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<document id="AF6E055311CEF8F327B950A116FE3D36" ID-DOI="10.11646/zootaxa.5505.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="13745912" ID-ZooBank="93F5E9F6-9EDC-4ED2-8F2B-1E6D0C18F893" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1726052234810" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2024" docId="03E7FF50FFD35025FF23FB5BFE5C409A" docLanguage="en" docName="zootaxa.5505.1.1.pdf" docOrigin="Zootaxa 5505 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5505.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Anhoraeomorphus lambomakandroi Jałoszyński 2024, comb. n." docType="treatment" docVersion="2" lastPageNumber="38" masterDocId="FFDE8728FFF05000FFB4FFCCFFC54473" masterDocTitle="The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="96" masterPageNumber="1" pageNumber="36" updateTime="1726053143081" updateUser="ExternalLinkService">
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<mods:title id="F7ABC7A1FDD54BBFEE9D20C1B193D552">The ‘ curse of Horaeomorphus’ (almost) lifted. Revision of misplaced species from Madagascar, Comoros and Mascarenes (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:date id="863585AC53A3E74B1A6540A763E09B47">2024</mods:date>
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<mods:number id="02FC2607605BCA34D8CA7E9278F893FB">2024-09-04</mods:number>
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<mods:number id="A2680DDEEF851EA6C5F482F5364C6D0D">5505</mods:number>
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<mods:classification id="471BF341BC2B98AF4E93D6CE4133D2FE">journal article</mods:classification>
<mods:identifier id="79B2096DB76648A571918E5F12BB713D" type="DOI">10.11646/zootaxa.5505.1.1</mods:identifier>
<mods:identifier id="8CB7458286FF2142FC33FD4E7602ECF6" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03E7FF50FFD35025FF23FB5BFE5C409A" ID-DOI="http://doi.org/10.5281/zenodo.13746061" ID-Zenodo-Dep="13746061" LSID="urn:lsid:plazi:treatment:03E7FF50FFD35025FF23FB5BFE5C409A" httpUri="http://treatment.plazi.org/id/03E7FF50FFD35025FF23FB5BFE5C409A" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36">
<subSubSection id="C3541DCDFFD35023FF23FB5BFCF440C1" box="[151,817,1175,1202]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8BF14E46FFD35023FF23FB5BFCF440C1" blockId="35.[151,817,1175,1238]" box="[151,817,1175,1202]" pageId="35" pageNumber="36">
<heading id="D0B9F92AFFD35023FF23FB5BFCF440C1" bold="true" box="[151,817,1175,1202]" fontSize="11" level="2" pageId="35" pageNumber="36" reason="4">
<emphasis id="B93A9254FFD35023FF23FB5BFCF440C1" bold="true" box="[151,817,1175,1202]" pageId="35" pageNumber="36">
<taxonomicName id="4C4E35C5FFD35023FF23FB5BFD7C40C2" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[151,697,1175,1202]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lambomakandroi" status="comb. nov.">
<emphasis id="B93A9254FFD35023FF23FB5BFD9740C1" bold="true" box="[151,594,1175,1202]" italics="true" pageId="35" pageNumber="36">Anhoraeomorphus lambomakandroi</emphasis>
(Franz)
</taxonomicName>
,
<taxonomicNameLabel id="A2092F2FFFD35023FD72FB54FCF440C1" box="[710,817,1176,1202]" pageId="35" pageNumber="36" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD35023FF23FB77FE8640A5" box="[151,323,1211,1238]" pageId="35" pageNumber="36" type="description">
<paragraph id="8BF14E46FFD35023FF23FB77FE8640A5" blockId="35.[151,817,1175,1238]" box="[151,323,1211,1238]" pageId="35" pageNumber="36">
(
<figureCitation id="137552C3FFD35023FF2BFB77FEFF40A5" box="[159,314,1211,1238]" captionStart="FIGURES 131–138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131–138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Figs 131–138</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD35023FF23FACCFD73416B" box="[151,694,1280,1304]" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="8BF14E46FFD35023FF23FACCFD73416B" blockId="35.[151,694,1280,1304]" box="[151,694,1280,1304]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="AB5E6968FFD35023FF23FACCFD73416B" box="[151,694,1280,1304]" pageId="35" pageNumber="36">
<treatmentCitation id="0AEF6857FFD35023FF23FACCFD77416B" author="Franz, H." box="[151,690,1280,1304]" page="154" pageId="35" pageNumber="36" year="1986">
<taxonomicName id="4C4E35C5FFD35023FF23FACCFD77416B" ID-CoL="3MNKG" authority="Franz, 1986 b: 154" authorityName="Franz" authorityPageNumber="154" authorityYear="1986" box="[151,690,1280,1304]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD35023FF23FACCFE35416B" box="[151,496,1280,1304]" italics="true" pageId="35" pageNumber="36">Horaeomorphus lambomakandroi</emphasis>
<bibRefCitation id="EFDF33B7FFD35023FE42FACCFD77416B" author="Franz, H." box="[502,690,1280,1304]" pageId="35" pageNumber="36" pagination="1 - 393" refId="ref53526" refString="Franz, H. (1986 b) Monographie der Scydmaeniden (Coleoptera) von Madagaskar (mitAusschluss der Cephenniini). Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 125, 1 - 393." type="journal article" year="1986">Franz, 1986b: 154</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD35023FF23FA8AFE67429E" pageId="35" pageNumber="36" type="materials_examined">
<paragraph id="8BF14E46FFD35023FF23FA8AFE6641BF" blockId="35.[151,1437,1350,2025]" pageId="35" pageNumber="36">
<emphasis id="B93A9254FFD35023FF23FA8AFE654113" bold="true" box="[151,416,1350,1376]" pageId="35" pageNumber="36">Type material studied.</emphasis>
<materialsCitation id="3B26441BFFD35023FE12FA8AFE6641BF" collectionCode="VII-, NHMW" country="Madagascar" determinerName="H. Franz" elevation="500" location="MADAGASCAR" pageId="35" pageNumber="36" specimenCount="58" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B93A9254FFD35023FE12FA8AFD7A4113" bold="true" box="[422,703,1350,1376]" pageId="35" pageNumber="36">
<typeStatus id="54F5F0E4FFD35023FE12FA8AFDD64113" box="[422,531,1350,1376]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
(Madagascar)
</emphasis>
:
<specimenCount id="9D4885CFFFD35023FD78FA8BFD264112" box="[716,739,1351,1377]" count="1" pageId="35" pageNumber="36" type="male">♂</specimenCount>
, four labels: “Madagascar Sud-Ouest / LAMBOMAKANDRO
<quantity id="4CB6E3A3FFD35023FF23FAA6FF1241F6" box="[151,215,1386,1413]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="35" pageNumber="36" unit="m" value="500.0">
<elevation id="0063A975FFD35023FF23FAA6FF1241F6" box="[151,215,1386,1413]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="35" pageNumber="36" unit="m" value="500.0">500m</elevation>
</quantity>
/ Tuléar / 18-
<collectionCode id="ED5FD683FFD35023FEC0FAA6FE6241F6" box="[372,423,1386,1413]" pageId="35" pageNumber="36">VII-</collectionCode>
<specimenCount id="9D4885CFFFD35023FE13FAA6FE2441F7" box="[423,481,1386,1413]" count="57" pageId="35" pageNumber="36" type="generic">57 P.</specimenCount>
Griv.” [white, printed], “INSTITUT / SCIENTIFIQUE /
<collectingCountry id="F3590ED6FFD35023FBDFFAA6FAF041F7" box="[1131,1333,1386,1412]" name="Madagascar" pageId="35" pageNumber="36">MADAGASCAR</collectingCountry>
” [bluish, printed], “
<taxonomicName id="4C4E35C5FFD35023FEB3FA42FE7B41DB" box="[263,446,1422,1448]" class="Insecta" family="Staphylinidae" genus="Horaeomorphus" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">Horaeomorphus</taxonomicName>
/ lambomakandroi m. / det.
<determinerName id="7E2DA282FFD35023FD45FA42FC8A41DB" box="[753,847,1422,1448]" pageId="35" pageNumber="36">H.Franz</determinerName>
” [yellowish, handwritten and printed], “Typus” [red, handwritten] (
<collectionCode id="ED5FD683FFD35023FE81FA7EFE5D41BF" box="[309,408,1458,1484]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="35" pageNumber="36" type="Museum">NHMW</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF14E46FFD35023FF73FA1AFE67429E" blockId="35.[151,1437,1350,2025]" pageId="35" pageNumber="36">
<emphasis id="B93A9254FFD35023FF73FA1AFE634183" bold="true" box="[199,422,1494,1520]" pageId="35" pageNumber="36">Revised diagnosis.</emphasis>
Body (
<figureCitation id="137552C3FFD35023FDB0FA1AFDAC4183" box="[516,617,1494,1520]" captionStart="FIGURES 131–138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131–138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 131</figureCitation>
) slender; head (
<figureCitation id="137552C3FFD35023FC9FFA1AFC094183" box="[811,972,1494,1520]" captionStart="FIGURES 131–138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131–138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Figs 132–133</figureCitation>
) round, with vertex posteriorly weakly rounded; pronotum (
<figureCitation id="137552C3FFD35023FECAFA36FE1B4267" box="[382,478,1530,1556]" captionStart="FIGURES 131–138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131–138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 131</figureCitation>
) bell-shaped, broadest near anterior third, with one pair of large, transverse antebasal pits; antennomere 3 (
<figureCitation id="137552C3FFD35023FE3EF9D2FE2E424B" box="[394,491,1566,1592]" captionStart="FIGURES 131–138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131–138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 134</figureCitation>
) about as long as broad, 4–7 each elongate, 8–10 each about as long as broad; all femora similarly slender; metatibiae in male modified, slightly recurved and broadened in distal region, broadened area with elongate outer impression bordered by sharp outer edge; aedeagus (
<figureCitation id="137552C3FFD35023FC5EF9AAFB4042F2" box="[1002,1157,1638,1665]" captionStart="FIGURES 131–138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131–138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Figs 135–138</figureCitation>
) in ventral view broadest near middle, apical region of dorsal wall in lateral view weakly curved dorsally, distolateral endophallic sclerites broadly separated, each elongate and with blunt subtriangular apex, each paramere with group of five short subapical setae distant from apex.
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD35025FF73F93AFE5B46BE" lastPageId="37" lastPageNumber="38" pageId="35" pageNumber="36" type="description">
<paragraph id="8BF14E46FFD35023FF73F93AFD914346" blockId="35.[151,1437,1350,2025]" pageId="35" pageNumber="36">
<emphasis id="B93A9254FFD35023FF73F93AFEB04363" bold="true" box="[199,373,1782,1808]" pageId="35" pageNumber="36">Redescription.</emphasis>
Body of male (
<figureCitation id="137552C3FFD35023FD87F93AFD524363" box="[563,663,1782,1808]" captionStart="FIGURES 131–138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131–138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 131</figureCitation>
) strongly convex, moderately dark brown, tarsi and palps slightly lighter; setae light brown; BL
<quantity id="4CB6E3A3FFD35023FE5CF8D6FD8A4346" box="[488,591,1818,1845]" metricMagnitude="-3" metricUnit="m" metricValue="2.43" pageId="35" pageNumber="36" unit="mm" value="2.43">2.43 mm</quantity>
.
</paragraph>
<paragraph id="8BF14E46FFD35024FF73F8F2FB0644A6" blockId="35.[151,1437,1350,2025]" lastBlockId="36.[151,1436,151,213]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
Head (
<figureCitation id="137552C3FFD35023FEA4F8F2FEAB432B" box="[272,366,1854,1880]" captionStart="FIGURES 131–138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131–138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 132</figureCitation>
) in anterodorsal view round, broadest at eyes, HL
<quantity id="4CB6E3A3FFD35023FC2EF8F2FC3B432A" box="[922,1022,1854,1881]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" pageId="35" pageNumber="36" unit="mm" value="0.45">0.45 mm</quantity>
, HW
<quantity id="4CB6E3A3FFD35023FB89F8F2FB67432A" box="[1085,1186,1854,1881]" metricMagnitude="-4" metricUnit="m" metricValue="4.8" pageId="35" pageNumber="36" unit="mm" value="0.48">0.48 mm</quantity>
; temple in lateral view (
<figureCitation id="137552C3FFD35023FF2BF8AEFEC4430F" box="[159,257,1890,1916]" captionStart="FIGURES 131–138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131–138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 133</figureCitation>
) slightly shorter than eye; vertex and frons confluent, together weakly convex and slightly transverse; vertex posteriorly weakly rounded; each eye large and weakly convex, nearly circular, weakly projecting from head silhouette. Punctures on vertex fine, inconspicuous; setae short, dense and suberect; frons and vertex densely covered with thick bristles directed posteriorly. Antenna (
<figureCitation id="137552C3FFD35023FCA9F802FCB8439B" box="[797,893,1998,2024]" captionStart="FIGURES 131–138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131–138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="35" pageNumber="36">Fig. 134</figureCitation>
) slender but short, as long as half BL, AnL
<quantity id="4CB6E3A3FFD35024FADFF802FF0044C2" lastPageId="36" lastPageNumber="37" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="35" pageNumber="36" unit="mm" value="1.25">1.25 mm</quantity>
, antennomeres 1–2 each elongate, 3 about as long as broad, 4–7 each elongate, 8–10 each about as long as broad, 11 slightly shorter than 9 and 10 combined, as wide as 10, nearly twice as long as broad.
</paragraph>
<caption id="DF311ECEFFD45024FF23F8E1FE2A43FE" ID-DOI="http://doi.org/10.5281/zenodo.13745967" ID-Zenodo-Dep="13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="36" pageNumber="37" startId="36.[151,264,1837,1861]" targetBox="[151,1423,237,1794]" targetPageId="36" targetType="figure">
<paragraph id="8BF14E46FFD45024FF23F8E1FE2A43FE" blockId="36.[151,1436,1837,1933]" pageId="36" pageNumber="37">
<emphasis id="B93A9254FFD45024FF23F8E1FEA84336" bold="true" box="[151,365,1837,1861]" pageId="36" pageNumber="37">FIGURES 131–138.</emphasis>
<taxonomicName id="4C4E35C5FFD45024FEC7F8E1FCF14336" authority="(Franz)" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[371,820,1837,1861]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD45024FEC7F8E1FD244336" box="[371,737,1837,1861]" italics="true" pageId="36" pageNumber="37">Anhoraeomorphus lambomakandroi</emphasis>
(Franz)
</taxonomicName>
, holotype. Habitus in dorsal view (
<emphasis id="B93A9254FFD45024FB20F8E1FB7F4336" bold="true" box="[1172,1210,1837,1861]" pageId="36" pageNumber="37">131</emphasis>
); head in anterodorsal view (
<emphasis id="B93A9254FFD45024FF63F89DFF38431A" bold="true" box="[215,253,1873,1897]" pageId="36" pageNumber="37">132</emphasis>
); head and prothorax in lateral view (only bristles on head are shown) (
<emphasis id="B93A9254FFD45024FC78F89DFC37431A" bold="true" box="[972,1010,1873,1897]" pageId="36" pageNumber="37">133</emphasis>
); antenna (
<emphasis id="B93A9254FFD45024FBD0F89DFB4F431A" bold="true" box="[1124,1162,1873,1897]" pageId="36" pageNumber="37">134</emphasis>
); aedeagus in ventral (
<emphasis id="B93A9254FFD45024FAC4F89DFF7843FE" bold="true" pageId="36" pageNumber="37">135, 137</emphasis>
) and lateral (
<emphasis id="B93A9254FFD45024FEF1F8B9FE5B43FE" bold="true" box="[325,414,1909,1933]" pageId="36" pageNumber="37">136, 138</emphasis>
) views.
</paragraph>
</caption>
<paragraph id="8BF14E46FFD55025FF73FF5BFEEA4532" blockId="37.[151,1437,150,1257]" pageId="37" pageNumber="38">
Pronotum bell-shaped, broadest near anterior third; PL
<quantity id="4CB6E3A3FFD55025FCE6FF5BFC7844C2" box="[850,957,151,177]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="37" pageNumber="38" unit="mm" value="0.63">0.63 mm</quantity>
, PW
<quantity id="4CB6E3A3FFD55025FBB6FF5AFBAB44C2" box="[1026,1134,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" pageId="37" pageNumber="38" unit="mm" value="0.53">0.53 mm</quantity>
. Anterior margin strongly rounded, lateral margins rounded in anterior half, weakly sinuate in posterior half; posterior corners blunt; posterior margin nearly straight. Base with one pair of large and strongly transverse pits, sublateral carinae short but distinct. Disc with fine, inconspicuous punctures; setae dense, short and suberect, lateral surface of pronotum with dense thick bristles.
</paragraph>
<paragraph id="8BF14E46FFD55025FF73FE87FACC45DE" blockId="37.[151,1437,150,1257]" pageId="37" pageNumber="38">
Elytra together oval, broadest distinctly in front of middle; EL
<quantity id="4CB6E3A3FFD55025FC39FE86FC314516" box="[909,1012,330,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" pageId="37" pageNumber="38" unit="mm" value="1.35">1.35 mm</quantity>
, EW
<quantity id="4CB6E3A3FFD55025FB87FE87FB5F4516" box="[1075,1178,331,357]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="38" unit="mm" value="0.9">0.90 mm</quantity>
, EI 1.50; humeral calli weakly elevated, basal elytral foveae barely discernible. Punctures small and shallow but distinct, those on anterior half separated by spaces slightly shorter than diameters of punctures; setae sparse, short and suberect.
</paragraph>
<paragraph id="8BF14E46FFD55025FF73FE7BFB164586" blockId="37.[151,1437,150,1257]" pageId="37" pageNumber="38">Legs conspicuously long and slender, unmodified except hind tibiae, which are recurved and slightly broadened in distal regions, with dorsal broadened area longitudinally impressed and with outer sharp edge.</paragraph>
<paragraph id="8BF14E46FFD55025FF73FE33FB7046DA" blockId="37.[151,1437,150,1257]" pageId="37" pageNumber="38">
Aedeagus (
<figureCitation id="137552C3FFD55025FEFDFE33FE2E466A" box="[329,491,510,537]" captionStart="FIGURES 131–138" captionStartId="36.[151,264,1837,1861]" captionTargetBox="[151,1423,237,1794]" captionTargetId="figure-67@36.[151,1435,237,1814]" captionTargetPageId="36" captionText="FIGURES 131–138. Anhoraeomorphus lambomakandroi (Franz), holotype. Habitus in dorsal view (131); head in anterodorsal view (132); head and prothorax in lateral view (only bristles on head are shown) (133); antenna (134); aedeagus in ventral (135, 137) and lateral (136, 138) views." figureDoi="http://doi.org/10.5281/zenodo.13745967" httpUri="https://zenodo.org/record/13745967/files/figure.png" pageId="37" pageNumber="38">Figs 135–138</figureCitation>
) slender, AeL
<quantity id="4CB6E3A3FFD55025FD29FE33FCCC466A" box="[669,777,511,537]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="37" pageNumber="38" unit="mm" value="0.4">0.40 mm</quantity>
; in ventral view suboval, broadest near middle, distally gradually narrowing and in subapical region rapidly narrowed to subtriangular apex; in lateral view apical region of dorsal wall weakly curved dorsally; endophallus with all sclerotized structures asymmetrical, distolateral sclerites broadly separated, each with subtriangular and blunt apical region; parameres broad and their apices not projecting beyond aedeagal apex, each paramere with subapical group of 5 short setae distant from apex.
</paragraph>
<paragraph id="8BF14E46FFD55025FF73FD7FFE5B46BE" blockId="37.[151,1437,150,1257]" box="[199,414,691,717]" pageId="37" pageNumber="38">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD55025FF73FD1AFC634682" box="[199,934,726,753]" pageId="37" pageNumber="38" type="distribution">
<paragraph id="8BF14E46FFD55025FF73FD1AFC634682" blockId="37.[151,1437,150,1257]" box="[199,934,726,753]" pageId="37" pageNumber="38">
<emphasis id="B93A9254FFD55025FF73FD1AFEA44683" bold="true" box="[199,353,726,752]" pageId="37" pageNumber="38">Distribution.</emphasis>
Southern
<collectingCountry id="F3590ED6FFD55025FE60FD1BFD9E4682" box="[468,603,727,753]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
(Atsimo-Andrefana Region).
</paragraph>
</subSubSection>
<subSubSection id="C3541DCDFFD55025FF73FD37FE5C409A" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8BF14E46FFD55025FF73FD37FE5C409A" blockId="37.[151,1437,150,1257]" pageId="37" pageNumber="38">
<emphasis id="B93A9254FFD55025FF73FD37FEFE4766" bold="true" box="[199,315,763,789]" pageId="37" pageNumber="38">Remarks.</emphasis>
Males of
<taxonomicName id="4C4E35C5FFD55025FE75FD37FC9B4766" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[449,862,763,789]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD55025FE75FD37FC9B4766" box="[449,862,763,789]" italics="true" pageId="37" pageNumber="38">Anhoraeomorphus lambomakandroi</emphasis>
</taxonomicName>
can be easily distinguished from all remaining congeners, except for one, by the body shape, slender legs with metatibiae recurved and slightly broadened in distal regions, with the broadened area longitudinally impressed and laterally delimited by an outer sharp edge, and by unique structures of the aedeagus. The exception is
<taxonomicName id="4C4E35C5FFD55025FD64FCABFCEF47F2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[720,810,871,897]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vellyi">
<emphasis id="B93A9254FFD55025FD64FCABFCEF47F2" box="[720,810,871,897]" italics="true" pageId="37" pageNumber="38">A. vellyi</emphasis>
</taxonomicName>
known to occur in Central
<collectingCountry id="F3590ED6FFD55025FBE0FCABFB1847F2" box="[1108,1245,871,897]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
, and these names may turn out to be synonyms. Externally, the
<typeStatus id="54F5F0E4FFD55025FD3AFC47FD3E47D6" box="[654,763,907,933]" pageId="37" pageNumber="38" type="holotype">holotypes</typeStatus>
of both species (each known from
<specimenCount id="9D4885CFFFD55025FBCBFC47FADC47D6" box="[1151,1305,907,933]" count="1" pageId="37" pageNumber="38" type="generic">one specimen</specimenCount>
) differ only in body length (
<taxonomicName id="4C4E35C5FFD55025FEFDFC63FDE347BA" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[329,550,943,969]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD55025FEFDFC63FDE347BA" box="[329,550,943,969]" italics="true" pageId="37" pageNumber="38">A. lambomakandroi</emphasis>
</taxonomicName>
<quantity id="4CB6E3A3FFD55025FD99FC63FD5647BA" box="[557,659,943,969]" metricMagnitude="-3" metricUnit="m" metricValue="2.43" pageId="37" pageNumber="38" unit="mm" value="2.43">2.43 mm</quantity>
,
<taxonomicName id="4C4E35C5FFD55025FD2AFC63FD3F47BA" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[670,762,943,969]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vellyi">
<emphasis id="B93A9254FFD55025FD2AFC63FD3F47BA" box="[670,762,943,969]" italics="true" pageId="37" pageNumber="38">A. vellyi</emphasis>
</taxonomicName>
<quantity id="4CB6E3A3FFD55025FCB4FC62FCAD47BA" box="[768,872,942,969]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" pageId="37" pageNumber="38" unit="mm" value="1.95">1.95 mm</quantity>
), relative lengths of antennae (
<taxonomicName id="4C4E35C5FFD55025FB0BFC63FA5947BA" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[1215,1436,943,969]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD55025FB0BFC63FA5947BA" box="[1215,1436,943,969]" italics="true" pageId="37" pageNumber="38">A. lambomakandroi</emphasis>
</taxonomicName>
AnL/Bl 0.52,
<taxonomicName id="4C4E35C5FFD55025FE8FFC1FFE5E479E" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[315,411,979,1005]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vellyi">
<emphasis id="B93A9254FFD55025FE8FFC1FFE5E479E" box="[315,411,979,1005]" italics="true" pageId="37" pageNumber="38">A. vellyi</emphasis>
</taxonomicName>
0.42), and in slightly (but rather indistinctively) different proportions of antennomeres. Unfortunately, the aedeagus of the
<typeStatus id="54F5F0E4FFD55025FDA9FC3BFDBA4062" box="[541,639,1015,1041]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C4E35C5FFD55025FD10FC3BFC444062" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[676,897,1015,1041]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD55025FD10FC3BFC444062" box="[676,897,1015,1041]" italics="true" pageId="37" pageNumber="38">A. lambomakandroi</emphasis>
</taxonomicName>
is partly erected, so the shapes and arrangement of twisted and displaced endophallic structures are difficult to compare with those in the
<typeStatus id="54F5F0E4FFD55025FBDFFBD7FB084046" box="[1131,1229,1051,1077]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C4E35C5FFD55025FB44FBD7FA8E4046" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[1264,1355,1051,1077]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vellyi">
<emphasis id="B93A9254FFD55025FB44FBD7FA8E4046" box="[1264,1355,1051,1077]" italics="true" pageId="37" pageNumber="38">A. vellyi</emphasis>
</taxonomicName>
, which are in repose (
<figureCitation id="137552C3FFD55025FE8EFBF3FE1E402A" box="[314,475,1087,1113]" captionStart="FIGURES 179–186" captionStartId="49.[151,264,1641,1665]" captionTargetBox="[151,1415,181,1602]" captionTargetId="figure-16@49.[151,1435,181,1617]" captionTargetPageId="49" captionText="FIGURES 179–186. Anhoraeomorphus vellyi (Franz), holotype. Habitus in dorsal view (179); head in anterodorsal view (180); head and prothorax in lateral view (only bristles on head are shown) (181); antenna (182); aedeagus in ventral (183, 185) and lateral (184, 186) views." figureDoi="http://doi.org/10.5281/zenodo.13745983" httpUri="https://zenodo.org/record/13745983/files/figure.png" pageId="37" pageNumber="38">Figs 183–186</figureCitation>
). Some of the endophallic structures seem identical, but comparison is inconclusive. The dorsal apical plate in the
<typeStatus id="54F5F0E4FFD55025FE45FBAFFD96400E" box="[497,595,1123,1149]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C4E35C5FFD55025FDC9FBAFFC99400E" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" box="[637,860,1123,1149]" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="lambomakandroi">
<emphasis id="B93A9254FFD55025FDC9FBAFFC99400E" box="[637,860,1123,1149]" italics="true" pageId="37" pageNumber="38">A. lambomakandroi</emphasis>
</taxonomicName>
is clearly more elongate, narrower than that in
<taxonomicName id="4C4E35C5FFD55025FA30FBAFFF1140D2" authorityName="Jałoszyński" authorityYear="2024" baseAuthorityName="Jałoszyński" baseAuthorityYear="2024" class="Insecta" family="Staphylinidae" genus="Anhoraeomorphus" kingdom="Animalia" order="Coleoptera" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="vellyi">
<emphasis id="B93A9254FFD55025FA30FBAFFF1140D2" italics="true" pageId="37" pageNumber="38">A. vellyi</emphasis>
</taxonomicName>
, but the distortion of median lobe does not allow for comparing other external aedeagal structures. With only
<specimenCount id="9D4885CFFFD55025FF23FB67FEF840B6" box="[151,317,1195,1221]" count="2" pageId="37" pageNumber="38" type="generic">two specimens</specimenCount>
at hand, I refrain from synonymizing these species names; more male specimens must be examined to clarify this problem.
</paragraph>
</subSubSection>
</treatment>
</document>
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