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<document id="8354F7975314A4D9E0F27FD9A3189B2C" ID-DOI="10.11646/zootaxa.4751.3.6" ID-GBIF-Dataset="0fd5ecf7-48de-4920-8ef5-93b6537f735e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3714753" ID-ZooBank="74A7039B-1B5B-4E67-8CF2-E4D57E44C06B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584523130345" checkinUser="plazi" docAuthor="Kirchner, Sandra, Kruckenhauser, Luise, Pichler, Arthur, Borkenhagen, Kai &amp; Freyhof, Jörg" docDate="2020" docId="03ED87FCBC58FF93FF41FF40FB418079" docLanguage="en" docName="zootaxa.4751.3.6.pdf" docOrigin="Zootaxa 4751 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Garra barreimiae Fowler &amp; Steinitz 1956" docType="treatment" docVersion="5" lastPageNumber="528" masterDocId="FFD4FF84BC5EFF94FFD6FFD9FFBD8570" masterDocTitle="Revision of the Garra species of the Hajar Mountains in Oman and the United Arab Emirates with the description of two new species (Teleostei: Cyprinidae)" masterLastPageNumber="545" masterPageNumber="521" pageNumber="527" updateTime="1698729799427" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="ADD0A85A37FF2A793F99CBE221953782" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9C81B012C54A85A49C1BB90E7D8E1ED4">Revision of the Garra species of the Hajar Mountains in Oman and the United Arab Emirates with the description of two new species (Teleostei: Cyprinidae)</mods:title>
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<mods:namePart id="95D27149F748D6723533973AB1D07B1B">Kirchner, Sandra</mods:namePart>
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<mods:namePart id="5B4FB709EEA1222BD83918981CC037AD">Kruckenhauser, Luise</mods:namePart>
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<mods:namePart id="02E863B6733364B294E3F2B8AC03D9B9">Pichler, Arthur</mods:namePart>
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<mods:namePart id="089D1A2A6056BC9CF368687F48E1BEC7">Borkenhagen, Kai</mods:namePart>
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<mods:namePart id="E7AE309B1B20DDDBB64B492B381CC886">Freyhof, Jörg</mods:namePart>
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<mods:title id="90BCBC7AAD96C576492498566ECB59A2">Zootaxa</mods:title>
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<mods:date id="2B902F12D5AB2126E2C49753A372035A">2020</mods:date>
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<mods:number id="3CFE375806EF2E395C6D5EF45C312393">2020-03-18</mods:number>
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<mods:number id="A28C46EAFDD1141E690260D14E7A9A79">4751</mods:number>
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<treatment id="03ED87FCBC58FF93FF41FF40FB418079" ID-DOI="http://doi.org/10.5281/zenodo.3718279" ID-GBIF-Taxon="162630018" ID-Zenodo-Dep="3718279" LSID="urn:lsid:plazi:treatment:03ED87FCBC58FF93FF41FF40FB418079" httpUri="http://treatment.plazi.org/id/03ED87FCBC58FF93FF41FF40FB418079" lastPageId="7" lastPageNumber="528" pageId="6" pageNumber="527">
<subSubSection id="C35E6561BC58FF92FF41FF40FD2785C3" box="[151,666,153,180]" pageId="6" pageNumber="527" type="nomenclature">
<paragraph id="8BFB36EABC58FF92FF41FF40FD2785C3" blockId="6.[151,666,153,215]" box="[151,666,153,180]" pageId="6" pageNumber="527">
<heading id="D0B38186BC58FF92FF41FF40FD2785C3" bold="true" box="[151,666,153,180]" fontSize="11" level="1" pageId="6" pageNumber="527" reason="1">
<taxonomicName id="4C444D69BC58FF92FF41FF40FD2785C3" ID-CoL="3FBGY" authority="Fowler &amp; Steinitz, 1956" authorityName="Fowler &amp; Steinitz" authorityYear="1956" box="[151,666,153,180]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="527" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="B930EAF8BC58FF92FF41FF40FD2785C3" bold="true" box="[151,666,153,180]" pageId="6" pageNumber="527">
<emphasis id="B930EAF8BC58FF92FF41FF40FED185C4" bold="true" box="[151,364,153,180]" italics="true" pageId="6" pageNumber="527">Garra barreimiae</emphasis>
Fowler &amp; Steinitz, 1956
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35E6561BC58FF92FF41FF64FF5685A7" box="[151,235,189,215]" pageId="6" pageNumber="527" type="description">
<paragraph id="8BFB36EABC58FF92FF41FF64FF5685A7" blockId="6.[151,666,153,215]" box="[151,235,189,215]" pageId="6" pageNumber="527">
(
<figureCitation id="137F2A6FBC58FF92FF76FF64FF5F85A7" box="[160,226,189,215]" captionStart="FIGURE 3" captionStartId="6.[151,250,1884,1909]" captionTargetBox="[151,1436,509,1858]" captionTargetId="figure@6.[151,1436,509,1858]" captionTargetPageId="6" captionText="FIGURE 3. Garra barreimiae, from top: FSJF 4077, 50 mm SL; Wadi Ajran, Oman; FSJF 4073, 49 mm SL; Al Juwayf, Oman; FSJF 4075, 74 mm SL; Wadi Qahfi, Oman." figureDoi="http://doi.org/10.5281/zenodo.3714757" httpUri="https://zenodo.org/record/3714757/files/figure.png" pageId="6" pageNumber="527">Fig. 3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35E6561BC58FF92FF41FEDBFC158427" pageId="6" pageNumber="527" type="reference_group">
<paragraph id="8BFB36EABC58FF92FF41FEDBFBE9846A" blockId="6.[151,1173,257,343]" box="[151,1108,257,282]" pageId="6" pageNumber="527">
<taxonomicName id="4C444D69BC58FF92FF41FEDBFDC8846A" ID-CoL="3FBGY" authority="Fowler &amp; Steinitz, 1956: 262" authorityName="Fowler &amp; Steinitz" authorityPageNumber="262" authorityYear="1956" box="[151,629,257,282]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="527" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="B930EAF8BC58FF92FF41FEDBFEF0846A" box="[151,333,258,282]" italics="true" pageId="6" pageNumber="527">Garra barreimiae</emphasis>
Fowler &amp; Steinitz, 1956:262
</taxonomicName>
(
<typeStatus id="54FF8848BC58FF92FD54FEDBFD12846A" box="[642,687,258,282]" pageId="6" pageNumber="527">type</typeStatus>
locality:
<collectingRegion id="4980F808BC58FF92FCD9FED8FCDF846A" box="[783,866,257,282]" country="Oman" name="Al Buraymi" pageId="6" pageNumber="527">Buraimi</collectingRegion>
[=
<collectingRegion id="4980F808BC58FF92FCABFED8FC4A846A" box="[893,1015,257,282]" country="Oman" name="Al Buraymi" pageId="6" pageNumber="527">Al Buraimi</collectingRegion>
],
<collectingCountry id="F353767ABC58FF92FBD1FEDBFBF4846A" box="[1031,1097,258,282]" name="Oman" pageId="6" pageNumber="527">Oman</collectingCountry>
);
</paragraph>
<paragraph id="8BFB36EABC58FF92FF41FEF8FB288448" blockId="6.[151,1173,257,343]" box="[151,1173,288,313]" pageId="6" pageNumber="527">
<taxonomicName id="4C444D69BC58FF92FF41FEF8FCB48449" ID-CoL="7K4PT" authority="Banister &amp; Clarke, 1977: 136" authorityName="Banister &amp; Clarke" authorityPageNumber="136" authorityYear="1977" box="[151,777,288,313]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="527" phylum="Chordata" rank="subSpecies" species="barreimiae" subSpecies="shawkahensis">
<emphasis id="B930EAF8BC58FF92FF41FEF8FE638448" box="[151,478,288,312]" italics="true" pageId="6" pageNumber="527">Garra barreimiae shawkahensis</emphasis>
<bibRefCitation id="EFD54B1BBC58FF92FE32FEF9FCB48449" author="Banister, K. E. &amp; Clarke, M. A." box="[484,777,288,313]" pageId="6" pageNumber="527" pagination="111 - 154" refId="ref11266" refString="Banister, K. E. &amp; Clarke, M. A. (1977) The freshwater fishes of the Arabian Peninsula. The Scientific Results of the Oman Flora and Fauna Survey 1975. Journal Oman Studies, 1, 111 - 154." type="journal article" year="1977">Banister &amp; Clarke, 1977:136</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54FF8848BC58FF92FCC1FEF9FCF98448" box="[791,836,288,312]" pageId="6" pageNumber="527">type</typeStatus>
locality: Wadi Shawkah,
<collectingCountry id="F353767ABC58FF92FB9EFEF9FB378448" box="[1096,1162,288,312]" name="Oman" pageId="6" pageNumber="527">Oman</collectingCountry>
);
</paragraph>
<paragraph id="8BFB36EABC58FF92FF41FEE6FC158427" blockId="6.[151,1173,257,343]" box="[151,936,318,343]" pageId="6" pageNumber="527">
<taxonomicName id="4C444D69BC58FF92FF41FEE6FEF08427" ID-CoL="3FBGY" authorityName="Fowler &amp; Steinitz" authorityYear="1956" box="[151,333,319,343]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="527" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="B930EAF8BC58FF92FF41FEE6FEF08427" box="[151,333,319,343]" italics="true" pageId="6" pageNumber="527">Garra barreimiae</emphasis>
</taxonomicName>
West Clade (
<bibRefCitation id="EFD54B1BBC58FF92FE05FEE7FD2C8427" author="Pichler, A. &amp; Ahnelt, H. &amp; Kirchner, S. &amp; Sattmann, H. &amp; Haring, E. &amp; Handschuh, S. &amp; Freyhof, J. &amp; Victor, R. &amp; Kruckenhauser, L." box="[467,657,318,343]" pageId="6" pageNumber="527" pagination="1053 - 1065" refId="ref13355" refString="Pichler, A., Ahnelt, H., Kirchner, S., Sattmann, H., Haring, E., Handschuh, S., Freyhof, J., Victor, R. &amp; Kruckenhauser, L. (2018) The morphological diversity of Garra barreimiae (Teleostei: Cyprinidae). Environmental Biology of Fishes, 101, 1053 - 1065. https: // doi. org / 10.1007 / s 10641 - 018 - 0758 - 7" type="journal article" year="2018">
Pichler
<emphasis id="B930EAF8BC58FF92FDF5FE99FDEB8427" box="[547,598,319,343]" italics="true" pageId="6" pageNumber="527">et al.</emphasis>
2018
</bibRefCitation>
, Kirchner
<emphasis id="B930EAF8BC58FF92FD2DFE99FC958427" box="[763,808,319,343]" italics="true" pageId="6" pageNumber="527">et al</emphasis>
. submitted).
</paragraph>
</subSubSection>
<subSubSection id="C35E6561BC58FF93FF41FE5CFE02877E" lastPageId="7" lastPageNumber="528" pageId="6" pageNumber="527" type="materials_examined">
<paragraph id="8BFB36EABC58FF92FF41FE5CFC808498" blockId="6.[151,1436,389,488]" pageId="6" pageNumber="527">
<emphasis id="B930EAF8BC58FF92FF41FE5CFE3C84D0" bold="true" box="[151,385,389,416]" pageId="6" pageNumber="527">Material examined.</emphasis>
<materialsCitation id="3B2C3CB7BC58FF92FE51FE5CFEBD84B4" ID-GBIF-Occurrence="2577370129" collectionCode="FSJF" country="Oman" latitude="24.024" location="Wadi Ajran" longLatPrecision="74" longitude="56.196" municipality="Ajran" pageId="6" pageNumber="527" specimenCode="FSJF 4077, 26" specimenCount="1">
<specimenCode id="DBE29E91BC58FF92FE51FE5CFDBE84EF" box="[391,515,389,415]" collectionCode="FSJF" pageId="6" pageNumber="527">FSJF 4077</specimenCode>
,
<specimenCode id="DBE29E91BC58FF92FDD8FE5CFD9184EF" box="[526,556,389,415]" collectionCode="FSJF" pageId="6" pageNumber="527">26</specimenCode>
,
<quantity id="4CBC9B0FBC58FF92FDEEFE5CFD0C84D0" box="[568,689,389,416]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="5.0" metricValueMin="2.5" pageId="6" pageNumber="527" unit="mm" value="37.5" valueMax="50.0" valueMin="25.0">25–50 mm</quantity>
SL;
<collectingCountry id="F353767ABC58FF92FD31FE5CFC8D84EF" box="[743,816,389,415]" name="Oman" pageId="6" pageNumber="527">Oman</collectingCountry>
:
<location id="8E9B6031BC58FF92FCECFE5CFC0784EF" LSID="urn:lsid:plazi:treatment:03ED87FCBC58FF93FF41FF40FB418079:8E9B6031BC58FF92FCECFE5CFC0784EF" box="[826,954,389,415]" country="Oman" latitude="24.024" longLatPrecision="74" longitude="56.196" municipality="Ajran" name="Wadi Ajran" pageId="6" pageNumber="527">Wadi Ajran</location>
about
<quantity id="4CBC9B0FBC58FF92FBD0FE5CFB8284EF" box="[1030,1087,389,415]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="6" pageNumber="527" unit="km" value="3.0">3 km</quantity>
south-east of
<collectingMunicipality id="6B9FAC90BC58FF92FB0DFE5CFAA384EF" box="[1243,1310,389,415]" pageId="6" pageNumber="527">Ajran</collectingMunicipality>
,
<geoCoordinate id="EE70502DBC58FF92FAFFFE5CFA2584EF" box="[1321,1432,389,415]" degrees="24.024" direction="north" orientation="latitude" pageId="6" pageNumber="527" precision="55" value="24.024">24.024°N</geoCoordinate>
,
<geoCoordinate id="EE70502DBC58FF92FF41FE70FEBD84B4" box="[151,256,425,452]" degrees="56.196" direction="east" orientation="longitude" pageId="6" pageNumber="527" precision="55" value="56.196">56.196°E</geoCoordinate>
</materialsCitation>
.—
<materialsCitation id="3B2C3CB7BC58FF92FEF7FE70FB4484B4" ID-GBIF-Occurrence="2577370123" box="[289,1273,425,452]" collectionCode="FSJF" country="Oman" latitude="24.549" location="Falaj" longLatPrecision="74" longitude="56.105" municipality="Al Juwayf" pageId="6" pageNumber="527" specimenCode="FSJF 4073, 4" specimenCount="1">
<specimenCode id="DBE29E91BC58FF92FEF7FE70FE2284B3" box="[289,415,425,452]" collectionCode="FSJF" pageId="6" pageNumber="527">FSJF 4073</specimenCode>
,
<specimenCode id="DBE29E91BC58FF92FE7FFE70FE0484B3" box="[425,441,425,451]" collectionCode="FSJF" pageId="6" pageNumber="527">4</specimenCode>
,
<quantity id="4CBC9B0FBC58FF92FE13FE70FD8084B4" box="[453,573,425,452]" metricMagnitude="-2" metricUnit="m" metricValue="4.45" metricValueMax="4.9" metricValueMin="4.0" pageId="6" pageNumber="527" unit="mm" value="44.5" valueMax="49.0" valueMin="40.0">40–49 mm</quantity>
SL;
<collectingCountry id="F353767ABC58FF92FDA4FE70FD0684B3" box="[626,699,425,451]" name="Oman" pageId="6" pageNumber="527">Oman</collectingCountry>
:
<location id="8E9B6031BC58FF92FD13FE70FD4084B3" LSID="urn:lsid:plazi:treatment:03ED87FCBC58FF93FF41FF40FB418079:8E9B6031BC58FF92FD13FE70FD4084B3" box="[709,765,425,451]" country="Oman" latitude="24.549" longLatPrecision="74" longitude="56.105" municipality="Al Juwayf" name="Falaj" pageId="6" pageNumber="527">Falaj</location>
at mosque in
<collectingMunicipality id="6B9FAC90BC58FF92FC40FE70FBB084B3" box="[918,1037,425,451]" pageId="6" pageNumber="527">Al Juwayf</collectingMunicipality>
,
<geoCoordinate id="EE70502DBC58FF92FBC1FE70FB3B84B4" box="[1047,1158,425,452]" degrees="24.549" direction="north" orientation="latitude" pageId="6" pageNumber="527" precision="55" value="24.549">24.549°N</geoCoordinate>
,
<geoCoordinate id="EE70502DBC58FF92FB47FE70FB4484B4" box="[1169,1273,425,452]" degrees="56.105" direction="east" orientation="longitude" pageId="6" pageNumber="527" precision="55" value="56.105">56.105°E</geoCoordinate>
</materialsCitation>
.—
<materialsCitation id="3B2C3CB7BC58FF92FACDFE70FC848498" ID-GBIF-Occurrence="2577370086" collectionCode="FSJF" country="Oman" latitude="24.713" location="Wadi Qahfi" longLatPrecision="74" longitude="56.186" pageId="6" pageNumber="527" specimenCode="FSJF 4075, 18" specimenCount="1">
<specimenCode id="DBE29E91BC58FF92FACDFE70FA2584B4" box="[1307,1432,425,452]" collectionCode="FSJF" pageId="6" pageNumber="527">FSJF 4075</specimenCode>
,
<specimenCode id="DBE29E91BC58FF92FF41FE14FF088497" box="[151,181,461,487]" collectionCode="FSJF" pageId="6" pageNumber="527">18</specimenCode>
,
<quantity id="4CBC9B0FBC58FF92FF17FE14FE878498" box="[193,314,461,488]" metricMagnitude="-2" metricUnit="m" metricValue="5.55" metricValueMax="7.4" metricValueMin="3.7" pageId="6" pageNumber="527" unit="mm" value="55.5" valueMax="74.0" valueMin="37.0">37–74 mm</quantity>
SL;
<collectingCountry id="F353767ABC58FF92FEA6FE14FE048497" box="[368,441,461,487]" name="Oman" pageId="6" pageNumber="527">Oman</collectingCountry>
:
<location id="8E9B6031BC58FF92FE15FE14FDFA8497" LSID="urn:lsid:plazi:treatment:03ED87FCBC58FF93FF41FF40FB418079:8E9B6031BC58FF92FE15FE14FDFA8497" box="[451,583,461,487]" country="Oman" latitude="24.713" longLatPrecision="74" longitude="56.186" name="Wadi Qahfi" pageId="6" pageNumber="527">Wadi Qahfi</location>
,
<geoCoordinate id="EE70502DBC58FF92FD84FE14FD7C8497" box="[594,705,461,487]" degrees="24.713" direction="north" orientation="latitude" pageId="6" pageNumber="527" precision="55" value="24.713">24.713°N</geoCoordinate>
,
<geoCoordinate id="EE70502DBC58FF92FD1AFE14FC848498" box="[716,825,461,488]" degrees="56.186" direction="east" orientation="longitude" pageId="6" pageNumber="527" precision="55" value="56.186">56.186°E</geoCoordinate>
</materialsCitation>
.
</paragraph>
<caption id="DF3B6662BC58FF92FF41F885FDED82E8" ID-DOI="http://doi.org/10.5281/zenodo.3714757" ID-Zenodo-Dep="3714757" httpUri="https://zenodo.org/record/3714757/files/figure.png" pageId="6" pageNumber="527" startId="6.[151,250,1884,1909]" targetBox="[151,1436,509,1858]" targetPageId="6">
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<emphasis id="B930EAF8BC58FF92FF41F885FEAF8205" bold="true" box="[151,274,1884,1909]" pageId="6" pageNumber="527">FIGURE 3.</emphasis>
<taxonomicName id="4C444D69BC58FF92FECEF884FE708204" authorityName="Fowler &amp; Steinitz" authorityYear="1956" box="[280,461,1884,1908]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="527" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="B930EAF8BC58FF92FECEF884FE708204" box="[280,461,1884,1908]" italics="true" pageId="6" pageNumber="527">Garra barreimiae</emphasis>
</taxonomicName>
, from top: FSJF 4077, 50 mm SL; Wadi Ajran, Oman; FSJF 4073, 49 mm SL; Al Juwayf, Oman; FSJF 4075, 74 mm SL; Wadi Qahfi, Oman.
</paragraph>
</caption>
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<emphasis id="B930EAF8BC59FF93FF11FF41FD7685C3" bold="true" box="[199,715,152,179]" pageId="7" pageNumber="528">Material used in molecular genetic analysis.</emphasis>
NMW-100045;
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3235;
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:
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about
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south-east of
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,
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(
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,
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,
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).— NMW-100036
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;
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(
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accession number:
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).—NMW-100037
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;
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:
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,
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,
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(
<collectorName id="26B1533CBC59FF93FDB3FED8FD6C846B" box="[613,721,257,283]" pageId="7" pageNumber="528">GenBank</collectorName>
accession number:
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).—NMW-100044
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;
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</materialsCitation>
;
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<collectingCountry id="F353767ABC59FF93FF35FEFDFE91844E" box="[227,300,292,318]" name="Oman" pageId="7" pageNumber="528">Oman</collectingCountry>
:
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at mosque in
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,
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,
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(
<collectorName id="26B1533CBC59FF93FC66FEFAFBA1844D" box="[944,1052,291,317]" pageId="7" pageNumber="528">GenBank</collectorName>
accession numbers:
<accessionNumber id="9417AB09BC59FF93FAC2FEFDFA25844E" box="[1300,1432,292,318]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN830407" pageId="7" pageNumber="528">MN830407</accessionNumber>
,
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,
<accessionNumber id="9417AB09BC59FF93FEE6FE9FFE098411" box="[304,436,326,353]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/MN830864" pageId="7" pageNumber="528">MN830864</accessionNumber>
).—NMW-100046
</materialsCitation>
;
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<collectingCountry id="F353767ABC59FF93FD4EFE9FFD5C8410" box="[664,737,326,352]" name="Oman" pageId="7" pageNumber="528">Oman</collectingCountry>
:
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,
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,
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(
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accession number:
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830406
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).—
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<specimenCode id="DBE29E91BC59FF93FE15FEB0FDF884F3" box="[451,581,361,387]" collectionCode="DNA-" pageId="7" pageNumber="528">DNA-3233</specimenCode>
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;
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<collectingCountry id="F353767ABC59FF93FD83FEB0FD2384F3" box="[597,670,361,387]" name="Oman" pageId="7" pageNumber="528">Oman</collectingCountry>
:
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,
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,
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(
<collectorName id="26B1533CBC59FF93FB92FEB0FB0D84F3" box="[1092,1200,361,387]" pageId="7" pageNumber="528">GenBank</collectorName>
accession numbers:
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,
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).—
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<collectingCountry id="F353767ABC59FF93FD7DFE55FD5484D6" box="[683,745,396,422]" name="United Arab Emirates" pageId="7" pageNumber="528">UAE</collectingCountry>
:
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, in
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of
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,
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,
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(
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accession numbers:
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,
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,
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,
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).—
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;
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<collectingCountry id="F353767ABC59FF93FF41FE08FF68849B" box="[151,213,465,491]" name="United Arab Emirates" pageId="7" pageNumber="528">UAE</collectingCountry>
:
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of Al-Fujaira,
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,
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. (
<collectorName id="26B1533CBC59FF93FC6EFE08FB99849B" box="[952,1060,465,491]" pageId="7" pageNumber="528">GenBank</collectorName>
accession numbers:
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,
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,
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="C35E6561BC59FF93FF11FDCCFAA28636" pageId="7" pageNumber="528" type="diagnosis">
<paragraph id="8BFB36EABC59FF93FF11FDCCFAA28636" blockId="7.[151,1437,152,1289]" pageId="7" pageNumber="528">
<emphasis id="B930EAF8BC59FF93FF11FDCCFEFF8740" bold="true" box="[199,322,533,560]" pageId="7" pageNumber="528">Diagnosis.</emphasis>
<taxonomicName id="4C444D69BC59FF93FE91FDCEFDB38740" authorityName="Fowler &amp; Steinitz" authorityYear="1956" box="[327,526,534,561]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="B930EAF8BC59FF93FE91FDCEFDB38740" box="[327,526,534,561]" italics="true" pageId="7" pageNumber="528">Garra barreimiae</emphasis>
</taxonomicName>
is distinguished from the other species of
<taxonomicName id="4C444D69BC59FF93FC08FDCEFB998741" box="[990,1060,535,561]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="genus">
<emphasis id="B930EAF8BC59FF93FC08FDCEFB998741" box="[990,1060,535,561]" italics="true" pageId="7" pageNumber="528">Garra</emphasis>
</taxonomicName>
in the Hajar Mountains by having 15–18 gill rakers on the lower limb of the first gill arch (vs. 10–14) and an orange spot present at the upper opercle (vs. absent). It also has a strongly mottled flank pattern with individual or series of orange midlateral scales (vs. orange scales absent in
<taxonomicName id="4C444D69BC59FF93FE73FDA6FD8087E8" authorityName="Krupp" authorityYear="1988" box="[421,573,638,665]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="gallagheri">
<emphasis id="B930EAF8BC59FF93FE73FDA6FD8087E8" box="[421,573,638,665]" italics="true" pageId="7" pageNumber="528">G. gallagheri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C444D69BC59FF93FD9AFDA6FD5387E8" authorityName="Banister &amp; Clarke" authorityYear="1977" box="[588,750,638,665]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="B930EAF8BC59FF93FD9AFDA6FD5387E8" box="[588,750,638,665]" italics="true" pageId="7" pageNumber="528">G. longipinnis</emphasis>
</taxonomicName>
and many individuals of
<taxonomicName id="4C444D69BC59FF93FBC4FDA6FB3B87E8" authorityName="Kirchner &amp; Kruckenhauser &amp; Pichler &amp; Borkenhagen &amp; Freyhof" authorityYear="2020" box="[1042,1158,638,665]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="shamal">
<emphasis id="B930EAF8BC59FF93FBC4FDA6FB3B87E8" box="[1042,1158,638,665]" italics="true" pageId="7" pageNumber="528">G. shamal</emphasis>
</taxonomicName>
), and bold black middle caudal-fin rays with white membranes (vs. middle caudal-fin rays and membranes slightly darker or the same colour as remaining caudal-fin rays and membranes in
<taxonomicName id="4C444D69BC59FF93FD78FD1DFC9C87AE" authorityName="Kirchner &amp; Kruckenhauser &amp; Pichler &amp; Borkenhagen &amp; Freyhof" authorityYear="2020" box="[686,801,708,734]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="shamal">
<emphasis id="B930EAF8BC59FF93FD78FD1DFC9C87AE" box="[686,801,708,734]" italics="true" pageId="7" pageNumber="528">G. shamal</emphasis>
</taxonomicName>
).
<taxonomicName id="4C444D69BC59FF93FCEEFD1DFC6887AE" authorityName="Fowler &amp; Steinitz" authorityYear="1956" box="[824,981,708,734]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="B930EAF8BC59FF93FCEEFD1DFC6887AE" box="[824,981,708,734]" italics="true" pageId="7" pageNumber="528">G. barreimiae</emphasis>
</taxonomicName>
is further distinguished by the dark margins of the flank scales below the lateral line being very indistinct or absent (vs. present in
<taxonomicName id="4C444D69BC59FF93FB4EFD3EFA858670" authorityName="Banister &amp; Clarke" authorityYear="1977" box="[1176,1336,742,769]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="B930EAF8BC59FF93FB4EFD3EFA858670" box="[1176,1336,742,769]" italics="true" pageId="7" pageNumber="528">G. longipinnis</emphasis>
</taxonomicName>
), having a white dorsal-fin tip (vs. evenly-coloured dorsal fin in
<taxonomicName id="4C444D69BC59FF93FCC1FCD0FC0D8653" authorityName="Krupp" authorityYear="1988" box="[791,944,777,803]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="gallagheri">
<emphasis id="B930EAF8BC59FF93FCC1FCD0FC0D8653" box="[791,944,777,803]" italics="true" pageId="7" pageNumber="528">G. gallagheri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C444D69BC59FF93FC16FCD0FBDF8653" authorityName="Banister &amp; Clarke" authorityYear="1977" box="[960,1122,777,803]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="B930EAF8BC59FF93FC16FCD0FBDF8653" box="[960,1122,777,803]" italics="true" pageId="7" pageNumber="528">G. longipinnis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C444D69BC59FF93FB4BFCD0FABC8653" authorityName="Kirchner &amp; Kruckenhauser &amp; Pichler &amp; Borkenhagen &amp; Freyhof" authorityYear="2020" box="[1181,1281,777,803]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="sharq">
<emphasis id="B930EAF8BC59FF93FB4BFCD0FABC8653" box="[1181,1281,777,803]" italics="true" pageId="7" pageNumber="528">G. sharq</emphasis>
</taxonomicName>
), and usually orange anal- and paired fins (vs. greenish, grey or black in
<taxonomicName id="4C444D69BC59FF93FCFDFCF5FC7F8636" authorityName="Krupp" authorityYear="1988" box="[811,962,812,838]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="gallagheri">
<emphasis id="B930EAF8BC59FF93FCFDFCF5FC7F8636" box="[811,962,812,838]" italics="true" pageId="7" pageNumber="528">G. gallagheri</emphasis>
</taxonomicName>
,
<taxonomicName id="4C444D69BC59FF93FC06FCF5FBCD8636" authorityName="Banister &amp; Clarke" authorityYear="1977" box="[976,1136,812,838]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="longipinnis">
<emphasis id="B930EAF8BC59FF93FC06FCF5FBCD8636" box="[976,1136,812,838]" italics="true" pageId="7" pageNumber="528">G. longipinnis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C444D69BC59FF93FB78FCF5FAB28636" authorityName="Kirchner &amp; Kruckenhauser &amp; Pichler &amp; Borkenhagen &amp; Freyhof" authorityYear="2020" box="[1198,1295,812,838]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="sharq">
<emphasis id="B930EAF8BC59FF93FB78FCF5FAB28636" box="[1198,1295,812,838]" italics="true" pageId="7" pageNumber="528">G. sharq</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C35E6561BC59FF93FF11FC94FACC86DE" pageId="7" pageNumber="528" type="distribution">
<paragraph id="8BFB36EABC59FF93FF11FC94FACC86DE" blockId="7.[151,1437,152,1289]" pageId="7" pageNumber="528">
<emphasis id="B930EAF8BC59FF93FF11FC94FEDC8618" bold="true" box="[199,353,845,872]" pageId="7" pageNumber="528">Distribution.</emphasis>
Found in wadis on both flanks of the northern Hajar Mountains in
<collectingCountry id="F353767ABC59FF93FBB8FC97FB158618" box="[1134,1192,846,872]" name="United Arab Emirates" pageId="7" pageNumber="528">UAE</collectingCountry>
and Northern
<collectingCountry id="F353767ABC59FF93FA87FC97FA258618" box="[1361,1432,846,872]" name="Oman" pageId="7" pageNumber="528">Oman</collectingCountry>
, with a range extending southwards from the southern edge of the Ru’us al-Jibal range (
<collectingRegion id="4980F808BC59FF93FBB5FCA8FB5C86FB" box="[1123,1249,881,907]" country="Oman" name="Musandam" pageId="7" pageNumber="528">Musandam</collectingRegion>
Peninsula) to Al Hayyal on the Gulf of
<collectingCountry id="F353767ABC59FF93FE42FC4DFE6586DE" box="[404,472,916,942]" name="Oman" pageId="7" pageNumber="528">Oman</collectingCountry>
coast and at least to Wadi al-Kabir and its tributaries on the interior flank (
<figureCitation id="137F2A6FBC59FF93FAC9FC4AFAD986DE" box="[1311,1380,915,942]" captionStart="FIGURE 2" captionStartId="5.[151,250,1410,1435]" captionTargetBox="[151,1436,181,1384]" captionTargetId="figure@5.[151,1436,181,1384]" captionTargetPageId="5" captionText="FIGURE 2. Records of Garra species from the Hajar Mountains based on material analysed in this study." figureDoi="http://doi.org/10.5281/zenodo.3714755" httpUri="https://zenodo.org/record/3714755/files/figure.png" pageId="7" pageNumber="528">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35E6561BC59FF93FF11FC6FFB418079" pageId="7" pageNumber="528" type="discussion">
<paragraph id="8BFB36EABC59FF93FF11FC6FFB418079" blockId="7.[151,1437,152,1289]" pageId="7" pageNumber="528">
<emphasis id="B930EAF8BC59FF93FF11FC6FFE8686A1" bold="true" box="[199,315,950,977]" pageId="7" pageNumber="528">Remarks.</emphasis>
When
<bibRefCitation id="EFD54B1BBC59FF93FE46FC6FFD0E86A0" author="Banister, K. E. &amp; Clarke, M. A." box="[400,691,950,976]" pageId="7" pageNumber="528" pagination="111 - 154" refId="ref11266" refString="Banister, K. E. &amp; Clarke, M. A. (1977) The freshwater fishes of the Arabian Peninsula. The Scientific Results of the Oman Flora and Fauna Survey 1975. Journal Oman Studies, 1, 111 - 154." type="journal article" year="1977">Banister &amp; Clarke (1977)</bibRefCitation>
described
<emphasis id="B930EAF8BC59FF93FCE7FC6EFBB086A0" box="[817,1037,950,977]" italics="true" pageId="7" pageNumber="528">G. b. shawkahensis</emphasis>
, they examined two juvenile
<typeStatus id="54FF8848BC59FF93FAB5FC6EFF6F8683" pageId="7" pageNumber="528" type="paratype">paratypes</typeStatus>
of
<taxonomicName id="4C444D69BC59FF93FF2EFC00FE2B8683" authorityName="Fowler &amp; Steinitz" authorityYear="1956" box="[248,406,985,1011]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="B930EAF8BC59FF93FF2EFC00FE2B8683" box="[248,406,985,1011]" italics="true" pageId="7" pageNumber="528">G. barreimiae</emphasis>
</taxonomicName>
(33, 34 mm SL), which were of “fragile condition” (p. 133) and not closely examined. The authors did examine fishes from
<collectingCountry id="F353767ABC59FF93FDC7FC25FDDE8166" box="[529,611,1020,1046]" name="Oman" pageId="7" pageNumber="528">Muscat</collectingCountry>
and Wadi Sahtan (in the coastal basin of
<collectingCountry id="F353767ABC59FF93FB95FC25FB348166" box="[1091,1161,1020,1046]" name="Oman" pageId="7" pageNumber="528">Oman</collectingCountry>
) in detail and identified them as
<emphasis id="B930EAF8BC59FF93FF22FBC6FE118148" box="[244,428,1054,1081]" italics="true" pageId="7" pageNumber="528">
G. b.
<taxonomicName id="4C444D69BC59FF93FEE7FBC7FE118148" authorityName="Fowler &amp; Steinitz" authorityYear="1956" box="[305,428,1054,1080]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="barreimiae">barreimiae</taxonomicName>
</emphasis>
since they possessed 11–14 gill rakers on the lower limb of the first gill arch (vs.
<quantity id="4CBC9B0FBC59FF93FAECFBC4FA218148" box="[1338,1436,1053,1080]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.572" metricValueMin="3.81" pageId="7" pageNumber="528" unit="in" value="16.5" valueMax="18.0" valueMin="15.0">15–18 in</quantity>
<emphasis id="B930EAF8BC59FF93FF41FB98FE3F812B" box="[151,386,1089,1115]" italics="true" pageId="7" pageNumber="528">G. b. shawkahensis).</emphasis>
However, the individuals that Banister and Clarke identified as
<emphasis id="B930EAF8BC59FF93FB8EFB98FAAE812B" box="[1112,1299,1089,1115]" italics="true" pageId="7" pageNumber="528">
G. b.
<taxonomicName id="4C444D69BC59FF93FB4EFB98FAAE812B" authorityName="Fowler &amp; Steinitz" authorityYear="1956" box="[1176,1299,1089,1115]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="barreimiae">barreimiae</taxonomicName>
</emphasis>
do not conform to those collected in the area of the type locality
<collectingRegion id="4980F808BC59FF93FD2AFBBAFCC6810D" box="[764,891,1123,1149]" country="Oman" name="Al Buraymi" pageId="7" pageNumber="528">Al Buraimi</collectingRegion>
(15–18 gill rakers), and they in fact redescribed
<taxonomicName id="4C444D69BC59FF93FF41FB5EFE8981D0" authorityName="Fowler &amp; Steinitz" authorityYear="1956" box="[151,308,1158,1185]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="B930EAF8BC59FF93FF41FB5EFE8981D0" box="[151,308,1158,1185]" italics="true" pageId="7" pageNumber="528">G. barreimiae</emphasis>
</taxonomicName>
which is conspecific with
<emphasis id="B930EAF8BC59FF93FD8BFB5EFC8981D0" box="[605,820,1158,1185]" italics="true" pageId="7" pageNumber="528">G. b. shawkahensis</emphasis>
. Moreover, the specimens Banister &amp; Clarke identified as
<emphasis id="B930EAF8BC59FF93FF60FB70FECC81B3" box="[182,369,1193,1219]" italics="true" pageId="7" pageNumber="528">
G. b.
<taxonomicName id="4C444D69BC59FF93FF20FB70FECC81B3" authorityName="Fowler &amp; Steinitz" authorityYear="1956" box="[246,369,1193,1219]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="barreimiae">barreimiae</taxonomicName>
</emphasis>
represent a distinct species, which is described below as
<taxonomicName id="4C444D69BC59FF93FC2BFB70FBCC81B3" authorityName="Kirchner &amp; Kruckenhauser &amp; Pichler &amp; Borkenhagen &amp; Freyhof" authorityYear="2020" box="[1021,1137,1193,1219]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="shamal">
<emphasis id="B930EAF8BC59FF93FC2BFB70FBCC81B3" box="[1021,1137,1193,1219]" italics="true" pageId="7" pageNumber="528">G. shamal</emphasis>
</taxonomicName>
. The description of
<taxonomicName id="4C444D69BC59FF93FA80FB70FE3E8196" authorityName="Khalaf" authorityYear="2009" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="subSpecies" species="barreimiae" subSpecies="wurayahi">
<emphasis id="B930EAF8BC59FF93FA80FB70FE3E8196" italics="true" pageId="7" pageNumber="528">Garra barreimiae wurayahi</emphasis>
</taxonomicName>
from the Emirate of Fujairah by
<bibRefCitation id="EFD54B1BBC59FF93FD27FB12FBC18196" author="Khalaf-Sakerfalke von Jaffa, N. A. B. A. T." box="[753,1148,1227,1254]" pageId="7" pageNumber="528" pagination="1 - 15" refId="ref12496" refString="Khalaf-Sakerfalke von Jaffa, N. A. B. A. T. (2009) Garra barreimiae wurayahi Khalaf, 2009: A new blind cave fish subspecies from Wadi Al Wurayah pools, Emirate of Fujairah, United Arab Emirates. Gazelle, 90, 1 - 15." type="journal article" year="2009">Khalaf-Sakerfalke von Jaffa (2009)</bibRefCitation>
does not fulfil the criteria of the ICZN, therefore the name is not available. This population is here identified as
<taxonomicName id="4C444D69BC59FF93FB81FB36FB498078" authorityName="Fowler &amp; Steinitz" authorityYear="1956" box="[1111,1268,1262,1289]" class="Actinopterygii" family="Cyprinidae" genus="Garra" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="528" phylum="Chordata" rank="species" species="barreimiae">
<emphasis id="B930EAF8BC59FF93FB81FB36FB498078" box="[1111,1268,1262,1289]" italics="true" pageId="7" pageNumber="528">G. barreimiae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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