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<document id="F86C905E8549551F0324C9E4A140E940" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFFE5010FF23F9DEFAF54062" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus procerus Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="17" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="15" updateTime="1726053143081" updateUser="ExternalLinkService">
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<mods:title id="45EE4D21E878C36B1ED6F6A0EEDE8A4D">The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:date id="6DB8A176E30EC151A9FB71C901CC5DAA">2024</mods:date>
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<mods:classification id="07ECC75C7D297F81C2CFCFB947323726">journal article</mods:classification>
<mods:identifier id="A59F7C986052EEA9C9ADFCDC0AFD0BDD" type="DOI">10.11646/zootaxa.5505.1.1</mods:identifier>
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<treatment id="03E7FF50FFFE5010FF23F9DEFAF54062" ID-DOI="http://doi.org/10.5281/zenodo.13746009" ID-Zenodo-Dep="13746009" LSID="urn:lsid:plazi:treatment:03E7FF50FFFE5010FF23F9DEFAF54062" httpUri="http://treatment.plazi.org/id/03E7FF50FFFE5010FF23F9DEFAF54062" lastPageId="16" lastPageNumber="17" pageId="14" pageNumber="15">
<subSubSection id="C3541DCDFFFE500EFF23F9DEFD09425E" box="[151,716,1554,1581]" pageId="14" pageNumber="15" type="nomenclature">
<paragraph id="8BF14E46FFFE500EFF23F9DEFD09425E" blockId="14.[151,716,1554,1617]" box="[151,716,1554,1581]" pageId="14" pageNumber="15">
<heading id="D0B9F92AFFFE500EFF23F9DEFD09425E" bold="true" box="[151,716,1554,1581]" fontSize="11" level="2" pageId="14" pageNumber="15" reason="4">
<emphasis id="B93A9254FFFE500EFF23F9DEFD09425E" bold="true" box="[151,716,1554,1581]" pageId="14" pageNumber="15">
<taxonomicName id="4C4E35C5FFFE500EFF23F9DEFD91425F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,596,1554,1581]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="procerus" status="comb. nov.">
<emphasis id="B93A9254FFFE500EFF23F9DEFE28425E" bold="true" box="[151,493,1554,1581]" italics="true" pageId="14" pageNumber="15">Anhoraeomorphus procerus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFFE500EFDD5F9DFFD09425E" box="[609,716,1555,1581]" pageId="14" pageNumber="15" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFE500EFF23F9FBFEE24222" box="[151,295,1591,1617]" pageId="14" pageNumber="15" type="description">
<paragraph id="8BF14E46FFFE500EFF23F9FBFEE24222" blockId="14.[151,716,1554,1617]" box="[151,295,1591,1617]" pageId="14" pageNumber="15">
(
<figureCitation id="137552C3FFFE500EFF2BF9FBFEDA4222" box="[159,287,1591,1617]" captionStart="FIGURES 36–43" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 36–43. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="14" pageNumber="15">Figs 36–43</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFE500EFF23F9B7FDA642E0" box="[151,611,1659,1683]" pageId="14" pageNumber="15" type="reference_group">
<paragraph id="8BF14E46FFFE500EFF23F9B7FDA642E0" blockId="14.[151,611,1659,1683]" box="[151,611,1659,1683]" pageId="14" pageNumber="15">
<treatmentCitationGroup id="AB5E6968FFFE500EFF23F9B7FDA642E0" box="[151,611,1659,1683]" pageId="14" pageNumber="15">
<treatmentCitation id="0AEF6857FFFE500EFF23F9B7FD9A42E0" author="Franz, H." box="[151,607,1659,1683]" page="166" pageId="14" pageNumber="15" year="1986">
<taxonomicName id="4C4E35C5FFFE500EFF23F9B7FD9A42E0" ID-CoL="3MNL6" authority="Franz, 1986 b: 166" authorityName="Franz" authorityPageNumber="166" authorityYear="1986" box="[151,607,1659,1683]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B93A9254FFFE500EFF23F9B7FE5842E0" box="[151,413,1659,1683]" italics="true" pageId="14" pageNumber="15">Horaeomorphus procerus</emphasis>
<bibRefCitation id="EFDF33B7FFFE500EFE17F9B0FD9A42E0" author="Franz, H." box="[419,607,1659,1683]" pageId="14" pageNumber="15" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 166</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFFE500FFF23F90DFADB448A" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="materials_examined">
<paragraph id="8BF14E46FFFE500EFF23F90DFEC84334" blockId="14.[151,1437,1729,2008]" pageId="14" pageNumber="15">
<emphasis id="B93A9254FFFE500EFF23F90DFE7042A8" bold="true" box="[151,437,1729,1755]" pageId="14" pageNumber="15">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFFE500EFE71F90DFEC84334" collectionCode="NHMW" country="Madagascar" determinerName="H. Franz" location="MADAGASCAR" pageId="14" pageNumber="15" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93A9254FFFE500EFE71F90DFD3742A8" bold="true" box="[453,754,1729,1755]" pageId="14" pageNumber="15">
<typeStatus id="54F5F0E4FFFE500EFE71F90DFDF742A8" box="[453,562,1729,1755]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
(Madagascar):
</emphasis>
<specimenCount id="9D4885CFFFFE500EFCB6F90EFCDC42AF" box="[770,793,1730,1756]" count="1" pageId="14" pageNumber="15" type="male">♂</specimenCount>
, three labels: “Madagascar Est / district Sambava / Marojejy / Ambinanitelo” [white, printed], “INSTITUT / SCIENTIFIQUE /
<collectingCountry id="F3590ED6FFFE500EFBB6F929FB0C428C" box="[1026,1225,1765,1791]" name="Madagascar" pageId="14" pageNumber="15">MADAGASCAR</collectingCountry>
” [bluish, printed], “
<taxonomicName id="4C4E35C5FFFE500EFF15F8C5FE9D4350" box="[161,344,1801,1827]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ procerus m. / det.
<determinerName id="7E2DA282FFFE500EFDF5F8C5FD5A4350" box="[577,671,1801,1827]" pageId="14" pageNumber="15">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFFE500EFF2BF8E1FEC74334" box="[159,258,1837,1863]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="15" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFFE500FFF73F89DFADB448A" blockId="14.[151,1437,1729,2008]" lastBlockId="15.[151,1436,151,249]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<emphasis id="B93A9254FFFE500EFF73F89DFE664318" bold="true" box="[199,419,1873,1899]" pageId="14" pageNumber="15">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFFE500EFE4EF89DFD884318" box="[506,589,1873,1899]" captionStart="FIGURES 36–43" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 36–43. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="14" pageNumber="15">Fig. 36</figureCitation>
) slender; head (
<figureCitation id="137552C3FFFE500EFCB1F89DFC9C4318" box="[773,857,1873,1899]" captionStart="FIGURES 36–43" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 36–43. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="14" pageNumber="15">Fig. 37</figureCitation>
) about as long as broad, with vertex transverse, in anterodorsal view its posterior margin truncate at middle; pronotum (
<figureCitation id="137552C3FFFE500EFC04F8B9FBC143FC" box="[944,1028,1909,1935]" captionStart="FIGURES 36–43" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 36–43. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="14" pageNumber="15">Fig. 36</figureCitation>
) bell-shaped, broadest near middle, with two pairs of deep antebasal pits: large and deep inner pits and barely discernible, small and shallow outer pits; antennomeres 1–2 elongate, 3 about as long as broad, 4–7 each elongate, 8–10 each about as long as broad; all femora similarly slender; metatibiae in males unmodified; aedeagus (
<figureCitation id="137552C3FFFF500FFC71FF5BFB8344C2" box="[965,1094,151,177]" captionStart="FIGURES 36–43" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 36–43. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="15" pageNumber="16">Figs 40–43</figureCitation>
) in ventral view broadest near base, with apical region of dorsal wall nearly straight, distolateral endophallic sclerites broadly separated, each with hook-like apex curved mesad, each paramere with one long and four subapical setae of various lengths.
</paragraph>
</subSubSection>
<caption id="DF311ECEFFFF500FFF23F943FEF5429C" ID-DOI="http://doi.org/10.5281/zenodo.13745935" ID-Zenodo-Dep="13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="15" pageNumber="16" startId="15.[151,264,1679,1703]" targetBox="[151,1418,273,1635]" targetPageId="15" targetType="figure">
<paragraph id="8BF14E46FFFF500FFF23F943FEF5429C" blockId="15.[151,1436,1679,1775]" pageId="15" pageNumber="16">
<emphasis id="B93A9254FFFF500FFF23F943FE9042D4" bold="true" box="[151,341,1679,1703]" pageId="15" pageNumber="16">FIGURES 36–43.</emphasis>
<taxonomicName id="4C4E35C5FFFF500FFEEFF943FD0F42D4" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[347,714,1679,1703]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B93A9254FFFF500FFEEFF943FDB242D4" box="[347,631,1679,1703]" italics="true" pageId="15" pageNumber="16">Anhoraeomorphus procerus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFFF500FFB9BF943FB8D42D4" bold="true" box="[1071,1096,1679,1703]" pageId="15" pageNumber="16">36</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFFF500FFAC7F943FA4942D4" bold="true" box="[1395,1420,1679,1703]" pageId="15" pageNumber="16">37</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFFF500FFCE3F97FFCB542B8" bold="true" box="[855,880,1715,1739]" pageId="15" pageNumber="16">38</emphasis>
); antenna (
<emphasis id="B93A9254FFFF500FFC57F97FFC3942B8" bold="true" box="[995,1020,1715,1739]" pageId="15" pageNumber="16">39</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFFF500FFB50F97FFAE642B8" bold="true" box="[1252,1315,1715,1739]" pageId="15" pageNumber="16">40, 42</emphasis>
) and lateral (
<emphasis id="B93A9254FFFF500FFF14F91BFF1A429C" bold="true" box="[160,223,1751,1775]" pageId="15" pageNumber="16">41, 43</emphasis>
) views.
</paragraph>
</caption>
<subSubSection id="C3541DCDFFFF5010FF73F88DFE5B472E" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="description">
<paragraph id="8BF14E46FFFF500FFF73F88DFF0F43F3" blockId="15.[151,1436,1857,2028]" pageId="15" pageNumber="16">
<emphasis id="B93A9254FFFF500FFF73F88DFEB04328" bold="true" box="[199,373,1857,1883]" pageId="15" pageNumber="16">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFFF500FFD91F88DFDB34328" box="[549,630,1857,1883]" captionStart="FIGURES 36–43" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 36–43. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="15" pageNumber="16">Fig. 36</figureCitation>
) light brown, tarsi and palps distinctly lighter, setae yellowish; BL
<quantity id="4CB6E3A3FFFF500FFADFF88DFF0343F3" metricMagnitude="-3" metricUnit="m" metricValue="1.53" pageId="15" pageNumber="16" unit="mm" value="1.53">1.53 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFFF5010FF73F845FB1B456E" blockId="15.[151,1436,1857,2028]" lastBlockId="16.[151,1437,151,1041]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
Head (
<figureCitation id="137552C3FFFF500FFEA0F845FE5243D0" box="[276,407,1929,1955]" captionStart="FIGURES 36–43" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 36–43. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="15" pageNumber="16">Figs 37–38</figureCitation>
) in anterodorsal view subquadrate, broadest at eyes, HL
<quantity id="4CB6E3A3FFFF500FFB9EF845FB5743D7" box="[1066,1170,1929,1956]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="16" unit="mm" value="0.25">0.25 mm</quantity>
, HW
<quantity id="4CB6E3A3FFFF500FFB6CF845FA8743D7" box="[1240,1346,1929,1956]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="15" pageNumber="16" unit="mm" value="0.28">0.28 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFFF500FFEFAF861FE6743B4" box="[334,418,1965,1991]" captionStart="FIGURES 36–43" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 36–43. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="15" pageNumber="16">Fig. 38</figureCitation>
) slightly longer than eye; vertex and frons confluent, together weakly convex and about as long as broad; posterior margin of vertex truncate at middle; each eye large and strongly convex, with straight posteromedian margin. Punctures on vertex fine, inconspicuous; setae short, sparse and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFE05010FC32FF77FC1C44A6" box="[902,985,187,213]" captionStart="FIGURES 36–43" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 36–43. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="16" pageNumber="17">Fig. 39</figureCitation>
) slender but short, slightly shorter than half BL, AnL
<quantity id="4CB6E3A3FFE05010FE81FF13FE5E448A" box="[309,411,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="7.3" pageId="16" pageNumber="17" unit="mm" value="0.73">0.73 mm</quantity>
, antennomeres 1–2 elongate, 3 about as long as broad, 4–7 each elongate, 8–10 each about as long as broad; 11 shorter than 9 and 10 combined, as wide as 10, nearly twice as long as broad.
</paragraph>
<paragraph id="8BF14E46FFE05010FF73FEEBFE6B4586" blockId="16.[151,1437,151,1041]" pageId="16" pageNumber="17">
Pronotum (
<figureCitation id="137552C3FFE05010FEFCFEEBFE584532" box="[328,413,295,321]" captionStart="FIGURES 28–35" captionStartId="13.[151,264,1909,1933]" captionTargetBox="[151,1434,614,1874]" captionTargetId="figure-99@13.[151,1435,614,1886]" captionTargetPageId="13" captionText="FIGURES 28–35. Anhoraeomorphus ivelonanus (Franz), holotype. Habitus in dorsal view (28); head in anterodorsal view (29); head and prothorax in lateral view (only bristles on head are shown) (30); antenna (31); aedeagus in ventral (32, 34) and lateral (33, 35) views." figureDoi="http://doi.org/10.5281/zenodo.13745931" httpUri="https://zenodo.org/record/13745931/files/figure.png" pageId="16" pageNumber="17">Fig. 28</figureCitation>
) bell-shaped, broadest near middle; PL
<quantity id="4CB6E3A3FFE05010FCC4FEEBFC1F4532" box="[880,986,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="16" pageNumber="17" unit="mm" value="0.43">0.43 mm</quantity>
, PW
<quantity id="4CB6E3A3FFE05010FBA9FEEBFB424532" box="[1053,1159,295,321]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="16" pageNumber="17" unit="mm" value="0.33">0.33 mm</quantity>
. Anterior margin nearly straight, lateral margins rounded, more strongly converging anteriorly than posteriorly; posterior corners blunt; posterior margin indistinctly convex. Base with two pairs of pits: inner pits conspicuously large, deep, outer pits barely discernible, small and shallow, and with barely discernible, extremely short vestiges of sublateral carinae. Disc with fine, inconspicuous punctures; setae sparse, short and suberect, lateral and lateroventral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFE05010FF73FE33FE7A4612" blockId="16.[151,1437,151,1041]" pageId="16" pageNumber="17">
Elytra (
<figureCitation id="137552C3FFE05010FEAEFE33FEAF466A" box="[282,362,511,537]" captionStart="FIGURES 36–43" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 36–43. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="16" pageNumber="17">Fig. 36</figureCitation>
) together oval, broadest slightly anterior to middle; EL
<quantity id="4CB6E3A3FFE05010FC65FE32FBF3466A" box="[977,1078,510,537]" metricMagnitude="-4" metricUnit="m" metricValue="8.5" pageId="16" pageNumber="17" unit="mm" value="0.85">0.85 mm</quantity>
, EW
<quantity id="4CB6E3A3FFE05010FBC6FE32FB12466A" box="[1138,1239,510,537]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="16" pageNumber="17" unit="mm" value="0.58">0.58 mm</quantity>
, EI 1.48; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures fine and inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFE05010FF73FDA7FBCC46F6" blockId="16.[151,1437,151,1041]" box="[199,1033,619,645]" pageId="16" pageNumber="17">Legs moderately long, slender, unmodified except slightly recurved tibiae.</paragraph>
<paragraph id="8BF14E46FFE05010FF73FD43FABF474A" blockId="16.[151,1437,151,1041]" pageId="16" pageNumber="17">
Aedeagus (
<figureCitation id="137552C3FFE05010FEF7FD43FE0746DA" box="[323,450,655,681]" captionStart="FIGURES 36–43" captionStartId="15.[151,264,1679,1703]" captionTargetBox="[151,1418,273,1635]" captionTargetId="figure-64@15.[151,1435,273,1655]" captionTargetPageId="15" captionText="FIGURES 36–43. Anhoraeomorphus procerus (Franz), holotype. Habitus in dorsal view (36); head in anterodorsal view (37); head and prothorax in lateral view (only bristles on head are shown) (38); antenna (39); aedeagus in ventral (40, 42) and lateral (41, 43) views." figureDoi="http://doi.org/10.5281/zenodo.13745935" httpUri="https://zenodo.org/record/13745935/files/figure.png" pageId="16" pageNumber="17">Figs 40–43</figureCitation>
) slender, AeL
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; in ventral view broadest near base and weakly narrowing towards step-wise delimited apical region, which is subtriangular with truncate apex; in lateral view apical region of dorsal plate nearly straight; endophallus with slightly asymmetrical and elongate distolateral sclerites and asymmetrical median structures, distolateral sclerites broadly separated, each with hook-like apex curved mesally; parameres thick, shorter than median lobe, each paramere with one long apical and four subapical setae of various lengths.
</paragraph>
<paragraph id="8BF14E46FFE05010FF73FC8FFE5B472E" blockId="16.[151,1437,151,1041]" box="[199,414,835,861]" pageId="16" pageNumber="17">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE05010FF73FCAAFCCF47F2" box="[199,778,870,897]" pageId="16" pageNumber="17" type="distribution">
<paragraph id="8BF14E46FFE05010FF73FCAAFCCF47F2" blockId="16.[151,1437,151,1041]" box="[199,778,870,897]" pageId="16" pageNumber="17">
<emphasis id="B93A9254FFE05010FF73FCAAFEA447F3" bold="true" box="[199,353,870,896]" pageId="16" pageNumber="17">Distribution.</emphasis>
Northern
<collectingCountry id="F3590ED6FFE05010FE60FCABFD9E47F2" box="[468,603,871,897]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
(Sava Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE05010FF73FC47FAF54062" pageId="16" pageNumber="17" type="discussion">
<paragraph id="8BF14E46FFE05010FF73FC47FB47479E" blockId="16.[151,1437,151,1041]" pageId="16" pageNumber="17">
<emphasis id="B93A9254FFE05010FF73FC47FEFE47D6" bold="true" box="[199,315,907,933]" pageId="16" pageNumber="17">Remarks.</emphasis>
<taxonomicName id="4C4E35C5FFE05010FEF1FC47FD4747D7" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[325,642,907,933]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B93A9254FFE05010FEF1FC47FD4747D7" box="[325,642,907,933]" italics="true" pageId="16" pageNumber="17">Anhoraeomorphus procerus</emphasis>
</taxonomicName>
was described based on
<specimenCount id="9D4885CFFFE05010FC10FC47FB9547D6" box="[932,1104,907,933]" count="2" pageId="16" pageNumber="17" type="generic">two specimens</specimenCount>
; only the
<typeStatus id="54F5F0E4FFE05010FB71FC47FAE247D6" box="[1221,1319,907,933]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
male was studied. The
<typeStatus id="54F5F0E4FFE05010FE9FFC63FE4E47BA" box="[299,395,943,969]" pageId="16" pageNumber="17" type="paratype">paratype</typeStatus>
male is deposited in Muséum National d'Histoire Naturelle, Paris and was not available for my study. The village Ambinanitelo was misspelled by
<bibRefCitation id="EFDF33B7FFE05010FCB0FC1FFC66479E" author="Franz, H." box="[772,931,979,1005]" pageId="16" pageNumber="17" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
as “Ambinaniteta”.
</paragraph>
<paragraph id="8BF14E46FFE05010FF73FC3BFAF54062" blockId="16.[151,1437,151,1041]" box="[199,1328,1015,1041]" pageId="16" pageNumber="17">
<taxonomicName id="4C4E35C5FFE05010FF73FC3BFDC74063" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[199,514,1015,1041]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="procerus">
<emphasis id="B93A9254FFE05010FF73FC3BFDC74063" box="[199,514,1015,1041]" italics="true" pageId="16" pageNumber="17">Anhoraeomorphus procerus</emphasis>
</taxonomicName>
is externally similar to
<taxonomicName id="4C4E35C5FFE05010FCBEFC3BFC674062" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[778,930,1015,1041]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="ivelonanus">
<emphasis id="B93A9254FFE05010FCBEFC3BFC674062" box="[778,930,1015,1041]" italics="true" pageId="16" pageNumber="17">A. ivelonanus</emphasis>
</taxonomicName>
, see Remarks for the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>
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