301 lines
47 KiB
XML
301 lines
47 KiB
XML
<document id="01ADFC764AC7C98B3286D71601F8BE92" ID-DOI="10.5281/zenodo.278442" ID-GBIF-Dataset="6b9505e0-ac6e-4baa-9066-6f729d38e9e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="278442" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460341493798" checkinUser="plazi" docAuthor="Hershler, Robert, Liu, Hsiu-Ping & Landye, Jerry" docDate="2011" docId="03E687C5BF66831CFF2E1595FA9D1A78" docLanguage="en" docName="zt03001p032.pdf" docOrigin="Zootaxa 3001" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tryonia taylori Hershler, Liu & Landye, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="FFDFFFBDBF77830EFFB9171AFFA4194F" masterDocTitle="New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot" masterLastPageNumber="32" masterPageNumber="1" pageNumber="18" updateTime="1698261240372" updateUser="plazi">
|
||
<mods:mods id="5F561901E024E1E923CDC16D300A9B35" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="028C7312C8241BAE57F4F67ED95C53BB">
|
||
<mods:title id="482BAC4F99C31D8CFA94E2EAD9EA3C84">New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="97F8D7F548C99CBC2CBDBCCA46101745" type="personal">
|
||
<mods:role id="FE663E8BA11A6175A1127AA1F282954B">
|
||
<mods:roleTerm id="479797C782B871BC5C985E1E00767110">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="75D78EE4F16AAC11822B9A1685AD20E7">Hershler, Robert</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="9F3FDE14BE2AEDF513E357780F06DABC" type="personal">
|
||
<mods:role id="57528DC86BCABC5502D461180BB7DF2B">
|
||
<mods:roleTerm id="CAB3ECF4F3704652BAE70A982672C875">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9C42A79338E184F495CA7ACF30F7E812">Liu, Hsiu-Ping</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="6446F8FF51BA2CB3452D5E6C8DF493DC" type="personal">
|
||
<mods:role id="6B8357FF3E071FFE51C82BC3F419A223">
|
||
<mods:roleTerm id="DD3E227F53DC141441E58CF6FCD53A19">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="34A3BF616AE423E0D76239EC301E8C6E">Landye, Jerry</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="1B1A64E9C8031EA91A6B87C52629FE0B">text</mods:typeOfResource>
|
||
<mods:relatedItem id="4666329AE4807ACB936737C7E712CD36" type="host">
|
||
<mods:titleInfo id="26A7959A6F5CF6CD2BCF9BE8E7BB3FA5">
|
||
<mods:title id="5DA5FEE052C21D7AAAE2FB6FE274F639">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="09E7CAD81AB513E2A8833E6551301969">
|
||
<mods:date id="E986DA36D56D52621DA64AB47E52AF9F">2011</mods:date>
|
||
<mods:detail id="39CEC416690B2C106F132A3BF85F9506" type="volume">
|
||
<mods:number id="01FEB9BC65875240EC74053EB41A3A8C">3001</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="28D32C7D8D0FD3E642ED38E24FD9881C" unit="page">
|
||
<mods:start id="4DAB06B3A83873124FB055B1BA1CFC53">1</mods:start>
|
||
<mods:end id="DF57D0B9BEA48CDF144C9E4DF4A96A28">32</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="54086B5178213A6D4BB2B2D0CCC58357">journal article</mods:classification>
|
||
<mods:identifier id="B6DA21063D48C3D54428717A2845B48A" type="DOI">10.5281/zenodo.278442</mods:identifier>
|
||
<mods:identifier id="F8ECB04DAA22F83A2FA1B9B1F68BB681" type="GBIF-Dataset">6b9505e0-ac6e-4baa-9066-6f729d38e9e2</mods:identifier>
|
||
<mods:identifier id="950B988A08758047113FAA7AEBB59DBB" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="1E61643CD294CAD9CBE829E2EB8B2068" type="Zenodo-Dep">278442</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03E687C5BF66831CFF2E1595FA9D1A78" ID-DOI="http://doi.org/10.5281/zenodo.5629203" ID-GBIF-Taxon="119406540" ID-Zenodo-Dep="5629203" LSID="urn:lsid:plazi:treatment:03E687C5BF66831CFF2E1595FA9D1A78" httpUri="http://treatment.plazi.org/id/03E687C5BF66831CFF2E1595FA9D1A78" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
|
||
<subSubSection id="C3556558BF66831FFF2E1595FAD21A79" pageId="17" pageNumber="18" type="nomenclature">
|
||
<paragraph id="8BF036D3BF66831FFF2E1595FE0B1BE6" blockId="17.[151,499,655,714]" box="[151,431,655,681]" pageId="17" pageNumber="18">
|
||
<heading id="D0B881BFBF66831FFF2E1595FE0B1BE6" bold="true" box="[151,431,655,681]" fontSize="11" level="1" pageId="17" pageNumber="18" reason="1">
|
||
<emphasis id="B93BEAC1BF66831FFF2E1595FE0B1BE6" bold="true" box="[151,431,655,681]" pageId="17" pageNumber="18">
|
||
<taxonomicName id="4C4F4D50BF66831FFF2E1595FEEF1BE6" box="[151,331,655,681]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="taylori" status="sp. nov.">
|
||
<emphasis id="B93BEAC1BF66831FFF2E1595FEEF1BE6" bold="true" box="[151,331,655,681]" italics="true" pageId="17" pageNumber="18">Tryonia taylori</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20857BABF66831FFEEB1595FE0B1BE6" box="[338,431,655,681]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF66831FFF2E15A8FE571B85" blockId="17.[151,499,655,714]" box="[151,499,690,714]" pageId="17" pageNumber="18">
|
||
(
|
||
<figureCitation id="13742A56BF66831FFF2615A8FF401B85" box="[159,228,690,714]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="17" pageNumber="18">Figs 5</figureCitation>
|
||
G–H, 9A–D, G–I, L–N)
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF66831FFF2E15E1FAD21A79" blockId="17.[151,1436,762,2010]" pageId="17" pageNumber="18">
|
||
<emphasis id="B93BEAC1BF66831FFF2E15E1FF7E1A5C" bold="true" box="[151,218,763,787]" pageId="17" pageNumber="18">
|
||
<typeStatus id="54F48871BF66831FFF2E15E1FF7E1A5C" box="[151,218,763,787]" pageId="17" pageNumber="18">Types</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<typeStatus id="54F48871BF66831FFF5515E0FEF31A5D" box="[236,343,762,786]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
|
||
, USNM 873254, ouflow of Ojo Caliente at Rancho Ojo Caliente, south of Highway 10, Chihuahua,
|
||
<geoCoordinate id="EE7B5014BF66831FFF741404FEC91A79" box="[205,365,798,822]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="29.902832">29º54'10.2" N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7B5014BF66831FFEC01404FD8F1A79" box="[377,555,798,822]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="-107.25753">107º15'27.1" W</geoCoordinate>
|
||
, leg. J.J.L.
|
||
<emphasis id="B93BEAC1BF66831FFD10143AFD7E1A79" box="[681,730,798,823]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
|
||
.,
|
||
<date id="FFF11013BF66831FFD561404FCDF1A79" box="[751,891,798,822]" pageId="17" pageNumber="18" value="1971-08-27">27/viii/1971</date>
|
||
.
|
||
<typeStatus id="54F48871BF66831FFC3C1405FC561A78" box="[901,1010,799,823]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
|
||
(from same lot), USNM 1153678.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF66831FFF7E1459FE9C1A31" pageId="17" pageNumber="18" type="etymology">
|
||
<paragraph id="8BF036D3BF66831FFF7E1459FE9C1A31" blockId="17.[151,1436,762,2010]" pageId="17" pageNumber="18">
|
||
<emphasis id="B93BEAC1BF66831FFF7E1459FEE21A14" bold="true" box="[199,326,835,859]" pageId="17" pageNumber="18">Etymology</emphasis>
|
||
. A patronym honoring the late Dwight W. Taylor for his pioneering taxonomic studies of
|
||
<taxonomicName id="4C4F4D50BF66831FFAFC145EFE901A31" authority="Taylor 1987" authorityName="Taylor" authorityYear="1987" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="17" pageNumber="18" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B93BEAC1BF66831FFAFC145EFA3E1A14" box="[1349,1434,836,859]" italics="true" pageId="17" pageNumber="18">Tryonia</emphasis>
|
||
(
|
||
<bibRefCitation id="EFDE4B22BF66831FFF27147CFE881A31" author="Taylor" box="[158,300,870,894]" pageId="17" pageNumber="18" refString="Taylor, D. W. (1987) Fresh-water molluscs from New Mexico and vicinity. New Mexico Bureau of Mines & Mineral Resources Bulletin, 116, 1 - 50." type="journal article" year="1987">Taylor 1987</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF66831FFF7E1490FEAE1D35" pageId="17" pageNumber="18" type="materials_examined">
|
||
<paragraph id="8BF036D3BF66831FFF7E1490FEAE1D35" blockId="17.[151,1436,762,2010]" pageId="17" pageNumber="18">
|
||
<emphasis id="B93BEAC1BF66831FFF7E1490FE3A1AEC" bold="true" box="[199,414,906,931]" pageId="17" pageNumber="18">Referred material</emphasis>
|
||
. CHIHUAHUA.
|
||
<collectionCode id="ED5EAE16BF66831FFDDD1490FD111AED" LSID="urn:lsid:biocol.org:col:34871" box="[612,693,906,930]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
|
||
873251, topotypes, leg. J.J.L.
|
||
<emphasis id="B93BEAC1BF66831FFBB51496FB991AEC" box="[1036,1085,907,931]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
|
||
.,
|
||
<date id="FFF11013BF66831FFBEB1490FB7B1AED" box="[1106,1247,906,930]" pageId="17" pageNumber="18" value="1971-08-27">27/viii/1971</date>
|
||
.
|
||
<collectionCode id="ED5EAE16BF66831FFB501490FA9E1AED" LSID="urn:lsid:biocol.org:col:34871" box="[1257,1338,906,930]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
|
||
874047, topotypes, leg. J.J.L. and P. Hines,
|
||
<date id="FFF11013BF66831FFD8B14B4FD0B1A89" box="[562,687,942,966]" pageId="17" pageNumber="18" value="1990-09-18">18/ix/1990</date>
|
||
.
|
||
<collectionCode id="ED5EAE16BF66831FFD0514B4FCA91A89" LSID="urn:lsid:biocol.org:col:34871" box="[700,781,942,966]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
|
||
1135493, topotypes, R.H. and J.J.L.,
|
||
<date id="FFF11013BF66831FFB7E14B4FA991A89" box="[1223,1341,942,966]" pageId="17" pageNumber="18" value="1998-12-07">7/xii/1998</date>
|
||
.
|
||
<collectionCode id="ED5EAE16BF66831FFAF214B4FA381A89" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,942,966]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
|
||
873283, El Ojo de Servin,
|
||
<quantity id="4CB79B36BF66831FFE6A14C8FD811AA5" box="[467,549,978,1002]" metricMagnitude="3" metricUnit="m" metricValue="5.6" pageId="17" pageNumber="18" unit="km" value="5.6">5.6 km</quantity>
|
||
west of Rancho Ojo Caliente,
|
||
<quantity id="4CB79B36BF66831FFC2B14C8FC401AA5" box="[914,996,978,1002]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="km" value="0.2">0.2 km</quantity>
|
||
north of Highway 10,
|
||
<geoCoordinate id="EE7B5014BF66831FFB4D14C8FA3C1AA5" box="[1268,1432,978,1002]" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="1" value="29.904778">29º54'17.2" N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE7B5014BF66831FFF2E14ECFEEC1D41" box="[151,328,1014,1038]" direction="west" orientation="longitude" pageId="17" pageNumber="18" precision="1" value="-107.307556">107º18'27.2" W</geoCoordinate>
|
||
, leg. J.J.L.,
|
||
<date id="FFF11013BF66831FFE7514ECFDED1D41" box="[460,585,1014,1038]" pageId="17" pageNumber="18" value="1973-04-14">14/iv/1973</date>
|
||
.
|
||
<collectionCode id="ED5EAE16BF66831FFDEA14ECFD001D41" LSID="urn:lsid:biocol.org:col:34871" box="[595,676,1014,1038]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
|
||
873252,
|
||
<collectionCode id="ED5EAE16BF66831FFCB514ECFCF91D41" LSID="urn:lsid:biocol.org:col:34871" box="[780,861,1014,1038]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
|
||
873297,
|
||
<emphasis id="B93BEAC1BF66831FFC7D14ECFC4B1D41" box="[964,1007,1014,1038]" italics="true" pageId="17" pageNumber="18">ibid</emphasis>
|
||
, leg. J.J.L.
|
||
<emphasis id="B93BEAC1BF66831FFBC914E2FB041D41" box="[1136,1184,1014,1039]" italics="true" pageId="17" pageNumber="18">et al</emphasis>
|
||
.,
|
||
<date id="FFF11013BF66831FFB0C14ECFAE51D41" box="[1205,1345,1014,1038]" pageId="17" pageNumber="18" value="1971-08-27">27/viii/1971</date>
|
||
.
|
||
<collectionCode id="ED5EAE16BF66831FFAF214ECFA381D41" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,1014,1038]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
|
||
874050,
|
||
<emphasis id="B93BEAC1BF66831FFF431301FE891D7C" box="[250,301,1051,1075]" italics="true" pageId="17" pageNumber="18">ibid.</emphasis>
|
||
, leg. J.J.L.,
|
||
<date id="FFF11013BF66831FFE011300FD911D7D" box="[440,565,1050,1074]" pageId="17" pageNumber="18" value="1990-09-18">18/ix/1990</date>
|
||
.
|
||
<collectionCode id="ED5EAE16BF66831FFDF91300FD351D7D" LSID="urn:lsid:biocol.org:col:34871" box="[576,657,1050,1074]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18">USNM</collectionCode>
|
||
892118,
|
||
<emphasis id="B93BEAC1BF66831FFD451301FC8B1D7C" box="[764,815,1051,1075]" italics="true" pageId="17" pageNumber="18">ibid.</emphasis>
|
||
, leg. R.H. and J.J.L.,
|
||
<date id="FFF11013BF66831FFB911300FB3A1D7D" box="[1064,1182,1050,1074]" pageId="17" pageNumber="18" value="1998-12-07">7/xii/1998</date>
|
||
.
|
||
<collectionCode id="ED5EAE16BF66831FFB131301FB541D7D" box="[1194,1264,1051,1074]" pageId="17" pageNumber="18">UTEP</collectionCode>
|
||
978,
|
||
<emphasis id="B93BEAC1BF66831FFA881301FAC01D7C" box="[1329,1380,1051,1075]" italics="true" pageId="17" pageNumber="18">ibid.</emphasis>
|
||
, leg. A. Metcalf and D.W. Taylor,
|
||
<date id="FFF11013BF66831FFE641324FDE81D19" box="[477,588,1086,1110]" pageId="17" pageNumber="18" value="1969-09-04">4/ix/1969</date>
|
||
. BellMNH 21172, BellMNH 21173,
|
||
<emphasis id="B93BEAC1BF66831FFC521324FBB91D19" box="[1003,1053,1086,1110]" italics="true" pageId="17" pageNumber="18">ibid.</emphasis>
|
||
, leg. D.W. Taylor and A. Metcalf,
|
||
<date id="FFF11013BF66831FFF2E1378FEA21D35" box="[151,262,1122,1146]" pageId="17" pageNumber="18" value="1969-09-04">4/ix/1969</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF66831FFF7E139CFE271DA9" pageId="17" pageNumber="18" type="diagnosis">
|
||
<paragraph id="8BF036D3BF66831FFF7E139CFE271DA9" blockId="17.[151,1436,762,2010]" pageId="17" pageNumber="18">
|
||
<emphasis id="B93BEAC1BF66831FFF7E139CFE9E1DD0" bold="true" box="[199,314,1158,1183]" pageId="17" pageNumber="18">Diagnosis</emphasis>
|
||
. Shell medium- to large-sized, conic or elongate-conic; penis having two distal papillae on inner edge and sometimes two–four basal papillae on inner edge and/or dorsal surface. Differentiated from closely similar
|
||
<taxonomicName id="4C4F4D50BF66831FFF0213CAFE8B1DA9" box="[187,303,1230,1255]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="17" pageNumber="18" phylum="Mollusca" rank="species" species="molinae">
|
||
<emphasis id="B93BEAC1BF66831FFF0213CAFE8B1DA9" box="[187,303,1230,1255]" italics="true" pageId="17" pageNumber="18">T. molinae</emphasis>
|
||
</taxonomicName>
|
||
above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF66831CFF7E13E8FC46182B" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
|
||
<paragraph id="8BF036D3BF66831FFF7E13E8FDE51CF1" blockId="17.[151,1436,762,2010]" pageId="17" pageNumber="18">
|
||
<emphasis id="B93BEAC1BF66831FFF7E13E8FEF51C44" bold="true" box="[199,337,1266,1291]" pageId="17" pageNumber="18">Description</emphasis>
|
||
. Shell (
|
||
<figureCitation id="13742A56BF66831FFE1E13E8FE4C1C45" box="[423,488,1266,1290]" captionStart="FIGURE 9" captionStartId="19.[151,244,1747,1768]" captionTargetBox="[151,1436,155,1723]" captionTargetId="figure@19.[151,1436,155,1723]" captionTargetPageId="19" captionText="FIGURE 9. Tryonia taylori sp. nov and T. metcalfi sp. nov. A. Holotype, T. taylori, USNM 873254. B. Male shell, T. taylori, USNM 1153678. C – D. Shells, T. taylori, USNM 873283 (D is a male). E. Holotype, T. metcalfi, USNM 1001931. F. Shell, T. metcalfi, USNM 1153672. G – I. Operculum (outer, inner, inner sides), T. taylori, USNM 1153678. J – K. Operculum (outer, inner sides), T. metcalfi, USNM 1153672. L. Portion of radular ribbon, T. taylori, USNM 1153678. M. Central radular teeth, T. taylori, USNM 1153678. N. Lateral tooth, T. taylori, USNM 1153678. O. Portion of radular ribbon, T. metcalfi, USNM 1153672. P. Central radular teeth, T. metcalfi, USNM 1153672. Q. Lateral teeth, T. metcalfi, USNM 1153672. Scale bars A – F = 1.0 mm; G – K = 250 µm; L, O = 25 µm, M – N, P – Q = 10 µm." httpUri="https://zenodo.org/record/278451/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
|
||
A–D) up to
|
||
<quantity id="4CB79B36BF66831FFDD713E8FD601C45" box="[622,708,1266,1290]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" pageId="17" pageNumber="18" unit="mm" value="6.7">6.7 mm</quantity>
|
||
tall, whorls ca. 5.50–8.75, spire height 135–220% width of shell, male shells smaller than those of females. Teleoconch whorls usually weak to medium convex, rarely high convex, evenly rounded or weakly shouldered with impressed or adpressed sutures. Aperture weakly or sharply angled adapically, parietal lip usually complete, usually adnate, rarely slightly disjunct, umbilicus narrow or absent. Outer lip sometimes slightly thickened adapically, orthocline or weakly prosocline. Sculpture of strong growth lines and weak spiral threads. Periostracum tan.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF66831FFF7E12D0FA881F65" blockId="17.[151,1436,762,2010]" pageId="17" pageNumber="18">
|
||
<emphasis id="B93BEAC1BF66831FFF7E12D0FE6D1CAC" bold="true" box="[199,457,1482,1507]" pageId="17" pageNumber="18">Shell measurements (</emphasis>
|
||
mean in parentheses): height
|
||
<quantity id="4CB79B36BF66831FFCA012D0FC1A1CAD" box="[793,958,1482,1506]" metricMagnitude="-3" metricUnit="m" metricValue="3.835" metricValueMax="4.25" metricValueMin="3.42" pageId="17" pageNumber="18" unit="mm" value="3.835" valueMax="4.25" valueMin="3.42">3.42–4.25 mm</quantity>
|
||
(3.74), width
|
||
<quantity id="4CB79B36BF66831FFBDA12D0FAAC1CAD" box="[1123,1288,1482,1506]" metricMagnitude="-3" metricUnit="m" metricValue="1.69" metricValueMax="1.84" metricValueMin="1.54" pageId="17" pageNumber="18" unit="mm" value="1.69" valueMax="1.84" valueMin="1.54">1.54–1.84 mm</quantity>
|
||
(1.67), body whorl height
|
||
<quantity id="4CB79B36BF66831FFE9412F4FE741F49" box="[301,464,1518,1542]" metricMagnitude="-3" metricUnit="m" metricValue="2.075" metricValueMax="2.22" metricValueMin="1.93" pageId="17" pageNumber="18" unit="mm" value="2.075" valueMax="2.22" valueMin="1.93">1.93–2.22 mm</quantity>
|
||
(2.09), body whorl width
|
||
<quantity id="4CB79B36BF66831FFD4012F4FC391F49" box="[761,925,1518,1542]" metricMagnitude="-3" metricUnit="m" metricValue="1.595" metricValueMax="1.73" metricValueMin="1.46" pageId="17" pageNumber="18" unit="mm" value="1.595" valueMax="1.73" valueMin="1.46">1.46–1.73 mm</quantity>
|
||
(1.59), aperture height
|
||
<quantity id="4CB79B36BF66831FFB1E12F4FAEE1F49" box="[1191,1354,1518,1542]" metricMagnitude="-3" metricUnit="m" metricValue="1.18" metricValueMax="1.33" metricValueMin="1.03" pageId="17" pageNumber="18" unit="mm" value="1.18" valueMax="1.33" valueMin="1.03">1.03–1.33 mm</quantity>
|
||
(1.22), aperture width
|
||
<quantity id="4CB79B36BF66831FFEF91108FE471F65" box="[320,483,1554,1578]" metricMagnitude="-4" metricUnit="m" metricValue="9.9" metricValueMax="10.9" metricValueMin="8.9" pageId="17" pageNumber="18" unit="mm" value="0.99" valueMax="1.09" valueMin="0.89">0.89–1.09 mm</quantity>
|
||
(1.00), total number of whorls 5.50–6.50 (6.00) (USNM 1153678,
|
||
<emphasis id="B93BEAC1BF66831FFB6D110EFB461F64" box="[1236,1250,1556,1579]" italics="true" pageId="17" pageNumber="18">n</emphasis>
|
||
= 15).
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF66831FFF7E112DFC671F3D" blockId="17.[151,1436,762,2010]" pageId="17" pageNumber="18">
|
||
<emphasis id="B93BEAC1BF66831FFF7E112DFDA71F00" bold="true" box="[199,515,1590,1615]" pageId="17" pageNumber="18">
|
||
Measurements of
|
||
<typeStatus id="54F48871BF66831FFE25112DFDA71F00" box="[412,515,1591,1615]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: height
|
||
<quantity id="4CB79B36BF66831FFDD9112CFD631F01" box="[608,711,1590,1614]" metricMagnitude="-3" metricUnit="m" metricValue="3.74" pageId="17" pageNumber="18" unit="mm" value="3.74">3.74 mm</quantity>
|
||
, width
|
||
<quantity id="4CB79B36BF66831FFCA0112CFC241F01" box="[793,896,1590,1614]" metricMagnitude="-3" metricUnit="m" metricValue="1.59" pageId="17" pageNumber="18" unit="mm" value="1.59">1.59 mm</quantity>
|
||
, body whorl height
|
||
<quantity id="4CB79B36BF66831FFBD8112CFB6D1F01" box="[1121,1225,1590,1614]" metricMagnitude="-3" metricUnit="m" metricValue="2.16" pageId="17" pageNumber="18" unit="mm" value="2.16">2.16 mm</quantity>
|
||
, body whorl width
|
||
<quantity id="4CB79B36BF66831FFF2E1140FF591F3D" box="[151,253,1626,1650]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="17" pageNumber="18" unit="mm" value="1.55">1.55 mm</quantity>
|
||
, aperture height
|
||
<quantity id="4CB79B36BF66831FFE011140FDBA1F3D" box="[440,542,1626,1650]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="17" pageNumber="18" unit="mm" value="1.24">1.24 mm</quantity>
|
||
, aperture width
|
||
<quantity id="4CB79B36BF66831FFD6A1140FC9D1F3D" box="[723,825,1626,1650]" metricMagnitude="-3" metricUnit="m" metricValue="1.01" pageId="17" pageNumber="18" unit="mm" value="1.01">1.01 mm</quantity>
|
||
, 6.0 whorls.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF66831FFF7E1164FDDC1E21" blockId="17.[151,1436,762,2010]" pageId="17" pageNumber="18">
|
||
Outer side of operculum smooth (
|
||
<figureCitation id="13742A56BF66831FFDE81164FD321FD9" box="[593,662,1662,1686]" captionStart="FIGURE 9" captionStartId="19.[151,244,1747,1768]" captionTargetBox="[151,1436,155,1723]" captionTargetId="figure@19.[151,1436,155,1723]" captionTargetPageId="19" captionText="FIGURE 9. Tryonia taylori sp. nov and T. metcalfi sp. nov. A. Holotype, T. taylori, USNM 873254. B. Male shell, T. taylori, USNM 1153678. C – D. Shells, T. taylori, USNM 873283 (D is a male). E. Holotype, T. metcalfi, USNM 1001931. F. Shell, T. metcalfi, USNM 1153672. G – I. Operculum (outer, inner, inner sides), T. taylori, USNM 1153678. J – K. Operculum (outer, inner sides), T. metcalfi, USNM 1153672. L. Portion of radular ribbon, T. taylori, USNM 1153678. M. Central radular teeth, T. taylori, USNM 1153678. N. Lateral tooth, T. taylori, USNM 1153678. O. Portion of radular ribbon, T. metcalfi, USNM 1153672. P. Central radular teeth, T. metcalfi, USNM 1153672. Q. Lateral teeth, T. metcalfi, USNM 1153672. Scale bars A – F = 1.0 mm; G – K = 250 µm; L, O = 25 µm, M – N, P – Q = 10 µm." httpUri="https://zenodo.org/record/278451/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
|
||
G) or with edges of last 0.5 whorl frilled; inner side of operculum nearly smooth (
|
||
<figureCitation id="13742A56BF66831FFEF111B8FE361FF5" box="[328,402,1698,1722]" captionStart="FIGURE 10" captionStartId="21.[151,244,1801,1822]" captionTargetBox="[151,1436,151,1774]" captionTargetId="figure@21.[151,1436,151,1775]" captionTargetPageId="21" captionText="FIGURE 10. Tryonia chuviscarae sp. nov. and Tryonia minckleyi sp. nov. A. Holotype, T. chuviscarae, USNM 873270. B. Male shell, T. chuviscarae, USNM 1153669. C. Male shell, T. chuviscarae, USNM 1001750. D. Holotype, T. minckleyi, USNM 1001751. E. Male shell, T. minckleyi, USNM 1153673. F – G. Operculum (outer, inner sides), T. chuviscarae, USNM 1001750. H – I. Operculum (outer, inner sides), T. minckleyi, USNM 1153673. J. Portion of radular ribbon, T. chuviscarae, USNM 0 0 1750. K. Central radular teeth, T. chuviscarae, USNM 1001750. L. Lateral and inner marginal teeth, T. chuviscarae, USNM 1001750. M. Portion of radular ribbon, T. minckleyi, USNM 1153673. N. Central radular teeth, T. minckleyi, USNM 1153673. O. Lateral tooth, T. minckleyi, USNM 1153673. Scale bars A – E = 1.0 mm, F – I = 100 µm, J – O = 10 µm." httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="17" pageNumber="18">Fig 10</figureCitation>
|
||
H) or with edges of muscle attachment scar thickened (
|
||
<figureCitation id="13742A56BF66831FFBBE11B8FBE81FF5" box="[1031,1100,1698,1722]" captionStart="FIGURE 9" captionStartId="19.[151,244,1747,1768]" captionTargetBox="[151,1436,155,1723]" captionTargetId="figure@19.[151,1436,155,1723]" captionTargetPageId="19" captionText="FIGURE 9. Tryonia taylori sp. nov and T. metcalfi sp. nov. A. Holotype, T. taylori, USNM 873254. B. Male shell, T. taylori, USNM 1153678. C – D. Shells, T. taylori, USNM 873283 (D is a male). E. Holotype, T. metcalfi, USNM 1001931. F. Shell, T. metcalfi, USNM 1153672. G – I. Operculum (outer, inner, inner sides), T. taylori, USNM 1153678. J – K. Operculum (outer, inner sides), T. metcalfi, USNM 1153672. L. Portion of radular ribbon, T. taylori, USNM 1153678. M. Central radular teeth, T. taylori, USNM 1153678. N. Lateral tooth, T. taylori, USNM 1153678. O. Portion of radular ribbon, T. metcalfi, USNM 1153672. P. Central radular teeth, T. metcalfi, USNM 1153672. Q. Lateral teeth, T. metcalfi, USNM 1153672. Scale bars A – F = 1.0 mm; G – K = 250 µm; L, O = 25 µm, M – N, P – Q = 10 µm." httpUri="https://zenodo.org/record/278451/files/figure.png" pageId="17" pageNumber="18">
|
||
Fig.
|
||
<date id="FFF11013BF66831FFB8511B8FBE81FF5" box="[1084,1100,1698,1722]" pageId="17" pageNumber="18">9</date>
|
||
</figureCitation>
|
||
I). Radula (
|
||
<figureCitation id="13742A56BF66831FFB7511B8FAB41FF5" box="[1228,1296,1698,1722]" captionStart="FIGURE 9" captionStartId="19.[151,244,1747,1768]" captionTargetBox="[151,1436,155,1723]" captionTargetId="figure@19.[151,1436,155,1723]" captionTargetPageId="19" captionText="FIGURE 9. Tryonia taylori sp. nov and T. metcalfi sp. nov. A. Holotype, T. taylori, USNM 873254. B. Male shell, T. taylori, USNM 1153678. C – D. Shells, T. taylori, USNM 873283 (D is a male). E. Holotype, T. metcalfi, USNM 1001931. F. Shell, T. metcalfi, USNM 1153672. G – I. Operculum (outer, inner, inner sides), T. taylori, USNM 1153678. J – K. Operculum (outer, inner sides), T. metcalfi, USNM 1153672. L. Portion of radular ribbon, T. taylori, USNM 1153678. M. Central radular teeth, T. taylori, USNM 1153678. N. Lateral tooth, T. taylori, USNM 1153678. O. Portion of radular ribbon, T. metcalfi, USNM 1153672. P. Central radular teeth, T. metcalfi, USNM 1153672. Q. Lateral teeth, T. metcalfi, USNM 1153672. Scale bars A – F = 1.0 mm; G – K = 250 µm; L, O = 25 µm, M – N, P – Q = 10 µm." httpUri="https://zenodo.org/record/278451/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
|
||
L-N): dorsal edge of central radular teeth concave, basal tongue rounded, median cusps elongate, distally pointed, lateral cusps four–seven, basal cusps one–three (innermost larger) (
|
||
<figureCitation id="13742A56BF66831FFD4411F0FCE41E4D" box="[765,832,1770,1794]" captionStart="FIGURE 9" captionStartId="19.[151,244,1747,1768]" captionTargetBox="[151,1436,155,1723]" captionTargetId="figure@19.[151,1436,155,1723]" captionTargetPageId="19" captionText="FIGURE 9. Tryonia taylori sp. nov and T. metcalfi sp. nov. A. Holotype, T. taylori, USNM 873254. B. Male shell, T. taylori, USNM 1153678. C – D. Shells, T. taylori, USNM 873283 (D is a male). E. Holotype, T. metcalfi, USNM 1001931. F. Shell, T. metcalfi, USNM 1153672. G – I. Operculum (outer, inner, inner sides), T. taylori, USNM 1153678. J – K. Operculum (outer, inner sides), T. metcalfi, USNM 1153672. L. Portion of radular ribbon, T. taylori, USNM 1153678. M. Central radular teeth, T. taylori, USNM 1153678. N. Lateral tooth, T. taylori, USNM 1153678. O. Portion of radular ribbon, T. metcalfi, USNM 1153672. P. Central radular teeth, T. metcalfi, USNM 1153672. Q. Lateral teeth, T. metcalfi, USNM 1153672. Scale bars A – F = 1.0 mm; G – K = 250 µm; L, O = 25 µm, M – N, P – Q = 10 µm." httpUri="https://zenodo.org/record/278451/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
|
||
M). Lateral teeth having two–five cusps on inner and three–seven cusps on outer side, length of outer wing 160–200% width of cutting edge, central cusp pointed (
|
||
<figureCitation id="13742A56BF66831FFAD71014FF011E05" captionStart="FIGURE 9" captionStartId="19.[151,244,1747,1768]" captionTargetBox="[151,1436,155,1723]" captionTargetId="figure@19.[151,1436,155,1723]" captionTargetPageId="19" captionText="FIGURE 9. Tryonia taylori sp. nov and T. metcalfi sp. nov. A. Holotype, T. taylori, USNM 873254. B. Male shell, T. taylori, USNM 1153678. C – D. Shells, T. taylori, USNM 873283 (D is a male). E. Holotype, T. metcalfi, USNM 1001931. F. Shell, T. metcalfi, USNM 1153672. G – I. Operculum (outer, inner, inner sides), T. taylori, USNM 1153678. J – K. Operculum (outer, inner sides), T. metcalfi, USNM 1153672. L. Portion of radular ribbon, T. taylori, USNM 1153678. M. Central radular teeth, T. taylori, USNM 1153678. N. Lateral tooth, T. taylori, USNM 1153678. O. Portion of radular ribbon, T. metcalfi, USNM 1153672. P. Central radular teeth, T. metcalfi, USNM 1153672. Q. Lateral teeth, T. metcalfi, USNM 1153672. Scale bars A – F = 1.0 mm; G – K = 250 µm; L, O = 25 µm, M – N, P – Q = 10 µm." httpUri="https://zenodo.org/record/278451/files/figure.png" pageId="17" pageNumber="18">Fig. 9</figureCitation>
|
||
N) or jagged-edged. Inner marginal teeth with 19–32 cusps, outer marginal teeth with 25–49 cusps. Radula data are from USNM 873283, USNM 1153678.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF66831CFF7E1060FC46182B" blockId="17.[151,1436,762,2010]" lastBlockId="18.[151,1436,151,823]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
|
||
Animal darkly pigmented. Penis (
|
||
<figureCitation id="13742A56BF66831FFDF81060FD271EDD" box="[577,643,1914,1938]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="17" pageNumber="18">Fig. 5</figureCitation>
|
||
G–H) usually having two distal papillae on inner edge (46 of
|
||
<specimenCount id="9D49FD5ABF66831FFA8F1060FF761EF9" pageId="17" pageNumber="18" type="generic">56 specimens</specimenCount>
|
||
from two samples), eight specimens differed in having three distal papillae and two differed in having a single distal papilla. Ojo Caliente specimens usually lacking basal penial papillae (25 of
|
||
<specimenCount id="9D49FD5ABF66831FFBED10D8FB541E95" box="[1108,1264,1986,2010]" pageId="17" pageNumber="18" type="generic">27 specimens</specimenCount>
|
||
), one specimen differed in having a basal papilla both on the inner and outer edges, one specimen differed in having two dorsally positioned basal papillae. Ojo de Servin specimens usually having two–three dorsally positioned basal penial papillae (27 of
|
||
<specimenCount id="9D49FD5ABF65831CFEB717C5FE0E19B8" box="[270,426,223,247]" pageId="18" pageNumber="19" type="generic">29 specimens</specimenCount>
|
||
) and sometimes also having a basal papilla on the inner edge (12 of
|
||
<specimenCount id="9D49FD5ABF65831CFB7917C5FAFB19B8" box="[1216,1375,223,247]" pageId="18" pageNumber="19" type="generic">29 specimens</specimenCount>
|
||
); one specimen had four dorsally positioned papillae and one specimen lacked basal papillae. Distal bulb of penis evenly rounded or expanded laterally on inner side, black; stylet small. Penial duct nearly straight to strongly undulating along most of length. Penial data are from USNM 873251, USNM 873252.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF65831CFF7E1675FC221B10" pageId="18" pageNumber="19" type="distribution">
|
||
<paragraph id="8BF036D3BF65831CFF7E1675FC221B10" blockId="18.[151,1436,151,823]" pageId="18" pageNumber="19">
|
||
<emphasis id="B93BEAC1BF65831CFF7E1675FE5218C7" bold="true" box="[199,502,367,392]" pageId="18" pageNumber="19">Distribution and habitat.</emphasis>
|
||
Narrowly endemic in the Río Santa Maria drainage (
|
||
<figureCitation id="13742A56BF65831CFBE41675FB0618C8" box="[1117,1186,367,391]" captionStart="FIGURE 1" captionStartId="2.[151,244,1795,1816]" captionTargetBox="[179,1402,157,1789]" captionTargetId="figure@2.[179,1423,125,1806]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of Tryonia species in the Chihuahuan Desert. 1. Tryonia hertleini, Las Palomas. 2. Tryonia peregrina sp. nov., Ojo Vareleño, northwest of Casas Grandes. 3. Tryonia peregrina sp. nov., Yepomera. 4. Tryonia zaragozae sp. nov., northeast of Ignacio Zaragoza. 5. Tryonia angostura sp. nov., T. ovata sp. nov., south of Galeana. 6. Tryonia molinae sp. nov., El Molino. 7. Tryonia taylori sp. nov., vicinity of Rancho Ojo Caliente. 8. Tryonia contrerasi sp. nov., vicinity of Rancho Nuevo. 9. Tryonia chuviscarae sp. nov., T. minckleyi sp. nov., Balneario de San Diego de Alcala. 10. Tryonia julimesensis sp. nov., south of Julimes. 11. Tryonia allendae sp. nov., southwest of Talamantes. 12. Tryonia metcalfi sp. nov., La Cienega, south of Candelaria. 13. Tryonia oasiensis sp. nov., Caroline Spring, Oasis Ranch. 14. Tryonia cheatumi, Phantom Lake Spring. 15. Tryonia circumstriata, Diamond Y Draw. 16. Tryonia seemani, Ojo de Agua del Obispo, Durango City. 17. Tryonia seemani, Nombre de Dios. 18. Tryonia seemani, Amado Nervo." httpUri="https://zenodo.org/record/278443/files/figure.png" pageId="18" pageNumber="19">Fig. 1</figureCitation>
|
||
, locality 7). The
|
||
<typeStatus id="54F48871BF65831CFAD5166AFA3818C7" box="[1388,1436,368,392]" pageId="18" pageNumber="19">type</typeStatus>
|
||
locality (Ojo Caliente) is a thermal spring (36–39°C) that is enclosed in a bath house (
|
||
<figureCitation id="13742A56BF65831CFBED168EFB3318E3" box="[1108,1175,404,428]" captionStart="FIGURE 6" captionStartId="11.[151,244,1710,1731]" captionTargetBox="[221,1371,156,1678]" captionTargetId="figure@11.[212,1381,149,1685]" captionTargetPageId="11" captionText="FIGURE 6. Photographs of habitats of species treated in this paper. A. Ojo Vareleño, Chihuahua, type locality of T. peregrina sp. nov. (photograph taken on 16 / ix / 1990). B. Main spring in Yepomera, Chihuahua, T. peregrina habitat (18 / ix / 1990). C. Springs northeast of Ignacio Zaragoza, Chihuaua, type locality of T. zaragozae sp. nov. (8 / xii / 1998). D. Western-most head spring, Ojos de Arrey, Chihuahua, type locality of T. angosturae sp. nov. and T. ovata sp. nov. (7 / xii / 1998). E. Spring in northern part of El Molino, Chihuahua, type locality of T. molinae sp. nov. (26 / iv / 2010). F. Ojo del Apache, Chihuahua, type locality of T. contrerasi sp. nov. (14 / xii / 1998). G. Ojo Carbonero, Chihuahua, T. contrerasi habitat (2 / iv / 1991). H. Outflow of Ojo Caliente, Chihuahua, type locality of T. taylori sp. nov. (7 / xii / 1998). I – J. El Ojo de Servin, Chihuahua, T. taylori habitat (27 - viii- 1971, 7 / xii / 1998, respectively). K. Seeps in La Cienega, Texas, type locality of T. metcalfi sp. nov. (6 / xii / 2001). L – M. Balneario de San Diego de Alcala, Chihuahua, type locality of T. chuviscarae sp. nov. and T. minckleyi sp. nov. (4 / xii / 2001). N. Springs southeast of Julimes, Chihuahua, type locality of T. julimesensis sp. nov. (5 / iv / 1991). O. El Ojito (outflow), southwest of Talamantes, Chihuahua, type locality of T. allendae sp. nov. (10 / xii / 1998). P. Caroline Spring, Oasis Ranch, Texas, type locality of T. oasiensis sp. nov. (30 / iii / 2009). Q. Spring in Nombre de Dios, Durango, T. seemani habitat (2 / ix / 1971). R. Spring in Amado Nervo, Durango, T. seemani habitat (7 / iv / 1991)." httpUri="https://zenodo.org/record/278448/files/figure.png" pageId="18" pageNumber="19">Fig. 6</figureCitation>
|
||
H).
|
||
<taxonomicName id="4C4F4D50BF65831CFB78168FFACE18E3" box="[1217,1386,404,428]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="taylori">
|
||
<emphasis id="B93BEAC1BF65831CFB78168FFACE18E3" box="[1217,1386,404,428]" italics="true" pageId="18" pageNumber="19">Tryonia taylori</emphasis>
|
||
</taxonomicName>
|
||
was found on mud, rocks and aquatic vegetation in the outflow 5–6 meters below the bath house. We did not discern a marked change in this spring from
|
||
<date id="FFF11013BF65831CFD9016C6FD0F18BB" box="[553,683,476,500]" pageId="18" pageNumber="19" value="1971" valueMax="1998">1971–1998</date>
|
||
. The second locality, El Ojo de Servin, is a warm spring complex (23–26°C) about
|
||
<quantity id="4CB79B36BF65831CFEE016E5FE351B58" box="[345,401,511,535]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="km" value="5.0">5 km</quantity>
|
||
west of Ojo Caliente.
|
||
<taxonomicName id="4C4F4D50BF65831CFD32151BFC901B57" box="[651,820,512,536]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="taylori">
|
||
<emphasis id="B93BEAC1BF65831CFD32151BFC901B57" box="[651,820,512,536]" italics="true" pageId="18" pageNumber="19">Tryonia taylori</emphasis>
|
||
</taxonomicName>
|
||
is extremely abundant on algae and
|
||
<taxonomicName id="4C4F4D50BF65831CFB73151AFAB61B57" box="[1226,1298,512,536]" class="Insecta" family="Noctuidae" genus="Chara" kingdom="Animalia" order="Lepidoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B93BEAC1BF65831CFB73151AFAB61B57" box="[1226,1298,512,536]" italics="true" pageId="18" pageNumber="19">Chara</emphasis>
|
||
</taxonomicName>
|
||
in the headspring area and somewhat less common in shallow portions of the spring runs on mud. This formerly lush cienega (as observed in 1971,
|
||
<figureCitation id="13742A56BF65831CFE34155DFE6B1B10" box="[397,463,583,607]" captionStart="FIGURE 6" captionStartId="11.[151,244,1710,1731]" captionTargetBox="[221,1371,156,1678]" captionTargetId="figure@11.[212,1381,149,1685]" captionTargetPageId="11" captionText="FIGURE 6. Photographs of habitats of species treated in this paper. A. Ojo Vareleño, Chihuahua, type locality of T. peregrina sp. nov. (photograph taken on 16 / ix / 1990). B. Main spring in Yepomera, Chihuahua, T. peregrina habitat (18 / ix / 1990). C. Springs northeast of Ignacio Zaragoza, Chihuaua, type locality of T. zaragozae sp. nov. (8 / xii / 1998). D. Western-most head spring, Ojos de Arrey, Chihuahua, type locality of T. angosturae sp. nov. and T. ovata sp. nov. (7 / xii / 1998). E. Spring in northern part of El Molino, Chihuahua, type locality of T. molinae sp. nov. (26 / iv / 2010). F. Ojo del Apache, Chihuahua, type locality of T. contrerasi sp. nov. (14 / xii / 1998). G. Ojo Carbonero, Chihuahua, T. contrerasi habitat (2 / iv / 1991). H. Outflow of Ojo Caliente, Chihuahua, type locality of T. taylori sp. nov. (7 / xii / 1998). I – J. El Ojo de Servin, Chihuahua, T. taylori habitat (27 - viii- 1971, 7 / xii / 1998, respectively). K. Seeps in La Cienega, Texas, type locality of T. metcalfi sp. nov. (6 / xii / 2001). L – M. Balneario de San Diego de Alcala, Chihuahua, type locality of T. chuviscarae sp. nov. and T. minckleyi sp. nov. (4 / xii / 2001). N. Springs southeast of Julimes, Chihuahua, type locality of T. julimesensis sp. nov. (5 / iv / 1991). O. El Ojito (outflow), southwest of Talamantes, Chihuahua, type locality of T. allendae sp. nov. (10 / xii / 1998). P. Caroline Spring, Oasis Ranch, Texas, type locality of T. oasiensis sp. nov. (30 / iii / 2009). Q. Spring in Nombre de Dios, Durango, T. seemani habitat (2 / ix / 1971). R. Spring in Amado Nervo, Durango, T. seemani habitat (7 / iv / 1991)." httpUri="https://zenodo.org/record/278448/files/figure.png" pageId="18" pageNumber="19">
|
||
Fig.
|
||
<date id="FFF11013BF65831CFE78155DFE6B1B10" box="[449,463,583,607]" pageId="18" pageNumber="19">6</date>
|
||
</figureCitation>
|
||
I) is now severely overgrazed (
|
||
<figureCitation id="13742A56BF65831CFC91155DFCC81B10" box="[808,876,583,607]" captionStart="FIGURE 6" captionStartId="11.[151,244,1710,1731]" captionTargetBox="[221,1371,156,1678]" captionTargetId="figure@11.[212,1381,149,1685]" captionTargetPageId="11" captionText="FIGURE 6. Photographs of habitats of species treated in this paper. A. Ojo Vareleño, Chihuahua, type locality of T. peregrina sp. nov. (photograph taken on 16 / ix / 1990). B. Main spring in Yepomera, Chihuahua, T. peregrina habitat (18 / ix / 1990). C. Springs northeast of Ignacio Zaragoza, Chihuaua, type locality of T. zaragozae sp. nov. (8 / xii / 1998). D. Western-most head spring, Ojos de Arrey, Chihuahua, type locality of T. angosturae sp. nov. and T. ovata sp. nov. (7 / xii / 1998). E. Spring in northern part of El Molino, Chihuahua, type locality of T. molinae sp. nov. (26 / iv / 2010). F. Ojo del Apache, Chihuahua, type locality of T. contrerasi sp. nov. (14 / xii / 1998). G. Ojo Carbonero, Chihuahua, T. contrerasi habitat (2 / iv / 1991). H. Outflow of Ojo Caliente, Chihuahua, type locality of T. taylori sp. nov. (7 / xii / 1998). I – J. El Ojo de Servin, Chihuahua, T. taylori habitat (27 - viii- 1971, 7 / xii / 1998, respectively). K. Seeps in La Cienega, Texas, type locality of T. metcalfi sp. nov. (6 / xii / 2001). L – M. Balneario de San Diego de Alcala, Chihuahua, type locality of T. chuviscarae sp. nov. and T. minckleyi sp. nov. (4 / xii / 2001). N. Springs southeast of Julimes, Chihuahua, type locality of T. julimesensis sp. nov. (5 / iv / 1991). O. El Ojito (outflow), southwest of Talamantes, Chihuahua, type locality of T. allendae sp. nov. (10 / xii / 1998). P. Caroline Spring, Oasis Ranch, Texas, type locality of T. oasiensis sp. nov. (30 / iii / 2009). Q. Spring in Nombre de Dios, Durango, T. seemani habitat (2 / ix / 1971). R. Spring in Amado Nervo, Durango, T. seemani habitat (7 / iv / 1991)." httpUri="https://zenodo.org/record/278448/files/figure.png" pageId="18" pageNumber="19">Fig. 6</figureCitation>
|
||
J).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF65831CFF7E1576FA9D1A78" pageId="18" pageNumber="19" type="discussion">
|
||
<paragraph id="8BF036D3BF65831CFF7E1576FA9D1A78" blockId="18.[151,1436,151,823]" pageId="18" pageNumber="19">
|
||
<emphasis id="B93BEAC1BF65831CFF7E1576FE901BCB" bold="true" box="[199,308,620,644]" pageId="18" pageNumber="19">Remarks</emphasis>
|
||
. The two populations of
|
||
<taxonomicName id="4C4F4D50BF65831CFDEC1577FD181BCB" box="[597,700,620,644]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="18" pageNumber="19" phylum="Mollusca" rank="species" species="taylori">
|
||
<emphasis id="B93BEAC1BF65831CFDEC1577FD181BCB" box="[597,700,620,644]" italics="true" pageId="18" pageNumber="19">T. taylori</emphasis>
|
||
</taxonomicName>
|
||
formed a strongly supported clade in the Bayesian tree (
|
||
<figureCitation id="13742A56BF65831CFAF11576FA2A1BCB" box="[1352,1422,620,644]" captionStart="FIGURE 2" captionStartId="5.[151,244,1938,1959]" captionTargetBox="[244,1345,828,1915]" captionTargetId="figure@5.[244,1346,828,1920]" captionTargetPageId="5" captionText="FIGURE 2. Bayesian tree based on the COI dataset. Posterior probabilities are provided when> 95 %. The names of the species treated herein are in bold face. Terminals and specimen codes are labeled as in Table 1. The two well supported clades composed of new species described herein are labeled A – B." httpUri="https://zenodo.org/record/278444/files/figure.png" pageId="18" pageNumber="19">Fig. 2</figureCitation>
|
||
). These populations differ in shell size (3.7 vs.
|
||
<quantity id="4CB79B36BF65831CFD2A1595FD4C1BE8" box="[659,744,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="18" pageNumber="19" unit="mm" value="5.8">5.8 mm</quantity>
|
||
mean shell height, t-test, t = -13.811, df = 18.6,
|
||
<emphasis id="B93BEAC1BF65831CFABB158BFAB71BE7" box="[1282,1299,657,680]" italics="true" pageId="18" pageNumber="19">P</emphasis>
|
||
<0.01, n=13 for the El Ojo de Servin sample), teleoconch whorl convexity, number of cusps on the outer marginal teeth, and the number and arrangement of basal papillae on the penis. However, they do not form reciprocally monophyletic clades in the Bayesian tree and their sequence divergence ranged from 0.3 to 1.7% (
|
||
<tableCitation id="C6CD0368BF65831CFBE915E6FB0C1A5B" box="[1104,1192,764,788]" captionStart="TABLE 2" captionStartId="18.[151,231,874,895]" captionTargetBox="[165,1425,927,1462]" captionTargetPageId="18" captionText="TABLE 2. Divergence of haplotypes detected in the Ojo Caliente (Try 7) and Ojo de Servin (Try 8) populations of T. taylori." httpUri="http://table.plazi.org/id/DF30665BBF65831CFF2E1470FA9F1A30" pageId="18" pageNumber="19" tableUuid="DF30665BBF65831CFF2E1470FA9F1A30">Table 2</tableCitation>
|
||
). We treat these morphologically distinctive populations as conspecific based on the evolutionary lineage concept used herein.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |