261 lines
42 KiB
XML
261 lines
42 KiB
XML
<document id="9187663439ED4B8C389A59AEA132099B" ID-DOI="10.5281/zenodo.4504166" ID-ISSN="2345-7600" ID-Zenodo-Dep="4504166" ID-ZooBank="924CD2C9-3007-4321-81E2-0237EE1359BA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612465749776" checkinUser="carolina" docAuthor="Schubart, Christoph D. & Ng, Peter K. L." docDate="2014" docId="03EA5318167CE35FAAB4F5C9B81C2FB4" docLanguage="en" docName="RafflesBZool.62.615-619.pdf" docOrigin="Raffles Bulletin of Zoology 62" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Geosesarma bintan Schubart & Ng 2014, new species" docType="treatment" docUuid="BBABCEDE-E066-443F-86DC-9CA238032073" docUuidSource="ZooBank" docVersion="5" lastPageNumber="616" masterDocId="FFD32B60167CE35EA978F24CBC422B21" masterDocTitle="Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia" masterLastPageNumber="619" masterPageNumber="615" pageNumber="615" updateTime="1698866020970" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F98C779ACF0581C638A6E51724DB5343">Two new species of land-dwelling crabs of the genus Geosesarma De Man, 1892 (Crustacea: Brachyura: Thoracotremata: Sesarmidae) from Bintan Island, Indonesia</mods:title>
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<mods:namePart id="A9F71B51863C69DFF1AD360A9B4BB87D">Schubart, Christoph D.</mods:namePart>
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<mods:namePart id="77BE5337A74153BB53D4193DC5350FC7">Ng, Peter K. L.</mods:namePart>
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<mods:title id="41A57B05199C0EF41E8D625469B1CCEA">Raffles Bulletin of Zoology</mods:title>
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<mods:date id="AA85F02A49A13B8342623D96DA2E14A8">2014</mods:date>
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<mods:number id="EC2F53311AE498506F0CAE95DFBDC827">2014-08-22</mods:number>
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<mods:number id="178BC16AFF5A9A91DDB92972BF424EA2">62</mods:number>
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<mods:identifier id="2CE96ABC9E6591E1876DEF18D2D2D6DE" type="DOI">10.5281/zenodo.4504166</mods:identifier>
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<treatment id="03EA5318167CE35FAAB4F5C9B81C2FB4" LSID="urn:lsid:zoobank.org:act:BBABCEDE-E066-443F-86DC-9CA238032073" httpUri="http://treatment.plazi.org/id/03EA5318167CE35FAAB4F5C9B81C2FB4" lastPageId="1" lastPageNumber="616" pageId="0" pageNumber="615">
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<subSubSection id="C359B185167CE35EAAB4F5C9B97A2CBD" box="[972,1336,1924,1949]" pageId="0" pageNumber="615" type="nomenclature">
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<paragraph id="8BFCE20E167CE35EAAB4F5C9B97A2CBD" blockId="0.[956,1351,1924,1981]" box="[972,1336,1924,1949]" pageId="0" pageNumber="615">
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<heading id="D0B45562167CE35EAAB4F5C9B97A2CBD" bold="true" box="[972,1336,1924,1949]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="615" reason="2">
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<emphasis id="B9373E1C167CE35EAAB4F5C9B97A2CBD" bold="true" box="[972,1336,1924,1949]" pageId="0" pageNumber="615">
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<taxonomicName id="4C43998D167CE35EAAB4F5C9B8E12CBD" ID-CoL="3FS8X" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" box="[972,1187,1924,1949]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="615" phylum="Arthropoda" rank="species" species="bintan" status="sp. nov.">
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<emphasis id="B9373E1C167CE35EAAB4F5C9B8E12CBD" bold="true" box="[972,1187,1924,1949]" italics="true" pageId="0" pageNumber="615">Geosesarma bintan</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2048367167CE35EADCAF5C9B97A2CBD" box="[1202,1336,1924,1949]" pageId="0" pageNumber="615" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C359B185167CE35EAAC4F5E9B9052C9C" box="[956,1351,1957,1981]" pageId="0" pageNumber="615" type="description">
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<paragraph id="8BFCE20E167CE35EAAC4F5E9B9052C9C" blockId="0.[956,1351,1924,1981]" box="[956,1351,1957,1981]" pageId="0" pageNumber="615">
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(
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<figureCitation id="1378FE8B167CE35EAABCF5E9B8042C9C" box="[964,1094,1957,1981]" captionStart="Fig" captionStartId="1.[113,148,1976,1998]" captionTargetBox="[113,752,1116,1953]" captionTargetId="figure-214@1.[113,752,1116,1953]" captionTargetPageId="1" captionText="Fig. 1. A, C, Geosesarma bintan, new species, holotype, male,11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, habitus; C, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504168" httpUri="https://zenodo.org/record/4504168/files/figure.png" pageId="0" pageNumber="615">Figs. 1A, C</figureCitation>
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,
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<figureCitation id="1378FE8B167CE35EAD2CF5E9B8D52C9C" box="[1108,1175,1957,1981]" captionStart="Fig" captionStartId="1.[812,847,1977,1999]" captionTargetBox="[812,1451,1572,1954]" captionTargetId="figure-266@1.[812,1451,1572,1954]" captionTargetPageId="1" captionText="Fig. 2.A, C, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, frontal view of carapace; C, D, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4504172" httpUri="https://zenodo.org/record/4504172/files/figure.png" pageId="0" pageNumber="615">2A, C</figureCitation>
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,
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<figureCitation id="1378FE8B167CE35EADDDF5E9B94B2C9D" box="[1189,1289,1957,1981]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="0" pageNumber="615">3A, C, E</figureCitation>
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, G–I)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C359B185167CE35FAA3BF5AFBE5F2BD1" lastPageId="1" lastPageNumber="616" pageId="0" pageNumber="615" type="materials_examined">
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<paragraph id="8BFCE20E167CE35EAA3BF5AFB9802313" blockId="0.[835,1474,2019,2099]" pageId="0" pageNumber="615">
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<emphasis id="B9373E1C167CE35EAA3BF5AFB8472CD8" bold="true" box="[835,1029,2019,2041]" pageId="0" pageNumber="615">Material examined</emphasis>
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.
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<materialsCitation id="3B2BE853167CE35EAD69F5AFB9802313" collectionCode="MZB, T" country="Indonesia" latitude="1.1666666" location="Bintan Island" longLatPrecision="15" longitude="104.3835" municipality="Archipelago" pageId="0" pageNumber="615" specimenCount="1" stateProvince="Riau" typeStatus="holotype">
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<typeStatus id="54F85CAC167CE35EAD69F5AFB8282CD8" box="[1041,1130,2019,2041]" pageId="0" pageNumber="615" type="holotype">Holotype</typeStatus>
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– male (11.2 ×
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<quantity id="4CBB4FEB167CE35EAC7BF5AFB90C2CD8" box="[1283,1358,2019,2041]" metricMagnitude="-3" metricUnit="m" metricValue="9.8" pageId="0" pageNumber="615" unit="mm" value="9.8">9.8 mm</quantity>
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) (
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<collectionCode id="ED527ACB167CE35EAC1AF5AFB9D52CD9" box="[1378,1431,2019,2040]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="0" pageNumber="615" type="Museum">MZB</collectionCode>
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Cru 3813),
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<collectingCountry id="F354A29E167CE35EAAF1FA4CBFA82337" box="[905,1002,2048,2070]" name="Indonesia" pageId="0" pageNumber="615">Indonesia</collectingCountry>
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:
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<collectingRegion id="49872CEC167CE35EAA8DFA4CB8602337" box="[1013,1058,2048,2070]" country="Indonesia" name="Riau" pageId="0" pageNumber="615">Riau</collectingRegion>
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<collectingMunicipality id="6B987874167CE35EAD52FA4CB8E12337" box="[1066,1187,2048,2070]" pageId="0" pageNumber="615">Archipelago</collectingMunicipality>
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,
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<location id="8E9CB4D5167CE35EADD7FA4CB9762337" LSID="urn:lsid:plazi:treatment:03EA5318167CE35FAAB4F5C9B81C2FB4:8E9CB4D5167CE35EADD7FA4CB9762337" box="[1199,1332,2048,2070]" country="Indonesia" latitude="1.1666666" longLatPrecision="15" longitude="104.3835" municipality="Archipelago" name="Bintan Island" pageId="0" pageNumber="615" stateProvince="Riau">Bintan Island</location>
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, near swampy lowland freshwater stream,
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<geoCoordinate id="EE7784C9167CE35EAD20FA51B8F12313" box="[1112,1203,2077,2098]" degrees="1" direction="north" minutes="10" orientation="latitude" pageId="0" pageNumber="615" precision="15" seconds="0" value="1.1666666">1°10'0"N</geoCoordinate>
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,
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<geoCoordinate id="EE7784C9167CE35EADB9FA51B9062312" box="[1217,1348,2077,2099]" degrees="104" direction="east" minutes="23" orientation="longitude" pageId="0" pageNumber="615" precision="1" seconds="0.6" value="104.3835">104°23'0.6"E</geoCoordinate>
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, coll.
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||
<collectionCode id="ED527ACB167CE35EACFCFA51B9D62313" box="[1412,1428,2077,2098]" name="Tavera, Department of Geology and Geophysics" pageId="0" pageNumber="615">T</collectionCode>
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.H.
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||
<collectionCode id="ED527ACB167CE35EACD6FA51B9FC2313" box="[1454,1470,2077,2098]" name="Tavera, Department of Geology and Geophysics" pageId="0" pageNumber="615">T</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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||
<paragraph id="8BFCE20E167DE35FA909F2F2BE5F2BD1" blockId="1.[113,752,189,240]" pageId="1" pageNumber="616">
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<materialsCitation id="3B2BE853167DE35FA909F2F2BE5B2BD1" collectingDate="1993-05-11" collectionCode="ZRC" pageId="1" pageNumber="616" specimenCode="ZRC 2014.0284" specimenCount="2" typeStatus="paratype">
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Tan et al.,
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<date id="FFFDC4CE167DE35FA9A5F2F1BD272BF3" box="[221,357,189,211]" pageId="1" pageNumber="616" value="1993-05-11">
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<collectingDate id="EFB93D26167DE35FA9A5F2F1BD272BF3" box="[221,357,189,211]" pageId="1" pageNumber="616" value="1993-05-11">11 May 1993</collectingDate>
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</date>
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||
.
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<typeStatus id="54F85CAC167DE35FA808F2F2BD862BF2" box="[368,452,190,211]" pageId="1" pageNumber="616" type="paratype">Paratype</typeStatus>
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– male (9.7 ×
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<quantity id="4CBB4FEB167DE35FAB25F2F1BEE92BF2" box="[605,683,189,211]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="616" unit="mm" value="8.5">8.5 mm</quantity>
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) (
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<specimenCode id="DBE54A75167DE35FABB9F2F1BC952BCE" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="616">ZRC 2014.0284</specimenCode>
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), same locality data as
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<typeStatus id="54F85CAC167DE35FA8BAF296BE5B2BD1" box="[450,537,218,240]" pageId="1" pageNumber="616" type="holotype">holotype</typeStatus>
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||
</materialsCitation>
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.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C359B185167DE35FA909F354BD4D2891" pageId="1" pageNumber="616" type="diagnosis">
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<paragraph id="8BFCE20E167DE35FA909F354BDAC2871" blockId="1.[112,754,280,848]" pageId="1" pageNumber="616">
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<emphasis id="B9373E1C167DE35FA909F354BCAF2A11" bold="true" box="[113,237,280,304]" pageId="1" pageNumber="616">Diagnosis.</emphasis>
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Carapace rectangular, distinctly wider than long, lateral margins gently diverging towards posterior carapace margin (
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<figureCitation id="1378FE8B167DE35FA845F314BDFF2A51" box="[317,445,344,368]" captionStart="Fig" captionStartId="1.[113,148,1976,1998]" captionTargetBox="[113,752,1116,1953]" captionTargetId="figure-214@1.[113,752,1116,1953]" captionTargetPageId="1" captionText="Fig. 1. A, C, Geosesarma bintan, new species, holotype, male,11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, habitus; C, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504168" httpUri="https://zenodo.org/record/4504168/files/figure.png" pageId="1" pageNumber="616">Fig. 1A, C</figureCitation>
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); dorsal surfaces generally smooth, except for rugose anterior regions; H-shaped median depression shallow; postfrontal cristae prominent, margins rounded; exorbital tooth triangular, directed obliquely, with additional low tooth behind it (
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<figureCitation id="1378FE8B167DE35FA8A9F394BE702AD1" box="[465,562,472,496]" captionStart="Fig" captionStartId="1.[113,148,1976,1998]" captionTargetBox="[113,752,1116,1953]" captionTargetId="figure-214@1.[113,752,1116,1953]" captionTargetPageId="1" captionText="Fig. 1. A, C, Geosesarma bintan, new species, holotype, male,11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, habitus; C, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504168" httpUri="https://zenodo.org/record/4504168/files/figure.png" pageId="1" pageNumber="616">Figs. 1C</figureCitation>
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||
,
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<figureCitation id="1378FE8B167DE35FAB3AF394BE252AD1" box="[578,615,472,496]" captionStart="Fig" captionStartId="1.[812,847,1977,1999]" captionTargetBox="[812,1451,1572,1954]" captionTargetId="figure-266@1.[812,1451,1572,1954]" captionTargetPageId="1" captionText="Fig. 2.A, C, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, frontal view of carapace; C, D, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4504172" httpUri="https://zenodo.org/record/4504172/files/figure.png" pageId="1" pageNumber="616">2A</figureCitation>
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||
). Exopod of third maxilliped relatively slender, with long flagellum (
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<figureCitation id="1378FE8B167DE35FABBDF3B4BCD52911" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="1" pageNumber="616">Fig. 3A</figureCitation>
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). Male chelae with long fingers, scattered granules, rugosities on dorsal and ventral surfaces of palm, as well as on proximal part of dactylus; dorsal margin of dactylus with 5 or 6 low, rounded granules on proximal half (
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||
<figureCitation id="1378FE8B167DE35FABBDF034BCD62991" captionStart="Fig" captionStartId="1.[812,847,1977,1999]" captionTargetBox="[812,1451,1572,1954]" captionTargetId="figure-266@1.[812,1451,1572,1954]" captionTargetPageId="1" captionText="Fig. 2.A, C, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, frontal view of carapace; C, D, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.4504172" httpUri="https://zenodo.org/record/4504172/files/figure.png" pageId="1" pageNumber="616">Fig. 2C</figureCitation>
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||
). Ambulatory legs with relatively broad meri, with sharp subdistal spine on dorsal margin, surface gently rugose; propodus subrectangular (
|
||
<figureCitation id="1378FE8B167DE35FA8E1F094BDBC29D1" box="[409,510,728,752]" captionStart="Fig" captionStartId="1.[113,148,1976,1998]" captionTargetBox="[113,752,1116,1953]" captionTargetId="figure-214@1.[113,752,1116,1953]" captionTargetPageId="1" captionText="Fig. 1. A, C, Geosesarma bintan, new species, holotype, male,11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, habitus; C, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504168" httpUri="https://zenodo.org/record/4504168/files/figure.png" pageId="1" pageNumber="616">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="1378FE8B167DE35FAB76F094BE7329D1" box="[526,561,728,752]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="1" pageNumber="616">3C</figureCitation>
|
||
). Male abdomen relatively broad; telson semicircular (
|
||
<figureCitation id="1378FE8B167DE35FAB71F0B4BE1C2831" box="[521,606,760,784]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="1" pageNumber="616">Fig. 3E</figureCitation>
|
||
). G1 slender; distal chitinous part elongate; spatuliform, subtruncate tip with weak indentation (
|
||
<figureCitation id="1378FE8B167DE35FA80BF174BDF12871" box="[371,435,824,848]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="1" pageNumber="616">Fig. 3</figureCitation>
|
||
G–I).
|
||
</paragraph>
|
||
<paragraph id="8BFCE20E167DE35FA909F134BD4D2891" blockId="1.[113,752,888,944]" pageId="1" pageNumber="616">
|
||
<emphasis id="B9373E1C167DE35FA909F134BC8428B1" bold="true" box="[113,198,888,912]" pageId="1" pageNumber="616">Colour.</emphasis>
|
||
In life, the specimens were brownish red, with pale yellow chelae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359B185167DE35FA909F194BD012F11" pageId="1" pageNumber="616" type="etymology">
|
||
<paragraph id="8BFCE20E167DE35FA909F194BD012F11" blockId="1.[113,752,984,1072]" pageId="1" pageNumber="616">
|
||
<emphasis id="B9373E1C167DE35FA909F194BCB028D1" bold="true" box="[113,242,984,1008]" pageId="1" pageNumber="616">Etymology.</emphasis>
|
||
The species is named after the island where it was discovered. The specific epithet,
|
||
<taxonomicName id="4C43998D167DE35FAB6EF1B4BE192F31" authorityName="Schubart & Ng" authorityYear="2014" box="[534,603,1016,1040]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="bintan">
|
||
<emphasis id="B9373E1C167DE35FAB6EF1B4BE192F31" box="[534,603,1016,1040]" italics="true" pageId="1" pageNumber="616">bintan</emphasis>
|
||
</taxonomicName>
|
||
, is used as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C359B185167DE35FAA55F2F1B81C2FB4" pageId="1" pageNumber="616" type="discussion">
|
||
<paragraph id="8BFCE20E167DE35FAA55F2F1BFBA2894" blockId="1.[811,1454,189,949]" pageId="1" pageNumber="616">
|
||
<emphasis id="B9373E1C167DE35FAA55F2F1BFD92BF4" bold="true" box="[813,923,189,213]" pageId="1" pageNumber="616">Remarks.</emphasis>
|
||
<taxonomicName id="4C43998D167DE35FAADBF2F2B8362BF4" authority="Schubart & Ng, 2014" authorityName="Schubart & Ng" authorityYear="2014" box="[931,1140,189,213]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="bintan" status="sp. nov.">
|
||
<emphasis id="B9373E1C167DE35FAADBF2F2B8362BF4" box="[931,1140,189,213]" italics="true" pageId="1" pageNumber="616">Geosesarma bintan</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A2048367167DE35FADFBF2F2B9452BF4" box="[1155,1287,189,213]" pageId="1" pageNumber="616" rank="species">new species</taxonomicNameLabel>
|
||
, belongs to the same species group as
|
||
<taxonomicName id="4C43998D167DE35FAD4CF292B9252BD4" authority="(Nobili, 1903)" baseAuthorityName="Nobili" baseAuthorityYear="1903" box="[1076,1383,221,245]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="B9373E1C167DE35FAD4CF292B8FF2BD4" box="[1076,1213,222,245]" italics="true" pageId="1" pageNumber="616">G. peraccae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD29FFF167DE35FADB7F291B9222BD4" author="Nobili G" box="[1231,1376,221,245]" pageId="1" pageNumber="616" pagination="1 - 39" refId="ref3972" refString="Nobili G (1903) Crostacei di Singapore. Bolletino dei Musei di Zoologia ed Anatomia comparata della R. Universita di Torino, 18 (455): 1 - 39, 1 pl." type="journal article" year="1903">Nobili, 1903</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in its carapace being distinctly broader than long, the presence of a flagellum on the exopod of the third maxilliped, the relatively broad meri of the ambulatory legs, and a G1 which is slender and its chitinous tip spatuliform (
|
||
<bibRefCitation id="EFD29FFF167DE35FAC7BF311B92E2A54" author="Ng PKL" box="[1283,1388,349,373]" pageId="1" pageNumber="616" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
). It is perhaps closest to
|
||
<taxonomicName id="4C43998D167DE35FAA94F332B82F2AB4" baseAuthorityName="Nobili" baseAuthorityYear="1903" box="[1004,1133,382,405]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="B9373E1C167DE35FAA94F332B82F2AB4" box="[1004,1133,382,405]" italics="true" pageId="1" pageNumber="616">G. peraccae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C43998D167DE35FADE6F332BF292A94" authority="(Tweedie, 1940)" baseAuthorityName="Tweedie" baseAuthorityYear="1940" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="penangense">
|
||
<emphasis id="B9373E1C167DE35FADE6F332B9792AB4" box="[1182,1339,382,405]" italics="true" pageId="1" pageNumber="616">G. penangense</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD29FFF167DE35FAC31F331BF212A94" author="Tweedie MWF" pageId="1" pageNumber="616" pagination="88 - 113" refId="ref4140" refString="Tweedie MWF (1940) New and interesting Malaysian species of Sesarma and Utica (Crustacea, Brachyura). Bulletin of the Raffles Museum, 16: 88 - 113, figs. 1 - 12, pl. XXIV." type="journal article" year="1940">Tweedie, 1940</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
in the general form of its carapace and gonopods. The new species can be separated from
|
||
<taxonomicName id="4C43998D167DE35FAD9FF3F2B92E2AF4" baseAuthorityName="Nobili" baseAuthorityYear="1903" box="[1255,1388,446,469]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="B9373E1C167DE35FAD9FF3F2B92E2AF4" box="[1255,1388,446,469]" italics="true" pageId="1" pageNumber="616">G. peraccae</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54F85CAC167DE35FAC04F3F1B9E92AF4" box="[1404,1451,445,469]" pageId="1" pageNumber="616">type</typeStatus>
|
||
locality:
|
||
<collectingCountry id="F354A29E167DE35FAAEBF391B8472AD4" box="[915,1029,477,501]" name="Singapore" pageId="1" pageNumber="616">Singapore</collectingCountry>
|
||
) by its relatively more slender third maxilliped meri (
|
||
<figureCitation id="1378FE8B167DE35FAA9AF3B1B8712934" box="[994,1075,509,533]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="1" pageNumber="616">Fig. 3A</figureCitation>
|
||
) (distinctly broader in
|
||
<taxonomicName id="4C43998D167DE35FAC5AF3B2B9E12934" baseAuthorityName="Nobili" baseAuthorityYear="1903" box="[1314,1443,510,533]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="B9373E1C167DE35FAC5AF3B2B9E12934" box="[1314,1443,510,533]" italics="true" pageId="1" pageNumber="616">G. peraccae</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EFD29FFF167DE35FAA2AF052BFFF2914" author="Ng PKL" box="[850,957,541,566]" pageId="1" pageNumber="616" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
: fig. 56C), relatively longer ambulatory legs (
|
||
<figureCitation id="1378FE8B167DE35FAA4DF071BFD42974" box="[821,918,573,597]" captionStart="Fig" captionStartId="1.[113,148,1976,1998]" captionTargetBox="[113,752,1116,1953]" captionTargetId="figure-214@1.[113,752,1116,1953]" captionTargetPageId="1" captionText="Fig. 1. A, C, Geosesarma bintan, new species, holotype, male,11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, habitus; C, D, dorsal view of carapace." figureDoi="http://doi.org/10.5281/zenodo.4504168" httpUri="https://zenodo.org/record/4504168/files/figure.png" pageId="1" pageNumber="616">Figs. 1A</figureCitation>
|
||
,
|
||
<figureCitation id="1378FE8B167DE35FAADEF071BF872974" box="[934,965,573,597]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="1" pageNumber="616">3C</figureCitation>
|
||
) (shorter in
|
||
<taxonomicName id="4C43998D167DE35FAD2CF072B8982974" baseAuthorityName="Nobili" baseAuthorityYear="1903" box="[1108,1242,574,597]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="B9373E1C167DE35FAD2CF072B8982974" box="[1108,1242,574,597]" italics="true" pageId="1" pageNumber="616">G. peraccae</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EFD29FFF167DE35FAC68F072B9392974" author="Ng PKL" box="[1296,1403,573,598]" pageId="1" pageNumber="616" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
: fig. 56A;
|
||
<bibRefCitation id="EFD29FFF167DE35FAA13F011BFBE2954" author="Serene R" box="[875,1020,605,629]" pageId="1" pageNumber="616" pagination="1084 - 1095" refId="ref4057" refString="Serene R (1968) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum naturelle d'Histoire National, Paris, (2) 39 (5): 1084 - 1095, pls. 1, 2." type="book chapter" year="1968">Serène, 1968</bibRefCitation>
|
||
: pl. 1 fig. 2), and relatively shorter G1, with the chitinised part proportionately much shorter (
|
||
<figureCitation id="1378FE8B167DE35FACF8F031BF7B2994" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="1" pageNumber="616">Fig. 3</figureCitation>
|
||
G–I) (proportionately longer with the chitinised part more elongate in
|
||
<taxonomicName id="4C43998D167DE35FAAD6F0F2B87229F4" baseAuthorityName="Nobili" baseAuthorityYear="1903" box="[942,1072,702,725]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="B9373E1C167DE35FAAD6F0F2B87229F4" box="[942,1072,702,725]" italics="true" pageId="1" pageNumber="616">G. peraccae</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EFD29FFF167DE35FAD1BF0F2B88E29F4" author="Ng PKL" box="[1123,1228,701,726]" pageId="1" pageNumber="616" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
: fig. 56D–F;
|
||
<bibRefCitation id="EFD29FFF167DE35FAC24F0F1BF2529D4" author="Serene R" pageId="1" pageNumber="616" pagination="1084 - 1095" refId="ref4057" refString="Serene R (1968) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum naturelle d'Histoire National, Paris, (2) 39 (5): 1084 - 1095, pls. 1, 2." type="book chapter" year="1968">Serène, 1968</bibRefCitation>
|
||
: fig. 3, 4). It can be separated from
|
||
<taxonomicName id="4C43998D167DE35FAC7FF092B9EE29D4" baseAuthorityName="Tweedie" baseAuthorityYear="1940" box="[1287,1452,734,757]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="penangense">
|
||
<emphasis id="B9373E1C167DE35FAC7FF092B9EE29D4" box="[1287,1452,734,757]" italics="true" pageId="1" pageNumber="616">G. penangense</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54F85CAC167DE35FAA4CF0B1BF212834" box="[820,867,765,789]" pageId="1" pageNumber="616">type</typeStatus>
|
||
locality: Penang) by its relatively more slender third maxilliped merus (
|
||
<figureCitation id="1378FE8B167DE35FAA81F151B80C2814" box="[1017,1102,797,821]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="1" pageNumber="616">Fig. 3A</figureCitation>
|
||
) (broader in
|
||
<taxonomicName id="4C43998D167DE35FAD99F152B9C22814" baseAuthorityName="Tweedie" baseAuthorityYear="1940" box="[1249,1408,798,821]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="penangense">
|
||
<emphasis id="B9373E1C167DE35FAD99F152B9C22814" box="[1249,1408,798,821]" italics="true" pageId="1" pageNumber="616">G. penangense</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EFD29FFF167DE35FAA57F172BFDB2874" author="Ng PKL" box="[815,921,829,854]" pageId="1" pageNumber="616" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
: fig. 58C) and relatively more slender G1, with the chitinised part subtruncate (
|
||
<figureCitation id="1378FE8B167DE35FAD05F111B8FE2854" box="[1149,1212,861,885]" captionStart="Fig" captionStartId="2.[136,171,1976,1998]" captionTargetBox="[183,1424,193,1952]" captionTargetId="figure-7@2.[171,1438,178,1963]" captionTargetPageId="2" captionText="Fig. 3. A, C, E, G–I, Geosesarma bintan, new species, holotype, male, 11.2 × 9.8 mm (MZB Cru 3813), Bintan Island; B, D, F, J–M, Geosesarma raj, new species, holotype, male, 11.8 × 10.9 mm (MZB Cru 3814, ex ZRC 2007.0676), Bintan Island. A, B, outer view of left third maxilliped; C, D, left fourth ambulatory leg, dorsal view; E, F, male abdomen; G, ventral view of left G1 (denuded); H, lateral view of left G1 (denuded); I, dorsal view of left G1 (denuded); J, ventral view of left G1 (denuded); K, dorsal view of left G1 (denuded); L, M, distal part of left G1 (denuded). Scale bars: A, B, E–I = 1.0 mm; C, D = 2.0 mm; J, K = 0.5 mm; L, M = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4504174" httpUri="https://zenodo.org/record/4504174/files/figure.png" pageId="1" pageNumber="616">Fig. 3</figureCitation>
|
||
G–I) (stouter, with the chitinised part gently tapering, in
|
||
<taxonomicName id="4C43998D167DE35FADD1F132B90E28B4" baseAuthorityName="Tweedie" baseAuthorityYear="1940" box="[1193,1356,894,917]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="penangense">
|
||
<emphasis id="B9373E1C167DE35FADD1F132B90E28B4" box="[1193,1356,894,917]" italics="true" pageId="1" pageNumber="616">G. penangense</emphasis>
|
||
</taxonomicName>
|
||
; cf.
|
||
<bibRefCitation id="EFD29FFF167DE35FACFCF132BF242894" author="Ng PKL" pageId="1" pageNumber="616" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
: fig. 58D–F).
|
||
</paragraph>
|
||
<paragraph id="8BFCE20E167DE35FAA55F191B81C2FB4" blockId="1.[812,1452,989,1173]" pageId="1" pageNumber="616">
|
||
The
|
||
<typeStatus id="54F85CAC167DE35FAA24F191BFFB28D4" box="[860,953,989,1013]" pageId="1" pageNumber="616" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C43998D167DE35FAAA1F192B87F28D4" authorityName="Schubart & Ng" authorityYear="2014" box="[985,1085,989,1013]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="bintan">
|
||
<emphasis id="B9373E1C167DE35FAAA1F192B87F28D4" box="[985,1085,989,1013]" italics="true" pageId="1" pageNumber="616">G. bintan</emphasis>
|
||
</taxonomicName>
|
||
, a male, is already mature, with its gonopods fully developed and chitinised, although its chelae are not enlarged, suggesting it can reach larger sizes. The same is known of the allied
|
||
<taxonomicName id="4C43998D167DE35FAD13F672B8B32F74" baseAuthorityName="Nobili" baseAuthorityYear="1903" box="[1131,1265,1086,1109]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="616" phylum="Arthropoda" rank="species" species="peraccae">
|
||
<emphasis id="B9373E1C167DE35FAD13F672B8B32F74" box="[1131,1265,1086,1109]" italics="true" pageId="1" pageNumber="616">G. peraccae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F354A29E167DE35FAC41F671B9EA2F74" box="[1337,1448,1085,1109]" name="Singapore" pageId="1" pageNumber="616">Singapore</collectingCountry>
|
||
, where only very large males have enlarged and brightly coloured chelae (
|
||
<bibRefCitation id="EFD29FFF167DE35FAA9FF631B8132FB4" author="Ng PKL" box="[999,1105,1149,1173]" pageId="1" pageNumber="616" pagination="1 - 156" refId="ref3613" refString="Ng PKL (1988) The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, i-viii, 1 - 156 pp., figs. 1 - 63, 4 pls." type="book chapter" year="1988">Ng, 1988</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |