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<document id="067EEAC0C563797A7DD36630BB6875E9" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFE05012FF23FBAEFA844586" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus sakarahanus Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="19" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="17" updateTime="1726053143081" updateUser="ExternalLinkService">
<mods:mods id="6CE1903D1ED8E0E2023B6247DD78DA0B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="E4CC5769CA7461936E8687E940A40646">The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:date id="839A8F8A6973D320F26252E230FA69A7">2024</mods:date>
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<mods:number id="68A445661FF9B60598B231E881073428">2024-09-04</mods:number>
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<mods:classification id="D7442D2841D56911BF304C934AAF86A7">journal article</mods:classification>
<mods:identifier id="1EDA61C8D6A838F13FB38EB296E6E77F" type="DOI">10.11646/zootaxa.5505.1.1</mods:identifier>
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<treatment id="03E7FF50FFE05012FF23FBAEFA844586" ID-DOI="http://doi.org/10.5281/zenodo.13746013" ID-Zenodo-Dep="13746013" LSID="urn:lsid:plazi:treatment:03E7FF50FFE05012FF23FBAEFA844586" httpUri="http://treatment.plazi.org/id/03E7FF50FFE05012FF23FBAEFA844586" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection id="C3541DCDFFE05010FF23FBAEFD3A400E" box="[151,767,1122,1149]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8BF14E46FFE05010FF23FBAEFD3A400E" blockId="16.[151,767,1122,1185]" box="[151,767,1122,1149]" pageId="16" pageNumber="17">
<heading id="D0B9F92AFFE05010FF23FBAEFD3A400E" bold="true" box="[151,767,1122,1149]" fontSize="11" level="2" pageId="16" pageNumber="17" reason="4">
<emphasis id="B93A9254FFE05010FF23FBAEFD3A400E" bold="true" box="[151,767,1122,1149]" pageId="16" pageNumber="17">
<taxonomicName id="4C4E35C5FFE05010FF23FBAEFD43400F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,646,1122,1149]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sakarahanus" status="comb. nov.">
<emphasis id="B93A9254FFE05010FF23FBAEFDDA400E" bold="true" box="[151,543,1122,1149]" italics="true" pageId="16" pageNumber="17">Anhoraeomorphus sakarahanus</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFE05010FD20FBAFFD3A400E" box="[660,767,1123,1149]" pageId="16" pageNumber="17" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE05010FF23FB4BFEE240D2" box="[151,295,1158,1185]" pageId="16" pageNumber="17" type="description">
<paragraph id="8BF14E46FFE05010FF23FB4BFEE240D2" blockId="16.[151,767,1122,1185]" box="[151,295,1158,1185]" pageId="16" pageNumber="17">
(
<figureCitation id="137552C3FFE05010FF2BFB4BFEDA40D2" box="[159,287,1158,1185]" captionStart="FIGURES 44–51" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 44–51. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Figs 44–51</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE05010FF23FB07FD4E4090" box="[151,651,1227,1251]" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="8BF14E46FFE05010FF23FB07FD4E4090" blockId="16.[151,651,1227,1251]" box="[151,651,1227,1251]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="AB5E6968FFE05010FF23FB07FD4E4090" box="[151,651,1227,1251]" pageId="16" pageNumber="17">
<treatmentCitation id="0AEF6857FFE05010FF23FB07FD424090" author="Franz, H." box="[151,647,1227,1251]" page="165" pageId="16" pageNumber="17" year="1986">
<taxonomicName id="4C4E35C5FFE05010FF23FB07FD424090" ID-CoL="3MNLF" authority="Franz, 1986 b: 165" authorityName="Franz" authorityPageNumber="165" authorityYear="1986" box="[151,647,1227,1251]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="sakarahanus">
<emphasis id="B93A9254FFE05010FF23FB07FE004090" box="[151,453,1227,1251]" italics="true" pageId="16" pageNumber="17">Horaeomorphus sakarahanus</emphasis>
<bibRefCitation id="EFDF33B7FFE05010FE7FFB00FD424090" author="Franz, H." box="[459,647,1227,1251]" pageId="16" pageNumber="17" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 165</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE05010FF23FADDFEE24238" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8BF14E46FFE05010FF23FADDFC5F4100" blockId="16.[151,1437,1297,1972]" pageId="16" pageNumber="17">
<emphasis id="B93A9254FFE05010FF23FADDFE6E4158" bold="true" box="[151,427,1297,1323]" pageId="16" pageNumber="17">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFE05010FE02FADDFC5F4100" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Foret de Sakahara" pageId="16" pageNumber="17" specimenCount="2" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B93A9254FFE05010FE02FADDFD184158" bold="true" box="[438,733,1297,1323]" pageId="16" pageNumber="17">
<typeStatus id="54F5F0E4FFE05010FE02FADDFDE64158" box="[438,547,1297,1323]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFE05010FD81FADDFD154158" box="[565,720,1297,1323]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
):
</emphasis>
<specimenCount id="9D4885CFFFE05010FD5CFADEFD3A415F" box="[744,767,1298,1324]" count="1" pageId="16" pageNumber="17" type="male">♂</specimenCount>
, four labels: “S-Madagaskar /
<location id="8E91189DFFE05010FBDDFADEFAF94158" LSID="urn:lsid:plazi:treatment:03E7FF50FFE05012FF23FBAEFA844586:8E91189DFFE05010FBDDFADEFAF94158" box="[1129,1340,1297,1324]" country="Madagascar" name="Foret de Sakahara" pageId="16" pageNumber="17">Foret de Sakahara</location>
{sic!} / lg.
<collectorName id="26BB2B90FFE05010FF05FAF9FECA413C" box="[177,271,1333,1359]" pageId="16" pageNumber="17">H.Franz</collectorName>
1969” [white, printed], “
<taxonomicName id="4C4E35C5FFE05010FD9AFAF9FD20413C" box="[558,741,1333,1359]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ sakarahanus m. / det.
<determinerName id="7E2DA282FFE05010FC5BFAF9FB89413C" box="[1007,1100,1333,1359]" pageId="16" pageNumber="17">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten], “
<specimenCount id="9D4885CFFFE05010FDF2FA95FD994100" box="[582,604,1369,1395]" count="1" pageId="16" pageNumber="17" type="male">♂</specimenCount>
” [white, printed] (
<collectionCode id="ED5FD683FFE05010FC98FA95FC4A4100" box="[812,911,1369,1395]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="16" pageNumber="17" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFE05010FF73FAB1FEE24238" blockId="16.[151,1437,1297,1972]" pageId="16" pageNumber="17">
<emphasis id="B93A9254FFE05010FF73FAB1FE6441E4" bold="true" box="[199,417,1405,1431]" pageId="16" pageNumber="17">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFE05010FE40FAB1FD8341E4" box="[500,582,1405,1431]" captionStart="FIGURES 44–51" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 44–51. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Fig. 44</figureCitation>
) stout; head (
<figureCitation id="137552C3FFE05010FD6BFAB1FCF441EB" box="[735,817,1405,1432]" captionStart="FIGURES 44–51" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 44–51. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Fig. 45</figureCitation>
) about as long as wide, with vertex posteriorly broadly rounded; pronotum (
<figureCitation id="137552C3FFE05010FECEFA6DFE0C41C8" box="[378,457,1441,1467]" captionStart="FIGURES 44–51" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 44–51. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Fig. 44</figureCitation>
) bell-shaped, broadest near middle, with one pair of distinct antebasal pits; antennomeres 3–10 not elongate; all femora similarly slender; metatibiae in males unmodified; aedeagus (
<figureCitation id="137552C3FFE05010FB15FA09FAE74193" box="[1185,1314,1477,1504]" captionStart="FIGURES 44–51" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 44–51. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Figs 48–51</figureCitation>
) in ventral view broadest near middle, with apical region of dorsal wall abruptly bent dorsally, distolateral endophallic sclerites widely separated and with rounded apices, each paramere with a row of subapical setae distributed on less than distal fourth.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE05012FF73F999FE5B45FA" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17" type="description">
<paragraph id="8BF14E46FFE05010FF73F999FF0F42E7" blockId="16.[151,1437,1297,1972]" pageId="16" pageNumber="17">
<emphasis id="B93A9254FFE05010FF73F999FEB0421C" bold="true" box="[199,373,1621,1647]" pageId="16" pageNumber="17">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFE05010FD93F999FDBF421C" box="[551,634,1621,1647]" captionStart="FIGURES 44–51" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 44–51. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Fig. 44</figureCitation>
) light brown, appendages indistinctly lighter, setae yellowish; BL
<quantity id="4CB6E3A3FFE05010FADFF999FF0342E7" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="16" pageNumber="17" unit="mm" value="1.4">1.40 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFE05010FF73F951FC8F43C0" blockId="16.[151,1437,1297,1972]" pageId="16" pageNumber="17">
Head (
<figureCitation id="137552C3FFE05010FEA4F951FE4A42CB" box="[272,399,1693,1720]" captionStart="FIGURES 44–51" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 44–51. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Figs 45–46</figureCitation>
) in anterodorsal view rounded, broadest at eyes, HL
<quantity id="4CB6E3A3FFE05010FC61F951FBFF42CB" box="[981,1082,1693,1720]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="17" unit="mm" value="0.28">0.28 mm</quantity>
, HW
<quantity id="4CB6E3A3FFE05010FBCDF951FB1B42CB" box="[1145,1246,1693,1720]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="17" unit="mm" value="0.28">0.28 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFE05010FF6AF90DFEF542A8" box="[222,304,1729,1755]" captionStart="FIGURES 44–51" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 44–51. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Fig. 46</figureCitation>
) subequal to length of eye; vertex and frons confluent, together weakly convex and about as long as broad; vertex posteriorly strongly rounded; each eye moderately large and weakly convex, with deep posteromedian emargination, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, sparse and suberect; posterior region of frons and entire vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFE05010FEB1F89DFE904318" box="[261,341,1873,1899]" captionStart="FIGURES 44–51" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 44–51. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="16" pageNumber="17">Fig. 47</figureCitation>
) slender but short, slightly shorter than half BL, AnL
<quantity id="4CB6E3A3FFE05010FC1DF89DFBCB431F" box="[937,1038,1873,1900]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" pageId="16" pageNumber="17" unit="mm" value="0.55">0.55 mm</quantity>
, antennomeres 1 and 2 each weakly elongate, 3 slightly transverse, 4–7 each about as long as broad, 8–10 each transverse; 11 slightly shorter than 9 and 10 combined, as wide as 10, about 1.8 times as long as broad.
</paragraph>
<caption id="DF311ECEFFE15011FF23FA3FFEB34227" ID-DOI="http://doi.org/10.5281/zenodo.13745937" ID-Zenodo-Dep="13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,264,1523,1547]" targetBox="[151,1422,181,1488]" targetPageId="17" targetType="figure">
<paragraph id="8BF14E46FFE15011FF23FA3FFEB34227" blockId="17.[151,1437,1523,1620]" pageId="17" pageNumber="18">
<emphasis id="B93A9254FFE15011FF23FA3FFE93427F" bold="true" box="[151,342,1523,1548]" pageId="17" pageNumber="18">FIGURES 44–51.</emphasis>
<taxonomicName id="4C4E35C5FFE15011FEE9FA38FD32427F" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[349,759,1524,1548]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="sakarahanus">
<emphasis id="B93A9254FFE15011FEE9FA38FD67427F" box="[349,674,1524,1548]" italics="true" pageId="17" pageNumber="18">Anhoraeomorphus sakarahanus</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFE15011FBD7FA38FBB9427F" bold="true" box="[1123,1148,1524,1548]" pageId="17" pageNumber="18">44</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFE15011FF14F9D4FF7C4243" bold="true" box="[160,185,1560,1584]" pageId="17" pageNumber="18">45</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFE15011FC23F9D4FC754243" bold="true" box="[919,944,1560,1584]" pageId="17" pageNumber="18">46</emphasis>
); antenna (
<emphasis id="B93A9254FFE15011FB91F9D4FBFB4243" bold="true" box="[1061,1086,1560,1584]" pageId="17" pageNumber="18">47</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFE15011FA9CF9D4FAAD4243" bold="true" box="[1320,1384,1560,1584]" pageId="17" pageNumber="18">48, 50</emphasis>
) and lateral (
<emphasis id="B93A9254FFE15011FF51F9F0FEE14227" bold="true" box="[229,292,1596,1620]" pageId="17" pageNumber="18">49, 51</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFE15011FF73F96AFCE84323" blockId="17.[151,1437,1702,2016]" pageId="17" pageNumber="18">
Pronotum (
<figureCitation id="137552C3FFE15011FEFCF96AFE5942B3" box="[328,412,1702,1728]" captionStart="FIGURES 44–51" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 44–51. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="17" pageNumber="18">Fig. 44</figureCitation>
) bell-shaped, broadest near middle; PL
<quantity id="4CB6E3A3FFE15011FCDFF96AFC1642B3" box="[875,979,1702,1728]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="17" pageNumber="18" unit="mm" value="0.38">0.38 mm</quantity>
, PW
<quantity id="4CB6E3A3FFE15011FBA1F96AFBBA42B3" box="[1045,1151,1702,1729]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="17" pageNumber="18" unit="mm" value="0.35">0.35 mm</quantity>
. Anterior margin weakly arcuate, lateral margins rounded, anteriorly converging stronger than posteriorly; posterior corners blunt; posterior margin indistinctly convex. Base with one pair of round and deep pits, and with barely discernible, extremely short vestiges of sublateral carinae. Disc with fine, inconspicuous punctures; setae sparse, short and suberect, lateral and lateroventral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFE15011FF73F896FE3B43CF" blockId="17.[151,1437,1702,2016]" pageId="17" pageNumber="18">
Elytra (
<figureCitation id="137552C3FFE15011FEAAF896FEB64307" box="[286,371,1882,1908]" captionStart="FIGURES 44–51" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 44–51. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="17" pageNumber="18">Fig. 44</figureCitation>
) together oval, broadest distinctly in front of middle; EL
<quantity id="4CB6E3A3FFE15011FBACF896FB474307" box="[1048,1154,1882,1909]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" pageId="17" pageNumber="18" unit="mm" value="0.75">0.75 mm</quantity>
, EW
<quantity id="4CB6E3A3FFE15011FB72F896FAF54307" box="[1222,1328,1882,1909]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="17" pageNumber="18" unit="mm" value="0.58">0.58 mm</quantity>
, EI 1.30; humeral calli weakly elevated, basal elytral foveae vestigial, barely discernible. Punctures fine and inconspicuous; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFE15011FF73F80AFBCC4393" blockId="17.[151,1437,1702,2016]" box="[199,1033,1990,2016]" pageId="17" pageNumber="18">Legs moderately long, slender, unmodified except slightly recurved tibiae.</paragraph>
<paragraph id="8BF14E46FFE25012FF73FF5BFE2F4516" blockId="18.[151,1437,150,501]" pageId="18" pageNumber="19">
Aedeagus (
<figureCitation id="137552C3FFE25012FEF3FF5BFE0F44C2" box="[327,458,150,177]" captionStart="FIGURES 44–51" captionStartId="17.[151,264,1523,1547]" captionTargetBox="[151,1422,181,1488]" captionTargetId="figure-16@17.[151,1435,181,1500]" captionTargetPageId="17" captionText="FIGURES 44–51. Anhoraeomorphus sakarahanus (Franz), holotype. Habitus in dorsal view (44); head in anterodorsal view (45); head and prothorax in lateral view (only bristles on head are shown) (46); antenna (47); aedeagus in ventral (48, 50) and lateral (49, 51) views." figureDoi="http://doi.org/10.5281/zenodo.13745937" httpUri="https://zenodo.org/record/13745937/files/figure.png" pageId="18" pageNumber="19">Figs 48–51</figureCitation>
) stout, AeL
<quantity id="4CB6E3A3FFE25012FDE8FF5BFD0344C2" box="[604,710,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="18" pageNumber="19" unit="mm" value="0.23">0.23 mm</quantity>
; in ventral view broadest near middle and distinctly narrowing towards truncate apex; in lateral view apical region of dorsal wall strongly and abruptly bent dorsally; endophallus with symmetrical and elongate distolateral sclerites and basilateral sclerites, and with asymmetrical weakly sclerotized median structures, distolateral sclerites broadly separated and each with rounded apex; parameres slender and their apices only slightly projecting beyond aedeagal apex, each paramere with a row of sparse and short setae occupying slightly less than distal fourth.
</paragraph>
<paragraph id="8BF14E46FFE25012FF73FEA3FE5B45FA" blockId="18.[151,1437,150,501]" box="[199,414,367,393]" pageId="18" pageNumber="19">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE25012FF73FE5EFC6345DE" box="[199,934,402,429]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="8BF14E46FFE25012FF73FE5EFC6345DE" blockId="18.[151,1437,150,501]" box="[199,934,402,429]" pageId="18" pageNumber="19">
<emphasis id="B93A9254FFE25012FF73FE5EFEA445DF" bold="true" box="[199,353,402,428]" pageId="18" pageNumber="19">Distribution.</emphasis>
Southern
<collectingCountry id="F3590ED6FFE25012FE60FE5FFD9E45DE" box="[468,603,403,429]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
(Atsimo-Andrefana Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFE25012FF73FE7BFA844586" pageId="18" pageNumber="19" type="discussion">
<paragraph id="8BF14E46FFE25012FF73FE7BFC3C45A2" blockId="18.[151,1437,150,501]" box="[199,1017,439,465]" pageId="18" pageNumber="19">
<emphasis id="B93A9254FFE25012FF73FE7BFEFE45A2" bold="true" box="[199,315,439,465]" pageId="18" pageNumber="19">Remarks.</emphasis>
Franz misspelled Sakaraha in the locality label as “Sakahara”.
</paragraph>
<paragraph id="8BF14E46FFE25012FF73FE17FA844586" blockId="18.[151,1437,150,501]" box="[199,1345,475,501]" pageId="18" pageNumber="19">
<taxonomicName id="4C4E35C5FFE25012FF73FE17FDEB4586" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[199,558,475,501]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="sakarahanus">
<emphasis id="B93A9254FFE25012FF73FE17FDEB4586" box="[199,558,475,501]" italics="true" pageId="18" pageNumber="19">Anhoraeomorphus sakarahanus</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C4E35C5FFE25012FD49FE17FC714586" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[765,948,475,501]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="ankaratranus">
<emphasis id="B93A9254FFE25012FD49FE17FC714586" box="[765,948,475,501]" italics="true" pageId="18" pageNumber="19">A. ankaratranus</emphasis>
</taxonomicName>
; see Remarks for the latter species.
</paragraph>
</subSubSection>
</treatment>
</document>
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