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<document id="813B4AF20036FCEC36AA99F098207CA3" ID-CLB-Dataset="8143" ID-DOI="10.2478/aemnp-2019-0020" ID-GBIF-Dataset="a03ecce2-b526-4f94-aa65-620016956099" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488918" ID-ZooBank="9F309FCC-A2ED-47B9-BC37-D0C4A3B482E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612219281920" checkinUser="felipe" docAuthor="Fikáček, Martin &amp; Liu, Hsing-Che" docDate="2019" docId="03F387C57A3DFFABFC3300A53552B25C" docLanguage="en" docName="ActaEntMusNatPra.59.1.229-252.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 59 (1)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Thysanarthria wadicola Fikáček &amp; Liu 2019, sp. nov." docType="treatment" docVersion="11" lastPageNumber="250" masterDocId="FFCAFFBD7A2EFFBEFF8901283431B709" masterDocTitle="A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)" masterLastPageNumber="252" masterPageNumber="229" pageNumber="248" updateTime="1698926077238" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
<mods:mods id="26DEDD76315C7DD183A37D33239F385F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="A53F402E7D3D8AA0DEDD241E4B443A3C">
<mods:title id="53BF209B42CEC22D09B21786A47C5BF8">A review of Thysanarthria with description of seven new species and comments on its relationship to Chaetarthria (Hydrophilidae: Chaetarthriini)</mods:title>
</mods:titleInfo>
<mods:name id="BDD9AE21E788EC639D0EE0E4F992E78A" type="personal">
<mods:role id="EC0B0F91FDFD6AE6D9D6B51694AAF4CC">
<mods:roleTerm id="49D5ADED0DCBE37584416C9F9F1C9656">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1E89F62B41E0F834B922C6F2EE307AF2">Fikáček, Martin</mods:namePart>
<mods:affiliation id="F8D6C247C675E078D1CCFC4C2768B69A">Department of Entomology, National Museum, Cirkusová 1740, CZ- 19100 Praha 9 – Horní Počernice, Czech Republic &amp; Department of Zoology, Faculty of Science, Charles University, Viničná 7, CZ- 12844 Praha 2, Czech Republic</mods:affiliation>
<mods:nameIdentifier id="2627A295D44BC7767D90A9248D141FB3" type="email">mfikacek@gmail.com</mods:nameIdentifier>
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<mods:name id="2CFA77AFA83D2241F28BECDA8CB8D45B" type="personal">
<mods:role id="CE5416E3F69DD1D50FB728345AB64032">
<mods:roleTerm id="5FBD5D6A0FD0687417159C7CAF625350">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="59340A86EE2047ED865B120DED6471FB">Liu, Hsing-Che</mods:namePart>
<mods:affiliation id="8BF411F1618DB1C020642AFA81E1EBDD">Department of Environmental Engineering and Management, Chaoyang University of Technology, No. 168, Jifeng E. Rd., Taichung City 413, Taiwan</mods:affiliation>
<mods:nameIdentifier id="CEF13BED4754FC1146271EEA3A6974C8" type="email">td965771@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="4A0BB36BE4ACCA6447C7B065C4CF2FC9">text</mods:typeOfResource>
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<mods:title id="F84D52F1454549B55CB17567F9DC645E">Acta Entomologica Musei Nationalis Pragae</mods:title>
</mods:titleInfo>
<mods:part id="3C5FEE5FBC2C0B33A4C888B21F7A61E1">
<mods:date id="C7B692018B925ADCFD848305C74A04E0">2019</mods:date>
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<mods:number id="B3D98FEF38E42C747D321C30C1C0206D">2019-06-20</mods:number>
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<mods:detail id="7165A78687084C61D1B7BA591BDCDEF7" type="volume">
<mods:number id="209A9770DDF9D2D4A6C19EAF4A0ACCDB">59</mods:number>
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<mods:classification id="A962CB1711B54AD5303437A8FEDEDF31">journal article</mods:classification>
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<mods:identifier id="A912BE6CE892FFD6B5C7F2DA6A840F24" type="DOI">10.2478/aemnp-2019-0020</mods:identifier>
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<mods:identifier id="C006DCAF0274FEAE0F6B4FF51D22AE1C" type="ISSN">1804-6487</mods:identifier>
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<treatment id="03F387C57A3DFFABFC3300A53552B25C" ID-DOI="http://doi.org/10.5281/zenodo.4548845" ID-GBIF-Taxon="177423413" ID-Zenodo-Dep="4548845" LSID="urn:lsid:plazi:treatment:03F387C57A3DFFABFC3300A53552B25C" httpUri="http://treatment.plazi.org/id/03F387C57A3DFFABFC3300A53552B25C" lastPageId="21" lastPageNumber="250" pageId="19" pageNumber="248">
<subSubSection id="C34065587A3DFFADFC3300A5312FB6AD" box="[954,1310,397,420]" pageId="19" pageNumber="248" type="nomenclature">
<paragraph id="8BE536D37A3DFFADFC3300A5312FB6AD" blockId="19.[954,1310,397,420]" box="[954,1310,397,420]" pageId="19" pageNumber="248">
<heading id="D0AD81BF7A3DFFADFC3300A5312FB6AD" box="[954,1310,397,420]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="248" reason="3">
<emphasis id="B92EEAC17A3DFFADFC3300A5312FB6AD" bold="true" box="[954,1310,397,420]" pageId="19" pageNumber="248">
<taxonomicName id="4C5A4D507A3DFFADFC3300A530F3B6AD" ID-CoL="8YBVY" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[954,1218,397,420]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="248" phylum="Arthropoda" rank="species" species="wadicola" status="sp. nov.">
<emphasis id="B92EEAC17A3DFFADFC3300A530F3B6AD" bold="true" box="[954,1218,397,420]" italics="true" pageId="19" pageNumber="248">Thysanarthria wadicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A3DFFADFB4100A5312FB6AD" box="[1224,1310,397,420]" pageId="19" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3DFFADFBAF00833083B6B7" box="[1062,1202,427,446]" pageId="19" pageNumber="248" type="description">
<paragraph id="8BE536D37A3DFFADFBAF00833083B6B7" blockId="19.[828,1437,427,762]" box="[1062,1202,427,446]" pageId="19" pageNumber="248">
(
<figureCitation id="13612A567A3DFFADFBA50083305DB6B7" box="[1068,1132,427,446]" captionStart="Fig" captionStartId="19.[151,182,2009,2028]" captionTargetBox="[159,1399,787,1981]" captionTargetId="figure-14@19.[151,1436,777,1989]" captionTargetPageId="19" captionText="Fig. 10. Male genitalia of Thysanarthria species, holotypes.A–E – T. persica sp. nov. (A–C – aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. wadicola sp. nov. (F–H – aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488944" httpUri="https://zenodo.org/record/4488944/files/figure.png" pageId="19" pageNumber="248">Figs 10</figureCitation>
F–J,
<figureCitation id="13612A567A3DFFADFB1F0083309AB6B7" box="[1174,1195,427,446]" captionStart="Fig" captionStartId="20.[151,182,1949,1968]" captionTargetBox="[151,1436,401,1931]" captionTargetId="figure-13@20.[151,1437,401,1931]" captionTargetPageId="20" captionText="Fig. 11. Distribution of Thysanarthria and American Chaetarthria in Asia (A–B) and Near East (C), and examples of habitats of Thysanarthria (D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of T. ceylonensis Hebauer, 2001; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of T. atriceps (Régimber, 1903), collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of T. chui sp. nov. (G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu." figureDoi="http://doi.org/10.5281/zenodo.4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="19" pageNumber="248">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3DFFADFCB500E537D7B5BC" pageId="19" pageNumber="248" type="reference_group">
<paragraph id="8BE536D37A3DFFADFCB500E5379CB51C" blockId="19.[828,1437,427,762]" pageId="19" pageNumber="248">
<treatmentCitationGroup id="AB4A11FD7A3DFFADFCB500E5379CB51C" pageId="19" pageNumber="248">
<taxonomicName id="4C5A4D507A3DFFADFCB500E537C8B6E9" authorityName=": HANSEN" authorityYear="1991" box="[828,1017,461,480]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="248" phylum="Arthropoda" rank="species" species="sulcata">
<emphasis id="B92EEAC17A3DFFADFCB500E537C8B6E9" box="[828,1017,461,480]" italics="true" pageId="19" pageNumber="248">Thysanarthria sulcata</emphasis>
</taxonomicName>
(misidentification):
<treatmentCitation id="0AFB10C27A3DFFADFB2F00E53164B6E9" author="HEBAUER F." box="[1190,1365,461,480]" page="267" pageId="19" pageNumber="248" year="1997">
<bibRefCitation id="EFCB4B227A3DFFADFB2F00E53164B6E9" author="HEBAUER" box="[1190,1365,461,480]" firstAuthor="HEBAUER" page="267" pageId="19" pageNumber="248" pagination="255 - 276" refId="ref18079" refString="HEBAUER F. 1997: Annotated checklist of the Hydrophilidae and Helophoridae (Insecta: Coleoptera) of the Arabian Peninsula with a description of a new genus and species. Fauna of Saudi Arabia 16: 255 - 276." type="journal article" year="1997">HEBAUER (1997: 267</bibRefCitation>
</treatmentCitation>
; records from
<collectingCountry id="F34D76437A3DFFADFC0E00C0378CB6F2" box="[903,957,488,507]" name="Oman" pageId="19" pageNumber="248">Oman</collectingCountry>
, illustration of genitalia);
<treatmentCitation id="0AFB10C27A3DFFADFB1800C03171B6F2" author="HEBAUER F." box="[1169,1344,488,507]" page="399" pageId="19" pageNumber="248" year="2001">
<bibRefCitation id="EFCB4B227A3DFFADFB1800C0310DB6F2" author="HEBAUER" box="[1169,1340,488,507]" firstAuthor="HEBAUER" page="399" pageId="19" pageNumber="248" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">HEBAUER (2001:399</bibRefCitation>
,
</treatmentCitation>
taxonomic revision).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536D37A3DFFADFCB503353799B542" blockId="19.[828,1437,427,762]" pageId="19" pageNumber="248">
<treatmentCitationGroup id="AB4A11FD7A3DFFADFCB503353799B542" pageId="19" pageNumber="248">
<taxonomicName id="4C5A4D507A3DFFADFCB503353029B539" authorityName=": HANSEN" authorityYear="1991" box="[828,1048,541,560]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="248" phylum="Arthropoda" rank="species" species="sulcata">
<emphasis id="B92EEAC17A3DFFADFCB503353787B539" box="[828,950,541,560]" italics="true" pageId="19" pageNumber="248">Thysanarthria</emphasis>
cf.
<emphasis id="B92EEAC17A3DFFADFC5303353029B539" box="[986,1048,541,560]" italics="true" pageId="19" pageNumber="248">sulcata</emphasis>
</taxonomicName>
(part):
<treatmentCitation id="0AFB10C27A3DFFADFBD20335310AB539" author="FIKACEK M. &amp; GENTILI E. &amp; SHORT A. E. Z." box="[1115,1339,541,560]" page="139" pageId="19" pageNumber="248" year="2010">
<bibRefCitation id="EFCB4B227A3DFFADFBD20335310AB539" author="FIKACEK" box="[1115,1339,541,560]" etAl="et al." firstAuthor="FIKACEK" page="139" pageId="19" pageNumber="248" pagination="135 - 165" refId="ref17868" refString="FIKACEK M., GENTILI E. &amp; SHORT A. E. Z. 2010: Order Coleoptera, family Hydrophilidae. Pp. 135 - 165. In: HARTEN A. VAN (ed.): Arthropod Fauna of the United Arab Emirates. Volume 3. Al Ummah Printing, Publishing, Distribution and Advertising, Abu Dhabi, 700 pp." type="book chapter" year="2010">FIKÁČEK et al. (2010: 139</bibRefCitation>
</treatmentCitation>
; misidentification).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536D37A3DFFADFCB5037A3793B589" blockId="19.[828,1437,427,762]" pageId="19" pageNumber="248">
<treatmentCitationGroup id="AB4A11FD7A3DFFADFCB5037A3793B589" pageId="19" pageNumber="248">
<taxonomicName id="4C5A4D507A3DFFADFCB5037A3787B56C" ID-CoL="8X7X2" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[828,950,594,613]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A3DFFADFCB5037A3787B56C" box="[828,950,594,613]" italics="true" pageId="19" pageNumber="248">Thysanarthria</emphasis>
</taxonomicName>
sp. (SLE127):
<treatmentCitation id="0AFB10C27A3DFFADFBB4037A313FB56C" author="SHORT A. E. Z. &amp; FIKACEK M. " box="[1085,1294,594,613]" pageId="19" pageNumber="248" year="2013">
<bibRefCitation id="EFCB4B227A3DFFADFBB4037A313FB56C" author="SHORT &amp; FIKACEK" box="[1085,1294,594,613]" firstAuthor="SHORT" pageId="19" pageNumber="248" pagination="723 - 752" refId="ref18724" refString="SHORT A. E. Z. &amp; FIKACEK M. 2013: Molecular phylogeny, evolution and classification of the Hydrophilidae (Coleoptera). Systematic Entomology 38: 723 - 752." type="journal article" year="2013">
SHORT &amp;
<subScript id="17DE34967A3DFFADFB06037A30E2B56D" attach="right" box="[1167,1235,594,613]" fontSize="6" pageId="19" pageNumber="248">FIKÁČEK</subScript>
(2013
</bibRefCitation>
</treatmentCitation>
; molecular phylogeny).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BE536D37A3DFFADFCB503A037D7B5BC" blockId="19.[828,1437,427,762]" pageId="19" pageNumber="248">
<treatmentCitationGroup id="AB4A11FD7A3DFFADFCB503A037D7B5BC" pageId="19" pageNumber="248">
<treatmentCitation id="0AFB10C27A3DFFADFCB503A030CCB592" ID-DOI="http://doi.org/10.5281/zenodo.4549039" ID-Zenodo-Dep="4549039" author="RIBERA I. &amp; HERNANDO C. &amp; CIESLAK A." box="[828,1277,648,667]" httpUri="http://treatment.plazi.org/id/03DD87A0FFD8FFDAFCF7FAADDD8AF886" page="264" pageId="19" pageNumber="248" year="2019">
<taxonomicName id="4C5A4D507A3DFFADFCB503A030CCB592" ID-CoL="8YBVY" authority=": RIBERA et al. (2019: 264" authorityName="RIBERA" authorityYear="2019" box="[828,1277,648,667]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="248" phylum="Arthropoda" rank="species" species="wadicola">
<emphasis id="B92EEAC17A3DFFADFCB503A03020B592" box="[828,1041,648,667]" italics="true" pageId="19" pageNumber="248">Thysanarthria wadicola</emphasis>
:
<bibRefCitation id="EFCB4B227A3DFFADFB9603A030CCB592" author="RIBERA" box="[1055,1277,648,667]" etAl="et al." firstAuthor="RIBERA" page="264" pageId="19" pageNumber="248" pagination="253 - 272" refId="ref18641" refString="RIBERA I., HERNANDO C. &amp; CIESLAK A. 2019: Aquatic Coleoptera of North Oman, with description of new species of Hydraenidae and Hydrophilidae. Acta Entomologica Musei Nationalis Pragae 59: 253 - 272." type="journal article" year="2019">RIBERA et al. (2019: 264</bibRefCitation>
</taxonomicName>
</treatmentCitation>
; review of
<collectingCountry id="F34D76437A3DFFADFAEE03A031ADB592" box="[1383,1436,648,667]" name="Oman" pageId="19" pageNumber="248">Oman</collectingCountry>
aquatic beetles).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3DFFAAFCB503E4305AB628" lastPageId="20" lastPageNumber="249" pageId="19" pageNumber="248" type="materials_examined">
<paragraph id="8BE536D37A3DFFAAFCB503E43599B635" blockId="19.[828,1437,427,762]" lastBlockId="20.[151,760,190,369]" lastPageId="20" lastPageNumber="249" pageId="19" pageNumber="248">
<emphasis id="B92EEAC17A3DFFADFCB503E437F3B5D6" bold="true" box="[828,962,716,735]" pageId="19" pageNumber="248">Type material.</emphasis>
<materialsCitation id="3B323C8E7A3DFFADFC4103E43132B5F3" ID-GBIF-Occurrence="3756360348" collectingDate="1985-04-17" collectingDateMax="1985-04-18" collectingDateMin="1985-04-17" collectionCode="NMPC" collectorName="C. Holzschuh" country="Oman" elevation="650" location="Wadi Andam" municipality="Wadi Andam" pageId="19" pageNumber="248" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54E188717A3DFFADFC4103E4302DB5D4" box="[968,1052,716,735]" pageId="19" pageNumber="248" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D5CFD5A7A3DFFADFBA103EE3009B5D6" box="[1064,1080,710,735]" pageId="19" pageNumber="248" type="male">♂</specimenCount>
(
<collectionCode id="ED4BAE167A3DFFADFBCC03E430B6B5D6" box="[1093,1159,716,735]" country="Czech Republic" name="National Museum Prague" pageId="19" pageNumber="248">NMPC</collectionCode>
):
<emphasis id="B92EEAC17A3DFFADFB1F03E430D0B5D6" bold="true" box="[1174,1249,716,735]" pageId="19" pageNumber="248">
<collectingCountry id="F34D76437A3DFFADFB1F03E430ECB5D6" box="[1174,1245,716,735]" name="Oman" pageId="19" pageNumber="248">OMAN</collectingCountry>
:
</emphasis>
<collectingMunicipality id="6B81ACA97A3DFFADFB6103E4316AB5D6" box="[1256,1371,716,735]" country="Oman" name="Wadi Andam" pageId="19" pageNumber="248">Wadi Andam</collectingMunicipality>
,
<locationDeviation id="9A572FF17A3DFFADFAEC03E437BCB5F3" pageId="19" pageNumber="248">
<quantity id="4CA29B367A3DFFADFAEC03E431ADB5D6" box="[1381,1436,716,735]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="248" unit="km" value="20.0">20 km</quantity>
N Samad
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,
<quantity id="4CA29B367A3DFFADFC1E03CF37E0B5F3" box="[919,977,743,762]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="248" unit="m" value="650.0">
<elevation id="0077D1E07A3DFFADFC1E03CF37E0B5F3" box="[919,977,743,762]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="19" pageNumber="248" unit="m" value="650.0">650 m</elevation>
</quantity>
,
<date id="FFE410137A3DFFADFC5303CF306AB5F3" box="[986,1115,743,762]" pageId="19" pageNumber="248" value="1985-04-17" valueMax="1985-04-18" valueMin="1985-04-17">
<collectingDate id="EFA0E9FB7A3DFFADFC5303CF306AB5F3" box="[986,1115,743,762]" pageId="19" pageNumber="248" value="1985-04-17" valueMax="1985-04-18" valueMin="1985-04-17">17.–18.iv.1985</collectingDate>
</date>
, lgt.
<collectorName id="26AF53057A3DFFADFB0E03CF30CEB5F3" box="[1159,1279,743,762]" pageId="19" pageNumber="248">C. Holzschuh</collectorName>
.
</materialsCitation>
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<typeStatus id="54E188717A3DFFADFA8103CF3151B5F1" box="[1288,1376,743,762]" pageId="19" pageNumber="248" type="paratype">PARATYPES</typeStatus>
: 2 ♂♂ (
<collectionCode id="ED4BAE167A3AFFAAFF14019634EDB7D8" box="[157,220,190,209]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34840" lsid="urn:lsid:biocol.org:col:34840" name="Staatliches Museum fuer Naturkund Stuttgart" pageId="20" pageNumber="249">SMNS</collectionCode>
): same data as the
<typeStatus id="54E188717A3AFFAAFE0B019635FEB7D8" box="[386,463,190,209]" pageId="20" pageNumber="249" type="holotype">holotype</typeStatus>
;
</materialsCitation>
<materialsCitation id="3B323C8E7A3AFFAAFE500196359FB60F" ID-GBIF-Occurrence="3756360307" latitude="23.496166" longitude="56.893" pageId="20" pageNumber="249">
<specimenCount id="9D5CFD5A7A3AFFAAFE5001963626B7D8" box="[473,535,190,209]" pageId="20" pageNumber="249" type="generic">3 spec.</specimenCount>
(
<collectionCode id="ED4BAE167A3AFFAAFDAB01963666B7D8" box="[546,599,190,209]" pageId="20" pageNumber="249">IBEB</collectionCode>
): Murri env., wadi Bani Ghafir,
<quantity id="4CA29B367A3AFFAAFE8501F13579B7E5" box="[268,328,217,236]" metricMagnitude="2" metricUnit="m" metricValue="7.59" pageId="20" pageNumber="249" unit="m" value="759.0">759 m</quantity>
, stream with pools,
<geoCoordinate id="EE6E50147A3AFFAAFD8801F13647B7E5" box="[513,630,217,236]" degrees="23" direction="north" minutes="29" orientation="latitude" pageId="20" pageNumber="249" precision="1" seconds="46.2" value="23.496166">23º29′46.2″N</geoCoordinate>
<geoCoordinate id="EE6E50147A3AFFAAFDF601F136C5B7E5" box="[639,756,217,236]" degrees="56" direction="east" minutes="53" orientation="longitude" pageId="20" pageNumber="249" precision="1" seconds="34.8" value="56.893">56º53′34.8″E</geoCoordinate>
,
<date id="FFE410137A3AFFAAFF1E01DB34D8B60F" box="[151,233,243,262]" pageId="20" pageNumber="249" value="2010-04-07">7.iv.2010</date>
, Ribera &amp; Cieslak lgt.;
</materialsCitation>
<materialsCitation id="3B323C8E7A3AFFAAFE3A01DB3599B635" ID-GBIF-Occurrence="3756360336" latitude="22.754223" longitude="58.015804" pageId="20" pageNumber="249">
<specimenCount id="9D5CFD5A7A3AFFAAFE3A01DB35C1B60F" box="[435,496,243,262]" pageId="20" pageNumber="249" type="generic">1 spec.</specimenCount>
(
<collectionCode id="ED4BAE167A3AFFAAFE7201DB3601B60F" box="[507,560,243,262]" pageId="20" pageNumber="249">IBEB</collectionCode>
): Said Bin Sahran env., wadi Indam, Rd.
<quantity id="4CA29B367A3AFFAAFEA0002635B1B628" box="[297,384,270,289]" metricMagnitude="4" metricUnit="m" metricValue="3.3463" pageId="20" pageNumber="249" unit="m" value="33463.0">33, 463 m</quantity>
, residual pools,
<geoCoordinate id="EE6E50147A3AFFAAFD800026364DB628" box="[521,636,270,289]" degrees="22" direction="north" minutes="45" orientation="latitude" pageId="20" pageNumber="249" precision="1" seconds="15.2" value="22.754223">22º45′15.2″N</geoCoordinate>
<geoCoordinate id="EE6E50147A3AFFAAFD08002636C5B628" box="[641,756,270,289]" degrees="58" direction="east" minutes="00" orientation="longitude" pageId="20" pageNumber="249" precision="1" seconds="56.9" value="58.015804">58º00′56.9″E</geoCoordinate>
,
<date id="FFE410137A3AFFAAFF1E000134D9B635" box="[151,232,297,316]" pageId="20" pageNumber="249" value="2010-04-08">8.iv.2010</date>
, Ribera &amp; Cieslak lgt.
</materialsCitation>
</paragraph>
<caption id="DF25665B7A3DFFADFF1E06F136BFBF28" ID-DOI="http://doi.org/10.5281/zenodo.4488944" ID-Zenodo-Dep="4488944" httpUri="https://zenodo.org/record/4488944/files/figure.png" pageId="19" pageNumber="248" startId="19.[151,182,2009,2028]" targetBox="[159,1399,787,1981]" targetPageId="19">
<paragraph id="8BE536D37A3DFFADFF1E06F136BFBF28" blockId="19.[151,1436,2009,2081]" pageId="19" pageNumber="248">
Fig. 10. Male genitalia of
<taxonomicName id="4C5A4D507A3DFFADFEFD06F135DFB0E5" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[372,494,2009,2028]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="248" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A3DFFADFEFD06F135DFB0E5" box="[372,494,2009,2028]" italics="true" pageId="19" pageNumber="248">Thysanarthria</emphasis>
</taxonomicName>
species, holotypes.A–E –
<taxonomicName id="4C5A4D507A3DFFADFD5A06F13716B0E5" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[723,807,2009,2028]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="248" phylum="Arthropoda" rank="species" species="persica" status="sp. nov.">
<emphasis id="B92EEAC17A3DFFADFD5A06F13716B0E5" box="[723,807,2009,2028]" italics="true" pageId="19" pageNumber="248">T. persica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A3DFFADFCA506F1375EB0E5" box="[812,879,2009,2028]" pageId="19" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
(A–C – aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J –
<taxonomicName id="4C5A4D507A3DFFADFD4306DC371FBF0E" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[714,814,2036,2055]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="248" phylum="Arthropoda" rank="species" species="wadicola" status="sp. nov.">
<emphasis id="B92EEAC17A3DFFADFD4306DC371FBF0E" box="[714,814,2036,2055]" italics="true" pageId="19" pageNumber="248">T. wadicola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A3DFFADFCBD06DC3746BF0E" box="[820,887,2036,2055]" pageId="19" pageNumber="248" rank="species">sp. nov.</taxonomicNameLabel>
(F–H – aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view).
</paragraph>
</caption>
<paragraph id="8BE536D37A3AFFAAFF1E006B305AB628" blockId="20.[151,760,190,369]" lastBlockId="20.[828,1436,190,289]" pageId="20" pageNumber="249">
<emphasis id="B92EEAC17A3AFFAAFF1E006B3583B65F" bold="true" box="[151,434,323,342]" pageId="20" pageNumber="249">Additional material examined.</emphasis>
<materialsCitation id="3B323C8E7A3AFFAAFE30006B36BFB678" ID-GBIF-Occurrence="3756360334" pageId="20" pageNumber="249">
<emphasis id="B92EEAC17A3AFFAAFE30006B36FFB65F" bold="true" box="[441,718,323,342]" pageId="20" pageNumber="249">
<collectingCountry id="F34D76437A3AFFAAFE30006B36FAB65F" box="[441,715,323,342]" name="United Arab Emirates" pageId="20" pageNumber="249">UNITED ARAB EMIRATES</collectingCountry>
:
</emphasis>
<specimenCount id="9D5CFD5A7A3AFFAAFD5F006B36C9B65E" box="[726,760,320,343]" pageId="20" pageNumber="249" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED4BAE167A3AFFAAFF14007634EEB678" box="[157,223,350,369]" country="Czech Republic" name="National Museum Prague" pageId="20" pageNumber="249">NMPC</collectionCode>
): Hatta, at light,
<date id="FFE410137A3AFFAAFEE7007635D4B678" box="[366,485,350,369]" pageId="20" pageNumber="249" value="2006-04-08" valueMax="2006-04-26" valueMin="2006-04-08">8.–26.iv.2006</date>
, lgt, A. van Harten.
</materialsCitation>
This female specimen was collected very close to the
<typeStatus id="54E188717A3AFFAAFB110196308CB7D8" box="[1176,1213,190,209]" pageId="20" pageNumber="249">type</typeStatus>
locality of the species and externally corresponds well with the
<collectingCountry id="F34D76437A3AFFAAFBF901F13095B7E5" box="[1136,1188,217,236]" name="Oman" pageId="20" pageNumber="249">Oman</collectingCountry>
specimen, we hence consider it conspecific. As no male is available from the locality, We are however not including it into the
<typeStatus id="54E188717A3AFFAAFB800026301FB628" box="[1033,1070,270,289]" pageId="20" pageNumber="249">type</typeStatus>
series.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3AFFABFCB500113581B49C" lastPageId="21" lastPageNumber="250" pageId="20" pageNumber="249" type="description">
<paragraph id="8BE536D37A3AFFABFCB500113581B49C" blockId="20.[828,1436,313,368]" lastBlockId="21.[151,760,190,1365]" lastPageId="21" lastPageNumber="250" pageId="20" pageNumber="249">
<emphasis id="B92EEAC17A3AFFAAFCB500113017B659" bold="true" box="[828,1062,313,336]" pageId="20" pageNumber="249">
Redescription.
<emphasis id="B92EEAC17A3AFFAAFC6400113017B659" bold="true" box="[1005,1062,313,336]" italics="true" pageId="20" pageNumber="249">Body</emphasis>
</emphasis>
length
<quantity id="4CA29B367A3AFFAAFBF500113131B659" box="[1148,1280,313,336]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="20" pageNumber="249" unit="mm" value="1.65" valueMax="1.7" valueMin="1.6">1.6–1.7 mm</quantity>
(
<typeStatus id="54E188717A3AFFAAFA9900113140B659" box="[1296,1393,313,336]" pageId="20" pageNumber="249" type="holotype">holotype</typeStatus>
<quantity id="4CA29B367A3AFFAAFAF30011375BB679" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="20" pageNumber="249" unit="mm" value="1.6">1.6 mm</quantity>
), maximum body width 0.9–1.0 mm (
<typeStatus id="54E188717A3AFFAAFA9900713140B679" box="[1296,1393,345,368]" pageId="20" pageNumber="249" type="holotype">holotype</typeStatus>
0.9 mm). Head and labrum black; pronotum uniformly dark brown; elytra uniformly dark brown; legs brown.
<emphasis id="B92EEAC17A3BFFABFD3501F636C9B7FC" bold="true" box="[700,760,222,245]" italics="true" pageId="21" pageNumber="250">Head</emphasis>
without microsculpture on interstices; punctation sparse, each puncture bearing pointed seta. Eyes separated by 3.3× the width of one eye in dorsal view.
<emphasis id="B92EEAC17A3BFFABFD0E001636C9B65C" bold="true" box="[647,760,318,341]" italics="true" pageId="21" pageNumber="250">Pronotum</emphasis>
with sparse setiferous punctation similar to that on head; interstices without microsculpture.
<emphasis id="B92EEAC17A3BFFABFD910056366CB69C" bold="true" box="[536,605,382,405]" italics="true" pageId="21" pageNumber="250">Elytra</emphasis>
with 10 striae sharply impressed except anteromedially (around scutellar shield) where neither striae nor serial punctures are visible; intervals weakly convex at midlength and near apex; interval punctation sparse, setiferous; interstices without microsculpture.
<emphasis id="B92EEAC17A3BFFABFE2B0336363EB53C" bold="true" box="[418,527,542,565]" italics="true" pageId="21" pageNumber="250">Aedeagus</emphasis>
(
<figureCitation id="13612A567A3BFFABFD950336365CB53C" box="[540,621,542,565]" captionStart="Fig" captionStartId="19.[151,182,2009,2028]" captionTargetBox="[159,1399,787,1981]" captionTargetId="figure-14@19.[151,1436,777,1989]" captionTargetPageId="19" captionText="Fig. 10. Male genitalia of Thysanarthria species, holotypes.A–E – T. persica sp. nov. (A–C – aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. wadicola sp. nov. (F–H – aedeagus in dorsal, lateral and ventral view; I–J – detail of median lobe and parameres in dorsal and ventral view)." figureDoi="http://doi.org/10.5281/zenodo.4488944" httpUri="https://zenodo.org/record/4488944/files/figure.png" pageId="21" pageNumber="250">Figs 10</figureCitation>
F–J)
<quantity id="4CA29B367A3BFFABFD2C033636C6B53C" box="[677,759,542,565]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="21" pageNumber="250" unit="mm" value="0.4">0.4 mm</quantity>
long. Phallobase slightly widened at bases of parameres, slightly narrower than parameres combined, weakly narrowing more basally in ventral/dorsal view, in lateral view phallobase c. as long as parameres, arcuate, and strongly constricted subbasally. Parameres wide basally, continually narrowing into prolonged pointed apex, outer face slightly sinuate, in lateral view very wide basally and very narrow apically. Median lobe very short, less than half the length of parameres, without distinct membranous part on the apex, without paired projections, gonopore circular, situated apically.
</paragraph>
</subSubSection>
<caption id="DF25665B7A3AFFAAFF1E06B53604BF13" ID-DOI="http://doi.org/10.5281/zenodo.4488946" ID-Zenodo-Dep="4488946" httpUri="https://zenodo.org/record/4488946/files/figure.png" pageId="20" pageNumber="249" startId="20.[151,182,1949,1968]" targetBox="[151,1436,401,1931]" targetPageId="20">
<paragraph id="8BE536D37A3AFFAAFF1E06B53604BF13" blockId="20.[151,1437,1949,2074]" pageId="20" pageNumber="249">
Fig. 11. Distribution of
<taxonomicName id="4C5A4D507A3AFFAAFEE806B535EAB0B9" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[353,475,1949,1968]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A3AFFAAFEE806B535EAB0B9" box="[353,475,1949,1968]" italics="true" pageId="20" pageNumber="249">Thysanarthria</emphasis>
</taxonomicName>
and American
<taxonomicName id="4C5A4D507A3AFFAAFDD406B536FDB0B9" authorityName="Stephens" authorityYear="1835" box="[605,716,1949,1968]" class="Insecta" family="Hydrophilidae" genus="Chaetarthria" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A3AFFAAFDD406B536FDB0B9" box="[605,716,1949,1968]" italics="true" pageId="20" pageNumber="249">Chaetarthria</emphasis>
</taxonomicName>
in Asia (A–B) and Near East (C), and examples of habitats of
<taxonomicName id="4C5A4D507A3AFFAAFB6A06B5316CB0B9" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[1251,1373,1949,1968]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="249" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEAC17A3AFFAAFB6A06B5316CB0B9" box="[1251,1373,1949,1968]" italics="true" pageId="20" pageNumber="249">Thysanarthria</emphasis>
</taxonomicName>
(D–F). Localities: D – India, Madhya Pradesh, Bhadhua Chora stream ca. 10 km E of Matkuli Jhir, habitat of
<taxonomicName id="4C5A4D507A3AFFAAFB85069F313BB0C3" authority="Hebauer, 2001" authorityName="Hebauer" authorityYear="2001" box="[1036,1290,1975,1994]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="249" phylum="Arthropoda" rank="species" species="ceylonensis">
<emphasis id="B92EEAC17A3AFFAAFB85069F30B5B0C3" box="[1036,1156,1975,1994]" italics="true" pageId="20" pageNumber="249">T. ceylonensis</emphasis>
<bibRefCitation id="EFCB4B227A3AFFAAFB03069F313BB0C3" author="Hebauer" box="[1162,1290,1975,1994]" firstAuthor="Hebauer" pageId="20" pageNumber="249" pagination="393 - 400" refId="ref18120" refString="HEBAUER F. 2001: The species of the genus Thysanarthria d'Orchymont, 1926 (Coleoptera: Hydrophilidae). Beitrage zur Entomologie 51: 393 - 400." type="journal article" year="2001">Hebauer, 2001</bibRefCitation>
</taxonomicName>
; E – Republic of South Africa, Western Cape, 8 km NEE of Stanford, habitat of
<taxonomicName id="4C5A4D507A3AFFAAFD2E06FA37AAB0EC" authority="(Regimber, 1903)" baseAuthorityName="Regimber" baseAuthorityYear="1903" box="[679,923,2002,2021]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="249" phylum="Arthropoda" rank="species" species="atriceps">
<emphasis id="B92EEAC17A3AFFAAFD2E06FA3731B0EC" box="[679,768,2002,2021]" italics="true" pageId="20" pageNumber="249">T. atriceps</emphasis>
(Régimber, 1903)
</taxonomicName>
, collecting spot marked by an arrow; F–G – Taiwan, Wufeng distr., 4.3 km SEE of Chaoyang University of Technology, habitat of
<taxonomicName id="4C5A4D507A3AFFAAFD6306C53714BF09" authority="Fikáček &amp; Liu, 2019" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[746,805,2029,2048]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="249" phylum="Arthropoda" rank="species" species="chui" status="sp. nov.">
<emphasis id="B92EEAC17A3AFFAAFD6306C53714BF09" box="[746,805,2029,2048]" italics="true" pageId="20" pageNumber="249">T. chui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21D57BA7A3AFFAAFCA306C5375FBF09" box="[810,878,2029,2048]" pageId="20" pageNumber="249" rank="species">sp. nov.</taxonomicNameLabel>
(G – detail of sandy shore with one alive beetle). Photo D by M. Jäch, photo E by M. Fikáček, F–G by H.-C. Liu.
</paragraph>
</caption>
<subSubSection id="C34065587A3BFFABFF1E02B63601B31C" pageId="21" pageNumber="250" type="diagnosis">
<paragraph id="8BE536D37A3BFFABFF1E02B63601B31C" blockId="21.[151,760,190,1365]" pageId="21" pageNumber="250">
<emphasis id="B92EEAC17A3BFFABFF1E02B635A6B4BC" bold="true" box="[151,407,926,949]" pageId="21" pageNumber="250">Differential diagnosis.</emphasis>
<taxonomicName id="4C5A4D507A3BFFABFE2802B63699B4BC" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[417,680,926,949]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="250" phylum="Arthropoda" rank="species" species="wadicola">
<emphasis id="B92EEAC17A3BFFABFE2802B63699B4BC" box="[417,680,926,949]" italics="true" pageId="21" pageNumber="250">Thysanarthria wadicola</emphasis>
</taxonomicName>
differs from most species of the genus except
<taxonomicName id="4C5A4D507A3BFFABFDB702963698B4DC" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[574,681,958,981]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="250" phylum="Arthropoda" rank="species" species="persica">
<emphasis id="B92EEAC17A3BFFABFDB702963698B4DC" box="[574,681,958,981]" italics="true" pageId="21" pageNumber="250">T. persica</emphasis>
</taxonomicName>
in uniformly dark brown pronotum and elytra. For differences from
<taxonomicName id="4C5A4D507A3BFFABFF5B02D6350AB31C" authorityName="Fikáček &amp; Liu" authorityYear="2019" box="[210,315,1022,1045]" class="Insecta" family="Hydrophilidae" genus="Thysanarthria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="250" phylum="Arthropoda" rank="species" species="persica">
<emphasis id="B92EEAC17A3BFFABFF5B02D6350AB31C" box="[210,315,1022,1045]" italics="true" pageId="21" pageNumber="250">T. persica</emphasis>
</taxonomicName>
see under that species.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3BFFABFF1E053635DDB39C" pageId="21" pageNumber="250" type="etymology">
<paragraph id="8BE536D37A3BFFABFF1E053635DDB39C" blockId="21.[151,760,190,1365]" pageId="21" pageNumber="250">
<emphasis id="B92EEAC17A3BFFABFF1E05363529B33C" bold="true" box="[151,280,1054,1077]" pageId="21" pageNumber="250">Etymology.</emphasis>
The species name is a combination of
<emphasis id="B92EEAC17A3BFFABFD4D053636C9B33C" box="[708,760,1054,1077]" italics="true" pageId="21" pageNumber="250">wadi</emphasis>
(an Arabic word for seasonally dried-up riverbed) and the suffix
<emphasis id="B92EEAC17A3BFFABFE810576350EB37C" box="[264,319,1118,1141]" italics="true" pageId="21" pageNumber="250">-cola</emphasis>
derived from the word
<emphasis id="B92EEAC17A3BFFABFDC2057636BFB37C" box="[587,654,1118,1141]" italics="true" pageId="21" pageNumber="250">incola</emphasis>
(Latin, = inhabitant). Noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3BFFABFF1E05B635E3B3FC" pageId="21" pageNumber="250" type="biology_ecology">
<paragraph id="8BE536D37A3BFFABFF1E05B635E3B3FC" blockId="21.[151,760,190,1365]" pageId="21" pageNumber="250">
<emphasis id="B92EEAC17A3BFFABFF1E05B634C2B3BC" bold="true" box="[151,243,1182,1205]" pageId="21" pageNumber="250">Biology.</emphasis>
Part of the specimens was collected at margins of small rocky pools at the side of a drying-up stream (see
<bibRefCitation id="EFCB4B227A3BFFABFF1E05F6356AB3FC" author="RIBERA" box="[151,347,1246,1269]" etAl="et al." firstAuthor="RIBERA" pageId="21" pageNumber="250" pagination="253 - 272" refId="ref18641" refString="RIBERA I., HERNANDO C. &amp; CIESLAK A. 2019: Aquatic Coleoptera of North Oman, with description of new species of Hydraenidae and Hydrophilidae. Acta Entomologica Musei Nationalis Pragae 59: 253 - 272." type="journal article" year="2019">RIBERA et al. 2019</bibRefCitation>
:
<figureCitation id="13612A567A3BFFABFEEC05F635F7B3FC" box="[357,454,1246,1269]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="14.[151,182,1334,1353]" captionStartId-1="16.[151,182,1976,1995]" captionTargetBox-0="[177,1428,199,1315]" captionTargetBox-1="[204,1420,202,1944]" captionTargetId-0="figure-13@14.[151,1436,186,1319]" captionTargetId-1="figure-108@16.[162,1439,189,1956]" captionTargetPageId-0="14" captionTargetPageId-1="16" captionText-0="Fig. 7. Male genitalia of Thysanarthria championi (Knisch, 1924).A–E – lectotype of Chaetarthria championi Knisch, 1924 (A–C – whole aedeagus in dorsal, lateral and ventral view; D – detail of median lobe and parameres in dorsal view; E – same after KOH treatment, with fully everted internal sac). F–J – lectotype of Chaetarthriomorphus sulcatus Chiesa, 1967 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view)." captionText-1="Fig. 8. Male genitalia of Thysanarthria species, holotypes.A–E – T. chui sp. nov.(A–C – whole aedeagus in dorsal, lateral and ventral view; D–E – detail of median lobe and parameres in dorsal and ventral view). F–J – T. hongsonensis Hebauer, 2001 (F–H – whole aedeagus in dorsal, lateral and ventral view; I–J – details of parameres and median lobe in dorsal and ventral view). K–O – T. madurensis Hebauer, 2001 (K–M – whole aedeagus in dorsal, lateral and ventral view; N–O – details of median lobe and parameres in dorsal and ventral view)." figureDoi-0="http://doi.org/10.5281/zenodo.4488938" figureDoi-1="http://doi.org/10.5281/zenodo.4488940" httpUri-0="https://zenodo.org/record/4488938/files/figure.png" httpUri-1="https://zenodo.org/record/4488940/files/figure.png" pageId="21" pageNumber="250">Figs 7–8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34065587A3BFFABFF1E05D63552B25C" pageId="21" pageNumber="250" type="distribution">
<paragraph id="8BE536D37A3BFFABFF1E05D63552B25C" blockId="21.[151,760,190,1365]" pageId="21" pageNumber="250">
<emphasis id="B92EEAC17A3BFFABFF1E05D6351BB21C" bold="true" box="[151,298,1278,1301]" pageId="21" pageNumber="250">Distribution.</emphasis>
The species is so far known only from few closely situated localities on the northeastern tip of the Arabian Peninsula.
</paragraph>
</subSubSection>
</treatment>
</document>
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