treatments-xml/data/03/ED/01/03ED0167FF90FFABFF1DF47EFA00FB95.xml
2024-06-21 12:22:17 +02:00

245 lines
47 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="2AD2AAE1AE5D995A2337D4F5397C3853" ID-CLB-Dataset="24935" ID-DOI="10.11646/zootaxa.4695.3.1" ID-GBIF-Dataset="256591b2-6d7b-465d-a818-9a158017ca09" ID-ISSN="1175-5326" ID-Zenodo-Dep="3532515" ID-ZooBank="32E17629-F29A-47C8-ACCB-69A32C17B147" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573197328991" checkinUser="plazi" docAuthor="Gunter, Nicole L. &amp; Weir, Thomas A." docDate="2019" docId="03ED0167FF90FFABFF1DF47EFA00FB95" docLanguage="en" docName="zootaxa.4695.3.1.pdf" docOrigin="Zootaxa 4695 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepanus monteithi in Matthews 1974" docType="key" docVersion="8" lastPageNumber="223" masterDocId="FFD4791FFF85FFBDFF8AF379FF9DFFA8" masterDocTitle="Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups" masterLastPageNumber="252" masterPageNumber="201" pageNumber="222" updateTime="1698764404900" updateUser="ExternalLinkService">
<mods:mods id="E3B53B07076A482EB0CA52BFD39883C5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="75687379F130F53B9C89D2D97684FE05">
<mods:title id="AE7A141D77D89134D9FC1FDED35FD7C7">Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups</mods:title>
</mods:titleInfo>
<mods:name id="FEB01DACEC3A90E5956B2D95FC5AF491" type="personal">
<mods:role id="8569A08FE8D089C6C6BB2727B47C2741">
<mods:roleTerm id="3BC07F7825A692995A21194FC360A221">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="91297A957A16CD7A2C6AB25473CBFD0B">Gunter, Nicole L.</mods:namePart>
</mods:name>
<mods:name id="27F42EC5AA399423C3BEE62F8FA4F254" type="personal">
<mods:role id="F0480C4CB14071231327E1A433248988">
<mods:roleTerm id="597268DF7515CB4D644B9B2C8031853B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B29DB92232EEEAA9B8207CF3924D8F65">Weir, Thomas A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1D56ADDA7FC9C8908C86174D1280CE14">text</mods:typeOfResource>
<mods:relatedItem id="A57169786AEAEED6FA0787607227407C" type="host">
<mods:titleInfo id="2B2642BE2BD084268A3F41EC229C8296">
<mods:title id="CD889195C84585F1FC903E6E170A5BE0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6CA6D3754A7E1A886C3F12FF7CF9CBF5">
<mods:date id="ABFB0CD27261D856F0086DB3DF561E07">2019</mods:date>
<mods:detail id="96FAAD80486FD947D8CC7C60D704EA79" type="pubDate">
<mods:number id="AAE2660E9BF473D829FC40000340D419">2019-11-08</mods:number>
</mods:detail>
<mods:detail id="921C4C6A415F7D4FC3BB7D2143613CC2" type="volume">
<mods:number id="4ED6666BD8FF0B20FBFEF71712C32363">4695</mods:number>
</mods:detail>
<mods:detail id="EBA531050179C38B1117E7DD76DDDBAF" type="issue">
<mods:number id="0A4803471CB4EFDCF56E32DE26D555F5">3</mods:number>
</mods:detail>
<mods:extent id="D1F13C673919ACEF715DA50F731B3BE9" unit="page">
<mods:start id="CFA4047D4456C8DC7D67B119FAC93D57">201</mods:start>
<mods:end id="548B07AF680916E5044D7A716BED860D">252</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="BF54FBEC54084513F1B3AEA962E27C6D">journal article</mods:classification>
<mods:identifier id="E6FBF1884524891D8F4EA056FC669682" type="CLB-Dataset">24935</mods:identifier>
<mods:identifier id="8E8F673CCA95CC79B7DE534BD51E497B" type="DOI">10.11646/zootaxa.4695.3.1</mods:identifier>
<mods:identifier id="62DADA76A77D59C35465AE13009ED5EE" type="GBIF-Dataset">256591b2-6d7b-465d-a818-9a158017ca09</mods:identifier>
<mods:identifier id="83FA14ECB1276FD1ED55D0E1F938F94B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="0646C8E435EE8919D6BEA664A9285B4F" type="Zenodo-Dep">3532515</mods:identifier>
<mods:identifier id="272625A54BF59855A39B1F3874D5E47A" type="ZooBank">32E17629-F29A-47C8-ACCB-69A32C17B147</mods:identifier>
</mods:mods>
<treatment id="03ED0167FF90FFABFF1DF47EFA00FB95" ID-GBIF-Taxon="160218005" LSID="urn:lsid:plazi:treatment:03ED0167FF90FFABFF1DF47EFA00FB95" httpUri="http://treatment.plazi.org/id/03ED0167FF90FFABFF1DF47EFA00FB95" lastPageId="22" lastPageNumber="223" pageId="21" pageNumber="222">
<subSubSection id="C35EE3FAFF90FFA8FF1DF47EFC95F889" box="[151,776,1798,1825]" pageId="21" pageNumber="222" type="nomenclature">
<paragraph id="8BFBB071FF90FFA8FF1DF47EFC95F889" blockId="21.[151,776,1798,1825]" box="[151,776,1798,1825]" pageId="21" pageNumber="222">
<heading id="D0B3071DFF90FFA8FF1DF47EFC95F889" bold="true" box="[151,776,1798,1825]" fontSize="11" level="1" pageId="21" pageNumber="222" reason="1">
<emphasis id="B9306C63FF90FFA8FF1DF47EFC95F889" bold="true" box="[151,776,1798,1825]" pageId="21" pageNumber="222">
Key to species of
<taxonomicName id="4C44CBF2FF90FFA8FEE4F47EFDCFF889" ID-CoL="3T8V4" authorityName="in Matthews" authorityYear="1974" box="[366,594,1798,1825]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B9306C63FF90FFA8FEE4F47EFDCFF889" bold="true" box="[366,594,1798,1825]" italics="true" pageId="21" pageNumber="222">Lepanus monteithi</emphasis>
</taxonomicName>
species group.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35EE3FAFF90FFABFF1DF432FA00FB95" lastPageId="22" lastPageNumber="223" pageId="21" pageNumber="222" type="key">
<keyStep id="30B0A8D4FF90FFABFF1DF432FA01FF64" lastPageId="22" lastPageNumber="223" pageId="21" pageNumber="222">
<paragraph id="8BFBB071FF90FFA8FF1DF432FA00F811" blockId="21.[151,1437,1867,2036]" pageId="21" pageNumber="222">
<keyLead id="30B51344FF90FFA8FF1DF432FA00F811" pageId="21" pageNumber="222">
1. Clypeal edge with a blunt projection lateral to clypeal teeth (
<figureCitation id="137FACF4FF90FFA8FCB0F432FC14F8C9" box="[826,905,1867,1889]" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" pageId="21" pageNumber="222">Fig. 8A</figureCitation>
); dorsal part of eyes wide, separated by 7 eye widths; medial lobe of metaventrite narrowly margined beside mesocoxae (
<figureCitation id="137FACF4FF90FFA8FCFFF411FC5EF8D6" box="[885,963,1896,1918]" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" pageId="21" pageNumber="222">Fig. 8D</figureCitation>
); male metatibiae without a notched inner edge; aedeagus as in
<figureCitation id="137FACF4FF90FFA8FEF8F4FFFE57F833" box="[370,458,1926,1948]" captionStart="FIGURE 10" captionStartId="25.[151,250,1924,1949]" captionTargetBox="[201,1386,175,1892]" captionTargetId="figure@25.[188,1400,166,1905]" captionTargetPageId="25" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532533" httpUri="https://zenodo.org/record/3532533/files/figure.png" pageId="21" pageNumber="222">Fig. 10D</figureCitation>
, with a small tuft of setae on apex of right paramere; northern
<collectingRegion id="49807E93FF90FFA8FBA8F4FFFB4EF834" box="[1058,1235,1926,1948]" country="Australia" name="Northern Territory" pageId="21" pageNumber="222">Northern Territory</collectingRegion>
...............................................................................
<emphasis id="B9306C63FF90FFA8FC56F4DAFA00F811" bold="true" box="[988,1437,1955,1977]" pageId="21" pageNumber="222">
<taxonomicName id="4C44CBF2FF90FFA8FC56F4DAFA81F811" ID-CoL="3T8VQ" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[988,1308,1955,1977]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="21" pageNumber="222" phylum="Arthropoda" rank="species" species="topend" status="sp. nov.">
<emphasis id="B9306C63FF90FFA8FC56F4DAFBE7F811" bold="true" box="[988,1146,1955,1977]" italics="true" pageId="21" pageNumber="222">Lepanus topend</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A203D118FF90FFA8FAACF4DAFA00F811" box="[1318,1437,1955,1977]" pageId="21" pageNumber="222" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFBB071FF90FFABFF1DF4B8FA01FF64" blockId="21.[151,1437,1867,2036]" lastBlockId="22.[151,1437,153,1085]" lastPageId="22" lastPageNumber="223" pageId="21" pageNumber="222">
<keyLead id="30B51344FF90FFABFF1DF4B8FA01FF64" lastPageId="22" lastPageNumber="223" pageId="21" pageNumber="222">
- Clypeal edge evenly rounded lateral to clypeal teeth (
<figureCitation id="137FACF4FF90FFA8FD61F4B9FCAAF87E" box="[747,823,1984,2006]" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" pageId="21" pageNumber="222">Fig. 8B</figureCitation>
); dorsal parts of eyes narrower, separated by more than 12 eye widths; medial lobe of metaventrite with margin beside mesocoxae either very narrowly margined or virtually absent (
<figureCitation id="137FACF4FF90FFABFAE6F4A7FF70FF18" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" lastPageId="22" lastPageNumber="223" pageId="21" pageNumber="222">Figs. 8</figureCitation>
HI); male metatibiae with or without a notch on inner edge; aedeagus with or without small tufts of setae on apices of parameres...............................................................................................
<emphasis id="B9306C63FF93FFABFA1AF3CEFA01FF64" bold="true" box="[1424,1436,183,204]" pageId="22" pageNumber="223">2</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B0A8D4FF93FFABFF1DF3ADFA01FED4" pageId="22" pageNumber="223">
<paragraph id="8BFBB071FF93FFABFF1DF3ADFA00FEEA" blockId="22.[151,1437,153,1085]" pageId="22" pageNumber="223">
<keyLead id="30B51344FF93FFABFF1DF3ADFA00FEEA" pageId="22" pageNumber="223">
2. Lateral lobe of metaventrite with a pit containing a tuft of setae (
<figureCitation id="137FACF4FF93FFABFCD8F3ADFC02FF42" box="[850,927,212,234]" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" pageId="22" pageNumber="223">Fig. 8G</figureCitation>
), virtually impunctate anteriorly, inner border beside mesocoxae not defined; head and disc of pronotum very finely punctate; aedeagus as in
<figureCitation id="137FACF4FF93FFABFBC2F388FB3CFEAF" box="[1096,1185,241,263]" captionStart="FIGURE 10" captionStartId="25.[151,250,1924,1949]" captionTargetBox="[201,1386,175,1892]" captionTargetId="figure@25.[188,1400,166,1905]" captionTargetPageId="25" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532533" httpUri="https://zenodo.org/record/3532533/files/figure.png" pageId="22" pageNumber="223">Fig. 10A</figureCitation>
, without tufts of setae on parameres; northern
<collectingRegion id="49807E93FF93FFABFE22F276FD81FE8D" box="[424,540,271,293]" country="Australia" name="Queensland" pageId="22" pageNumber="223">Queensland</collectingRegion>
, Iron Range area north to tip of Cape
<collectingRegion id="49807E93FF93FFABFC0CF276FC28FE8D" box="[902,949,271,293]" country="United Kingdom" name="York" pageId="22" pageNumber="223">York</collectingRegion>
Peninsula................................................................................................
<emphasis id="B9306C63FF93FFABFC6CF255FA00FEEA" bold="true" box="[998,1437,300,322]" pageId="22" pageNumber="223">
<taxonomicName id="4C44CBF2FF93FFABFC6CF255FA81FEEA" ID-CoL="6P7XZ" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[998,1308,300,322]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="geoffi" status="sp. nov.">
<emphasis id="B9306C63FF93FFABFC6CF255FBE7FEEA" bold="true" box="[998,1146,300,322]" italics="true" pageId="22" pageNumber="223">Lepanus geoffi</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A203D118FF93FFABFAACF255FA00FEEA" box="[1318,1437,300,322]" pageId="22" pageNumber="223" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFBB071FF93FFABFF1DF233FA01FED4" blockId="22.[151,1437,153,1085]" pageId="22" pageNumber="223">
<keyLead id="30B51344FF93FFABFF1DF233FA01FED4" pageId="22" pageNumber="223">
- Lateral lobe of metaventrite without a setose pit, punctate anteriorly, inner border beside mesocoxae well defined; head and disc of pronotum with punctation fine or not; aedeagus with or without tufts of setae on apices of parameres.................
<emphasis id="B9306C63FF93FFABFA1AF21EFA01FED4" bold="true" box="[1424,1436,359,380]" pageId="22" pageNumber="223">3</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B0A8D4FF93FFABFF1DF2FDFA01FDA7" pageId="22" pageNumber="223">
<paragraph id="8BFBB071FF93FFABFF1DF2FDFA01FE7C" blockId="22.[151,1437,153,1085]" pageId="22" pageNumber="223">
<keyLead id="30B51344FF93FFABFF1DF2FDFA01FE7C" pageId="22" pageNumber="223">
3. Outer edge of protibiae with only 2 distinct teeth, 3
<superScript id="7C311D39FF93FFABFD42F2FBFD49FE26" attach="left" box="[712,724,386,398]" fontSize="5" pageId="22" pageNumber="223">rd</superScript>
or basal tooth reduced to little more than a denticle (
<figureCitation id="137FACF4FF93FFABFB43F2FDFA8EFE32" box="[1225,1299,388,410]" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" pageId="22" pageNumber="223">Fig. 8E</figureCitation>
); male metatibiae with a notch on inner edge (eg
<figureCitation id="137FACF4FF93FFABFDB0F2D8FD1FFE10" box="[570,642,417,440]" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" pageId="22" pageNumber="223">Fig. 8F</figureCitation>
); aedeagus with small tufts of setae on apices of both right and left parameres... ...................................................................................................
<emphasis id="B9306C63FF93FFABFA1AF2C6FA01FE7C" bold="true" box="[1424,1436,447,468]" pageId="22" pageNumber="223">4</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BFBB071FF93FFABFF1DF2A4FA01FDA7" blockId="22.[151,1437,153,1085]" pageId="22" pageNumber="223">
<keyLead id="30B51344FF93FFABFF1DF2A4FA01FDA7" pageId="22" pageNumber="223">
- Outer edge of protibiae with 3 distinct teeth; male metatibiae with or without a notch on inner edge; aedeagus with or without small tufts of setae on apices of parameres.................................................................
<emphasis id="B9306C63FF93FFABFA1AF283FA01FDA7" bold="true" box="[1424,1436,506,527]" pageId="22" pageNumber="223">5</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B0A8D4FF93FFABFF1DF16EFA01FCBE" pageId="22" pageNumber="223">
<paragraph id="8BFBB071FF93FFABFF1DF16EFA00FD2C" blockId="22.[151,1437,153,1085]" pageId="22" pageNumber="223">
<keyLead id="30B51344FF93FFABFF1DF16EFA00FD2C" pageId="22" pageNumber="223">
4. Head without a basal carina; head and disc of pronotum very finely punctate; elytral intervals virtually impunctate; lateral lobe of metaventrite with inner border beside mesocoxae anteriorly curving to meet lateral border (
<figureCitation id="137FACF4FF93FFABFBEAF14DFB31FDE2" box="[1120,1196,564,586]" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" pageId="22" pageNumber="223">Fig. 8H</figureCitation>
); pygidium not reticulate basally and virtually impunctate (
<figureCitation id="137FACF4FF93FFABFDACF128FDE0FDC0" box="[550,637,593,616]" captionStart="FIGURE 11" captionStartId="26.[151,250,1247,1272]" captionTargetBox="[152,1428,188,1216]" captionTargetId="figure@26.[151,1435,174,1228]" captionTargetPageId="26" captionText="FIGURE 11. Pygidial configuration of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532535" httpUri="https://zenodo.org/record/3532535/files/figure.png" pageId="22" pageNumber="223">Fig. 11C</figureCitation>
); aedeagus as in
<figureCitation id="137FACF4FF93FFABFCA8F128FCE4FDCF" box="[802,889,593,615]" captionStart="FIGURE 10" captionStartId="25.[151,250,1924,1949]" captionTargetBox="[201,1386,175,1892]" captionTargetId="figure@25.[188,1400,166,1905]" captionTargetPageId="25" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532533" httpUri="https://zenodo.org/record/3532533/files/figure.png" pageId="22" pageNumber="223">Fig. 10C</figureCitation>
, with small tufts of setae on apices of both right and left parameres, left paramere pointed; northern
<collectingRegion id="49807E93FF93FFABFD08F116FD6BFD2D" box="[642,758,623,645]" country="Australia" name="Queensland" pageId="22" pageNumber="223">Queensland</collectingRegion>
, Wet Tropics......
<emphasis id="B9306C63FF93FFABFC3CF116FAFCFD2D" bold="true" box="[950,1377,622,645]" pageId="22" pageNumber="223">
<taxonomicName id="4C44CBF2FF93FFABFC3CF116FB7CFD2D" ID-CoL="3T8VL" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[950,1249,622,645]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="reidi" status="sp. nov.">
<emphasis id="B9306C63FF93FFABFC3CF116FBA3FD2C" bold="true" box="[950,1086,622,644]" italics="true" pageId="22" pageNumber="223">Lepanus reidi</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A203D118FF93FFABFB61F116FAFCFD2D" box="[1259,1377,623,645]" pageId="22" pageNumber="223" rank="species">new species</taxonomicNameLabel>
</emphasis>
[part]
</keyLead>
</paragraph>
<paragraph id="8BFBB071FF93FFABFF1DF1F4FA01FCBE" blockId="22.[151,1437,153,1085]" pageId="22" pageNumber="223">
<keyLead id="30B51344FF93FFABFF1DF1F4FA01FCBE" pageId="22" pageNumber="223">
- Head with short basal carina present only adjacent to eyes (
<figureCitation id="137FACF4FF93FFABFCACF1F5FCEFFD0A" box="[806,882,652,674]" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" pageId="22" pageNumber="223">Fig. 8C</figureCitation>
); head and disc of pronotum with punctation not so fine; elytral intervals distinctly punctate; lateral lobe of metaventrite with inner border bedside mesocoxae anteriorly straight, lateral border virtually absent (
<figureCitation id="137FACF4FF93FFABFE46F1BEFD8DFD75" box="[460,528,711,733]" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" pageId="22" pageNumber="223">
Fig.
<date id="FFFA96B1FF93FFABFE70F1BEFD8DFD75" box="[506,528,711,733]" pageId="22" pageNumber="223">8I</date>
</figureCitation>
); pygidium often finely reticulate basally and with distinct punctures (
<figureCitation id="137FACF4FF93FFABFB34F1BEFA8BFD75" box="[1214,1302,711,733]" captionStart="FIGURE 11" captionStartId="26.[151,250,1247,1272]" captionTargetBox="[152,1428,188,1216]" captionTargetId="figure@26.[151,1435,174,1228]" captionTargetPageId="26" captionText="FIGURE 11. Pygidial configuration of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532535" httpUri="https://zenodo.org/record/3532535/files/figure.png" pageId="22" pageNumber="223">Fig. 11B</figureCitation>
); aedeagus as in
<figureCitation id="137FACF4FF93FFABFF76F19DFECEFD51" box="[252,339,740,762]" captionStart="FIGURE 10" captionStartId="25.[151,250,1924,1949]" captionTargetBox="[201,1386,175,1892]" captionTargetId="figure@25.[188,1400,166,1905]" captionTargetPageId="25" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532533" httpUri="https://zenodo.org/record/3532533/files/figure.png" pageId="22" pageNumber="223">Fig. 10B</figureCitation>
, with small tufts of setae on apices of both right and left parameres, left paramere somewhat spatulate; northern
<collectingRegion id="49807E93FF93FFABFF68F078FECBFCBF" box="[226,342,769,791]" country="Australia" name="Queensland" pageId="22" pageNumber="223">Queensland</collectingRegion>
, Iron Range area................................................
<taxonomicName id="4C44CBF2FF93FFABFBB1F07BFA01FCBE" ID-CoL="3T8V4" authority="Matthews, 1974" authorityName="in Matthews" authorityYear="1974" box="[1083,1436,769,791]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="monteithi">
<emphasis id="B9306C63FF93FFABFBB1F07BFA01FCBE" bold="true" box="[1083,1436,769,791]" pageId="22" pageNumber="223">
<emphasis id="B9306C63FF93FFABFBB1F07BFB68FCBF" bold="true" box="[1083,1269,769,791]" italics="true" pageId="22" pageNumber="223">Lepanus monteithi</emphasis>
<bibRefCitation id="EFD5CD80FF93FFABFB71F07BFA01FCBE" author="Matthews, E. G." box="[1275,1436,769,791]" pageId="22" pageNumber="223" pagination="1 - 211" refId="ref56271" refString="Matthews, E. G. (1974) A revision of the scarabaeine dung beetles of Australia. II. Tribe Scarabaeini, Australian Journal of Zoology, Supplementary Series, 24, 1 - 211. https: // doi. org / 10.1071 / AJZS 024" type="journal article" year="1974">Matthews, 1974</bibRefCitation>
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30B0A8D4FF93FFABFF1DF066FA00FB95" pageId="22" pageNumber="223">
<paragraph id="8BFBB071FF93FFABFF1DF066FA00FC01" blockId="22.[151,1437,153,1085]" pageId="22" pageNumber="223">
<keyLead id="30B51344FF93FFABFF1DF066FA00FC01" pageId="22" pageNumber="223">
5. Basal margin of pronotum without a row of small punctures; head and disc of pronotum very finely punctate; elytral striae obsolete; lateral lobe of metaventrite with inner border beside mesocoxae anteriorly curving forward to meet lateral border (
<figureCitation id="137FACF4FF93FFABFAFFF045FE9FFCD8" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" pageId="22" pageNumber="223">Fig. 8H</figureCitation>
); pygidium not reticulate basally and virtually impunctate (
<figureCitation id="137FACF4FF93FFABFCB7F020FC0EFCD8" box="[829,915,857,880]" captionStart="FIGURE 11" captionStartId="26.[151,250,1247,1272]" captionTargetBox="[152,1428,188,1216]" captionTargetId="figure@26.[151,1435,174,1228]" captionTargetPageId="26" captionText="FIGURE 11. Pygidial configuration of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532535" httpUri="https://zenodo.org/record/3532535/files/figure.png" pageId="22" pageNumber="223">Fig. 11C</figureCitation>
); male metatibiae with a notch on the inner edge (
<figureCitation id="137FACF4FF93FFABFAFFF020FF63FC25" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" pageId="22" pageNumber="223">Fig. 8F</figureCitation>
); aedeagus as in
<figureCitation id="137FACF4FF93FFABFE15F00EFE69FC24" box="[415,500,887,909]" captionStart="FIGURE 10" captionStartId="25.[151,250,1924,1949]" captionTargetBox="[201,1386,175,1892]" captionTargetId="figure@25.[188,1400,166,1905]" captionTargetPageId="25" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532533" httpUri="https://zenodo.org/record/3532533/files/figure.png" pageId="22" pageNumber="223">Fig. 10C</figureCitation>
, with small tufts of setae on the apices of both right and left parameres; northern
<collectingRegion id="49807E93FF93FFABFB70F00EFAF3FC25" box="[1274,1390,887,909]" country="Australia" name="Queensland" pageId="22" pageNumber="223">Queensland</collectingRegion>
, Wet Tropics.....................................................
<emphasis id="B9306C63FF93FFABFC3CF0EDFAFCFC02" bold="true" box="[950,1377,916,938]" pageId="22" pageNumber="223">
<taxonomicName id="4C44CBF2FF93FFABFC3CF0EDFB7CFC02" ID-CoL="3T8VL" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[950,1249,916,938]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="reidi" status="sp. nov.">
<emphasis id="B9306C63FF93FFABFC3CF0EDFBA3FC02" bold="true" box="[950,1086,916,938]" italics="true" pageId="22" pageNumber="223">Lepanus reidi</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A203D118FF93FFABFB61F0EDFAFCFC02" box="[1259,1377,916,938]" pageId="22" pageNumber="223" rank="species">new species</taxonomicNameLabel>
</emphasis>
[part]
</keyLead>
</paragraph>
<paragraph id="8BFBB071FF93FFABFF1DF0CBFA00FB95" blockId="22.[151,1437,153,1085]" pageId="22" pageNumber="223">
<keyLead id="30B51344FF93FFABFF1DF0CBFA00FB95" pageId="22" pageNumber="223">
- Basal margin of pronotum with a distinct row of small punctures; head and disc of pronotum with punctation not so fine, easily discernable; elytral striae clearly visible; lateral lobe of metaventrite with inner border beside mesocoxae anteriorly straight, lateral border virtually absent (as in
<figureCitation id="137FACF4FF93FFABFDCCF095FD17FBAA" box="[582,650,1004,1026]" captionStart="FIGURE 8" captionStartId="23.[151,250,1718,1743]" captionTargetBox="[188,1400,169,1698]" captionTargetId="figure@23.[188,1400,166,1698]" captionTargetPageId="23" captionText="FIGURE 8. Some diagnostic characters of the Lepanus monteithi species group. A, Head of Lepanus topend, black arrow points to blunt projection lateral to clypeal teeth; B, Head of Lepanus reidi example of rounded clypeal edge, white arrow showing absence of basal carina; C, Head of Lepanus monteithi with dotted lines showing short basal carina; D, Metaventrite of Lepanus topend, white arrow points to narrow margin of medial lobe beside mesocoxae; E, protibia of Lepanus monteithi showing only two distinct teeth; F, male metatibiae of Lepanus reidi, black arrow and dotted line indicates notch on the inner edge; G, ventrolateral view of Lepanus geoffi highlighting pit containing a tuft of setae on lateral lobe of metaventrite; H, ventrolateral view of Lepanus reidi white arrow points to very narrow margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly curved inner border of lateral lobe; I, ventrolateral view of Lepanus monteithi white arrow points to virtually absent margin of medial lobe of metaventrite beside mesocoxa, dotted line showing anteriorly straight inner border of lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.3532529" httpUri="https://zenodo.org/record/3532529/files/figure.png" pageId="22" pageNumber="223">
Fig.
<date id="FFFA96B1FF93FFABFDFEF095FD17FBAA" box="[628,650,1004,1026]" pageId="22" pageNumber="223">8I</date>
</figureCitation>
); pygidium finely reticulate basally and with distinct punctures (
<figureCitation id="137FACF4FF93FFABFA89F095FAC7FBAA" box="[1283,1370,1004,1026]" captionStart="FIGURE 11" captionStartId="26.[151,250,1247,1272]" captionTargetBox="[152,1428,188,1216]" captionTargetId="figure@26.[151,1435,174,1228]" captionTargetPageId="26" captionText="FIGURE 11. Pygidial configuration of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532535" httpUri="https://zenodo.org/record/3532535/files/figure.png" pageId="22" pageNumber="223">Fig. 11E</figureCitation>
); male metatibiae without a notch on inner edge; aedeagus as in
<figureCitation id="137FACF4FF93FFABFC8AF770FCC9FBB7" box="[768,852,1033,1055]" captionStart="FIGURE 10" captionStartId="25.[151,250,1924,1949]" captionTargetBox="[201,1386,175,1892]" captionTargetId="figure@25.[188,1400,166,1905]" captionTargetPageId="25" captionText="FIGURE 10. Left, dorsal, and right view of aedeagi of species within the Lepanus monteithi species group. A, Lepanus geoffi; B, Lepanus monteithi; C, Lepanus reidi; D, Lepanus topend; E, Lepanus yorkensis. All specimens in comparative scale. Scale bar 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3532533" httpUri="https://zenodo.org/record/3532533/files/figure.png" pageId="22" pageNumber="223">Fig. 10E</figureCitation>
, without small tufts of setae on apices of parameres; northern
<collectingRegion id="49807E93FF93FFABFF68F75EFECBFB95" box="[226,342,1063,1085]" country="Australia" name="Queensland" pageId="22" pageNumber="223">Queensland</collectingRegion>
, Iron Range area north to Torres Strait.....................
<emphasis id="B9306C63FF93FFABFC4BF75EFA00FB95" bold="true" box="[961,1437,1062,1085]" pageId="22" pageNumber="223">
<taxonomicName id="4C44CBF2FF93FFABFC4BF75EFA81FB95" ID-CoL="3T8VV" authority="Gunter &amp; Weir" authorityName="Gunter &amp; Weir" authorityYear="2019" box="[961,1308,1062,1085]" class="Insecta" family="Scarabaeidae" genus="Lepanus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="223" phylum="Arthropoda" rank="species" species="yorkensis" status="sp. nov.">
<emphasis id="B9306C63FF93FFABFC4BF75EFBE7FB94" bold="true" box="[961,1146,1062,1084]" italics="true" pageId="22" pageNumber="223">Lepanus yorkensis</emphasis>
Gunter &amp; Weir
</taxonomicName>
,
<taxonomicNameLabel id="A203D118FF93FFABFAACF75EFA00FB95" box="[1318,1437,1063,1085]" pageId="22" pageNumber="223" rank="species">new species</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>