263 lines
32 KiB
XML
263 lines
32 KiB
XML
<document id="AE7D1B225B56BA95EB3091E54F3B94C0" ID-DOI="10.11646/zootaxa.3736.5.8" ID-GBIF-Dataset="c231e1f1-ac35-406c-871b-a5ad87995af9" ID-ISSN="1175-5326" ID-Zenodo-Dep="249099" ID-ZooBank="D961FDF8-B2FB-4B63-A01A-45A1978FC6D4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460699311845" checkinUser="plazi" docAuthor="Giłka, Wojciech, Zakrzewska, Marta, Dominiak, Patrycja & Urbanek, Aleksandra" docDate="2013" docId="03E687CEEA1CFFF9FF0BFB1D4C4AFC25" docLanguage="en" docName="zt03736p586.pdf" docOrigin="Zootaxa 3736 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rheotanytarsus alliciens Gilka et Zakrzewska, sp. nov." docType="treatment" docVersion="9" lastPageNumber="581" masterDocId="FFDFFFB6EA16FFF5FF9CFFA6482BFFF9" masterDocTitle="Non-biting midges of the tribe Tanytarsini in Eocene amber from the Rovno region (Ukraine): a pioneer systematic study with notes on the phylogeny (Diptera: Chironomidae)" masterLastPageNumber="586" masterPageNumber="569" pageNumber="579" updateTime="1698354172026" updateUser="plazi">
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<mods:mods id="68CC55EE05B3DFC39E137A4DF0D60B16" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="1BB838DC07B5B396DFF1D162C78B88F1">
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<mods:title id="C3A8EF320FA1441B876C25E51EF5A25F">Non-biting midges of the tribe Tanytarsini in Eocene amber from the Rovno region (Ukraine): a pioneer systematic study with notes on the phylogeny (Diptera: Chironomidae)</mods:title>
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<mods:roleTerm id="BC3C19E16A92F8EA20A6A86DE6F87E6C">Author</mods:roleTerm>
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<mods:namePart id="72C76035AE15EA5F796713FFC645E307">Giłka, Wojciech</mods:namePart>
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</mods:name>
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<mods:roleTerm id="2AB9A1564CFA3D3B64C7CFE67187C2EF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="D62951DB484C2DA42F785962CBCDEA60">Zakrzewska, Marta</mods:namePart>
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</mods:name>
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<mods:roleTerm id="EDDFD2DC26AD75BB616003A1E1888673">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="45398D0E12992AB37B3D26DED6111F09">Dominiak, Patrycja</mods:namePart>
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</mods:name>
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<mods:roleTerm id="613683F385C9EA0C28C3A9072A9BF873">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8AEE374A918D41CBCF348856F1E91035">Urbanek, Aleksandra</mods:namePart>
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<mods:typeOfResource id="03C75A07E237BB7A6CF68AB721969D9C">text</mods:typeOfResource>
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<mods:title id="C7C3A6130D794467065CBD5474AC7D8B">Zootaxa</mods:title>
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<mods:part id="50EE32434E36A1A876DA9EDDFB4E4A6A">
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<mods:date id="382FD052005BD02C49C0A78B6276780E">2013</mods:date>
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<mods:detail id="700E905A234426E1EEAD01674B955E23" type="volume">
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<mods:number id="D8B1D36592FCABFB00CB429C3B58F807">3736</mods:number>
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<mods:detail id="FA498DAF2AAA630E0003D8A35E70997B" type="issue">
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<mods:number id="B132FF6329F536577F0F97EE26E4006A">5</mods:number>
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<mods:classification id="0845C1C714D1AB12C292BB8E60A0ACDE">journal article</mods:classification>
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<mods:identifier id="64A49270BDB84FC710E4B969579F2D02" type="DOI">10.11646/zootaxa.3736.5.8</mods:identifier>
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<treatment id="03E687CEEA1CFFF9FF0BFB1D4C4AFC25" ID-DOI="http://doi.org/10.5281/zenodo.6161854" ID-GBIF-Taxon="119582678" ID-Zenodo-Dep="6161854" LSID="urn:lsid:plazi:treatment:03E687CEEA1CFFF9FF0BFB1D4C4AFC25" httpUri="http://treatment.plazi.org/id/03E687CEEA1CFFF9FF0BFB1D4C4AFC25" lastPageId="12" lastPageNumber="581" pageId="10" pageNumber="579">
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<subSubSection id="C3556553EA1CFFFFFF0BFB1D4B1EFB2C" box="[151,821,1211,1237]" pageId="10" pageNumber="579" type="nomenclature">
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<paragraph id="8BF036D8EA1CFFFFFF0BFB1D4B1EFB2C" blockId="10.[151,821,1211,1237]" box="[151,821,1211,1237]" pageId="10" pageNumber="579">
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<heading id="D0B881B4EA1CFFFFFF0BFB1D4B1EFB2C" bold="true" box="[151,821,1211,1237]" fontSize="11" level="1" pageId="10" pageNumber="579" reason="1">
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<emphasis id="B93BEACAEA1CFFFFFF0BFB1D4B1EFB2C" bold="true" box="[151,821,1211,1237]" pageId="10" pageNumber="579">
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<taxonomicName id="4C4F4D5BEA1CFFFFFF0BFB1D4AE0FB2C" authority="Gilka et Zakrzewska" authorityName="Gilka et Zakrzewska" box="[151,715,1211,1237]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="579" phylum="Arthropoda" rank="species" species="alliciens" status="sp. nov.">
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<emphasis id="B93BEACAEA1CFFFFFF0BFB1D4994FB2C" bold="true" box="[151,447,1211,1237]" italics="true" pageId="10" pageNumber="579">Rheotanytarsus alliciens</emphasis>
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Giłka
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<emphasis id="B93BEACAEA1CFFFFFD88FB1D4A02FB2C" bold="true" box="[532,553,1211,1237]" italics="true" pageId="10" pageNumber="579">et</emphasis>
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Zakrzewska
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20857B1EA1CFFFFFD44FB1D4B1EFB2C" box="[728,821,1211,1237]" pageId="10" pageNumber="579" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3556553EA1CFFFFFF0BFAA44CE6FAC6" pageId="10" pageNumber="579" type="materials_examined">
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<paragraph id="8BF036D8EA1CFFFFFF0BFAA44CE6FAC6" blockId="10.[151,1437,1282,1999]" pageId="10" pageNumber="579">
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<emphasis id="B93BEACAEA1CFFFFFF0BFAA44961FAE2" bold="true" box="[151,330,1282,1307]" pageId="10" pageNumber="579">
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<typeStatus id="54F4887AEA1CFFFFFF0BFAA448F9FAE2" box="[151,210,1282,1307]" pageId="10" pageNumber="579">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54F4887AEA1CFFFFFECAFAA449E9FAE2" box="[342,450,1282,1307]" pageId="10" pageNumber="579" type="holotype">Holotype</typeStatus>
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. Adult male (distal flagellomeres, palpi and tarsi missing or broken and separated) preserved in 12 x
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<date id="FFF11018EA1CFFFFFEC3FA8149A9FAC7" box="[351,386,1319,1342]" pageId="10" pageNumber="579">7 x</date>
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<quantity id="4CB79B3DEA1CFFFFFE15FA8149E1FAC7" box="[393,458,1319,1343]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="579" unit="mm" value="2.0">2 mm</quantity>
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piece of amber (SIZ 155,
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<figureCitation id="13742A5DEA1CFFFFFD6DFA804B18FAC6" box="[753,819,1318,1343]" captionStart="FIGURE 6" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[246,1343,157,1924]" captionTargetId="figure@11.[246,1344,157,1924]" captionTargetPageId="11" captionText="FIGURE 6. Rheotanytarsus alliciens sp. nov., male. A— holotype (inclusion); B— habitus; C— head with antenna; D – wing; E – H— hypopygium: E, F— dorsal aspect; G— superior volsella, digitus and median volsella; H— median volsella (magnified c. three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249105/files/figure.png" pageId="10" pageNumber="579">Fig. 6</figureCitation>
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A; syninclusion: Acari,
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<specimenCount id="9D49FD51EA1CFFFFFBA2FA814CE9FAC6" box="[1086,1218,1318,1343]" pageId="10" pageNumber="579" type="generic">1 specimen</specimenCount>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3556553EA1CFFFFFF5BFAEC4ADDFA7E" pageId="10" pageNumber="579" type="etymology">
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<paragraph id="8BF036D8EA1CFFFFFF5BFAEC4ADDFA7E" blockId="10.[151,1437,1282,1999]" pageId="10" pageNumber="579">
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<emphasis id="B93BEACAEA1CFFFFFF5BFAEC498CFA9A" bold="true" box="[199,423,1354,1379]" pageId="10" pageNumber="579">Derivatio nominis.</emphasis>
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From Latin, meaning ‘interesting, engaging’. This is the oldest
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<taxonomicName id="4C4F4D5BEA1CFFFFFB0CFAEC4D15FA9A" box="[1168,1342,1354,1379]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="579" phylum="Arthropoda" rank="genus">
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<emphasis id="B93BEACAEA1CFFFFFB0CFAEC4D15FA9A" box="[1168,1342,1354,1379]" italics="true" pageId="10" pageNumber="579">Rheotanytarsus</emphasis>
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</taxonomicName>
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, known from male with a fully-developed hypopygial digitus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556553EA1CFFFFFF5BFA344B6AF9C2" pageId="10" pageNumber="579" type="diagnosis">
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<paragraph id="8BF036D8EA1CFFFFFF5BFA344B6AF9C2" blockId="10.[151,1437,1282,1999]" pageId="10" pageNumber="579">
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<emphasis id="B93BEACAEA1CFFFFFF5BFA34496AFA52" bold="true" box="[199,321,1426,1451]" pageId="10" pageNumber="579">Diagnosis.</emphasis>
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Tibial combs of mid and hind legs with short spurs at most twice the length of the comb. Gonostylus distinctly narrowed in distal half, much longer than gonocoxite. Anal point long, triangular, tapering to slender pointed apex, bearing two long crests; spinulae absent. Superior volsella roundish, with slightly projecting posterior part. Digitus long, distinctly extending beyond superior volsella. Stem of median volsella stout and short, with several pectinate or leaf-shaped lamellae fused at bases.
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3556553EA1CFFF9FF5BF9E04977FD61" lastPageId="12" lastPageNumber="581" pageId="10" pageNumber="579" type="description">
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<paragraph id="8BF036D8EA1CFFFFFF5BF9E04C04F9A7" blockId="10.[151,1437,1282,1999]" box="[199,1071,1606,1631]" pageId="10" pageNumber="579">
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<emphasis id="B93BEACAEA1CFFFFFF5BF9E049CEF9A6" bold="true" box="[199,485,1606,1631]" pageId="10" pageNumber="579">Description. Adult male</emphasis>
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(n = 1). Total length
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<quantity id="4CB79B3DEA1CFFFFFD4FF9E14B07F9A7" box="[723,812,1607,1631]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="10" pageNumber="579" unit="mm" value="2.3">2.3 mm</quantity>
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; wing length 1315 Μm.
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</paragraph>
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<paragraph id="8BF036D8EA1CFFFFFF5BF9CC4A68F95E" blockId="10.[151,1437,1282,1999]" pageId="10" pageNumber="579">
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<emphasis id="B93BEACAEA1CFFFFFF5BF9CC492FF97A" box="[199,260,1642,1667]" italics="true" pageId="10" pageNumber="579">Head</emphasis>
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(
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<figureCitation id="13742A5DEA1CFFFFFE88F9CC4973F97A" box="[276,344,1642,1667]" captionStart="FIGURE 6" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[246,1343,157,1924]" captionTargetId="figure@11.[246,1344,157,1924]" captionTargetPageId="11" captionText="FIGURE 6. Rheotanytarsus alliciens sp. nov., male. A— holotype (inclusion); B— habitus; C— head with antenna; D – wing; E – H— hypopygium: E, F— dorsal aspect; G— superior volsella, digitus and median volsella; H— median volsella (magnified c. three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249105/files/figure.png" pageId="10" pageNumber="579">Fig. 6</figureCitation>
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B, C). Eyes bare, with dorsomedian extensions. Frontal tubercles not observed. Antenna with 13 flagellomeres. Clypeals unobservable.
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</paragraph>
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<paragraph id="8BF036D8EA1CFFFFFF5BF91448CCF916" blockId="10.[151,1437,1282,1999]" pageId="10" pageNumber="579">
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<emphasis id="B93BEACAEA1CFFFFFF5BF9144933F932" box="[199,280,1714,1739]" italics="true" pageId="10" pageNumber="579">Thorax</emphasis>
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(
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<figureCitation id="13742A5DEA1CFFFFFEB5F9144945F932" box="[297,366,1714,1739]" captionStart="FIGURE 6" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[246,1343,157,1924]" captionTargetId="figure@11.[246,1344,157,1924]" captionTargetPageId="11" captionText="FIGURE 6. Rheotanytarsus alliciens sp. nov., male. A— holotype (inclusion); B— habitus; C— head with antenna; D – wing; E – H— hypopygium: E, F— dorsal aspect; G— superior volsella, digitus and median volsella; H— median volsella (magnified c. three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249105/files/figure.png" pageId="10" pageNumber="579">Fig. 6</figureCitation>
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B). At least several acrostichals present; Dc at least 7 on each side, Pa 2 on each side, Scts at least 5.
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</paragraph>
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<paragraph id="8BF036D8EA1CFFFFFF5BF95D4C9EF8A4" blockId="10.[151,1437,1282,1999]" pageId="10" pageNumber="579">
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<emphasis id="B93BEACAEA1CFFFFFF5BF95D48D4F8EB" box="[199,255,1787,1810]" italics="true" pageId="10" pageNumber="579">Wing</emphasis>
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(
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<figureCitation id="13742A5DEA1CFFFFFE8DF95C497EF8EA" box="[273,341,1786,1811]" captionStart="FIGURE 6" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[246,1343,157,1924]" captionTargetId="figure@11.[246,1344,157,1924]" captionTargetPageId="11" captionText="FIGURE 6. Rheotanytarsus alliciens sp. nov., male. A— holotype (inclusion); B— habitus; C— head with antenna; D – wing; E – H— hypopygium: E, F— dorsal aspect; G— superior volsella, digitus and median volsella; H— median volsella (magnified c. three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249105/files/figure.png" pageId="10" pageNumber="579">Fig. 6</figureCitation>
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D). Slender, with anal lobe weak, broadest at 2/3 length, width: 380 Μm, length/width ratio 3.46. Venation observable only in part. FCu placed distally of RM; VR Cu non-measurable. Visible veins ending as follows (in order from base to tip): An, Cu1, M3+4, R4+5, M1+2. Wing macrotrichia unobservable.
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</paragraph>
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<paragraph id="8BF036D8EA1CFFFFFF5BF8CB4C17F836" blockId="10.[151,1437,1282,1999]" pageId="10" pageNumber="579">
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||
<emphasis id="B93BEACAEA1CFFFFFF5BF8CB48D7F87D" box="[199,252,1901,1924]" italics="true" pageId="10" pageNumber="579">Legs</emphasis>
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. Apices of fore leg tibiae poorly preserved. Tibial combs of mid and hind legs with short spurs at most twice the length of the comb. Sensilla chaetica on ta1 of p2 not observed. Pulvilli not observed (distal tarsomeres missing or poorly preserved). For length of leg segments and leg ratios see
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<tableCitation id="C6CD0363EA1CFFFFFC7EF8104C1CF836" box="[994,1079,1974,1999]" captionStart="TABLE 5" captionStartId="12.[151,239,152,175]" captionTargetBox="[159,1379,202,359]" captionTargetPageId="12" captionText="TABLE 5. Leg segment lengths (µm) and leg ratios of male Rheotanytarsus alliciens sp. nov." httpUri="http://table.plazi.org/id/DF306650EA1AFFF9FF0BFF3E4CABFF56" pageId="10" pageNumber="579" tableUuid="DF306650EA1AFFF9FF0BFF3E4CABFF56">Table 5</tableCitation>
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.
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</paragraph>
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<caption id="DF306650EA1DFFFEFF0BF83A4A4DF816" httpUri="https://zenodo.org/record/249105/files/figure.png" pageId="11" pageNumber="580" targetBox="[246,1343,157,1924]" targetPageId="11">
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<paragraph id="8BF036D8EA1DFFFEFF0BF83A4A4DF816" blockId="11.[151,1436,1948,2031]" pageId="11" pageNumber="580">
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<emphasis id="B93BEACAEA1DFFFEFF0BF83A4938F848" bold="true" box="[151,275,1948,1970]" pageId="11" pageNumber="580">FIGURE 6.</emphasis>
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<taxonomicName id="4C4F4D5BEA1DFFFEFE86F83A4A3EF84B" box="[282,533,1948,1970]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="alliciens" status="sp. nov.">
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<emphasis id="B93BEACAEA1DFFFEFE86F83A4A3EF84B" box="[282,533,1948,1970]" italics="true" pageId="11" pageNumber="580">Rheotanytarsus alliciens</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20857B1EA1DFFFEFD80F83A4A46F848" box="[540,621,1948,1969]" pageId="11" pageNumber="580" rank="species">
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<emphasis id="B93BEACAEA1DFFFEFD80F83A4A46F848" bold="true" box="[540,621,1948,1969]" pageId="11" pageNumber="580">sp. nov.</emphasis>
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</taxonomicNameLabel>
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, male.
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<emphasis id="B93BEACAEA1DFFFEFD25F83A4ACEF848" bold="true" box="[697,741,1948,1969]" pageId="11" pageNumber="580">A—</emphasis>
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holotype (inclusion);
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<emphasis id="B93BEACAEA1DFFFEFC5CF83A4BC1F848" bold="true" box="[960,1002,1948,1969]" pageId="11" pageNumber="580">B—</emphasis>
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habitus;
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<emphasis id="B93BEACAEA1DFFFEFBDEF83A4C45F84B" bold="true" box="[1090,1134,1948,1970]" pageId="11" pageNumber="580">C—</emphasis>
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head with antenna;
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<emphasis id="B93BEACAEA1DFFFEFAA9F83A4D70F848" bold="true" box="[1333,1371,1948,1969]" pageId="11" pageNumber="580">D –</emphasis>
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wing;
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<emphasis id="B93BEACAEA1DFFFEFF0BF81C48CAF836" bold="true" box="[151,225,1978,1999]" pageId="11" pageNumber="580">E–H—</emphasis>
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hypopygium:
|
||
<emphasis id="B93BEACAEA1DFFFEFEECF81C4993F836" bold="true" box="[368,440,1978,1999]" pageId="11" pageNumber="580">E, F—</emphasis>
|
||
dorsal aspect;
|
||
<emphasis id="B93BEACAEA1DFFFEFDD7F81C4A53F829" bold="true" box="[587,632,1978,2000]" pageId="11" pageNumber="580">G—</emphasis>
|
||
superior volsella, digitus and median volsella;
|
||
<emphasis id="B93BEACAEA1DFFFEFBCBF81C4CAFF836" bold="true" box="[1111,1156,1978,1999]" pageId="11" pageNumber="580">H—</emphasis>
|
||
median volsella (magnified c. three times relative to hypopygium drawn).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF306650EA1AFFF9FF0BFF3E4CABFF56" ID-Table-UUID="DF306650EA1AFFF9FF0BFF3E4CABFF56" box="[151,1152,152,175]" httpUri="http://table.plazi.org/id/DF306650EA1AFFF9FF0BFF3E4CABFF56" pageId="12" pageNumber="581" targetBox="[159,1379,202,359]" targetIsTable="true" targetPageId="12">
|
||
<paragraph id="8BF036D8EA1AFFF9FF0BFF3E4CABFF56" blockId="12.[151,1152,152,175]" box="[151,1152,152,175]" pageId="12" pageNumber="581">
|
||
<emphasis id="B93BEACAEA1AFFF9FF0BFF3E4929FF56" bold="true" box="[151,258,152,175]" pageId="12" pageNumber="581">TABLE 5</emphasis>
|
||
. Leg segment lengths (µm) and leg ratios of male
|
||
<taxonomicName id="4C4F4D5BEA1AFFF9FC81FF3E4C08FF56" box="[797,1059,152,175]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="581" phylum="Arthropoda" rank="species" species="alliciens" status="sp. nov.">
|
||
<emphasis id="B93BEACAEA1AFFF9FC81FF3E4C08FF56" box="[797,1059,152,175]" italics="true" pageId="12" pageNumber="581">Rheotanytarsus alliciens</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20857B1EA1AFFF9FBB6FF3E4CABFF56" box="[1066,1152,152,175]" pageId="12" pageNumber="581" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF036D8EA1AFFF9FF4BFF6C4D48FE9D" pageId="12" pageNumber="581">
|
||
<table id="F94FC478EA1A000AFF03FF6C4D48FE9E" box="[159,1379,202,359]" gridcols="6" gridrows="4" pageId="12" pageNumber="581">
|
||
<tr id="357F349AEA1A000AFF03FF6C4D48FF1D" box="[159,1379,202,228]" gridrow="0" pageId="12" pageNumber="581">
|
||
<th id="76AE5DE6EA1A000AFF03FF6C48D6FF1D" box="[159,253,202,228]" gridcol="0" gridrow="0" pageId="12" pageNumber="581">fe</th>
|
||
<th id="76AE5DE6EA1A000AFEF3FF6C4AEFFF1D" box="[367,708,202,228]" gridcol="1" gridrow="0" pageId="12" pageNumber="581">ti ta1 ta2</th>
|
||
<th id="76AE5DE6EA1A000AFCAAFF6C4B77FF1D" box="[822,860,202,228]" gridcol="2" gridrow="0" pageId="12" pageNumber="581">ta3</th>
|
||
<th id="76AE5DE6EA1A000AFC52FF6C4BD8FF1D" box="[974,1011,202,228]" gridcol="3" gridrow="0" pageId="12" pageNumber="581">ta4</th>
|
||
<th id="76AE5DE6EA1A000AFBFAFF6C4C54FF1D" box="[1126,1151,202,228]" gridcol="4" gridrow="0" pageId="12" pageNumber="581">ta5</th>
|
||
<th id="76AE5DE6EA1A000AFB62FF6C4D48FF1D" box="[1278,1379,202,228]" gridcol="5" gridrow="0" pageId="12" pageNumber="581">LR</th>
|
||
</tr>
|
||
<tr id="357F349AEA1A000AFF03FF504D48FEE9" box="[159,1379,246,272]" gridrow="1" pageId="12" pageNumber="581">
|
||
<th id="76AE5DE6EA1A000AFF03FF5048D6FEE9" box="[159,253,246,272]" gridcol="0" gridrow="1" pageId="12" pageNumber="581">P1 -</th>
|
||
<td id="76AE5DE6EA1A000AFEF3FF504AEFFEE9" box="[367,708,246,272]" gridcol="1" gridrow="1" pageId="12" pageNumber="581">305–310 - -</td>
|
||
<td id="76AE5DE6EA1A000AFCAAFF504B77FEE9" box="[822,860,246,272]" gridcol="2" gridrow="1" pageId="12" pageNumber="581">-</td>
|
||
<td id="76AE5DE6EA1A000AFC52FF504BD8FEE9" box="[974,1011,246,272]" gridcol="3" gridrow="1" pageId="12" pageNumber="581">-</td>
|
||
<td id="76AE5DE6EA1A000AFBFAFF504C54FEE9" box="[1126,1151,246,272]" gridcol="4" gridrow="1" pageId="12" pageNumber="581">-</td>
|
||
<td id="76AE5DE6EA1A000AFB62FF504D48FEE9" box="[1278,1379,246,272]" gridcol="5" gridrow="1" pageId="12" pageNumber="581">-</td>
|
||
</tr>
|
||
<tr id="357F349AEA1A000AFF03FE854D48FEC5" box="[159,1379,291,316]" gridrow="2" pageId="12" pageNumber="581">
|
||
<th id="76AE5DE6EA1A000AFF03FE8548D6FEC5" box="[159,253,291,316]" gridcol="0" gridrow="2" pageId="12" pageNumber="581">P2 -</th>
|
||
<td id="76AE5DE6EA1A000AFEF3FE854AEFFEC5" box="[367,708,291,316]" gridcol="1" gridrow="2" pageId="12" pageNumber="581">- - -</td>
|
||
<td id="76AE5DE6EA1A000AFCAAFE854B77FEC5" box="[822,860,291,316]" gridcol="2" gridrow="2" pageId="12" pageNumber="581">-</td>
|
||
<td id="76AE5DE6EA1A000AFC52FE854BD8FEC5" box="[974,1011,291,316]" gridcol="3" gridrow="2" pageId="12" pageNumber="581">-</td>
|
||
<td id="76AE5DE6EA1A000AFBFAFE854C54FEC5" box="[1126,1151,291,316]" gridcol="4" gridrow="2" pageId="12" pageNumber="581">-</td>
|
||
<td id="76AE5DE6EA1A000AFB62FE854D48FEC5" box="[1278,1379,291,316]" gridcol="5" gridrow="2" pageId="12" pageNumber="581">-</td>
|
||
</tr>
|
||
<tr id="357F349AEA1A000AFF03FEE84D48FE9E" box="[159,1379,334,359]" gridrow="3" pageId="12" pageNumber="581">
|
||
<th id="76AE5DE6EA1A000AFF03FEE848D6FE9E" box="[159,253,334,359]" gridcol="0" gridrow="3" pageId="12" pageNumber="581">P3 670</th>
|
||
<td id="76AE5DE6EA1A000AFEF3FEE84AEFFE9E" box="[367,708,334,359]" gridcol="1" gridrow="3" pageId="12" pageNumber="581">595–605 380–400 255</td>
|
||
<td id="76AE5DE6EA1A000AFCAAFEE84B77FE9E" box="[822,860,334,359]" gridcol="2" gridrow="3" pageId="12" pageNumber="581">195</td>
|
||
<td id="76AE5DE6EA1A000AFC52FEE84BD8FE9E" box="[974,1011,334,359]" gridcol="3" gridrow="3" pageId="12" pageNumber="581">110</td>
|
||
<td id="76AE5DE6EA1A000AFBFAFEE84C54FE9E" box="[1126,1151,334,359]" gridcol="4" gridrow="3" pageId="12" pageNumber="581">-</td>
|
||
<td id="76AE5DE6EA1A000AFB62FEE84D48FE9E" box="[1278,1379,334,359]" gridcol="5" gridrow="3" pageId="12" pageNumber="581">0.64–0.66</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BF036D8EA1AFFF9FF5BFE0E4977FD61" blockId="12.[151,1437,423,988]" pageId="12" pageNumber="581">
|
||
<emphasis id="B93BEACAEA1AFFF9FF5BFE0E497DFE46" box="[199,342,424,447]" italics="true" pageId="12" pageNumber="581">Hypopygium</emphasis>
|
||
(
|
||
<figureCitation id="13742A5DEA1AFFF9FEFAFE014982FE39" box="[358,425,423,448]" captionStart="FIGURE 6" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[246,1343,157,1924]" captionTargetId="figure@11.[246,1344,157,1924]" captionTargetPageId="11" captionText="FIGURE 6. Rheotanytarsus alliciens sp. nov., male. A— holotype (inclusion); B— habitus; C— head with antenna; D – wing; E – H— hypopygium: E, F— dorsal aspect; G— superior volsella, digitus and median volsella; H— median volsella (magnified c. three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249105/files/figure.png" pageId="12" pageNumber="581">Fig. 6</figureCitation>
|
||
E–H). Gonostylus cylindrical, slightly curved, distinctly narrowed in distal half, c. 90 Μm long, much longer than gonocoxite. Anal point long, triangular, wide at base, tapering to slender pointed apex, bearing two long crests, spinulae absent, posterolateral setae not observed (
|
||
<figureCitation id="13742A5DEA1AFFF9FC78FE494C0DFDF1" box="[996,1062,495,520]" captionStart="FIGURE 6" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[246,1343,157,1924]" captionTargetId="figure@11.[246,1344,157,1924]" captionTargetPageId="11" captionText="FIGURE 6. Rheotanytarsus alliciens sp. nov., male. A— holotype (inclusion); B— habitus; C— head with antenna; D – wing; E – H— hypopygium: E, F— dorsal aspect; G— superior volsella, digitus and median volsella; H— median volsella (magnified c. three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249105/files/figure.png" pageId="12" pageNumber="581">Fig. 6</figureCitation>
|
||
E, F). Superior volsella roundish, with slightly projecting posterior part; digitus long, distinctly extending beyond superior volsella (
|
||
<figureCitation id="13742A5DEA1AFFF9FA8EFDB54D73FDD5" box="[1298,1368,531,556]" captionStart="FIGURE 6" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[246,1343,157,1924]" captionTargetId="figure@11.[246,1344,157,1924]" captionTargetPageId="11" captionText="FIGURE 6. Rheotanytarsus alliciens sp. nov., male. A— holotype (inclusion); B— habitus; C— head with antenna; D – wing; E – H— hypopygium: E, F— dorsal aspect; G— superior volsella, digitus and median volsella; H— median volsella (magnified c. three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249105/files/figure.png" pageId="12" pageNumber="581">Fig. 6</figureCitation>
|
||
E–G). Stem of median volsella stout but short, c. 5 Μm, with several pectinate or leaf-shaped lamellae fused at bases (
|
||
<figureCitation id="13742A5DEA1AFFF9FAF2FD91488FFD8D" captionStart="FIGURE 6" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[246,1343,157,1924]" captionTargetId="figure@11.[246,1344,157,1924]" captionTargetPageId="11" captionText="FIGURE 6. Rheotanytarsus alliciens sp. nov., male. A— holotype (inclusion); B— habitus; C— head with antenna; D – wing; E – H— hypopygium: E, F— dorsal aspect; G— superior volsella, digitus and median volsella; H— median volsella (magnified c. three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249105/files/figure.png" pageId="12" pageNumber="581">Fig. 6</figureCitation>
|
||
G, H). Inferior volsella reaching 1/3 length of gonostylus, somewhat S-shaped, with slightly swollen head-like apex (
|
||
<figureCitation id="13742A5DEA1AFFF9FF47FDD94936FD61" box="[219,285,639,664]" captionStart="FIGURE 6" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[246,1343,157,1924]" captionTargetId="figure@11.[246,1344,157,1924]" captionTargetPageId="11" captionText="FIGURE 6. Rheotanytarsus alliciens sp. nov., male. A— holotype (inclusion); B— habitus; C— head with antenna; D – wing; E – H— hypopygium: E, F— dorsal aspect; G— superior volsella, digitus and median volsella; H— median volsella (magnified c. three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249105/files/figure.png" pageId="12" pageNumber="581">Fig. 6</figureCitation>
|
||
E, F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556553EA1AFFF9FF5BFD054C4AFC25" pageId="12" pageNumber="581" type="discussion">
|
||
<paragraph id="8BF036D8EA1AFFF9FF5BFD054C4AFC25" blockId="12.[151,1437,423,988]" pageId="12" pageNumber="581">
|
||
<emphasis id="B93BEACAEA1AFFF9FF5BFD054910FD45" bold="true" box="[199,315,675,700]" pageId="12" pageNumber="581">Remarks.</emphasis>
|
||
Adult males of
|
||
<taxonomicName id="4C4F4D5BEA1AFFF9FD9BFD054A9EFD45" box="[519,693,675,700]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="581" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93BEACAEA1AFFF9FD9BFD054A9EFD45" box="[519,693,675,700]" italics="true" pageId="12" pageNumber="581">Rheotanytarsus</emphasis>
|
||
</taxonomicName>
|
||
are distinguishable by the long and usually distally narrowed gonostylus and by the elongate anal tergite point and its crests. Their lamellae are more or less fused into a plate on the stem of the median volsella. They are also known to lack the digitus. In rare cases, the lamellae are separate or fused only at their bases, and the digitus is vestigial and only exceptionally fully developed (Kyerematen & Saether 2000, Giłka 2011a, Moubayed-Breil
|
||
<emphasis id="B93BEACAEA1AFFF9FDA8FC924A4EFCB5" box="[564,613,819,844]" italics="true" pageId="12" pageNumber="581">et al</emphasis>
|
||
. 2012). We recognize that the fusion of the lamellae and the formation of the plate, as well as the reduction of the digitus are distinct trends in
|
||
<taxonomicName id="4C4F4D5BEA1AFFF9FC2DFCF14C74FC89" box="[945,1119,855,880]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="581" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93BEACAEA1AFFF9FC2DFCF14C74FC89" box="[945,1119,855,880]" italics="true" pageId="12" pageNumber="581">Rheotanytarsus</emphasis>
|
||
</taxonomicName>
|
||
; in contrast, the lack of the plate and the presence of the digitus indicate plesiomorphy as displayed by
|
||
<taxonomicName id="4C4F4D5BEA1AFFF9FC78FCDA4C49FC6D" box="[996,1122,891,916]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="581" phylum="Arthropoda" rank="species" species="alliciens">
|
||
<emphasis id="B93BEACAEA1AFFF9FC78FCDA4C49FC6D" box="[996,1122,891,916]" italics="true" pageId="12" pageNumber="581">R. alliciens</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13742A5DEA1AFFF9FBEDFCDD4C99FC6D" box="[1137,1202,891,916]" captionStart="FIGURE 6" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[246,1343,157,1924]" captionTargetId="figure@11.[246,1344,157,1924]" captionTargetPageId="11" captionText="FIGURE 6. Rheotanytarsus alliciens sp. nov., male. A— holotype (inclusion); B— habitus; C— head with antenna; D – wing; E – H— hypopygium: E, F— dorsal aspect; G— superior volsella, digitus and median volsella; H— median volsella (magnified c. three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249105/files/figure.png" pageId="12" pageNumber="581">Fig. 6</figureCitation>
|
||
E–H) and few extant
|
||
<taxonomicName id="4C4F4D5BEA1AFFF9FF0BFC39496EFC41" box="[151,325,927,952]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="581" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93BEACAEA1AFFF9FF0BFC39496EFC41" box="[151,325,927,952]" italics="true" pageId="12" pageNumber="581">Rheotanytarsus</emphasis>
|
||
</taxonomicName>
|
||
(op. cit.). The adult male of
|
||
<taxonomicName id="4C4F4D5BEA1AFFF9FD0AFC064B3DFC41" box="[662,790,927,952]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="581" phylum="Arthropoda" rank="species" species="alliciens">
|
||
<emphasis id="B93BEACAEA1AFFF9FD0AFC064B3DFC41" box="[662,790,927,952]" italics="true" pageId="12" pageNumber="581">R. alliciens</emphasis>
|
||
</taxonomicName>
|
||
is peculiar in the shape of its superior volsella which is roundish, but has the slight posterior projection typical of
|
||
<taxonomicName id="4C4F4D5BEA1AFFF9FCBDFC654BE4FC25" box="[801,975,963,988]" class="Insecta" family="Chironomidae" genus="Rheotanytarsus" kingdom="Animalia" order="Diptera" pageId="12" pageNumber="581" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93BEACAEA1AFFF9FCBDFC654BE4FC25" box="[801,975,963,988]" italics="true" pageId="12" pageNumber="581">Rheotanytarsus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13742A5DEA1AFFF9FC42FC654C0BFC25" box="[990,1056,963,988]" captionStart="FIGURE 6" captionStartId="11.[151,250,1948,1970]" captionTargetBox="[246,1343,157,1924]" captionTargetId="figure@11.[246,1344,157,1924]" captionTargetPageId="11" captionText="FIGURE 6. Rheotanytarsus alliciens sp. nov., male. A— holotype (inclusion); B— habitus; C— head with antenna; D – wing; E – H— hypopygium: E, F— dorsal aspect; G— superior volsella, digitus and median volsella; H— median volsella (magnified c. three times relative to hypopygium drawn)." httpUri="https://zenodo.org/record/249105/files/figure.png" pageId="12" pageNumber="581">Fig. 6</figureCitation>
|
||
F, G).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |