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<document ID-DOI="10.1371/journal.pone.0179555" ID-GBIF-Dataset="6ea2cc5c-857b-4b47-8135-8bff7efbd1fc" ID-PMC="PMC5526618" ID-PubMed="28742835" ID-Zenodo-Dep="4251352" approvedBy="poa" checkinTime="1604676189431" checkinUser="tatiana" docAuthor="Voon-Ching Lim, Rosli Ramli, Subha Bhassu &amp; John-James Wilson" docDate="2017" docId="03F3F77FFFACFFB1FDB4D505BDAEFE78" docLanguage="en" docName="PLoSOne.12.7.e0179555.pdf.imf" docOrigin="PLoS ONE 12 (7)" docStyle="DocumentStyle{}" docTitle="Myotis ater" docType="treatment" docVersion="7" lastPageNumber="52" masterDocId="FFCA8F07FF9EFF82FF81D072BF4BFFA2" masterDocTitle="A checklist of the bats of Peninsular Malaysia and progress towards a DNA barcode reference library" masterLastPageNumber="65" masterPageNumber="1" pageNumber="51" updateTime="1668143238971" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo>
<mods:title>A checklist of the bats of Peninsular Malaysia and progress towards a DNA barcode reference library</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Voon-Ching Lim</mods:namePart>
<mods:affiliation>Institute of Biological Sciences, Faculty of Science, University of Malaya, Kuala Lumpur, Malaysia</mods:affiliation>
<mods:nameIdentifier type="email">voonchinglim@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rosli Ramli</mods:namePart>
<mods:affiliation>Institute of Biological Sciences, Faculty of Science, University of Malaya, Kuala Lumpur, Malaysia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Subha Bhassu</mods:namePart>
<mods:affiliation>Institute of Biological Sciences, Faculty of Science, University of Malaya, Kuala Lumpur, Malaysia,</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>John-James Wilson</mods:namePart>
<mods:affiliation>International College Beijing, China Agricultural University, Beijing, P. R. China</mods:affiliation>
<mods:nameIdentifier type="email">wilso04@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>PLoS ONE</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="pubDate">
<mods:number>2017-07-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>12</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>7</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>65</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1371/journal.pone.0179555</mods:identifier>
<mods:identifier type="GBIF-Dataset">6ea2cc5c-857b-4b47-8135-8bff7efbd1fc</mods:identifier>
<mods:identifier type="PMC">PMC5526618</mods:identifier>
<mods:identifier type="PubMed">28742835</mods:identifier>
<mods:identifier type="Zenodo-Dep">4251352</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.4343587" ID-GBIF-Taxon="168816085" ID-Zenodo-Dep="4343587" LSID="urn:lsid:plazi:treatment:03F3F77FFFACFFB1FDB4D505BDAEFE78" httpUri="http://treatment.plazi.org/id/03F3F77FFFACFFB1FDB4D505BDAEFE78" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
<subSubSection box="[565,839,1399,1424]" pageId="50" pageNumber="51" type="nomenclature">
<paragraph blockId="50.[533,1536,1399,1840]" box="[565,839,1399,1424]" pageId="50" pageNumber="51">
<heading box="[565,833,1399,1424]" pageId="50" pageNumber="51">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1866" box="[565,688,1399,1423]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="ater">
<emphasis bold="true" box="[565,688,1399,1423]" italics="true" pageId="50" pageNumber="51">Myotis ater</emphasis>
</taxonomicName>
[Peters, 1866
</heading>
]
</paragraph>
</subSubSection>
<subSubSection pageId="50" pageNumber="51" type="reference_group">
<paragraph blockId="50.[533,1536,1399,1840]" pageId="50" pageNumber="51">
<taxonomicName authority="Peters, 1866: 18" authorityName="Peters" authorityPageNumber="18" authorityYear="1866" box="[565,895,1434,1458]" class="Mammalia" family="Vespertilionidae" genus="Vespertilio" kingdom="Animalia" order="Chiroptera" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="ater">
<emphasis box="[565,722,1434,1458]" italics="true" pageId="50" pageNumber="51">Vespertilio ater</emphasis>
<materialsCitation ID-GBIF-Occurrence="2960680396" box="[729,895,1434,1458]" country="Indonesia" location="Ternate Island" municipality="Moluccas" pageId="50" pageNumber="51" specimenCount="1">Peters, 1866: 18</materialsCitation>
</taxonomicName>
;
<location LSID="urn:lsid:plazi:treatment:03F3F77FFFACFFB1FDB4D505BDAEFE78:8E8510B2FFACFFB0FC08D5E8BB6EFA10" box="[905,1061,1434,1458]" country="Indonesia" municipality="Moluccas" name="Ternate Island" pageId="50" pageNumber="51">Ternate Island</location>
,
<collectingMunicipality box="[1072,1176,1434,1458]" pageId="50" pageNumber="51">Moluccas</collectingMunicipality>
,
<collectingCountry box="[1185,1328,1434,1458]" name="Indonesia" pageId="50" pageNumber="51">INDONESIA</collectingCountry>
(Collector unknown;
<typeStatus box="[648,701,1469,1493]" pageId="50" pageNumber="51">Type</typeStatus>
unknown) [
<bibRefCitation author="Peters W." box="[834,872,1469,1493]" journalOrPublisher="Monatsberichte der Koniglichen Preussische Akademie der Wissenschaften zu Berlin" pageId="50" pageNumber="51" pagination="16 - 24" part="1866" refId="ref46595" refString="212. Peters W. Uber einige neue oder weniger bekannte Flederthiere. Monatsberichte der Koniglichen Preussische Akademie der Wissenschaften zu Berlin, 1866; 1866. pp. 16 - 24." title="Uber einige neue oder weniger bekannte Flederthiere" type="journal article" year="1866">212</bibRefCitation>
]
</paragraph>
<paragraph blockId="50.[533,1536,1399,1840]" box="[565,726,1504,1528]" pageId="50" pageNumber="51">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1866" box="[565,682,1504,1527]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="ater">
<emphasis box="[565,682,1504,1527]" italics="true" pageId="50" pageNumber="51">Myotis ater</emphasis>
</taxonomicName>
[
<bibRefCitation author="Corbet GB &amp; Hill JE" box="[698,711,1504,1528]" journalOrPublisher="Oxford: Oxford University Press" pageId="50" pageNumber="51" refId="ref38080" refString="9. Corbet GB, Hill JE. The mammals of the Indomalayan region: a systematic review. Oxford: Oxford University Press; 1992." title="The mammals of the Indomalayan region: a systematic review" type="book" year="1992">9</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection box="[565,1000,1538,1562]" pageId="50" pageNumber="51" type="vernacular_names">
<paragraph blockId="50.[533,1536,1399,1840]" box="[565,1000,1538,1562]" pageId="50" pageNumber="51">
<emphasis bold="true" box="[565,834,1538,1562]" pageId="50" pageNumber="51">Common English name:</emphasis>
Peterss Myotis
</paragraph>
</subSubSection>
<subSubSection pageId="50" pageNumber="51" type="description">
<paragraph blockId="50.[533,1536,1399,1840]" pageId="50" pageNumber="51">
<emphasis bold="true" box="[565,832,1573,1597]" pageId="50" pageNumber="51">Barcode Index Number:</emphasis>
A DNA barcode recorded as
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1866" box="[1144,1250,1573,1597]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="ater">
<emphasis box="[1144,1167,1573,1596]" italics="true" pageId="50" pageNumber="51">M</emphasis>
. cf.
<emphasis box="[1210,1250,1573,1596]" italics="true" pageId="50" pageNumber="51">ater</emphasis>
</taxonomicName>
(BM487-04) is from Peninsular
<collectingCountry box="[613,707,1608,1632]" name="Malaysia" pageId="50" pageNumber="51">Malaysia</collectingCountry>
but is not associated with any BIN due to its short sequence length (
<emphasis box="[1428,1449,1608,1632]" italics="true" pageId="50" pageNumber="51">&lt;</emphasis>
500 bp). Other DNA barcodes recorded as
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1866" box="[895,970,1642,1666]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="ater">
<emphasis box="[895,918,1643,1666]" italics="true" pageId="50" pageNumber="51">M</emphasis>
.
<emphasis box="[930,970,1643,1666]" italics="true" pageId="50" pageNumber="51">ater</emphasis>
</taxonomicName>
are associated with the BIN, BOLD:AAA8748.
</paragraph>
</subSubSection>
<subSubSection lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51" type="discussion">
<paragraph blockId="50.[533,1536,1399,1840]" lastBlockId="51.[533,1509,207,474]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
<emphasis bold="true" box="[565,669,1677,1701]" pageId="50" pageNumber="51">Remarks:</emphasis>
The only BIN (BOLD:AAA8748) that is associated with
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1866" box="[1268,1343,1677,1701]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="ater">
<emphasis box="[1268,1291,1677,1700]" italics="true" pageId="50" pageNumber="51">M</emphasis>
.
<emphasis box="[1303,1343,1677,1700]" italics="true" pageId="50" pageNumber="51">ater</emphasis>
</taxonomicName>
also contains DNA barcodes recorded as
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1846" box="[826,954,1711,1736]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="muricola">
<emphasis box="[826,849,1712,1735]" italics="true" pageId="50" pageNumber="51">M</emphasis>
.
<emphasis box="[861,954,1711,1735]" italics="true" pageId="50" pageNumber="51">muricola</emphasis>
</taxonomicName>
(
<figureCitation box="[969,1021,1712,1736]" captionStart="Fig 6" captionStartId="36.[470,502,1797,1816]" captionTargetBox="[470,1535,208,1781]" captionTargetId="figure-1@36.[470,1536,208,1781]" captionTargetPageId="36" captionText="Fig 6. Neighbour-joining tree showing all available DNA barcodes for species in family Vespertilionidae reported from Peninsular Malaysia. The percentage of pseudoreplicate trees (±70%) in which the DNA barcodes clustered together in the bootstrap test (500 pseudoreplicates) are shown above the branches. Abbreviation as follows: PM = Peninsular Malaysia,VN = Vietnam, BN = Borneo (including Sabah &amp; Sarawak of East Malaysia, Brunei and Kalimantan Indonesia), TH = Thailand, LA = Laos, SM = Sumatera Indonesia, JV = Java Indonesia, CH = China, CM = Cambodia. https://doi.org/10.1371/journal.pone.0179555.g006" figureDoi="http://doi.org/10.5281/zenodo.4251370" httpUri="https://zenodo.org/record/4251370/files/figure.png" pageId="50" pageNumber="51">Fig 6</figureCitation>
). These taxa were previously considered to be conspecific [
<bibRefCitation author="Hill JE" box="[667,705,1746,1770]" pageId="50" pageNumber="51" pagination="119 - 137" part="139" refId="ref46628" refString="213. Hill JE. Notes on some insectivores and bats from Upper Burma. In Proceedings of the Zoological Society of London 139 (1); 1962. pp. 119 - 137." title="Notes on some insectivores and bats from Upper Burma" type="book chapter" volumeTitle="Proceedings of the Zoological Society of London" year="1962">213</bibRefCitation>
] but Francis and Hill [
<bibRefCitation author="Francis CM &amp; Hill JE" box="[947,985,1746,1770]" journalOrPublisher="Mammalia." pageId="50" pageNumber="51" pagination="241 - 252" part="62" refId="ref46664" refString="214. Francis CM, Hill JE. New records and a new species of Myotis (Chiroptera, Vespertilionidae) from Malaysia. Mammalia. 1998; 62: 241 - 252." title="New records and a new species of Myotis (Chiroptera, Vespertilionidae) from Malaysia" type="journal article" year="1998">214</bibRefCitation>
] recognised two species occuring sympatrically in
<collectingCountry box="[533,629,1781,1805]" name="Malaysia" pageId="50" pageNumber="51">Malaysia</collectingCountry>
, and commented that specimens from Peninsular
<collectingCountry box="[1162,1256,1781,1805]" name="Malaysia" pageId="50" pageNumber="51">Malaysia</collectingCountry>
recorded as
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1866" box="[1390,1465,1781,1805]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="ater">
<emphasis box="[1390,1413,1781,1804]" italics="true" pageId="50" pageNumber="51">M</emphasis>
.
<emphasis box="[1425,1465,1781,1804]" italics="true" pageId="50" pageNumber="51">ater</emphasis>
</taxonomicName>
are larger than
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1866" box="[655,730,1816,1840]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="ater">
<emphasis box="[655,678,1816,1839]" italics="true" pageId="50" pageNumber="51">M</emphasis>
.
<emphasis box="[690,730,1816,1839]" italics="true" pageId="50" pageNumber="51">ater</emphasis>
</taxonomicName>
from elsewhere. As
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1846" box="[945,1073,1815,1840]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="50" pageNumber="51" phylum="Chordata" rank="species" species="muricola">
<emphasis box="[945,968,1816,1839]" italics="true" pageId="50" pageNumber="51">M</emphasis>
.
<emphasis box="[980,1073,1815,1839]" italics="true" pageId="50" pageNumber="51">muricola</emphasis>
</taxonomicName>
is putatively a species complex (refer to remarks on
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1846" box="[661,789,207,232]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="muricola">
<emphasis box="[661,684,208,231]" italics="true" pageId="51" pageNumber="52">M</emphasis>
.
<emphasis box="[696,789,207,231]" italics="true" pageId="51" pageNumber="52">muricola</emphasis>
</taxonomicName>
), further studies of
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1866" box="[997,1072,208,232]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="ater">
<emphasis box="[997,1020,208,231]" italics="true" pageId="51" pageNumber="52">M</emphasis>
.
<emphasis box="[1032,1072,208,231]" italics="true" pageId="51" pageNumber="52">ater</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Gray" baseAuthorityYear="1846" box="[1125,1253,207,232]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="muricola">
<emphasis box="[1125,1148,208,231]" italics="true" pageId="51" pageNumber="52">M</emphasis>
.
<emphasis box="[1160,1253,207,231]" italics="true" pageId="51" pageNumber="52">muricola</emphasis>
</taxonomicName>
are required to resolve the relationship between these taxa.
</paragraph>
<paragraph blockId="51.[533,1509,207,474]" box="[565,870,277,301]" pageId="51" pageNumber="52">
<emphasis bold="true" box="[565,710,277,301]" pageId="51" pageNumber="52">IUCN status:</emphasis>
Least Concern
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="distribution">
<paragraph blockId="51.[533,1509,207,474]" pageId="51" pageNumber="52">
<emphasis bold="true" box="[565,794,312,336]" pageId="51" pageNumber="52">
Recorded at:
<collectingRegion box="[711,794,312,336]" country="Malaysia" name="Pahang" pageId="51" pageNumber="52">Pahang</collectingRegion>
</emphasis>
: Krau Wildlife Reserve [
<bibRefCitation author="Kingston T &amp; Lim BL &amp; Akbar Z." box="[1054,1079,312,336]" journalOrPublisher="Penerbit Universiti Kebangsaan Malaysia" pageId="51" pageNumber="52" refId="ref38130" refString="11. Kingston T, Lim BL, Akbar Z. Bats of Krau wildlife reserve. Penerbit Universiti Kebangsaan Malaysia; 2006." title="Bats of Krau wildlife reserve" type="book" year="2006">11</bibRefCitation>
], Cameron
<collectingRegion box="[1207,1308,312,336]" country="United Kingdom" name="Highland" pageId="51" pageNumber="52">Highland</collectingRegion>
[
<bibRefCitation author="Shahfiz MA &amp; Sahrul-Anuar MS &amp; Shukor MN &amp; Zalipah MN &amp; Pan KA &amp; Muin MA" box="[1323,1348,312,336]" editor="Rahman A &amp; Koh HL &amp; Mohd-Paiz K &amp; Muhamad A &amp; Latiff A" journalOrPublisher="Department of Wildlife and National Parks, Malaysia" pageId="51" pageNumber="52" pagination="293 - 298" refId="ref40430" refString="60. Shahfiz MA, Sahrul-Anuar MS, Shukor MN, Zalipah MN, Pan KA, Muin MA, et al. Survey of small mammals at two forest reserves in Cameron Highlands, Pahang, Peninsular Malaysia. In: Rahman A, Koh HL, Mohd-Paiz K, Muhamad A, Latiff A, editors. Proceedings for Forestry Department of Peninsular Malaysia, 3 - 5 August 2008. Department of Wildlife and National Parks, Malaysia; 2008. pp. 293 - 298." title="Survey of small mammals at two forest reserves in Cameron Highlands, Pahang, Peninsular Malaysia" type="book chapter" volumeTitle="Proceedings for Forestry Department of Peninsular Malaysia" year="2008">60</bibRefCitation>
];
<emphasis bold="true" box="[1369,1433,312,336]" pageId="51" pageNumber="52">
<collectingRegion box="[1369,1433,312,336]" country="Malaysia" name="Perak" pageId="51" pageNumber="52">Perak</collectingRegion>
</emphasis>
: Bukit Jerneh Cave and Tumang Lembing Cave [
<bibRefCitation author="Douangboubpha B &amp; Bumrungsri S &amp; Soisook P &amp; Murray SW &amp; Puechmaille SJ &amp; Satasook C" box="[979,1004,346,370]" journalOrPublisher="Acta Chiropt." pageId="51" pageNumber="52" pagination="29 - 50" part="12" refId="ref39009" refString="30. Douangboubpha B, Bumrungsri S, Soisook P, Murray SW, Puechmaille SJ, Satasook C, et al. A taxonomic review of Hipposideros halophyllus, with additional information on H. ater and H. cineraceus (Chiroptera: Hipposideridae) from Thailand and Myanmar. Acta Chiropt. 2010; 12 (1): 29 - 50." title="A taxonomic review of Hipposideros halophyllus, with additional information on H. ater and H. cineraceus (Chiroptera: Hipposideridae) from Thailand and Myanmar" type="journal article" year="2010">30</bibRefCitation>
];
<emphasis bold="true" box="[1026,1098,346,370]" pageId="51" pageNumber="52">
<collectingRegion box="[1026,1098,346,370]" country="Malaysia" name="Kedah" pageId="51" pageNumber="52">Kedah</collectingRegion>
</emphasis>
: Ulu Muda Forest Reserve [
<bibRefCitation author="Norsham Y &amp; Shariff F &amp; Norhayati A &amp; Nordin M &amp; Lim BL" box="[1394,1419,346,370]" journalOrPublisher="Journal of Wildlife and Parks." pageId="51" pageNumber="52" pagination="28 - 43" part="17" refId="ref40272" refString="57. Norsham Y, Shariff F, Norhayati A, Nordin M, Lim BL. Pre-logging survey of mammal fauna at Sungai Weng sub-catchment, Ulu Muda Forest Reserve Forest Reserve, Kedah. Journal of Wildlife and Parks. 1999; 17: 28 - 43." title="Pre-logging survey of mammal fauna at Sungai Weng sub-catchment, Ulu Muda Forest Reserve Forest Reserve, Kedah" type="journal article" year="1999">57</bibRefCitation>
].
</paragraph>
</subSubSection>
<subSubSection pageId="51" pageNumber="52" type="discussion">
<paragraph blockId="51.[533,1509,207,474]" pageId="51" pageNumber="52">
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1866" box="[565,640,381,405]" class="Mammalia" family="Vespertilionidae" genus="Myotis" kingdom="Animalia" order="Chiroptera" pageId="51" pageNumber="52" phylum="Chordata" rank="species" species="ater">
<emphasis box="[565,588,382,405]" italics="true" pageId="51" pageNumber="52">M</emphasis>
.
<emphasis box="[600,640,382,405]" italics="true" pageId="51" pageNumber="52">ater</emphasis>
</taxonomicName>
has been reported roosting in caves, either solitarily or in small colonies and recorded foraging in open areas such as gaps created by fallen trees, midstorey openings and forest edge [
<bibRefCitation author="Kingston T &amp; Lim BL &amp; Akbar Z." box="[663,688,450,474]" journalOrPublisher="Penerbit Universiti Kebangsaan Malaysia" pageId="51" pageNumber="52" refId="ref38130" refString="11. Kingston T, Lim BL, Akbar Z. Bats of Krau wildlife reserve. Penerbit Universiti Kebangsaan Malaysia; 2006." title="Bats of Krau wildlife reserve" type="book" year="2006">11</bibRefCitation>
,
<bibRefCitation author="Francis CM" box="[700,725,450,474]" journalOrPublisher="New Holland Publishers" pageId="51" pageNumber="52" refId="ref38235" refString="14. Francis CM. A field guide to the mammals of South-East Asia. New Holland Publishers; 2008." title="A field guide to the mammals of South-East Asia" type="book" year="2008">14</bibRefCitation>
].
</paragraph>
</subSubSection>
</treatment>
</document>