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<document id="D834B65C18A9BD6DA038B5306F26C8B9" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFDA502BFF23FAF6FC2A43A7" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus nudipennis Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="44" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="43" updateTime="1726053143081" updateUser="ExternalLinkService">
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<mods:title id="0E8159668E8185B2688FCBCE7F59CD91">The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:date id="CA58994D9DC0406B9635D64166C3608E">2024</mods:date>
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<mods:number id="3E69AFA628A3B5CCA62BC87357ACCE44">2024-09-04</mods:number>
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<mods:number id="DE006A604F7175870A535C71388EFD9E">5505</mods:number>
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<mods:identifier id="7C2B130FBB5DA95A7B7710D7047D9B76" type="DOI">10.11646/zootaxa.5505.1.1</mods:identifier>
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<treatment id="03E7FF50FFDA502BFF23FAF6FC2A43A7" ID-DOI="http://doi.org/10.5281/zenodo.13746073" ID-Zenodo-Dep="13746073" LSID="urn:lsid:plazi:treatment:03E7FF50FFDA502BFF23FAF6FC2A43A7" httpUri="http://treatment.plazi.org/id/03E7FF50FFDA502BFF23FAF6FC2A43A7" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
<subSubSection id="C3541DCDFFDA502AFF23FAF6FD2C4126" box="[151,745,1338,1365]" pageId="42" pageNumber="43" type="nomenclature">
<paragraph id="8BF14E46FFDA502AFF23FAF6FD2C4126" blockId="42.[151,745,1338,1401]" box="[151,745,1338,1365]" pageId="42" pageNumber="43">
<heading id="D0B9F92AFFDA502AFF23FAF6FD2C4126" bold="true" box="[151,745,1338,1365]" fontSize="11" level="2" pageId="42" pageNumber="43" reason="4">
<emphasis id="B93A9254FFDA502AFF23FAF6FD2C4126" bold="true" box="[151,745,1338,1365]" pageId="42" pageNumber="43">
<taxonomicName id="4C4E35C5FFDA502AFF23FAF6FDB44127" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,625,1338,1365]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="nudipennis" status="comb. nov.">
<emphasis id="B93A9254FFDA502AFF23FAF6FDCF4126" bold="true" box="[151,522,1338,1365]" italics="true" pageId="42" pageNumber="43">Anhoraeomorphus nudipennis</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFDA502AFDCBFAF7FD2C4126" box="[639,745,1339,1365]" pageId="42" pageNumber="43" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDA502AFF23FA93FE86410A" box="[151,323,1374,1401]" pageId="42" pageNumber="43" type="description">
<paragraph id="8BF14E46FFDA502AFF23FA93FE86410A" blockId="42.[151,745,1338,1401]" box="[151,323,1374,1401]" pageId="42" pageNumber="43">
(
<figureCitation id="137552C3FFDA502AFF2BFA93FEFE410A" box="[159,315,1374,1401]" captionStart="FIGURES 159–162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159–162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Figs 159–162</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDA502AFF23FA6FFDBC41C8" box="[151,633,1443,1467]" pageId="42" pageNumber="43" type="reference_group">
<paragraph id="8BF14E46FFDA502AFF23FA6FFDBC41C8" blockId="42.[151,633,1443,1467]" box="[151,633,1443,1467]" pageId="42" pageNumber="43">
<treatmentCitationGroup id="AB5E6968FFDA502AFF23FA6FFDBC41C8" box="[151,633,1443,1467]" pageId="42" pageNumber="43">
<treatmentCitation id="0AEF6857FFDA502AFF23FA6FFDB041C8" author="Franz, H." box="[151,629,1443,1467]" page="148" pageId="42" pageNumber="43" year="1986">
<taxonomicName id="4C4E35C5FFDA502AFF23FA6FFDB041C8" ID-CoL="3MNKZ" authority="Franz, 1986 b: 148" authorityName="Franz" authorityPageNumber="148" authorityYear="1986" box="[151,629,1443,1467]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="species" species="nudipennis">
<emphasis id="B93A9254FFDA502AFF23FA6FFE7641C8" box="[151,435,1443,1467]" italics="true" pageId="42" pageNumber="43">Horaeomorphus nudipennis</emphasis>
<bibRefCitation id="EFDF33B7FFDA502AFE0DFA68FDB041C8" author="Franz, H." box="[441,629,1443,1467]" pageId="42" pageNumber="43" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 148</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDA502AFF23FA25FDC0428C" pageId="42" pageNumber="43" type="materials_examined">
<paragraph id="8BF14E46FFDA502AFF23FA25FE1B4238" blockId="42.[151,1436,1513,2008]" pageId="42" pageNumber="43">
<emphasis id="B93A9254FFDA502AFF23FA25FE614270" bold="true" box="[151,420,1513,1539]" pageId="42" pageNumber="43">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFDA502AFE1FFA25FE1B4238" collectionCode="NHMW" collectorName="H. Franz" country="Madagascar" determinerName="H. Franz" location="Madagascar" pageId="42" pageNumber="43" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B93A9254FFDA502AFE1FFA25FD0A4270" bold="true" box="[427,719,1513,1539]" pageId="42" pageNumber="43">
<typeStatus id="54F5F0E4FFDA502AFE1FFA25FDDD4270" box="[427,536,1513,1539]" pageId="42" pageNumber="43" type="holotype">Holotype</typeStatus>
(
<collectingCountry id="F3590ED6FFDA502AFD93FA25FD074270" box="[551,706,1513,1539]" name="Madagascar" pageId="42" pageNumber="43">Madagascar</collectingCountry>
):
</emphasis>
<specimenCount id="9D4885CFFFDA502AFD62FA26FD2E4277" box="[726,747,1514,1540]" count="1" pageId="42" pageNumber="43" type="female">♀</specimenCount>
, three labels: “Madagaskar / Umg.Périnet / lg.
<collectorName id="26BB2B90FFDA502AFB46FA25FA954270" box="[1266,1360,1513,1539]" pageId="42" pageNumber="43">H.Franz</collectorName>
1969” [white, printed], “
<taxonomicName id="4C4E35C5FFDA502AFED7F9C1FDDF4254" box="[355,538,1549,1575]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="42" pageNumber="43" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ nudipennis m. / det.
<determinerName id="7E2DA282FFDA502AFCAFF9C1FCBC4254" box="[795,889,1549,1575]" pageId="42" pageNumber="43">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFDA502AFEC4F9FDFE164238" box="[368,467,1585,1611]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="42" pageNumber="43" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFDA502AFF73F999FDC0428C" blockId="42.[151,1436,1513,2008]" pageId="42" pageNumber="43">
<emphasis id="B93A9254FFDA502AFF73F999FE62421C" bold="true" box="[199,423,1621,1647]" pageId="42" pageNumber="43">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFDA502AFDB2F999FDA94203" box="[518,620,1621,1648]" captionStart="FIGURES 159–162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159–162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Fig. 159</figureCitation>
) moderately stout; head (
<figureCitation id="137552C3FFDA502AFC15F999FB86421C" box="[929,1091,1621,1647]" captionStart="FIGURES 159–162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159–162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Figs 160–161</figureCitation>
) sub-pentagonal, with vertex posteriorly weakly rounded; pronotum (
<figureCitation id="137552C3FFDA502AFDEDF9B5FD7F42E7" box="[601,698,1657,1684]" captionStart="FIGURES 159–162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159–162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Fig. 159</figureCitation>
) bell-shaped, broadest slightly in front of anterior third, with one pair of large subcircular antebasal pits; antennomeres 1–6 (
<figureCitation id="137552C3FFDA502AFCF5F951FC6642C4" box="[833,931,1693,1719]" captionStart="FIGURES 159–162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159–162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Fig. 162</figureCitation>
) each elongate, 7–10 each about as long as broad; all femora similarly slender and densely setose in lateroventral distal region; metatibiae in female with flattened and matt dorsal surface
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDA502BFF73F8C5FE4042E3" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43" type="description">
<paragraph id="8BF14E46FFDA502AFF73F8C5FF0F433B" blockId="42.[151,1436,1513,2008]" pageId="42" pageNumber="43">
<emphasis id="B93A9254FFDA502AFF73F8C5FEB04350" bold="true" box="[199,373,1801,1827]" pageId="42" pageNumber="43">Redescription.</emphasis>
Body of female (
<figureCitation id="137552C3FFDA502AFDFCF8C5FD6E4357" box="[584,683,1801,1828]" captionStart="FIGURES 159–162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159–162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Fig. 159</figureCitation>
) moderately convex, yellowish brown; setae yellowish; BL
<quantity id="4CB6E3A3FFDA502AFADFF8C5FF03433B" metricMagnitude="-3" metricUnit="m" metricValue="2.55" pageId="42" pageNumber="43" unit="mm" value="2.55">2.55 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFDA502BFF73F89DFF1A456E" blockId="42.[151,1436,1513,2008]" lastBlockId="43.[151,1436,151,285]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
Head (
<figureCitation id="137552C3FFDA502AFEA6F89DFEB74318" box="[274,370,1873,1899]" captionStart="FIGURES 159–162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159–162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Fig. 160</figureCitation>
) in anterodorsal view sub-pentagonal, broadest at eyes, HL
<quantity id="4CB6E3A3FFDA502AFBA3F89DFBB8431F" box="[1047,1149,1873,1900]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="42" pageNumber="43" unit="mm" value="0.43">0.43 mm</quantity>
, HW
<quantity id="4CB6E3A3FFDA502AFB0BF89DFAE2431F" box="[1215,1319,1873,1900]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="42" pageNumber="43" unit="mm" value="0.45">0.45 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFDA502AFE9AF8B9FE4A43FC" box="[302,399,1909,1935]" captionStart="FIGURES 159–162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159–162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="42" pageNumber="43">Fig. 161</figureCitation>
) slightly shorter than eye; vertex and frons confluent, together weakly convex and weakly transverse; vertex posteriorly weakly rounded; each eye large and weakly convex, oval and posteriorly barely noticeably emarginate, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFDB502BFA86FF5BFA5144C2" box="[1330,1428,151,177]" captionStart="FIGURES 159–162" captionStartId="43.[151,264,1115,1139]" captionTargetBox="[151,1412,309,1092]" captionTargetId="figure-76@43.[151,1435,309,1092]" captionTargetPageId="43" captionText="FIGURES 159–162. Anhoraeomorphus nudipennis (Franz), holotype. Habitus in dorsal view (159); head in anterodorsal view (160); head and prothorax in lateral view (only bristles on head are shown) (161); antenna (162)." figureDoi="http://doi.org/10.5281/zenodo.13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="43" pageNumber="44">Fig. 162</figureCitation>
) slender but short, indistinctly shorter than half BL, AnL
<quantity id="4CB6E3A3FFDB502BFCB6FF77FCA244A6" box="[770,871,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.23" pageId="43" pageNumber="44" unit="mm" value="1.23">1.23 mm</quantity>
, antennomeres 1–6 each elongate, 7–10 each about as long as broad, 11 much shorter than 9 and 10 combined, slightly narrower than 10, about 1.7 times as long as broad.
</paragraph>
<caption id="DF311ECEFFDB502BFF23FB97FBAF40EB" ID-DOI="http://doi.org/10.5281/zenodo.13745977" ID-Zenodo-Dep="13745977" httpUri="https://zenodo.org/record/13745977/files/figure.png" pageId="43" pageNumber="44" startId="43.[151,264,1115,1139]" targetBox="[151,1412,309,1092]" targetPageId="43" targetType="figure">
<paragraph id="8BF14E46FFDB502BFF23FB97FBAF40EB" blockId="43.[151,1436,1115,1176]" pageId="43" pageNumber="44">
<emphasis id="B93A9254FFDB502BFF23FB97FEAB4007" bold="true" box="[151,366,1115,1140]" pageId="43" pageNumber="44">FIGURES 159–162.</emphasis>
<taxonomicName id="4C4E35C5FFDB502BFEC7FB90FD3C4007" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[371,761,1116,1140]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="nudipennis">
<emphasis id="B93A9254FFDB502BFEC7FB90FD604007" box="[371,677,1116,1140]" italics="true" pageId="43" pageNumber="44">Anhoraeomorphus nudipennis</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFDB502BFBE8FB90FB474007" bold="true" box="[1116,1154,1116,1140]" pageId="43" pageNumber="44">159</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFDB502BFF14FB4CFF0340EB" bold="true" box="[160,198,1152,1176]" pageId="43" pageNumber="44">160</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFDB502BFC28FB4CFC0740EB" bold="true" box="[924,962,1152,1176]" pageId="43" pageNumber="44">161</emphasis>
); antenna (
<emphasis id="B93A9254FFDB502BFB82FB4CFB9940EB" bold="true" box="[1078,1116,1152,1176]" pageId="43" pageNumber="44">162</emphasis>
).
</paragraph>
</caption>
<paragraph id="8BF14E46FFDB502BFF73FB27FDE741E7" blockId="43.[151,1437,1258,2005]" pageId="43" pageNumber="44">
Pronotum bell-shaped, broadest slightly in front of anterior third; PL
<quantity id="4CB6E3A3FFDB502BFC66FB26FBFC4176" box="[978,1081,1258,1285]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="43" pageNumber="44" unit="mm" value="0.68">0.68 mm</quantity>
, PW
<quantity id="4CB6E3A3FFDB502BFBC2FB26FB1B4176" box="[1142,1246,1258,1285]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="43" pageNumber="44" unit="mm" value="0.6">0.60 mm</quantity>
. Anterior margin strongly rounded, lateral margins strongly rounded in anterior half, sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large subcircular pits, sublateral carinae conspicuously long and sharply marked. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFDB502BFF73FA52FF354256" blockId="43.[151,1437,1258,2005]" pageId="43" pageNumber="44">
Elytra together oval, moderately convex, dorsally distinctly impressed along suture in anterior third, broadest slightly in front of middle; EL
<quantity id="4CB6E3A3FFDB502BFDBFFA0EFDB041AE" box="[523,629,1474,1501]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="43" pageNumber="44" unit="mm" value="1.45">1.45 mm</quantity>
, EW
<quantity id="4CB6E3A3FFDB502BFD08FA0EFCE341AE" box="[700,806,1474,1501]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="43" pageNumber="44" unit="mm" value="1.05">1.05 mm</quantity>
, EI 1.38; humeral calli weakly elevated, basal elytral foveae barely discernible. Punctures small and shallow, inconspicuous; setae extremely short, so that elytra appear asetose.
</paragraph>
<paragraph id="8BF14E46FFDB502BFF73F9E3FCF5421E" blockId="43.[151,1437,1258,2005]" pageId="43" pageNumber="44">Legs long and slender, all femora densely setose in lateroventral distal region; all tibiae slightly recurved, metatibiae with dorsal surface distinctly flattened and matt.</paragraph>
<paragraph id="8BF14E46FFDB502BFF73F9BAFE4042E3" blockId="43.[151,1437,1258,2005]" box="[199,389,1654,1680]" pageId="43" pageNumber="44">Male. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDB502BFF73F956FC4042C7" box="[199,901,1690,1716]" pageId="43" pageNumber="44" type="distribution">
<paragraph id="8BF14E46FFDB502BFF73F956FC4042C7" blockId="43.[151,1437,1258,2005]" box="[199,901,1690,1716]" pageId="43" pageNumber="44">
<emphasis id="B93A9254FFDB502BFF73F956FEA442C7" bold="true" box="[199,353,1690,1716]" pageId="43" pageNumber="44">Distribution.</emphasis>
Central
<collectingCountry id="F3590ED6FFDB502BFE75F956FD8D42C7" box="[449,584,1690,1716]" name="Madagascar" pageId="43" pageNumber="44">Madagascar</collectingCountry>
(Alaotra-Mangoro Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFDB502BFF73F972FC2A43A7" pageId="43" pageNumber="44" type="discussion">
<paragraph id="8BF14E46FFDB502BFF73F972FEF44353" blockId="43.[151,1437,1258,2005]" pageId="43" pageNumber="44">
<emphasis id="B93A9254FFDB502BFF73F972FEFE42AB" bold="true" box="[199,315,1726,1752]" pageId="43" pageNumber="44">Remarks.</emphasis>
Périnet (= Analamazaotra Special Reserve, often referred to as Andasibe) was a part of the large Mantadia National Park, but currently is a separated component of Andasibe-Mantadia NP divided by deforestation.
</paragraph>
<paragraph id="8BF14E46FFDB502BFF73F8E6FC2A43A7" blockId="43.[151,1437,1258,2005]" pageId="43" pageNumber="44">
The only known female of
<taxonomicName id="4C4E35C5FFDB502BFE40F8E6FD484337" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[500,653,1834,1860]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="nudipennis">
<emphasis id="B93A9254FFDB502BFE40F8E6FD484337" box="[500,653,1834,1860]" italics="true" pageId="43" pageNumber="44">A. nudipennis</emphasis>
</taxonomicName>
has the elytra so finely setose that at magnification 80 times it appears virtually asetose. All femora have a dense group of setae in the lateroventral distal region (this feature is not as distinct as in illustration in
<bibRefCitation id="EFDF33B7FFDB502BFE67F8BFFDBD43FF" author="Franz, H." box="[467,632,1906,1933]" pageId="43" pageNumber="44" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz (1986b)</bibRefCitation>
: fig. 130), and the metatibiae have distinctly flattened and matt dorsal surface, suggesting that in males they may be more conspicuously modified. The conspicuously short antennomere
<quantity id="4CB6E3A3FFDB502BFF23F876FF0A43A7" box="[151,207,1978,2004]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="43" pageNumber="44" unit="in" value="11.0">11 in</quantity>
relation to 9 and 10 combined is another unique feature of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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