" captionStartId="10.[144,223,1342,1361]" captionTargetBox="[305,1261,197,1311]" captionTargetId="figure-168@10.[303,1263,195,1313]" captionTargetPageId="10" captionText="Figure 5. Distribution maps (Sabah). A, Everettia subconsul in the east and west coast lowland, Everettia jucunda in the west coast, and Everettia occidentalis sp. nov. in the western mountain range; B, Everettia klemmantanica in the central mountain ranges, Everettia interior sp. nov. in the interior area, Everettia paulbasintali sp. nov. in the east coast lowland, Everettia themis in the central mountain ranges; C, Everettia lapidini sp. nov. in the central mountain ranges, Everettia jucundior sp. nov. in the east coast lowland; D, Everettia monticola sp. nov. in the central mountain ranges, Everettia planispira sp. nov. in the eastern and northern parts of Sabah." figureDoi="http://doi.org/10.5281/zenodo.5443033" httpUri="https://zenodo.org/record/5443033/files/figure.png" pageId="22" pageNumber="537">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8BEF369CFF99E8140F058B0F22A1FEFD" blockId="22.[806,1422,1820,1903]" lastBlockId="23.[164,779,198,312]" lastPageId="23" lastPageNumber="538" pageId="22" pageNumber="537">
<emphasis id="B924EA8EFF99E8150F058B0F20F7F8FB" box="[806,914,1820,1841]" italics="true" pageId="22" pageNumber="537">Remarks:</emphasis>
Some of the populations on the left bank of the Segama river (area around Imbak Valley, Danum Valley, and Kinabatangan) have coarser radial threads and these are cut by more closely placed spiral grooves than in the populations with typical sculpture from the right bank of the Segama river (Tabin and Tawau area).
</paragraph>
</subSubSection>
</treatment>
</document>
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