189 lines
29 KiB
XML
189 lines
29 KiB
XML
<document id="9071AF380CE141550293D715BC7FBD68" ID-DOI="10.5281/zenodo.278442" ID-GBIF-Dataset="6b9505e0-ac6e-4baa-9066-6f729d38e9e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="278442" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460341493798" checkinUser="plazi" docAuthor="Hershler, Robert, Liu, Hsiu-Ping & Landye, Jerry" docDate="2011" docId="03E687C5BF798301FF2E139DFAF71B58" docLanguage="en" docName="zt03001p032.pdf" docOrigin="Zootaxa 3001" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tryonia molinae Hershler, Liu & Landye, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="16" masterDocId="FFDFFFBDBF77830EFFB9171AFFA4194F" masterDocTitle="New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot" masterLastPageNumber="32" masterPageNumber="1" pageNumber="15" updateTime="1698261240372" updateUser="plazi">
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<mods:titleInfo id="23F4B958963615740AF8B9630FC20DAE">
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<mods:title id="F1C8F8221BF1A0CF9F7AF7D2687CE5C7">New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot</mods:title>
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</mods:titleInfo>
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<mods:name id="7ADDF81C47262E0560097454880A7AA0" type="personal">
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<mods:namePart id="E1C68A7669D27FB6BDF72906E70101B4">Hershler, Robert</mods:namePart>
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<mods:name id="C12374DDE784466A94633A1C03D277C0" type="personal">
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<mods:roleTerm id="CF1B55282E3AD1271A54CE8C43FAAFCF">Author</mods:roleTerm>
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<mods:namePart id="FBA5A8C561EBF538335D9516C94915E4">Liu, Hsiu-Ping</mods:namePart>
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</mods:name>
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<mods:name id="7B911D86DCE5552CF09DC590824EA992" type="personal">
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<mods:roleTerm id="445DCCF7FA2E8BD9F7724F0B50917E3E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6B2FA85C52DA95E32981CB6C6AD17CE6">Landye, Jerry</mods:namePart>
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<mods:typeOfResource id="C47CF96E883EB46C7C53F18B01032C89">text</mods:typeOfResource>
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<mods:title id="138D5AC3F2CDCD72806AEFFD689D5A3D">Zootaxa</mods:title>
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<mods:part id="7B518D41C63A272E907AD3D8E2C71244">
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<mods:date id="DFB5719B69008C8D6CF867B2A949FF86">2011</mods:date>
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<mods:detail id="16BE73E43019650095F103A25CE710CB" type="volume">
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<mods:number id="A620A48F42F1EEF138CFAD3711484AA4">3001</mods:number>
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<mods:classification id="2E5AE53A6FD7FDDDA38E06F8A20A5D4D">journal article</mods:classification>
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<mods:identifier id="AFAE8D8E5C4966E6B0182F60AE42C67E" type="DOI">10.5281/zenodo.278442</mods:identifier>
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<mods:identifier id="852B1BD403FC7D3F8A2F116FB42789E6" type="GBIF-Dataset">6b9505e0-ac6e-4baa-9066-6f729d38e9e2</mods:identifier>
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<mods:identifier id="B862F30FEF221EA99A4259CDAB727B23" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="929CAFEE2EE4B04CD6268E7BD0C1D626" type="Zenodo-Dep">278442</mods:identifier>
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<treatment id="03E687C5BF798301FF2E139DFAF71B58" ID-DOI="http://doi.org/10.5281/zenodo.5629197" ID-GBIF-Taxon="119406530" ID-Zenodo-Dep="5629197" LSID="urn:lsid:plazi:treatment:03E687C5BF798301FF2E139DFAF71B58" httpUri="http://treatment.plazi.org/id/03E687C5BF798301FF2E139DFAF71B58" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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<subSubSection id="C3556558BF798300FF2E139DFF5B1C1D" pageId="14" pageNumber="15" type="nomenclature">
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<paragraph id="8BF036D3BF798300FF2E139DFE601DEE" blockId="14.[151,479,1159,1218]" box="[151,452,1159,1185]" pageId="14" pageNumber="15">
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<heading id="D0B881BFBF798300FF2E139DFE601DEE" bold="true" box="[151,452,1159,1185]" fontSize="11" level="1" pageId="14" pageNumber="15" reason="1">
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<emphasis id="B93BEAC1BF798300FF2E139DFE601DEE" bold="true" box="[151,452,1159,1185]" pageId="14" pageNumber="15">
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<taxonomicName id="4C4F4D50BF798300FF2E139DFEFB1DEE" box="[151,351,1159,1185]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="molinae" status="sp. nov.">
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<emphasis id="B93BEAC1BF798300FF2E139DFEFB1DEE" bold="true" box="[151,351,1159,1185]" italics="true" pageId="14" pageNumber="15">Tryonia molinae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20857BABF798300FEDF139DFE601DEE" box="[358,452,1159,1185]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF036D3BF798300FF2E13B0FE7B1D8D" blockId="14.[151,479,1159,1218]" box="[151,479,1194,1218]" pageId="14" pageNumber="15">
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(
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<figureCitation id="13742A56BF798300FF2613B0FF411D8D" box="[159,229,1194,1218]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="14" pageNumber="15">Figs 5</figureCitation>
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E, 8A–C, G–H, K–M)
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</paragraph>
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<paragraph id="8BF036D3BF798300FF2E13E9FF5B1C1D" blockId="14.[151,1436,1266,2010]" pageId="14" pageNumber="15">
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<emphasis id="B93BEAC1BF798300FF2E13E9FF7E1C44" bold="true" box="[151,218,1267,1291]" pageId="14" pageNumber="15">
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<typeStatus id="54F48871BF798300FF2E13E9FF7E1C44" box="[151,218,1267,1291]" pageId="14" pageNumber="15">Types</typeStatus>
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</emphasis>
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.
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<typeStatus id="54F48871BF798300FF5413E8FEFC1C45" box="[237,344,1266,1290]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
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, USNM 1139278, unnamed spring in northern part of El Molino, ca.
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<quantity id="4CB79B36BF798300FBC213E8FB601C45" box="[1147,1220,1266,1290]" metricMagnitude="2" metricUnit="m" metricValue="1.25" pageId="14" pageNumber="15" unit="m" value="125.0">125 m</quantity>
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east of Chihuahua Highway 15, Chihuahua,
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<geoCoordinate id="EE7B5014BF798300FE07120CFDEF1C61" box="[446,587,1302,1326]" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="15" value="29.203056">29º12'11" N</geoCoordinate>
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,
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<geoCoordinate id="EE7B5014BF798300FDE0120CFD5D1C61" box="[601,761,1302,1326]" direction="west" orientation="longitude" pageId="14" pageNumber="15" precision="15" value="-107.40667">107º24'24" W</geoCoordinate>
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, leg. J.J.L.,
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<date id="FFF11013BF798300FC3F120CFBA71C61" box="[902,1027,1302,1326]" pageId="14" pageNumber="15" value="2010-04-26">26/iv/2010</date>
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.
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<typeStatus id="54F48871BF798300FBA9120DFBD91C60" box="[1040,1149,1303,1327]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
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(from same lot), USNM 1153674.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF798300FF7E1245FCFE1C39" box="[199,858,1374,1399]" pageId="14" pageNumber="15" type="etymology">
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<paragraph id="8BF036D3BF798300FF7E1245FCFE1C39" blockId="14.[151,1436,1266,2010]" box="[199,858,1374,1399]" pageId="14" pageNumber="15">
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<emphasis id="B93BEAC1BF798300FF7E1245FEE21C38" bold="true" box="[199,326,1375,1399]" pageId="14" pageNumber="15">Etymology</emphasis>
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. A geographical epithet referring to El Molino.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF798300FF7E1298FEF91CAD" pageId="14" pageNumber="15" type="diagnosis">
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<paragraph id="8BF036D3BF798300FF7E1298FEF91CAD" blockId="14.[151,1436,1266,2010]" pageId="14" pageNumber="15">
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<emphasis id="B93BEAC1BF798300FF7E1298FE9E1CD4" bold="true" box="[199,314,1410,1435]" pageId="14" pageNumber="15">Diagnosis</emphasis>
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. Shell medium-sized, elongate-conic; penis having two distal and one basal papillae on the inner edge. Differentiated from closely similar
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<taxonomicName id="4C4F4D50BF798300FDD012B2FD6A1CF1" box="[617,718,1446,1471]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="15" phylum="Mollusca" rank="species" species="taylori">
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<emphasis id="B93BEAC1BF798300FDD012B2FD6A1CF1" box="[617,718,1446,1471]" italics="true" pageId="14" pageNumber="15">T. taylori</emphasis>
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</taxonomicName>
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(described below) by having a single basal papilla on the inner edge of the penis.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF798301FF7E12F4FE31182B" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
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<paragraph id="8BF036D3BF798300FF7E12F4FC451FD9" blockId="14.[151,1436,1266,2010]" pageId="14" pageNumber="15">
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<emphasis id="B93BEAC1BF798300FF7E12F4FEF51F48" bold="true" box="[199,337,1518,1543]" pageId="14" pageNumber="15">Description</emphasis>
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. Shell (
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<figureCitation id="13742A56BF798300FE0D12F4FE5E1F49" box="[436,506,1518,1542]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
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A–C) up to
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<quantity id="4CB79B36BF798300FD2A12F4FD4B1F49" box="[659,751,1518,1542]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="14" pageNumber="15" unit="mm" value="3.9">3.9 mm</quantity>
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tall, large females having 4.75–6.25 whorls, spire height 133–167% width of shell, male shells smaller than those of females. Teleoconch whorls weak to medium convex, evenly rounded, with impressed sutures. Aperture weakly angled and sometimes slightly thickened adapically, parietal lip complete, sometimes thickened, adnate, umbilicus narrow. Outer lip orthocoline or weakly prosocline. Sculpture of strong growth lines and weak spiral threads. Periostracum tan.
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</paragraph>
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<paragraph id="8BF036D3BF798300FF7E11B8FA881E4D" blockId="14.[151,1436,1266,2010]" pageId="14" pageNumber="15">
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<emphasis id="B93BEAC1BF798300FF7E11B8FE6D1FF4" bold="true" box="[199,457,1698,1723]" pageId="14" pageNumber="15">Shell measurements (</emphasis>
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mean in parentheses): height
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<quantity id="4CB79B36BF798300FCA011B8FC1A1FF5" box="[793,958,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="3.38" metricValueMax="3.92" metricValueMin="2.84" pageId="14" pageNumber="15" unit="mm" value="3.38" valueMax="3.92" valueMin="2.84">2.84–3.92 mm</quantity>
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(3.35), width
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<quantity id="4CB79B36BF798300FBDA11B8FAAC1FF5" box="[1123,1288,1698,1722]" metricMagnitude="-3" metricUnit="m" metricValue="1.56" metricValueMax="1.77" metricValueMin="1.35" pageId="14" pageNumber="15" unit="mm" value="1.56" valueMax="1.77" valueMin="1.35">1.35–1.77 mm</quantity>
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(1.54), body whorl height
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<quantity id="4CB79B36BF798300FE9411DCFE741F91" box="[301,464,1734,1758]" metricMagnitude="-3" metricUnit="m" metricValue="1.87" metricValueMax="2.04" metricValueMin="1.7" pageId="14" pageNumber="15" unit="mm" value="1.87" valueMax="2.04" valueMin="1.7">1.70–2.04 mm</quantity>
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(1.85), body whorl width
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<quantity id="4CB79B36BF798300FD4011DCFC391F91" box="[761,925,1734,1758]" metricMagnitude="-3" metricUnit="m" metricValue="1.44" metricValueMax="1.63" metricValueMin="1.25" pageId="14" pageNumber="15" unit="mm" value="1.44" valueMax="1.63" valueMin="1.25">1.25–1.63 mm</quantity>
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(1.44), aperture height
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<quantity id="4CB79B36BF798300FB1E11DCFAEE1F91" box="[1191,1354,1734,1758]" metricMagnitude="-3" metricUnit="m" metricValue="1.075" metricValueMax="1.19" metricValueMin="0.96" pageId="14" pageNumber="15" unit="mm" value="1.075" valueMax="1.19" valueMin="0.96">0.96–1.19 mm</quantity>
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(1.06), aperture width
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<quantity id="4CB79B36BF798300FEF911F0FE471E4D" box="[320,483,1770,1794]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="9.6" metricValueMin="7.9" pageId="14" pageNumber="15" unit="mm" value="0.875" valueMax="0.96" valueMin="0.79">0.79–0.96 mm</quantity>
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(0.87), total number of whorls 4.75–6.25 (5.73) (USNM 1153674,
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<emphasis id="B93BEAC1BF798300FB6D11F6FB461E4C" box="[1236,1250,1772,1795]" italics="true" pageId="14" pageNumber="15">n</emphasis>
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= 10).
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</paragraph>
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<paragraph id="8BF036D3BF798300FF7E1015FC751E05" blockId="14.[151,1436,1266,2010]" pageId="14" pageNumber="15">
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<emphasis id="B93BEAC1BF798300FF7E1015FDA71E68" bold="true" box="[199,515,1806,1831]" pageId="14" pageNumber="15">
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Measurements of
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<typeStatus id="54F48871BF798300FE251015FDA71E68" box="[412,515,1807,1831]" pageId="14" pageNumber="15" type="holotype">holotype</typeStatus>
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</emphasis>
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: height
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<quantity id="4CB79B36BF798300FDD91014FD631E69" box="[608,711,1806,1830]" metricMagnitude="-3" metricUnit="m" metricValue="3.26" pageId="14" pageNumber="15" unit="mm" value="3.26">3.26 mm</quantity>
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, width
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<quantity id="4CB79B36BF798300FCA01014FC241E69" box="[793,896,1806,1830]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="14" pageNumber="15" unit="mm" value="1.55">1.55 mm</quantity>
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, body whorl height
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<quantity id="4CB79B36BF798300FBD81014FB6D1E69" box="[1121,1225,1806,1830]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="14" pageNumber="15" unit="mm" value="1.9">1.90 mm</quantity>
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, body whorl width
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<quantity id="4CB79B36BF798300FF2E1028FF591E05" box="[151,253,1842,1866]" metricMagnitude="-3" metricUnit="m" metricValue="1.48" pageId="14" pageNumber="15" unit="mm" value="1.48">1.48 mm</quantity>
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, aperture height
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<quantity id="4CB79B36BF798300FE011028FDBA1E05" box="[440,542,1842,1866]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="14" pageNumber="15" unit="mm" value="1.05">1.05 mm</quantity>
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, aperture width
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<quantity id="4CB79B36BF798300FD6A1028FC9D1E05" box="[723,825,1842,1866]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="14" pageNumber="15" unit="mm" value="0.9">0.90 mm</quantity>
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, 5.75 whorls.
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</paragraph>
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<paragraph id="8BF036D3BF798301FF7E104DFD47199B" blockId="14.[151,1436,1266,2010]" lastBlockId="15.[151,1436,151,536]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
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Inner and outer sides of operculum smooth (
|
||
<figureCitation id="13742A56BF798300FD7D104CFCA31E21" box="[708,775,1878,1902]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
|
||
G–H). Radula (
|
||
<figureCitation id="13742A56BF798300FC01104CFC5F1E21" box="[952,1019,1878,1902]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
|
||
K-M): dorsal edge of central radular teeth concave, basal tongue V-shaped, median cusps elongate, distally pointed or jagged-edged, parallel-sided proximally, lateral cusps four–seven, basal cusps two–three (innermost larger) (
|
||
<figureCitation id="13742A56BF798300FB841084FB201EF9" box="[1085,1156,1950,1974]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="14" pageNumber="15">Fig. 8</figureCitation>
|
||
L). Lateral teeth having two–six cusps on inner and three–six cusps on outer side, length of outer wing 167–200% width of cutting edge, central cusp pointed or jagged-edged (
|
||
<figureCitation id="13742A56BF788301FDF0178DFD2F19E0" box="[585,651,151,175]" captionStart="FIGURE 8" captionStartId="16.[151,244,1854,1875]" captionTargetBox="[151,1436,151,1835]" captionTargetId="figure@16.[151,1436,149,1835]" captionTargetPageId="16" captionText="FIGURE 8. Tryonia molinae sp. nov. and Tryonia contrerasi sp. nov. A. Holotype, T. molinae, USNM 1139278. B – C. Shells, T. molinae, USNM 1153674 (C is a male). D. Holotype, T. contrerasi, USNM 874121. E – F. Shells, T. contrerasi, USNM 874128 (F is a male). G – H. Operculum (outer, inner sides), T. molinae, USNM 1153674. I – J. Operculum (outer, inner sides), T. contrerasi, USNM 1153670. K. Portion of radular ribbon, T. molinae, USNM 1153674. L. Central radular teeth, T. molinae, USNM 1153670. M. Lateral and inner marginal teeth, T. molinae, USNM 1153670. N. Portion of radular ribbon, T. contrerasi, USNM 1153670. Q. Central radular teeth, T. contrerasi, USNM 1153670. R. Lateral tooth, T. contrerasi, USNM 1153670. Scale bars A – F = 1.0 mm; G – J = 250 µm; K, N = 25 µm, L – M, O – P = 10 µm." httpUri="https://zenodo.org/record/278450/files/figure.png" pageId="15" pageNumber="16">Fig. 8</figureCitation>
|
||
M). Inner marginal teeth with 19–31 cusps, outer marginal teeth with 27–35 cusps. Radula data are from USNM 1153674.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF788301FF7E17C5FE31182B" blockId="15.[151,1436,151,536]" pageId="15" pageNumber="16">
|
||
Animal darkly pigmented. Penis (
|
||
<figureCitation id="13742A56BF788301FDF817C5FD2619B8" box="[577,642,223,247]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="15" pageNumber="16">Fig. 5</figureCitation>
|
||
E) having two distal and one basal papillae on the inner edge (30 of
|
||
<specimenCount id="9D49FD5ABF788301FA3917C5FEA81853" pageId="15" pageNumber="16" type="generic">30 specimens</specimenCount>
|
||
examined). Basal papilla often small, sometimes positioned ventrally near inner edge. Distal bulb of penis expanded laterally on inner side, lightly pigmented; stylet small. Penial duct nearly straight. Penial data are from USNM 1153674.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF788301FF7E1675FD5D18BB" pageId="15" pageNumber="16" type="distribution">
|
||
<paragraph id="8BF036D3BF788301FF7E1675FD5D18BB" blockId="15.[151,1436,151,536]" pageId="15" pageNumber="16">
|
||
<emphasis id="B93BEAC1BF788301FF7E1675FE5418C7" bold="true" box="[199,496,367,392]" pageId="15" pageNumber="16">Distribution and habitat.</emphasis>
|
||
<taxonomicName id="4C4F4D50BF788301FE4F166BFD0A18C7" box="[502,686,368,392]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="molinae">
|
||
<emphasis id="B93BEAC1BF788301FE4F166BFD0A18C7" box="[502,686,368,392]" italics="true" pageId="15" pageNumber="16">Tryonia molinae</emphasis>
|
||
</taxonomicName>
|
||
is endemic to a single locality in the Río Santa Maria basin (
|
||
<figureCitation id="13742A56BF788301FAEA1675FA3118C8" box="[1363,1429,367,391]" captionStart="FIGURE 1" captionStartId="2.[151,244,1795,1816]" captionTargetBox="[179,1402,157,1789]" captionTargetId="figure@2.[179,1423,125,1806]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of Tryonia species in the Chihuahuan Desert. 1. Tryonia hertleini, Las Palomas. 2. Tryonia peregrina sp. nov., Ojo Vareleño, northwest of Casas Grandes. 3. Tryonia peregrina sp. nov., Yepomera. 4. Tryonia zaragozae sp. nov., northeast of Ignacio Zaragoza. 5. Tryonia angostura sp. nov., T. ovata sp. nov., south of Galeana. 6. Tryonia molinae sp. nov., El Molino. 7. Tryonia taylori sp. nov., vicinity of Rancho Ojo Caliente. 8. Tryonia contrerasi sp. nov., vicinity of Rancho Nuevo. 9. Tryonia chuviscarae sp. nov., T. minckleyi sp. nov., Balneario de San Diego de Alcala. 10. Tryonia julimesensis sp. nov., south of Julimes. 11. Tryonia allendae sp. nov., southwest of Talamantes. 12. Tryonia metcalfi sp. nov., La Cienega, south of Candelaria. 13. Tryonia oasiensis sp. nov., Caroline Spring, Oasis Ranch. 14. Tryonia cheatumi, Phantom Lake Spring. 15. Tryonia circumstriata, Diamond Y Draw. 16. Tryonia seemani, Ojo de Agua del Obispo, Durango City. 17. Tryonia seemani, Nombre de Dios. 18. Tryonia seemani, Amado Nervo." httpUri="https://zenodo.org/record/278443/files/figure.png" pageId="15" pageNumber="16">Fig. 1</figureCitation>
|
||
, locality 6) that is locally known as “El Ojo.” The outflow of this thermal (28°C) spring is ponded by a small retention dam and used for recreation (
|
||
<figureCitation id="13742A56BF788301FDA616ADFDC01880" box="[543,612,439,463]" captionStart="FIGURE 6" captionStartId="11.[151,244,1710,1731]" captionTargetBox="[221,1371,156,1678]" captionTargetId="figure@11.[212,1381,149,1685]" captionTargetPageId="11" captionText="FIGURE 6. Photographs of habitats of species treated in this paper. A. Ojo Vareleño, Chihuahua, type locality of T. peregrina sp. nov. (photograph taken on 16 / ix / 1990). B. Main spring in Yepomera, Chihuahua, T. peregrina habitat (18 / ix / 1990). C. Springs northeast of Ignacio Zaragoza, Chihuaua, type locality of T. zaragozae sp. nov. (8 / xii / 1998). D. Western-most head spring, Ojos de Arrey, Chihuahua, type locality of T. angosturae sp. nov. and T. ovata sp. nov. (7 / xii / 1998). E. Spring in northern part of El Molino, Chihuahua, type locality of T. molinae sp. nov. (26 / iv / 2010). F. Ojo del Apache, Chihuahua, type locality of T. contrerasi sp. nov. (14 / xii / 1998). G. Ojo Carbonero, Chihuahua, T. contrerasi habitat (2 / iv / 1991). H. Outflow of Ojo Caliente, Chihuahua, type locality of T. taylori sp. nov. (7 / xii / 1998). I – J. El Ojo de Servin, Chihuahua, T. taylori habitat (27 - viii- 1971, 7 / xii / 1998, respectively). K. Seeps in La Cienega, Texas, type locality of T. metcalfi sp. nov. (6 / xii / 2001). L – M. Balneario de San Diego de Alcala, Chihuahua, type locality of T. chuviscarae sp. nov. and T. minckleyi sp. nov. (4 / xii / 2001). N. Springs southeast of Julimes, Chihuahua, type locality of T. julimesensis sp. nov. (5 / iv / 1991). O. El Ojito (outflow), southwest of Talamantes, Chihuahua, type locality of T. allendae sp. nov. (10 / xii / 1998). P. Caroline Spring, Oasis Ranch, Texas, type locality of T. oasiensis sp. nov. (30 / iii / 2009). Q. Spring in Nombre de Dios, Durango, T. seemani habitat (2 / ix / 1971). R. Spring in Amado Nervo, Durango, T. seemani habitat (7 / iv / 1991)." httpUri="https://zenodo.org/record/278448/files/figure.png" pageId="15" pageNumber="16">Fig. 6</figureCitation>
|
||
E). Snails were abundant on all substrates (including rocks, rootlets and algae) in the headspring and upper portion of the pool.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF788301FF7E151AFAF71B58" box="[199,1363,511,536]" pageId="15" pageNumber="16" type="discussion">
|
||
<paragraph id="8BF036D3BF788301FF7E151AFAF71B58" blockId="15.[151,1436,151,536]" box="[199,1363,511,536]" pageId="15" pageNumber="16">
|
||
<emphasis id="B93BEAC1BF788301FF7E151AFE901B57" bold="true" box="[199,308,512,536]" pageId="15" pageNumber="16">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C4F4D50BF788301FEFB151BFE5E1B57" box="[322,506,512,536]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="15" pageNumber="16" phylum="Mollusca" rank="species" species="molinae">
|
||
<emphasis id="B93BEAC1BF788301FEFB151BFE5E1B57" box="[322,506,512,536]" italics="true" pageId="15" pageNumber="16">Tryonia molinae</emphasis>
|
||
</taxonomicName>
|
||
was delineated as a well supported lineage in the Bayesian analysis (
|
||
<figureCitation id="13742A56BF788301FAB816E5FAE21B58" box="[1281,1350,511,535]" captionStart="FIGURE 2" captionStartId="5.[151,244,1938,1959]" captionTargetBox="[244,1345,828,1915]" captionTargetId="figure@5.[244,1346,828,1920]" captionTargetPageId="5" captionText="FIGURE 2. Bayesian tree based on the COI dataset. Posterior probabilities are provided when> 95 %. The names of the species treated herein are in bold face. Terminals and specimen codes are labeled as in Table 1. The two well supported clades composed of new species described herein are labeled A – B." httpUri="https://zenodo.org/record/278444/files/figure.png" pageId="15" pageNumber="16">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |