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<document id="1ABAB5BB7A083A76951F374A02A923E8" ID-DOI="10.11646/zootaxa.5443.4.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="11064413" ID-ZooBank="AD120933-D237-4B1B-BDD5-39490B6376B6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1714029068808" checkinUser="plazi" docAuthor="Tan, Ming Kai, Liu, Chunxiang, Ingrisch, Sigfrid, Japir, Razy &amp; Chung, Arthur Y. C." docDate="2024" docId="03E687F4AB2CA644ACC3F96DFBC4FD0A" docLanguage="en" docName="zootaxa.5443.4.1.pdf" docOrigin="Zootaxa 5443 (4)" docSource="http://dx.doi.org/10.11646/zootaxa.5443.4.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tapiena incisa Karny 1923" docType="treatment" docVersion="2" lastPageNumber="482" masterDocId="FFDFFF8CAB32A65BAC54FFFAFFE5FF94" masterDocTitle="An account on the Phaneropterinae (Tettigonioidea: Tettigoniidae) from the highlands of western Sabah" masterLastPageNumber="494" masterPageNumber="451" pageNumber="481" updateTime="1714030066638" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="FC7C09F0A511FBED82D16545C72008A5" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E1F42DBDE8C84EA2D05B4AE9C693DEAB">
<mods:title id="758C81F290A8BC05E2C8EFFABB94A4BE">An account on the Phaneropterinae (Tettigonioidea: Tettigoniidae) from the highlands of western Sabah</mods:title>
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<mods:name id="4702ACC4442A03A3DBF1914C58E5CC43" type="personal">
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<mods:namePart id="AB03757AEE3645B666F9795B560FFDD7">Tan, Ming Kai</mods:namePart>
<mods:affiliation id="DB65A0670AAC982BDBB4E6B11A6E8168">Block 207 A, Woodleigh Link, Singapore 361207, Republic of Singapore.</mods:affiliation>
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<mods:namePart id="0EFBC5564E63CE20E5C0DD81471848E6">Liu, Chunxiang</mods:namePart>
<mods:affiliation id="7F9DC19BDF8BB07A447F85A10EFD6DC0">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="92FE6432C60AC9FDE8D910C94169AE72">Ingrisch, Sigfrid</mods:namePart>
<mods:affiliation id="59F645DD0AF6389C0472137033BDB552">Leibniz Institute for the Analysis of Biodiversity Change; Museum Koenig, Adenauerallee 160, D- 53113 Bonn, Germany.</mods:affiliation>
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<mods:roleTerm id="FFA53BCF97F8D4531BF594A1BADC08EC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD3A4D0316767FDB7E9EA148597A7E5F">Japir, Razy</mods:namePart>
<mods:nameIdentifier id="FA9837A4F056F2148E7FA9E2C81E8066" type="ORCID">0009-0004-1879-6550</mods:nameIdentifier>
<mods:affiliation id="C5BA52496CB2188C98E6E42AAF2A1810">Block 207 A, Woodleigh Link, Singapore 361207, Republic of Singapore. &amp; Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia.</mods:affiliation>
<mods:nameIdentifier id="0969F1D07B86A3DB84D661EA3A2D58AB" type="email">razy.japir@sabah.gov.my</mods:nameIdentifier>
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</mods:role>
<mods:namePart id="D4BDD7314B206C8EA6B48D80419042FF">Chung, Arthur Y. C.</mods:namePart>
<mods:nameIdentifier id="2C06E06AA0129B52E0C1844F234F9ECC" type="ORCID">0000-0002-9529-4114</mods:nameIdentifier>
<mods:affiliation id="A05FDB2B254DC55C9896F639CA112243">Block 207 A, Woodleigh Link, Singapore 361207, Republic of Singapore. &amp; Forest Research Centre (Sepilok), Sabah Forestry Department, PO Box 1407, 90715 Sandakan, Sabah, Malaysia. &amp; Block 207 A, Woodleigh Link, Singapore 361207, Republic of Singapore. &amp; Block 207 A, Woodleigh Link, Singapore 361207, Republic of Singapore.</mods:affiliation>
<mods:nameIdentifier id="C3FAE9616DF5C47A30E30A89C46485CD" type="email">arthur.chung@sabah.gov.my</mods:nameIdentifier>
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<mods:title id="600D29DEFD343E4B5C25D1F8005296E1">Zootaxa</mods:title>
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<mods:date id="9244C961340B05888FB5287731FCA5DF">2024</mods:date>
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<mods:number id="4C33DC5D7007F000E8676B32C0F369CB">2024-04-24</mods:number>
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<mods:classification id="CD9CBB1C24032F5793A3E2F7D44D15DD">journal article</mods:classification>
<mods:identifier id="C356B25F15645608E896CE172AA69628" type="DOI">10.11646/zootaxa.5443.4.1</mods:identifier>
<mods:identifier id="1A04BB08C098FAE9A79BA48833407182" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E687F4AB2CA644ACC3F96DFBC4FD0A" ID-DOI="http://doi.org/10.5281/zenodo.11064600" ID-Zenodo-Dep="11064600" LSID="urn:lsid:plazi:treatment:03E687F4AB2CA644ACC3F96DFBC4FD0A" httpUri="http://treatment.plazi.org/id/03E687F4AB2CA644ACC3F96DFBC4FD0A" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481">
<subSubSection id="C3556569AB2CA645ACC3F96DFE13F925" box="[151,502,1687,1714]" pageId="30" pageNumber="481" type="nomenclature">
<paragraph id="8BF036E2AB2CA645ACC3F96DFE13F925" blockId="30.[151,502,1687,1750]" box="[151,502,1687,1714]" pageId="30" pageNumber="481">
<heading id="D0B8818EAB2CA645ACC3F96DFE13F925" bold="true" box="[151,502,1687,1714]" fontSize="11" level="1" pageId="30" pageNumber="481" reason="1">
<taxonomicName id="4C4F4D61AB2CA645ACC3F96DFE13F925" ID-CoL="54SPK" authority="Karny, 1923" authorityName="Karny" authorityYear="1923" box="[151,502,1687,1714]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="incisa">
<emphasis id="B93BEAF0AB2CA645ACC3F96DFE13F925" bold="true" box="[151,502,1687,1714]" pageId="30" pageNumber="481">
<emphasis id="B93BEAF0AB2CA645ACC3F96DFEB3F925" bold="true" box="[151,342,1687,1713]" italics="true" pageId="30" pageNumber="481">Tapiena?incisa</emphasis>
<bibRefCitation id="EFDE4B13AB2CA645AD09F962FE13F925" author="Karny, H. H." box="[349,502,1687,1714]" pageId="30" pageNumber="481" pagination="117 - 193" refId="ref14905" refString="Karny, H. H. (1923) On Malaysian katydids (Gryllacridae and Tettigoniidae) from the Raffles Museum, Singapore. Journal of the Malaysian Branch of the Royal Asiatic Society, 1, 117 - 193." type="journal article" year="1923">Karny, 1923</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3556569AB2CA645ACC3F941FEC2F942" box="[151,295,1723,1750]" pageId="30" pageNumber="481" type="description">
<paragraph id="8BF036E2AB2CA645ACC3F941FEC2F942" blockId="30.[151,502,1687,1750]" box="[151,295,1723,1750]" pageId="30" pageNumber="481">
(
<figureCitation id="13742A67AB2CA645ACCBF941FF13F942" box="[159,246,1723,1750]" captionStart="FIGURE 32" captionStartId="31.[151,250,1574,1598]" captionTargetBox="[151,1435,694,1550]" captionTargetId="figure-238@31.[151,1435,694,1550]" captionTargetPageId="31" captionText="FIGURE 32. Tapiena?incisa Karny, 1923 ♀ attracted to light at Mount Trus Madi." figureDoi="http://doi.org/10.5281/zenodo.11064495" httpUri="https://zenodo.org/record/11064495/files/figure.png" pageId="30" pageNumber="481">Figs 32</figureCitation>
,
<figureCitation id="13742A67AB2CA645AD56F946FEFAF942" box="[258,287,1724,1750]" captionStart="FIGURE 33" captionStartId="32.[151,250,1982,2006]" captionTargetBox="[151,1436,590,1958]" captionTargetId="figure-122@32.[151,1436,590,1958]" captionTargetPageId="32" captionText="FIGURE 33. Tapiena?incisa Karny, 1923 ♂: stridulatory area on the left (A) and right (B) tegmina in dorsal view; abdominal apex in lateral (C) and dorsal (D) views; subgenital plate in ventral view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064497" httpUri="https://zenodo.org/record/11064497/files/figure.png" pageId="30" pageNumber="481">33</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3556569AB2CA645ACC3F8FBFEEBF8A2" pageId="30" pageNumber="481" type="reference_group">
<paragraph id="8BF036E2AB2CA645ACC3F8FBFEEBF8A2" blockId="30.[151,1437,1792,1846]" pageId="30" pageNumber="481">
<treatmentCitationGroup id="AB5F11CCAB2CA645ACC3F8FBFEEBF8A2" pageId="30" pageNumber="481">
<treatmentCitation id="0AEE10F3AB2CA645ACC3F8FBFE05F88C" author="Karny, H. H." box="[151,480,1792,1816]" page="141" pageId="30" pageNumber="481" year="1923">
<taxonomicName id="4C4F4D61AB2CA645ACC3F8FBFE05F88C" ID-CoL="54SPK" authority="Karny, 1923: 141" authorityName="Karny" authorityPageNumber="141" authorityYear="1923" box="[151,480,1792,1816]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="30" pageNumber="481" phylum="Arthropoda" rank="species" species="incisa">
<emphasis id="B93BEAF0AB2CA645ACC3F8FBFEC9F88C" box="[151,300,1793,1816]" italics="true" pageId="30" pageNumber="481">Tapiena incisa</emphasis>
<bibRefCitation id="EFDE4B13AB2CA645AD66F8FAFE05F88C" author="Karny, H. H." box="[306,480,1792,1816]" pageId="30" pageNumber="481" pagination="117 - 193" refId="ref14905" refString="Karny, H. H. (1923) On Malaysian katydids (Gryllacridae and Tettigoniidae) from the Raffles Museum, Singapore. Journal of the Malaysian Branch of the Royal Asiatic Society, 1, 117 - 193." type="journal article" year="1923">Karny, 1923: 141</bibRefCitation>
</taxonomicName>
</treatmentCitation>
—
<treatmentCitation id="0AEE10F3AB2CA645ADAFF8FAFC8EF88C" author="Tan, M. K. &amp; Liu, C. &amp; Artchawakom, T." box="[507,875,1792,1816]" page="42" pageId="30" pageNumber="481" year="2015">
<bibRefCitation id="EFDE4B13AB2CA645ADAFF8FAFC8EF88C" author="Tan, M. K. &amp; Liu, C. &amp; Artchawakom, T." box="[507,875,1792,1816]" pageId="30" pageNumber="481" pagination="40 - 50" refId="ref15357" refString="Tan, M. K., Liu, C. &amp; Artchawakom, T. (2015) Taxonomic review of Tapiena (Orthoptera: Phaneropteridae: Phaneropterinae), with key to species and new species from Thailand. Zootaxa, 3920 (1), 40 - 50. https: // doi. org / 10.11646 / zootaxa. 3920.1.2" type="journal article" year="2015">Tan, Liu &amp; Artchawakom, 2015: 42</bibRefCitation>
</treatmentCitation>
, 43 (key);
<treatmentCitation id="0AEE10F3AB2CA645AF8CF8FAFAF1F88C" author="Tan, M. K. &amp; Japir, R. &amp; Chung, A. Y. C." box="[984,1300,1792,1816]" httpUri="http://treatment.plazi.org/id/039F8783FFF7AD3B10EF62ECFF076A8E" page="328" pageId="30" pageNumber="481" year="2019">
<bibRefCitation id="EFDE4B13AB2CA645AF8CF8FAFAF1F88C" author="Tan, M. K. &amp; Japir, R. &amp; Chung, A. Y. C." box="[984,1300,1792,1816]" pageId="30" pageNumber="481" pagination="301 - 349" refId="ref15290" refString="Tan, M. K., Japir, R. &amp; Chung, A. Y. C. (2019) Uncovering the Grylloidea and Tettigonioidea (Orthoptera: Ensifera) in the Forest Research Center (Sepilok) Entomological Collection. Zootaxa, 4701 (4), 301 - 349. https: // doi. org / 10.11646 / zootaxa. 4701.4.1" type="journal article" year="2019">Tan, Japir &amp; Chung, 2019: 328</bibRefCitation>
</treatmentCitation>
(see remarks below)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3556569AB2CA644ACC3F89EFC51FF40" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481" type="materials_examined">
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<emphasis id="B93BEAF0AB2CA645ACC3F89EFE67F8EA" bold="true" box="[151,386,1892,1918]" pageId="30" pageNumber="481">Material examined.</emphasis>
<collectionCode id="ED5EAE27AB2CA645ADDFF89EFE35F8EA" box="[395,464,1892,1918]" pageId="30" pageNumber="481">EAST</collectionCode>
<materialsCitation id="3B273CBFAB2CA644AD8DF89EFC51FF40" collectingDate="2023-10-31" collectionCode="FRC" collectorName="M. K. Tan &amp; Tan, J. L &amp; A. Y. C. Chung &amp; Trusmadi Entomology Camp" country="Malaysia" county="Mount Trus Madi" elevation="5" lastPageId="31" lastPageNumber="482" latitude="5.44307" location="Mount Trus Madi" longLatPrecision="1" longitude="116.45155" municipality="Trusmadi Entomology Camp" pageId="30" pageNumber="481" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Sabah">
<collectingCountry id="F3587672AB2CA645AD8DF89EFD8AF8EA" box="[473,623,1892,1918]" name="Malaysia" pageId="30" pageNumber="481">MALAYSIA</collectingCountry>
:
<collectingRegion id="498BF800AB2CA645AE2FF89EFCE5F8EA" box="[635,768,1892,1918]" country="Malaysia" name="Sabah" pageId="30" pageNumber="481">Sabah State</collectingRegion>
•
<specimenCount id="9D49FD6BAB2CA645AF48F89EFCA1F8EA" box="[796,836,1892,1918]" count="1" pageId="30" pageNumber="481" type="male">1♂</specimenCount>
<specimenCount id="9D49FD6BAB2CA645AF10F89EFC86F8EA" box="[836,867,1892,1918]" count="1" pageId="30" pageNumber="481" type="female">1♀</specimenCount>
;
<collectingCounty id="62914E6EAB2CA645AF27F89EFBDAF8EA" box="[883,1087,1892,1918]" pageId="30" pageNumber="481">Mount Trus Madi</collectingCounty>
,
<collectingMunicipality id="6B94AC98AB2CA645A81FF89EFA7DF8EA" box="[1099,1432,1892,1918]" pageId="30" pageNumber="481">Trusmadi Entomology Camp</collectingMunicipality>
;
<geoCoordinate id="EE7B5025AB2CA645ACC3F87DFEEFF836" box="[151,266,1927,1954]" degrees="5.44174" direction="north" orientation="latitude" pageId="30" pageNumber="481" precision="1" value="5.44174">N5.44174</geoCoordinate>
,
<geoCoordinate id="EE7B5025AB2CA645AD42F87DFE45F836" box="[278,416,1927,1954]" degrees="116.45074" direction="east" orientation="longitude" pageId="30" pageNumber="481" precision="1" value="116.45074">E116.45074</geoCoordinate>
, 1189.0±
<quantity id="4CB79B07AB2CA645AE5CF87DFDB5F835" box="[520,592,1927,1954]" metricMagnitude="0" metricUnit="m" metricValue="5.2" pageId="30" pageNumber="481" unit="m" value="5.2">
<elevation id="0062D1D1AB2CA645AE5CF87DFDB5F835" box="[520,592,1927,1954]" metricMagnitude="0" metricUnit="m" metricValue="5.2" pageId="30" pageNumber="481" unit="m" value="5.2">5.2 m</elevation>
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.a.s.l.;
<date id="FFF11022AB2CA645AEC1F87DFCB2F835" box="[661,855,1927,1953]" pageId="30" pageNumber="481" value="2023-10-31">
<collectingDate id="EFB5E9CAAB2CA645AEC1F87DFCB2F835" box="[661,855,1927,1953]" pageId="30" pageNumber="481" value="2023-10-31">31 October 2023</collectingDate>
</date>
, 19h58–20h23;
<collectingMethod id="520E4EF5AB2CA645A85AF87DFB18F836" box="[1038,1277,1927,1954]" pageId="30" pageNumber="481">attracted to light trap</collectingMethod>
at ridge; coll.
<collectorName id="26BA5334AB2CA645ACC3F851FEE2F851" box="[151,263,1963,1989]" pageId="30" pageNumber="481">M.K. Tan</collectorName>
, J.L. Yukang &amp;
<collectorName id="26BA5334AB2CA645AD91F850FD8CF850" box="[453,617,1962,1988]" pageId="30" pageNumber="481">A.Y.C. Chung</collectorName>
; SBH.23.14, 28 (
<collectionCode id="ED5EAE27AB2CA645AF66F850FC8FF850" box="[818,874,1962,1988]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="30" pageNumber="481">FRC</collectionCode>
) •
<specimenCount id="9D49FD6BAB2CA645AFDBF850FC56F850" box="[911,947,1962,1988]" count="1" pageId="30" pageNumber="481" type="female">1♀</specimenCount>
;
<location id="8E906039AB2CA645AF97F851FB74F850" LSID="urn:lsid:plazi:treatment:03E687F4AB2CA644ACC3F96DFBC4FD0A:8E906039AB2CA645AF97F851FB74F850" box="[963,1169,1962,1989]" country="Malaysia" county="Mount Trus Madi" latitude="5.44307" longLatPrecision="1" longitude="116.45155" municipality="Trusmadi Entomology Camp" name="Mount Trus Madi" pageId="30" pageNumber="481" stateProvince="Sabah">Mount Trus Madi</location>
,
<collectorName id="26BA5334AB2CA644A8CAF850FF38FF25" lastPageId="31" lastPageNumber="482" pageId="30" pageNumber="481">Trusmadi Entomology Camp</collectorName>
;
<geoCoordinate id="EE7B5025AB2DA644ACB8FF6CFEBBFF25" box="[236,350,150,177]" degrees="5.44307" direction="north" orientation="latitude" pageId="31" pageNumber="482" precision="1" value="5.44307">N5.44307</geoCoordinate>
,
<geoCoordinate id="EE7B5025AB2DA644AD38FF6CFE13FF25" box="[364,502,150,177]" degrees="116.45155" direction="east" orientation="longitude" pageId="31" pageNumber="482" precision="1" value="116.45155">E116.45155</geoCoordinate>
, 1192.0±
<quantity id="4CB79B07AB2DA644AE34FF6CFD4FFF25" box="[608,682,150,177]" metricMagnitude="0" metricUnit="m" metricValue="5.6" pageId="31" pageNumber="482" unit="m" value="5.6">
<elevation id="0062D1D1AB2DA644AE34FF6CFD4FFF25" box="[608,682,150,177]" metricMagnitude="0" metricUnit="m" metricValue="5.6" pageId="31" pageNumber="482" unit="m" value="5.6">5.6 m</elevation>
</quantity>
.a.s.l.;
<date id="FFF11022AB2DA644AEBBFF6DFC50FF25" box="[751,949,151,177]" pageId="31" pageNumber="482" value="2023-10-31">
<collectingDate id="EFB5E9CAAB2DA644AEBBFF6DFC50FF25" box="[751,949,151,177]" pageId="31" pageNumber="482" value="2023-10-31">31 October 2023</collectingDate>
</date>
, 20h33–20h49;
<collectingMethod id="520E4EF5AB2DA644A83BFF6DFA87FF25" box="[1135,1378,151,177]" pageId="31" pageNumber="482">attracted to light trap</collectingMethod>
near camp; coll.
<collectorName id="26BA5334AB2DA644AD4FFF40FE6DFF40" box="[283,392,186,212]" pageId="31" pageNumber="482">M.K. Tan</collectorName>
, J.L. Yukang &amp;
<collectorName id="26BA5334AB2DA644AE6FFF40FD38FF40" box="[571,733,186,212]" pageId="31" pageNumber="482">A.Y.C. Chung</collectorName>
; SBH.23.36 (
<collectionCode id="ED5EAE27AB2DA644AF20FF40FC49FF40" box="[884,940,186,212]" country="USA" httpUri="http://grbio.org/cool/z9c1-g2b0" name="Fusarium Research Center" pageId="31" pageNumber="482">FRC</collectionCode>
)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3556569AB2DA644AC93FF27FD93FDC3" pageId="31" pageNumber="482" type="discussion">
<paragraph id="8BF036E2AB2DA644AC93FF27FCC2FD85" blockId="31.[151,1437,150,671]" pageId="31" pageNumber="482">
<emphasis id="B93BEAF0AB2DA644AC93FF27FEDEFF63" bold="true" box="[199,315,221,247]" pageId="31" pageNumber="482">Remarks.</emphasis>
Our specimen resembles most
<taxonomicName id="4C4F4D61AB2DA644AEFBFF24FC15FF63" authority="Karny, 1923" authorityName="Karny" authorityYear="1923" box="[687,1008,221,248]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="incisa">
<emphasis id="B93BEAF0AB2DA644AEFBFF24FCB3FF63" box="[687,854,222,247]" italics="true" pageId="31" pageNumber="482">Tapiena incisa</emphasis>
<bibRefCitation id="EFDE4B13AB2DA644AF36FF24FC15FF63" author="Karny, H. H." box="[866,1008,221,248]" pageId="31" pageNumber="482" pagination="117 - 193" refId="ref14905" refString="Karny, H. H. (1923) On Malaysian katydids (Gryllacridae and Tettigoniidae) from the Raffles Museum, Singapore. Journal of the Malaysian Branch of the Royal Asiatic Society, 1, 117 - 193." type="journal article" year="1923">Karny, 1923</bibRefCitation>
</taxonomicName>
previously described from
<collectingCountry id="F3587672AB2DA644A961FF27FA7DFF63" box="[1333,1432,221,247]" name="Malaysia" pageId="31" pageNumber="482">Sarawak</collectingCountry>
, particularly by the shape of the male cercus (
<figureCitation id="13742A67AB2DA644AECCFEFAFCE5FE8E" box="[664,768,256,282]" captionStart="FIGURE 33" captionStartId="32.[151,250,1982,2006]" captionTargetBox="[151,1436,590,1958]" captionTargetId="figure-122@32.[151,1436,590,1958]" captionTargetPageId="32" captionText="FIGURE 33. Tapiena?incisa Karny, 1923 ♂: stridulatory area on the left (A) and right (B) tegmina in dorsal view; abdominal apex in lateral (C) and dorsal (D) views; subgenital plate in ventral view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064497" httpUri="https://zenodo.org/record/11064497/files/figure.png" pageId="31" pageNumber="482">Fig. 33C</figureCitation>
). However, our male specimen also differs in an important character: the male tenth abdominal tergite is truncated excised in the middle (and not triangularly excised) (
<figureCitation id="13742A67AB2DA644A93BFED9FF2EFEF5" captionStart="FIGURE 33" captionStartId="32.[151,250,1982,2006]" captionTargetBox="[151,1436,590,1958]" captionTargetId="figure-122@32.[151,1436,590,1958]" captionTargetPageId="32" captionText="FIGURE 33. Tapiena?incisa Karny, 1923 ♂: stridulatory area on the left (A) and right (B) tegmina in dorsal view; abdominal apex in lateral (C) and dorsal (D) views; subgenital plate in ventral view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064497" httpUri="https://zenodo.org/record/11064497/files/figure.png" pageId="31" pageNumber="482">Fig. 33D</figureCitation>
). This species is also similar to
<taxonomicName id="4C4F4D61AB2DA644AE60FEBDFC13FEF5" authority="Tan &amp; Wahab, 2018" authorityName="Tan &amp; Wahab" authorityYear="2018" box="[564,1014,327,353]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="paraincisa">
<emphasis id="B93BEAF0AB2DA644AE60FEBDFCE8FEF4" box="[564,781,327,352]" italics="true" pageId="31" pageNumber="482">Tapiena paraincisa</emphasis>
Tan &amp; Wahab, 2018
</taxonomicName>
from
<collectingCountry id="F3587672AB2DA644A869FEBDFAF0FEF5" box="[1085,1301,327,353]" name="Brunei" pageId="31" pageNumber="482">Brunei Darussalam</collectingCountry>
(not too far from Mount Trus Madi) by its stout body, red (or purple) fore femora and tympanal area (
<figureCitation id="13742A67AB2DA644A8DDFE90FB3EFE10" box="[1161,1243,362,388]" captionStart="FIGURE 32" captionStartId="31.[151,250,1574,1598]" captionTargetBox="[151,1435,694,1550]" captionTargetId="figure-238@31.[151,1435,694,1550]" captionTargetPageId="31" captionText="FIGURE 32. Tapiena?incisa Karny, 1923 ♀ attracted to light at Mount Trus Madi." figureDoi="http://doi.org/10.5281/zenodo.11064495" httpUri="https://zenodo.org/record/11064495/files/figure.png" pageId="31" pageNumber="482">Fig. 32</figureCitation>
) and stridulatory areas on both tegmina (
<figureCitation id="13742A67AB2DA644ADC1FE77FDDCFE33" box="[405,569,397,423]" captionStart="FIGURE 33" captionStartId="32.[151,250,1982,2006]" captionTargetBox="[151,1436,590,1958]" captionTargetId="figure-122@32.[151,1436,590,1958]" captionTargetPageId="32" captionText="FIGURE 33. Tapiena?incisa Karny, 1923 ♂: stridulatory area on the left (A) and right (B) tegmina in dorsal view; abdominal apex in lateral (C) and dorsal (D) views; subgenital plate in ventral view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064497" httpUri="https://zenodo.org/record/11064497/files/figure.png" pageId="31" pageNumber="482">Figs 33A, 33B</figureCitation>
), but also differs distinctly by the cercus at its apex broad and flattened (instead of tapering into two tooth-like processes) (
<figureCitation id="13742A67AB2DA644AE2EFE4AFD07FE5E" box="[634,738,432,458]" captionStart="FIGURE 33" captionStartId="32.[151,250,1982,2006]" captionTargetBox="[151,1436,590,1958]" captionTargetId="figure-122@32.[151,1436,590,1958]" captionTargetPageId="32" captionText="FIGURE 33. Tapiena?incisa Karny, 1923 ♂: stridulatory area on the left (A) and right (B) tegmina in dorsal view; abdominal apex in lateral (C) and dorsal (D) views; subgenital plate in ventral view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064497" httpUri="https://zenodo.org/record/11064497/files/figure.png" pageId="31" pageNumber="482">Fig. 33C</figureCitation>
), the tenth abdominal tergite being broadly and truncated and excised in middle (instead of narrowly excised) (
<figureCitation id="13742A67AB2DA644AEE6FE29FCF3FE7A" box="[690,790,467,494]" captionStart="FIGURE 33" captionStartId="32.[151,250,1982,2006]" captionTargetBox="[151,1436,590,1958]" captionTargetId="figure-122@32.[151,1436,590,1958]" captionTargetPageId="32" captionText="FIGURE 33. Tapiena?incisa Karny, 1923 ♂: stridulatory area on the left (A) and right (B) tegmina in dorsal view; abdominal apex in lateral (C) and dorsal (D) views; subgenital plate in ventral view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064497" httpUri="https://zenodo.org/record/11064497/files/figure.png" pageId="31" pageNumber="482">Fig. 33D</figureCitation>
) and the subgenital plate having much shorter apical lobes but longer styli (
<figureCitation id="13742A67AB2DA644AD04FE0DFE57FD85" box="[336,434,503,529]" captionStart="FIGURE 33" captionStartId="32.[151,250,1982,2006]" captionTargetBox="[151,1436,590,1958]" captionTargetId="figure-122@32.[151,1436,590,1958]" captionTargetPageId="32" captionText="FIGURE 33. Tapiena?incisa Karny, 1923 ♂: stridulatory area on the left (A) and right (B) tegmina in dorsal view; abdominal apex in lateral (C) and dorsal (D) views; subgenital plate in ventral view (E). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.11064497" httpUri="https://zenodo.org/record/11064497/files/figure.png" pageId="31" pageNumber="482">Fig. 33E</figureCitation>
) (see also
<bibRefCitation id="EFDE4B13AB2DA644AE7EFE0DFCFEFD85" author="Tan, M. K. &amp; Wahab, R. A." box="[554,795,503,529]" pageId="31" pageNumber="482" pagination="193 - 196" refId="ref15480" refString="Tan, M. K. &amp; Wahab, R. A. (2018 a) A new species of Tapiena (Orthoptera: Tettigonioidea: Phaneropterinae) from Brunei Darussalam (Borneo, Southeast Asia). Zootaxa, 4413 (1), 193 - 196. https: // doi. org / 10.11646 / zootaxa. 4413.1.9" type="journal article" year="2018">Tan &amp; Wahab, 2018a</bibRefCitation>
).
</paragraph>
<paragraph id="8BF036E2AB2DA644AC93FDE0FD93FDC3" blockId="31.[151,1437,150,671]" pageId="31" pageNumber="482">
Based on new material from
<collectingRegion id="498BF800AB2DA644AE50FDE0FDAEFDA0" box="[516,587,538,564]" country="Malaysia" name="Sabah" pageId="31" pageNumber="482">Sabah</collectingRegion>
, it appears that
<bibRefCitation id="EFDE4B13AB2DA644AEA0FDE0FC4FFDA0" author="Tan, M. K. &amp; Japir, R. &amp; Chung, A. Y. C." box="[756,938,538,564]" pageId="31" pageNumber="482" pagination="301 - 349" refId="ref15290" refString="Tan, M. K., Japir, R. &amp; Chung, A. Y. C. (2019) Uncovering the Grylloidea and Tettigonioidea (Orthoptera: Ensifera) in the Forest Research Center (Sepilok) Entomological Collection. Zootaxa, 4701 (4), 301 - 349. https: // doi. org / 10.11646 / zootaxa. 4701.4.1" type="journal article" year="2019">
Tan
<emphasis id="B93BEAF0AB2DA644AF77FDE1FCB6FDA0" box="[803,851,538,564]" italics="true" pageId="31" pageNumber="482">et al</emphasis>
. (2019)
</bibRefCitation>
may have misidentified the
<taxonomicName id="4C4F4D61AB2DA644A8B4FDE1FADCFDA0" authorityName="Bolivar" authorityYear="1906" box="[1248,1337,539,564]" class="Insecta" family="Tettigoniidae" genus="Tapiena" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAF0AB2DA644A8B4FDE1FADCFDA0" box="[1248,1337,539,564]" italics="true" pageId="31" pageNumber="482">Tapiena</emphasis>
</taxonomicName>
reported in Lungmanis (near Lahad Datu in
<collectingRegion id="498BF800AB2DA644AE75FDC7FD8FFDC3" box="[545,618,573,599]" country="Malaysia" name="Sabah" pageId="31" pageNumber="482">Sabah</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C3556569AB2DA644AC93FD9AFD62FDEE" box="[199,647,608,635]" pageId="31" pageNumber="482" type="distribution">
<paragraph id="8BF036E2AB2DA644AC93FD9AFD62FDEE" blockId="31.[151,1437,150,671]" box="[199,647,608,635]" pageId="31" pageNumber="482">
<emphasis id="B93BEAF0AB2DA644AC93FD9AFE84FDEE" bold="true" box="[199,353,608,634]" pageId="31" pageNumber="482">Distribution.</emphasis>
Borneo (
<collectingRegion id="498BF800AB2DA644AD9DFD9AFDF7FDEE" box="[457,530,608,634]" country="Malaysia" name="Sabah" pageId="31" pageNumber="482">Sabah</collectingRegion>
,
<collectingCountry id="F3587672AB2DA644AE49FD9AFD9AFDEE" box="[541,639,608,634]" name="Malaysia" pageId="31" pageNumber="482">Sarawak</collectingCountry>
)
</paragraph>
</subSubSection>
<subSubSection id="C3556569AB2DA644AC93FD7EFBC4FD0A" box="[199,1057,644,671]" pageId="31" pageNumber="482" type="materials_examined">
<paragraph id="8BF036E2AB2DA644AC93FD7EFBC4FD0A" blockId="31.[151,1437,150,671]" box="[199,1057,644,671]" pageId="31" pageNumber="482">
<materialsCitation id="3B273CBFAB2DA644AC93FD7EFBC4FD0A" box="[199,1057,644,671]" collectionCode="EAST" country="Malaysia" location="Gunung Tamabo" municipality="Baram River" pageId="31" pageNumber="482" specimenCount="1" stateProvince="Sarawak" typeStatus="holotype">
<emphasis id="B93BEAF0AB2DA644AC93FD7EFE83FD0A" bold="true" box="[199,358,644,670]" pageId="31" pageNumber="482">
<typeStatus id="54F48840AB2DA644AC93FD7EFEE7FD0A" box="[199,258,644,670]" pageId="31" pageNumber="482">Type</typeStatus>
locality.
</emphasis>
<collectionCode id="ED5EAE27AB2DA644AD38FD7EFE54FD0A" box="[364,433,644,670]" pageId="31" pageNumber="482">EAST</collectionCode>
<collectingCountry id="F3587672AB2DA644ADEDFD7EFDAAFD0A" box="[441,591,644,670]" name="Malaysia" pageId="31" pageNumber="482">MALAYSIA</collectingCountry>
:
<collectingRegion id="498BF800AB2DA644AE0DFD7EFD5BFD0A" box="[601,702,644,670]" country="Malaysia" name="Sarawak" pageId="31" pageNumber="482">Sarawak</collectingRegion>
:
<collectingMunicipality id="6B94AC98AB2DA644AE9CFD7FFCB8FD0A" box="[712,861,644,671]" pageId="31" pageNumber="482">Baram River</collectingMunicipality>
:
<location id="8E906039AB2DA644AF33FD7EFBC4FD0A" LSID="urn:lsid:plazi:treatment:03E687F4AB2CA644ACC3F96DFBC4FD0A:8E906039AB2DA644AF33FD7EFBC4FD0A" box="[871,1057,644,670]" country="Malaysia" municipality="Baram River" name="Gunung Tamabo" pageId="31" pageNumber="482" stateProvince="Sarawak">Gunung Tamabo</location>
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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