314 lines
44 KiB
XML
314 lines
44 KiB
XML
<document id="23E7100B34BB797B25A492FB897C52BC" ID-DOI="10.5281/zenodo.278442" ID-GBIF-Dataset="6b9505e0-ac6e-4baa-9066-6f729d38e9e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="278442" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460341493798" checkinUser="plazi" docAuthor="Hershler, Robert, Liu, Hsiu-Ping & Landye, Jerry" docDate="2011" docId="03E687C5BF638318FF2E13BEFAA01A30" docLanguage="en" docName="zt03001p032.pdf" docOrigin="Zootaxa 3001" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tryonia chuviscarae Hershler, Liu & Landye, 2011, sp. nov." docType="treatment" docVersion="8" lastPageNumber="23" masterDocId="FFDFFFBDBF77830EFFB9171AFFA4194F" masterDocTitle="New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot" masterLastPageNumber="32" masterPageNumber="1" pageNumber="21" updateTime="1698261240372" updateUser="plazi">
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<mods:titleInfo id="C033DBD728BA8FD4FA910DEEAD4B5DDF">
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<mods:title id="210C82A168107A019AA208DBD0D27569">New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot</mods:title>
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<mods:name id="C5C68C21D577437885219481B2080A1E" type="personal">
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<mods:namePart id="782FFBCB9295DA6924BD3CD1D05BA159">Hershler, Robert</mods:namePart>
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<mods:name id="5938AE21B979F5F3EE6106797448B547" type="personal">
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<mods:roleTerm id="552C2CB4366875A15F032AD0F4EC0E34">Author</mods:roleTerm>
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<mods:namePart id="BEAEB707A68478F20C4485B432592DD1">Liu, Hsiu-Ping</mods:namePart>
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</mods:name>
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<mods:name id="8E7604039CD6E822D5CEB6646679BF9B" type="personal">
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<mods:roleTerm id="C1704A4207BEF74AF05EA92C229F2FE2">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="1A5FB0DE61E4AF3BC377EDBD7E18BDCC">Landye, Jerry</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E64957DAE24C9C10414AC6F9D80F2665">text</mods:typeOfResource>
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<mods:title id="EC35D4534FA2245B25CB57971642C165">Zootaxa</mods:title>
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<mods:part id="35657923F731853EE7217C974A36C79C">
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<mods:date id="B47BEBAB5F1F6B4187C06A3648930C6D">2011</mods:date>
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<mods:detail id="896FD53713850628D757214DA60250A7" type="volume">
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<mods:number id="FF5564FE1D35023F3EA73795FF325F5F">3001</mods:number>
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<mods:classification id="4D36DA190D4E840BF520CDE1923747A3">journal article</mods:classification>
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<mods:identifier id="36E82C0FFC01736E8A28C260DBE4AC1E" type="DOI">10.5281/zenodo.278442</mods:identifier>
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<mods:identifier id="29FD44E9BDC329084CFA47EAC34BE9C9" type="GBIF-Dataset">6b9505e0-ac6e-4baa-9066-6f729d38e9e2</mods:identifier>
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<mods:identifier id="D54949942C5B008122DE751E64748CA9" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D6B7AA95077D5D821CB061AF9DE0EEC6" type="Zenodo-Dep">278442</mods:identifier>
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<treatment id="03E687C5BF638318FF2E13BEFAA01A30" ID-DOI="http://doi.org/10.5281/zenodo.5629207" ID-GBIF-Taxon="119406542" ID-Zenodo-Dep="5629207" LSID="urn:lsid:plazi:treatment:03E687C5BF638318FF2E13BEFAA01A30" httpUri="http://treatment.plazi.org/id/03E687C5BF638318FF2E13BEFAA01A30" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21">
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<subSubSection id="C3556558BF63831AFF2E13BEFB571C0B" pageId="20" pageNumber="21" type="nomenclature">
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<paragraph id="8BF036D3BF63831AFF2E13BEFE571DF2" blockId="20.[151,499,1187,1246]" box="[151,499,1187,1214]" pageId="20" pageNumber="21">
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<heading id="D0B881BFBF63831AFF2E13BEFE571DF2" bold="true" box="[151,499,1187,1214]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
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<emphasis id="B93BEAC1BF63831AFF2E13BEFE571DF2" bold="true" box="[151,499,1187,1214]" pageId="20" pageNumber="21">
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<taxonomicName id="4C4F4D50BF63831AFF2E13BEFE2A1DF1" box="[151,398,1188,1214]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="chuviscarae" status="sp. nov.">
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<emphasis id="B93BEAC1BF63831AFF2E13BEFE2A1DF1" bold="true" box="[151,398,1188,1214]" italics="true" pageId="20" pageNumber="21">Tryonia chuviscarae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20857BABF63831AFE2C13B9FE571DF2" box="[405,499,1187,1213]" pageId="20" pageNumber="21" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF036D3BF63831AFF2E13DCFE691D91" blockId="20.[151,499,1187,1246]" box="[151,461,1222,1246]" pageId="20" pageNumber="21">
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(
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<figureCitation id="13742A56BF63831AFF2613DCFF411D91" box="[159,229,1222,1246]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="20" pageNumber="21">Figs 5</figureCitation>
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J, 10A–C, F–G, J–L)
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</paragraph>
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<paragraph id="8BF036D3BF63831AFF2E1212FB571C0B" blockId="20.[151,1436,1287,2032]" pageId="20" pageNumber="21">
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<emphasis id="B93BEAC1BF63831AFF2E1212FF7E1C6F" bold="true" box="[151,218,1288,1312]" pageId="20" pageNumber="21">
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<typeStatus id="54F48871BF63831AFF2E1212FF7E1C6F" box="[151,218,1288,1312]" pageId="20" pageNumber="21">Types</typeStatus>
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</emphasis>
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.
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<typeStatus id="54F48871BF63831AFF52121DFEF21C50" box="[235,342,1287,1311]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
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, USNM 873270, Balneario de San Diego de Alcala, Chihuahua,
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<geoCoordinate id="EE7B5014BF63831AFB8E121DFB7D1C50" box="[1079,1241,1287,1311]" direction="north" orientation="latitude" pageId="20" pageNumber="21" precision="1" value="28.588223">28º35'17.6" N</geoCoordinate>
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,
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<geoCoordinate id="EE7B5014BF63831AFB5C121DFA3C1C50" box="[1253,1432,1287,1311]" direction="west" orientation="longitude" pageId="20" pageNumber="21" precision="1" value="-105.54745">105º32'50.8" W</geoCoordinate>
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, leg. J.J.L.
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<emphasis id="B93BEAC1BF63831AFEB31237FE9F1C0B" box="[266,315,1324,1348]" italics="true" pageId="20" pageNumber="21">et al</emphasis>
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.,
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<date id="FFF11013BF63831AFEE91236FE791C0B" box="[336,477,1324,1348]" pageId="20" pageNumber="21" value="1971-08-29">29/viii/1971</date>
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.
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<typeStatus id="54F48871BF63831AFE5E1237FDF01C0A" box="[487,596,1325,1349]" pageId="20" pageNumber="21" type="paratype">Paratypes</typeStatus>
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(from same lot), USNM 873294, USNM 905265, 1153669.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF63831AFF7E124AFAAF1C28" box="[199,1291,1359,1384]" pageId="20" pageNumber="21" type="etymology">
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<paragraph id="8BF036D3BF63831AFF7E124AFAAF1C28" blockId="20.[151,1436,1287,2032]" box="[199,1291,1359,1384]" pageId="20" pageNumber="21">
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<emphasis id="B93BEAC1BF63831AFF7E124AFEE21C27" bold="true" box="[199,326,1360,1384]" pageId="20" pageNumber="21">Etymology</emphasis>
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. Epithet refers to the Río Chuviscar, which is close to the
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<typeStatus id="54F48871BF63831AFC6B124AFBA61C28" box="[978,1026,1360,1383]" pageId="20" pageNumber="21">type</typeStatus>
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locality of this species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF63831AFF7E1269FCCB1CE0" pageId="20" pageNumber="21" type="materials_examined">
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<paragraph id="8BF036D3BF63831AFF7E1269FCCB1CE0" blockId="20.[151,1436,1287,2032]" pageId="20" pageNumber="21">
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<emphasis id="B93BEAC1BF63831AFF7E1269FE001CC3" bold="true" box="[199,420,1395,1420]" pageId="20" pageNumber="21">Referred material</emphasis>
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. CHIHUAHUA.
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<collectionCode id="ED5EAE16BF63831AFDC1126EFD6D1CC3" LSID="urn:lsid:biocol.org:col:34871" box="[632,713,1396,1420]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21">USNM</collectionCode>
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854941, topotypes, leg. R.H. and J.J.L.,
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<date id="FFF11013BF63831AFB7A126EFA9D1CC3" box="[1219,1337,1396,1420]" pageId="20" pageNumber="21" value="1998-12-09">9/xii/1998</date>
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.
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<collectionCode id="ED5EAE16BF63831AFAF2126EFA381CC3" LSID="urn:lsid:biocol.org:col:34871" box="[1355,1436,1396,1420]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21">USNM</collectionCode>
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1001750,
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<collectionCode id="ED5EAE16BF63831AFEBE128DFEFC1CE0" LSID="urn:lsid:biocol.org:col:34871" box="[263,344,1431,1455]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="20" pageNumber="21">USNM</collectionCode>
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1001756,
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<emphasis id="B93BEAC1BF63831AFE761282FDA61CFF" box="[463,514,1432,1456]" italics="true" pageId="20" pageNumber="21">ibid.</emphasis>
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, leg. R.H. and J.J.L.,
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<date id="FFF11013BF63831AFD4C128DFCC81CE0" box="[757,876,1431,1455]" pageId="20" pageNumber="21" value="2001-12-04">4/xii/2001</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF63831AFF7E12A1FBD61FE3" pageId="20" pageNumber="21" type="diagnosis">
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<paragraph id="8BF036D3BF63831AFF7E12A1FBD61FE3" blockId="20.[151,1436,1287,2032]" pageId="20" pageNumber="21">
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<emphasis id="B93BEAC1BF63831AFF7E12A1FE9E1C9B" bold="true" box="[199,314,1467,1492]" pageId="20" pageNumber="21">Diagnosis</emphasis>
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. Shell small, broadly- or ovate-conic; penis having two distal papillae on the inner edge and one–two basal papillae on the outer edge. Differentiated from sympatric
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<taxonomicName id="4C4F4D50BF63831AFC7B12FBFBEC1CB7" box="[962,1096,1504,1528]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="minckleyi">
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<emphasis id="B93BEAC1BF63831AFC7B12FBFBEC1CB7" box="[962,1096,1504,1528]" italics="true" pageId="20" pageNumber="21">T. minckleyi</emphasis>
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</taxonomicName>
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(described next) in its smaller shells (t-test for shell height, t = -10.699, df = 24.7,
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<emphasis id="B93BEAC1BF63831AFD54111FFD5A1F53" box="[749,766,1541,1564]" italics="true" pageId="20" pageNumber="21">P</emphasis>
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<0.01), thickened inner lip of shell, larger umbilicus, more convex dorsal edge of central radular teeth, V-shaped basal tongue of central radular teeth, narrower central radular teeth, and square-shaped face of lateral teeth; and from conchologically similar and closely related
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<taxonomicName id="4C4F4D50BF63831AFB4E1157FA381F2B" box="[1271,1436,1612,1636]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="20" pageNumber="21" phylum="Mollusca" rank="species" species="julimesensis">
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<emphasis id="B93BEAC1BF63831AFB4E1157FA381F2B" box="[1271,1436,1612,1636]" italics="true" pageId="20" pageNumber="21">T. julimesensis</emphasis>
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</taxonomicName>
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(described below) in its larger shells (t-test for shell height, t = -6.666, df = 23.8,
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<emphasis id="B93BEAC1BF63831AFB91116BFB9D1FC7" box="[1064,1081,1649,1672]" italics="true" pageId="20" pageNumber="21">P</emphasis>
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<0.01), in having basal papillae on the penis, and in having a larger number of cusps on the outer marginal radular teeth.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF638318FF7E11ADFC921B73" lastPageId="22" lastPageNumber="23" pageId="20" pageNumber="21" type="description">
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<paragraph id="8BF036D3BF63831AFF7E11ADFD971ECB" blockId="20.[151,1436,1287,2032]" pageId="20" pageNumber="21">
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<emphasis id="B93BEAC1BF63831AFF7E11ADFEF51F9F" bold="true" box="[199,337,1719,1744]" pageId="20" pageNumber="21">Description</emphasis>
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. Shell (
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<figureCitation id="13742A56BF63831AFE0B11ADFDA31F80" box="[434,519,1719,1743]" captionStart="FIGURE 10" captionStartId="21.[151,244,1801,1822]" captionTargetBox="[151,1436,151,1774]" captionTargetId="figure@21.[151,1436,151,1775]" captionTargetPageId="21" captionText="FIGURE 10. Tryonia chuviscarae sp. nov. and Tryonia minckleyi sp. nov. A. Holotype, T. chuviscarae, USNM 873270. B. Male shell, T. chuviscarae, USNM 1153669. C. Male shell, T. chuviscarae, USNM 1001750. D. Holotype, T. minckleyi, USNM 1001751. E. Male shell, T. minckleyi, USNM 1153673. F – G. Operculum (outer, inner sides), T. chuviscarae, USNM 1001750. H – I. Operculum (outer, inner sides), T. minckleyi, USNM 1153673. J. Portion of radular ribbon, T. chuviscarae, USNM 0 0 1750. K. Central radular teeth, T. chuviscarae, USNM 1001750. L. Lateral and inner marginal teeth, T. chuviscarae, USNM 1001750. M. Portion of radular ribbon, T. minckleyi, USNM 1153673. N. Central radular teeth, T. minckleyi, USNM 1153673. O. Lateral tooth, T. minckleyi, USNM 1153673. Scale bars A – E = 1.0 mm, F – I = 100 µm, J – O = 10 µm." httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="20" pageNumber="21">Fig. 10</figureCitation>
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A–C) up to
|
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<quantity id="4CB79B36BF63831AFD2511ADFD531F80" box="[668,759,1719,1743]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="mm" value="2.5">2.5 mm</quantity>
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tall, large females having 3.75–5.00 whorls, spire height 75–95% width of shell, male shells smaller than those of females. Teleoconch whorls high convex, frequently shouldered, sutures impressed, last 0.5 whorl sometimes loosened. Aperture usually rounded adapically, rarely weakly angled, inner lip complete, usually thickened, adnate or disjunct, umbilicus narrow or open. Outer lip thin, orthocline or prosocline, weakly sinuate. Sculpture of strong growth lines, weak spiral threads also sometimes present. Periostracum light tan or absent.
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</paragraph>
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<paragraph id="8BF036D3BF63831AFF7E1095FA891EBF" blockId="20.[151,1436,1287,2032]" pageId="20" pageNumber="21">
|
||
<emphasis id="B93BEAC1BF63831AFF7E1095FE6D1EE7" bold="true" box="[199,457,1935,1960]" pageId="20" pageNumber="21">Shell measurements (</emphasis>
|
||
mean in parentheses): height
|
||
<quantity id="4CB79B36BF63831AFCA01095FC1A1EE8" box="[793,958,1935,1959]" metricMagnitude="-3" metricUnit="m" metricValue="1.9849999999999999" metricValueMax="2.17" metricValueMin="1.8" pageId="20" pageNumber="21" unit="mm" value="1.9849999999999999" valueMax="2.17" valueMin="1.8">1.80–2.17 mm</quantity>
|
||
(1.98), width
|
||
<quantity id="4CB79B36BF63831AFBDA1095FAAC1EE8" box="[1123,1288,1935,1959]" metricMagnitude="-3" metricUnit="m" metricValue="1.3199999999999998" metricValueMax="1.43" metricValueMin="1.21" pageId="20" pageNumber="21" unit="mm" value="1.3199999999999998" valueMax="1.43" valueMin="1.21">1.21–1.43 mm</quantity>
|
||
(1.31), body whorl height
|
||
<quantity id="4CB79B36BF63831AFE9410AEFE741E83" box="[301,464,1972,1996]" metricMagnitude="-3" metricUnit="m" metricValue="1.485" metricValueMax="1.61" metricValueMin="1.36" pageId="20" pageNumber="21" unit="mm" value="1.485" valueMax="1.61" valueMin="1.36">1.36–1.61 mm</quantity>
|
||
(1.48), body whorl width
|
||
<quantity id="4CB79B36BF63831AFD4010AEFC391E83" box="[761,925,1972,1996]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.24" metricValueMin="1.06" pageId="20" pageNumber="21" unit="mm" value="1.15" valueMax="1.24" valueMin="1.06">1.06–1.24 mm</quantity>
|
||
(1.15), aperture height
|
||
<quantity id="4CB79B36BF63831AFB1E10AEFAEE1E83" box="[1191,1354,1972,1996]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" metricValueMax="9.9" metricValueMin="7.9" pageId="20" pageNumber="21" unit="mm" value="0.89" valueMax="0.99" valueMin="0.79">0.79–0.99 mm</quantity>
|
||
(0.88), aperture width
|
||
<quantity id="4CB79B36BF63831AFEF910C2FE471EBF" box="[320,483,2008,2032]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" metricValueMax="8.5" metricValueMin="7.1" pageId="20" pageNumber="21" unit="mm" value="0.78" valueMax="0.85" valueMin="0.71">0.71–0.85 mm</quantity>
|
||
(0.77), total number of whorls 3.75–4.25 (4.13) (USNM 1001750,
|
||
<emphasis id="B93BEAC1BF63831AFB6C10C3FB471EBF" box="[1237,1251,2009,2032]" italics="true" pageId="20" pageNumber="21">n</emphasis>
|
||
= 12).
|
||
</paragraph>
|
||
<caption id="DF30665BBF62831BFF2E1013FCAB1EAC" httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="21" pageNumber="22" targetBox="[151,1436,151,1774]" targetPageId="21">
|
||
<paragraph id="8BF036D3BF62831BFF2E1013FCAB1EAC" blockId="21.[151,1436,1801,2019]" pageId="21" pageNumber="22">
|
||
<emphasis id="B93BEAC1BF62831BFF2E1013FEB11E51" bold="true" box="[151,277,1801,1822]" pageId="21" pageNumber="22">FIGURE 10</emphasis>
|
||
.
|
||
<taxonomicName id="4C4F4D50BF62831BFE9A1010FE4E1E50" box="[291,490,1802,1823]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="chuviscarae" status="sp. nov.">
|
||
<emphasis id="B93BEAC1BF62831BFE9A1010FE4E1E50" box="[291,490,1802,1823]" italics="true" pageId="21" pageNumber="22">Tryonia chuviscarae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20857BABF62831BFE4A1013FDE51E50" box="[499,577,1801,1823]" pageId="21" pageNumber="22" rank="species">
|
||
<emphasis id="B93BEAC1BF62831BFE4A1013FD981E51" bold="true" box="[499,572,1801,1822]" pageId="21" pageNumber="22">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C4F4D50BF62831BFDCC1010FC801E50" box="[629,804,1802,1823]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="minckleyi" status="sp. nov.">
|
||
<emphasis id="B93BEAC1BF62831BFDCC1010FC801E50" box="[629,804,1802,1823]" italics="true" pageId="21" pageNumber="22">Tryonia minckleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20857BABF62831BFC951013FCDF1E50" box="[812,891,1801,1823]" pageId="21" pageNumber="22" rank="species">
|
||
<emphasis id="B93BEAC1BF62831BFC951013FCD21E51" bold="true" box="[812,886,1801,1822]" pageId="21" pageNumber="22">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
A. Holotype,
|
||
<taxonomicName id="4C4F4D50BF62831BFBB51010FB331E50" box="[1036,1175,1802,1823]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="chuviscarae">
|
||
<emphasis id="B93BEAC1BF62831BFBB51010FB331E50" box="[1036,1175,1802,1823]" italics="true" pageId="21" pageNumber="22">T. chuviscarae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 873270. B. Male shell,
|
||
<taxonomicName id="4C4F4D50BF62831BFF691036FEFF1E0E" box="[208,347,1836,1857]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="chuviscarae">
|
||
<emphasis id="B93BEAC1BF62831BFF691036FEFF1E0E" box="[208,347,1836,1857]" italics="true" pageId="21" pageNumber="22">T. chuviscarae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153669. C. Male shell,
|
||
<taxonomicName id="4C4F4D50BF62831BFD221036FC831E0E" box="[667,807,1836,1857]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="chuviscarae">
|
||
<emphasis id="B93BEAC1BF62831BFD221036FC831E0E" box="[667,807,1836,1857]" italics="true" pageId="21" pageNumber="22">T. chuviscarae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1001750. D. Holotype,
|
||
<taxonomicName id="4C4F4D50BF62831BFBE71036FB751E0E" box="[1118,1233,1836,1857]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="minckleyi">
|
||
<emphasis id="B93BEAC1BF62831BFBE71036FB751E0E" box="[1118,1233,1836,1857]" italics="true" pageId="21" pageNumber="22">T. minckleyi</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1001751. E. Male shell,
|
||
<taxonomicName id="4C4F4D50BF62831BFEB51056FE261E2E" box="[268,386,1868,1889]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="minckleyi">
|
||
<emphasis id="B93BEAC1BF62831BFEB51056FE261E2E" box="[268,386,1868,1889]" italics="true" pageId="21" pageNumber="22">T. minckleyi</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153673. F–G. Operculum (outer, inner sides),
|
||
<taxonomicName id="4C4F4D50BF62831BFC091056FB981E2E" box="[944,1084,1868,1889]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="chuviscarae">
|
||
<emphasis id="B93BEAC1BF62831BFC091056FB981E2E" box="[944,1084,1868,1889]" italics="true" pageId="21" pageNumber="22">T. chuviscarae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1001750. H–I. Operculum (outer, inner sides),
|
||
<taxonomicName id="4C4F4D50BF62831BFEE41076FE761ECE" box="[349,466,1900,1921]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="minckleyi">
|
||
<emphasis id="B93BEAC1BF62831BFEE41076FE761ECE" box="[349,466,1900,1921]" italics="true" pageId="21" pageNumber="22">T. minckleyi</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153673. J. Portion of radular ribbon,
|
||
<taxonomicName id="4C4F4D50BF62831BFC121076FB9C1ECE" box="[939,1080,1900,1921]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="chuviscarae">
|
||
<emphasis id="B93BEAC1BF62831BFC121076FB9C1ECE" box="[939,1080,1900,1921]" italics="true" pageId="21" pageNumber="22">T. chuviscarae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 0 0 1750. K. Central radular teeth,
|
||
<taxonomicName id="4C4F4D50BF62831BFF681096FEFE1EEE" box="[209,346,1932,1953]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="chuviscarae">
|
||
<emphasis id="B93BEAC1BF62831BFF681096FEFE1EEE" box="[209,346,1932,1953]" italics="true" pageId="21" pageNumber="22">T. chuviscarae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1001750. L. Lateral and inner marginal teeth,
|
||
<taxonomicName id="4C4F4D50BF62831BFCD01096FC501EEE" box="[873,1012,1932,1953]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="chuviscarae">
|
||
<emphasis id="B93BEAC1BF62831BFCD01096FC501EEE" box="[873,1012,1932,1953]" italics="true" pageId="21" pageNumber="22">T. chuviscarae</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1001750. M. Portion of radular ribbon,
|
||
<taxonomicName id="4C4F4D50BF62831BFF7010B6FE9B1E8E" box="[201,319,1964,1985]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="minckleyi">
|
||
<emphasis id="B93BEAC1BF62831BFF7010B6FE9B1E8E" box="[201,319,1964,1985]" italics="true" pageId="21" pageNumber="22">T. minckleyi</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153673. N. Central radular teeth,
|
||
<taxonomicName id="4C4F4D50BF62831BFD4D10B6FCCE1E8E" box="[756,874,1964,1985]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="minckleyi">
|
||
<emphasis id="B93BEAC1BF62831BFD4D10B6FCCE1E8E" box="[756,874,1964,1985]" italics="true" pageId="21" pageNumber="22">T. minckleyi</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153673. O. Lateral tooth,
|
||
<taxonomicName id="4C4F4D50BF62831BFB6A10B6FAEC1E8E" box="[1235,1352,1964,1985]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="21" pageNumber="22" phylum="Mollusca" rank="species" species="minckleyi">
|
||
<emphasis id="B93BEAC1BF62831BFB6A10B6FAEC1E8E" box="[1235,1352,1964,1985]" italics="true" pageId="21" pageNumber="22">T. minckleyi</emphasis>
|
||
</taxonomicName>
|
||
, USNM 1153673. Scale bars A–E = 1.0 mm, F–I = 100 µm, J–O = 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF036D3BF618318FF7E1782FC67199B" blockId="22.[151,1436,151,896]" pageId="22" pageNumber="23">
|
||
<emphasis id="B93BEAC1BF618318FF7E1782FDA719FF" bold="true" box="[199,515,151,176]" pageId="22" pageNumber="23">
|
||
Measurements of
|
||
<typeStatus id="54F48871BF618318FE251782FDA719FF" box="[412,515,152,176]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: height
|
||
<quantity id="4CB79B36BF618318FDD9178DFD6319E0" box="[608,711,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="2.28" pageId="22" pageNumber="23" unit="mm" value="2.28">2.28 mm</quantity>
|
||
, width
|
||
<quantity id="4CB79B36BF618318FCA0178DFC2419E0" box="[793,896,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" pageId="22" pageNumber="23" unit="mm" value="1.45">1.45 mm</quantity>
|
||
, body whorl height
|
||
<quantity id="4CB79B36BF618318FBD8178DFB6D19E0" box="[1121,1225,151,175]" metricMagnitude="-3" metricUnit="m" metricValue="1.66" pageId="22" pageNumber="23" unit="mm" value="1.66">1.66 mm</quantity>
|
||
, body whorl width
|
||
<quantity id="4CB79B36BF618318FF2E17A6FF59199B" box="[151,253,188,212]" metricMagnitude="-3" metricUnit="m" metricValue="1.31" pageId="22" pageNumber="23" unit="mm" value="1.31">1.31 mm</quantity>
|
||
, aperture height
|
||
<quantity id="4CB79B36BF618318FE0117A6FDBA199B" box="[440,542,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="9.8" pageId="22" pageNumber="23" unit="mm" value="0.98">0.98 mm</quantity>
|
||
, aperture width
|
||
<quantity id="4CB79B36BF618318FD6A17A6FC9D199B" box="[723,825,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="22" pageNumber="23" unit="mm" value="0.79">0.79 mm</quantity>
|
||
, 4.5 whorls.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF618318FF7E17C5FBBB18E3" blockId="22.[151,1436,151,896]" pageId="22" pageNumber="23">
|
||
Outer side of operculum smooth (
|
||
<figureCitation id="13742A56BF618318FDF817C5FD3719B8" box="[577,659,223,247]" captionStart="FIGURE 10" captionStartId="21.[151,244,1801,1822]" captionTargetBox="[151,1436,151,1774]" captionTargetId="figure@21.[151,1436,151,1775]" captionTargetPageId="21" captionText="FIGURE 10. Tryonia chuviscarae sp. nov. and Tryonia minckleyi sp. nov. A. Holotype, T. chuviscarae, USNM 873270. B. Male shell, T. chuviscarae, USNM 1153669. C. Male shell, T. chuviscarae, USNM 1001750. D. Holotype, T. minckleyi, USNM 1001751. E. Male shell, T. minckleyi, USNM 1153673. F – G. Operculum (outer, inner sides), T. chuviscarae, USNM 1001750. H – I. Operculum (outer, inner sides), T. minckleyi, USNM 1153673. J. Portion of radular ribbon, T. chuviscarae, USNM 0 0 1750. K. Central radular teeth, T. chuviscarae, USNM 1001750. L. Lateral and inner marginal teeth, T. chuviscarae, USNM 1001750. M. Portion of radular ribbon, T. minckleyi, USNM 1153673. N. Central radular teeth, T. minckleyi, USNM 1153673. O. Lateral tooth, T. minckleyi, USNM 1153673. Scale bars A – E = 1.0 mm, F – I = 100 µm, J – O = 10 µm." httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="22" pageNumber="23">Fig. 10</figureCitation>
|
||
F), portion of attachment scar margin slightly thickened on inner side (
|
||
<figureCitation id="13742A56BF618318FF26161EFF501853" box="[159,244,260,284]" captionStart="FIGURE 10" captionStartId="21.[151,244,1801,1822]" captionTargetBox="[151,1436,151,1774]" captionTargetId="figure@21.[151,1436,151,1775]" captionTargetPageId="21" captionText="FIGURE 10. Tryonia chuviscarae sp. nov. and Tryonia minckleyi sp. nov. A. Holotype, T. chuviscarae, USNM 873270. B. Male shell, T. chuviscarae, USNM 1153669. C. Male shell, T. chuviscarae, USNM 1001750. D. Holotype, T. minckleyi, USNM 1001751. E. Male shell, T. minckleyi, USNM 1153673. F – G. Operculum (outer, inner sides), T. chuviscarae, USNM 1001750. H – I. Operculum (outer, inner sides), T. minckleyi, USNM 1153673. J. Portion of radular ribbon, T. chuviscarae, USNM 0 0 1750. K. Central radular teeth, T. chuviscarae, USNM 1001750. L. Lateral and inner marginal teeth, T. chuviscarae, USNM 1001750. M. Portion of radular ribbon, T. minckleyi, USNM 1153673. N. Central radular teeth, T. minckleyi, USNM 1153673. O. Lateral tooth, T. minckleyi, USNM 1153673. Scale bars A – E = 1.0 mm, F – I = 100 µm, J – O = 10 µm." httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="22" pageNumber="23">Fig. 10</figureCitation>
|
||
G). Radula (
|
||
<figureCitation id="13742A56BF618318FE3C161EFE7D1853" box="[389,473,260,284]" captionStart="FIGURE 10" captionStartId="21.[151,244,1801,1822]" captionTargetBox="[151,1436,151,1774]" captionTargetId="figure@21.[151,1436,151,1775]" captionTargetPageId="21" captionText="FIGURE 10. Tryonia chuviscarae sp. nov. and Tryonia minckleyi sp. nov. A. Holotype, T. chuviscarae, USNM 873270. B. Male shell, T. chuviscarae, USNM 1153669. C. Male shell, T. chuviscarae, USNM 1001750. D. Holotype, T. minckleyi, USNM 1001751. E. Male shell, T. minckleyi, USNM 1153673. F – G. Operculum (outer, inner sides), T. chuviscarae, USNM 1001750. H – I. Operculum (outer, inner sides), T. minckleyi, USNM 1153673. J. Portion of radular ribbon, T. chuviscarae, USNM 0 0 1750. K. Central radular teeth, T. chuviscarae, USNM 1001750. L. Lateral and inner marginal teeth, T. chuviscarae, USNM 1001750. M. Portion of radular ribbon, T. minckleyi, USNM 1153673. N. Central radular teeth, T. minckleyi, USNM 1153673. O. Lateral tooth, T. minckleyi, USNM 1153673. Scale bars A – E = 1.0 mm, F – I = 100 µm, J – O = 10 µm." httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="22" pageNumber="23">Fig. 10</figureCitation>
|
||
J-L): dorsal edge of central radular teeth strongly concave, basal tongue V-shaped, median cusps distally pointed, parallel-sided proximally, lateral cusps four–six, basal cusps two–three (outermost larger) (
|
||
<figureCitation id="13742A56BF618318FF491656FEE6182B" box="[240,322,332,356]" captionStart="FIGURE 10" captionStartId="21.[151,244,1801,1822]" captionTargetBox="[151,1436,151,1774]" captionTargetId="figure@21.[151,1436,151,1775]" captionTargetPageId="21" captionText="FIGURE 10. Tryonia chuviscarae sp. nov. and Tryonia minckleyi sp. nov. A. Holotype, T. chuviscarae, USNM 873270. B. Male shell, T. chuviscarae, USNM 1153669. C. Male shell, T. chuviscarae, USNM 1001750. D. Holotype, T. minckleyi, USNM 1001751. E. Male shell, T. minckleyi, USNM 1153673. F – G. Operculum (outer, inner sides), T. chuviscarae, USNM 1001750. H – I. Operculum (outer, inner sides), T. minckleyi, USNM 1153673. J. Portion of radular ribbon, T. chuviscarae, USNM 0 0 1750. K. Central radular teeth, T. chuviscarae, USNM 1001750. L. Lateral and inner marginal teeth, T. chuviscarae, USNM 1001750. M. Portion of radular ribbon, T. minckleyi, USNM 1153673. N. Central radular teeth, T. minckleyi, USNM 1153673. O. Lateral tooth, T. minckleyi, USNM 1153673. Scale bars A – E = 1.0 mm, F – I = 100 µm, J – O = 10 µm." httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="22" pageNumber="23">Fig. 10</figureCitation>
|
||
K). Lateral teeth having two–four cusps on inner and three–six cusps on outer sides, length of outer wing about 140% width of cutting edge, central cusp pointed (
|
||
<figureCitation id="13742A56BF618318FCEA1675FC0018C8" box="[851,932,367,391]" captionStart="FIGURE 10" captionStartId="21.[151,244,1801,1822]" captionTargetBox="[151,1436,151,1774]" captionTargetId="figure@21.[151,1436,151,1775]" captionTargetPageId="21" captionText="FIGURE 10. Tryonia chuviscarae sp. nov. and Tryonia minckleyi sp. nov. A. Holotype, T. chuviscarae, USNM 873270. B. Male shell, T. chuviscarae, USNM 1153669. C. Male shell, T. chuviscarae, USNM 1001750. D. Holotype, T. minckleyi, USNM 1001751. E. Male shell, T. minckleyi, USNM 1153673. F – G. Operculum (outer, inner sides), T. chuviscarae, USNM 1001750. H – I. Operculum (outer, inner sides), T. minckleyi, USNM 1153673. J. Portion of radular ribbon, T. chuviscarae, USNM 0 0 1750. K. Central radular teeth, T. chuviscarae, USNM 1001750. L. Lateral and inner marginal teeth, T. chuviscarae, USNM 1001750. M. Portion of radular ribbon, T. minckleyi, USNM 1153673. N. Central radular teeth, T. minckleyi, USNM 1153673. O. Lateral tooth, T. minckleyi, USNM 1153673. Scale bars A – E = 1.0 mm, F – I = 100 µm, J – O = 10 µm." httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="22" pageNumber="23">Fig. 10</figureCitation>
|
||
L). Inner marginal teeth having 18–24 cusps, outer marginal teeth having 20–28 cusps. Radula data are from USNM 1001750.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF618318FF7E16ADFC921B73" blockId="22.[151,1436,151,896]" pageId="22" pageNumber="23">
|
||
Animal darkly pigmented. Penis (
|
||
<figureCitation id="13742A56BF618318FD8716ADFD241880" box="[574,640,439,463]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="22" pageNumber="23">Fig. 5</figureCitation>
|
||
J) having two distal papillae on the inner edge and one–two basal papillae on the outer edge (27 of
|
||
<specimenCount id="9D49FD5ABF618318FE6916C6FDC818BB" box="[464,620,476,500]" pageId="22" pageNumber="23" type="generic">30 specimens</specimenCount>
|
||
), two specimens differed in also having a basal papilla on the inner edge. Distal bulb of penis expanded laterally on inner side, black; stylet large. Penial duct straight or weakly undulating along most of length. Penial data are from USNM 1001756.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF618318FF7E155DFD3D1A5B" pageId="22" pageNumber="23" type="distribution">
|
||
<paragraph id="8BF036D3BF618318FF7E155DFD3D1A5B" blockId="22.[151,1436,151,896]" pageId="22" pageNumber="23">
|
||
<emphasis id="B93BEAC1BF618318FF7E155DFE551B2F" bold="true" box="[199,497,583,608]" pageId="22" pageNumber="23">Distribution and habitat.</emphasis>
|
||
Endemic to the
|
||
<typeStatus id="54F48871BF618318FD111552FD7C1B2F" box="[680,728,584,608]" pageId="22" pageNumber="23">type</typeStatus>
|
||
locality in the Río Conchos basin (
|
||
<figureCitation id="13742A56BF618318FBDD155DFB031B10" box="[1124,1191,583,607]" captionStart="FIGURE 1" captionStartId="2.[151,244,1795,1816]" captionTargetBox="[179,1402,157,1789]" captionTargetId="figure@2.[179,1423,125,1806]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of Tryonia species in the Chihuahuan Desert. 1. Tryonia hertleini, Las Palomas. 2. Tryonia peregrina sp. nov., Ojo Vareleño, northwest of Casas Grandes. 3. Tryonia peregrina sp. nov., Yepomera. 4. Tryonia zaragozae sp. nov., northeast of Ignacio Zaragoza. 5. Tryonia angostura sp. nov., T. ovata sp. nov., south of Galeana. 6. Tryonia molinae sp. nov., El Molino. 7. Tryonia taylori sp. nov., vicinity of Rancho Ojo Caliente. 8. Tryonia contrerasi sp. nov., vicinity of Rancho Nuevo. 9. Tryonia chuviscarae sp. nov., T. minckleyi sp. nov., Balneario de San Diego de Alcala. 10. Tryonia julimesensis sp. nov., south of Julimes. 11. Tryonia allendae sp. nov., southwest of Talamantes. 12. Tryonia metcalfi sp. nov., La Cienega, south of Candelaria. 13. Tryonia oasiensis sp. nov., Caroline Spring, Oasis Ranch. 14. Tryonia cheatumi, Phantom Lake Spring. 15. Tryonia circumstriata, Diamond Y Draw. 16. Tryonia seemani, Ojo de Agua del Obispo, Durango City. 17. Tryonia seemani, Nombre de Dios. 18. Tryonia seemani, Amado Nervo." httpUri="https://zenodo.org/record/278443/files/figure.png" pageId="22" pageNumber="23">Fig. 1</figureCitation>
|
||
, locality 9). This site, which is also known as Baños de San Diego de Alcala, is a warm spring complex (headspring temperatures ca. 42–45°C) that has long been used for recreation. Numerous springs discharge from a travertine mound (where a large bathhouse is situated) on the western edge of the complex (
|
||
<figureCitation id="13742A56BF618318FCD015AEFC081B83" box="[873,940,692,716]" captionStart="FIGURE 6" captionStartId="11.[151,244,1710,1731]" captionTargetBox="[221,1371,156,1678]" captionTargetId="figure@11.[212,1381,149,1685]" captionTargetPageId="11" captionText="FIGURE 6. Photographs of habitats of species treated in this paper. A. Ojo Vareleño, Chihuahua, type locality of T. peregrina sp. nov. (photograph taken on 16 / ix / 1990). B. Main spring in Yepomera, Chihuahua, T. peregrina habitat (18 / ix / 1990). C. Springs northeast of Ignacio Zaragoza, Chihuaua, type locality of T. zaragozae sp. nov. (8 / xii / 1998). D. Western-most head spring, Ojos de Arrey, Chihuahua, type locality of T. angosturae sp. nov. and T. ovata sp. nov. (7 / xii / 1998). E. Spring in northern part of El Molino, Chihuahua, type locality of T. molinae sp. nov. (26 / iv / 2010). F. Ojo del Apache, Chihuahua, type locality of T. contrerasi sp. nov. (14 / xii / 1998). G. Ojo Carbonero, Chihuahua, T. contrerasi habitat (2 / iv / 1991). H. Outflow of Ojo Caliente, Chihuahua, type locality of T. taylori sp. nov. (7 / xii / 1998). I – J. El Ojo de Servin, Chihuahua, T. taylori habitat (27 - viii- 1971, 7 / xii / 1998, respectively). K. Seeps in La Cienega, Texas, type locality of T. metcalfi sp. nov. (6 / xii / 2001). L – M. Balneario de San Diego de Alcala, Chihuahua, type locality of T. chuviscarae sp. nov. and T. minckleyi sp. nov. (4 / xii / 2001). N. Springs southeast of Julimes, Chihuahua, type locality of T. julimesensis sp. nov. (5 / iv / 1991). O. El Ojito (outflow), southwest of Talamantes, Chihuahua, type locality of T. allendae sp. nov. (10 / xii / 1998). P. Caroline Spring, Oasis Ranch, Texas, type locality of T. oasiensis sp. nov. (30 / iii / 2009). Q. Spring in Nombre de Dios, Durango, T. seemani habitat (2 / ix / 1971). R. Spring in Amado Nervo, Durango, T. seemani habitat (7 / iv / 1991)." httpUri="https://zenodo.org/record/278448/files/figure.png" pageId="22" pageNumber="23">Fig. 6</figureCitation>
|
||
L–M).
|
||
<taxonomicName id="4C4F4D50BF618318FC4315AFFB7B1B83" box="[1018,1247,692,716]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="chuviscarae">
|
||
<emphasis id="B93BEAC1BF618318FC4315AFFB7B1B83" box="[1018,1247,692,716]" italics="true" pageId="22" pageNumber="23">Tryonia chuviscarae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4F4D50BF618318FAAC15AFFA381B83" box="[1301,1436,692,716]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="minckleyi">
|
||
<emphasis id="B93BEAC1BF618318FAAC15AFFA381B83" box="[1301,1436,692,716]" italics="true" pageId="22" pageNumber="23">T. minckleyi</emphasis>
|
||
</taxonomicName>
|
||
were found throughout the spring complex (except in the headspring vents) on mud and gravel; the highest temperature recorded for this snail habitat was 41°C.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF618318FF7E143AFAA01A30" pageId="22" pageNumber="23" type="discussion">
|
||
<paragraph id="8BF036D3BF618318FF7E143AFAA01A30" blockId="22.[151,1436,151,896]" pageId="22" pageNumber="23">
|
||
<emphasis id="B93BEAC1BF618318FF7E143AFE901A77" bold="true" box="[199,308,800,824]" pageId="22" pageNumber="23">Remarks</emphasis>
|
||
. The occurrence of undescribed
|
||
<taxonomicName id="4C4F4D50BF618318FD0F143BFCAF1A77" box="[694,779,801,824]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="23" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B93BEAC1BF618318FD0F143BFCAF1A77" box="[694,779,801,824]" italics="true" pageId="22" pageNumber="23">Tryonia</emphasis>
|
||
</taxonomicName>
|
||
in the Balneario de San Diego de Alcala was noted by
|
||
<bibRefCitation id="EFDE4B22BF618318FF2E145EFE4C1A13" author="Minckley" box="[151,488,836,860]" pageId="22" pageNumber="23" refString="Minckley, W. L. & Minckley, C. O. (1986) Cyprinodon pachycephalus, a new species of pupfish (Cyprinodontidae) from the Chihuahuan Desert of northern Mexico. Copeia, 1986, 184 - 192." type="journal article" year="1986">Minckley & Minckley (1986)</bibRefCitation>
|
||
, however it is not clear whether they were referring to
|
||
<taxonomicName id="4C4F4D50BF618318FBE3145FFB5A1A13" box="[1114,1278,836,860]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="chuviscarae">
|
||
<emphasis id="B93BEAC1BF618318FBE3145FFB5A1A13" box="[1114,1278,836,860]" italics="true" pageId="22" pageNumber="23">T. chuviscarae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C4F4D50BF618318FAB4145FFA311A13" box="[1293,1429,836,860]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="minckleyi">
|
||
<emphasis id="B93BEAC1BF618318FAB4145FFA311A13" box="[1293,1429,836,860]" italics="true" pageId="22" pageNumber="23">T. minckleyi</emphasis>
|
||
</taxonomicName>
|
||
, or both.
|
||
<taxonomicName id="4C4F4D50BF618318FF4C1473FE7E1ACF" box="[245,474,872,896]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="chuviscarae">
|
||
<emphasis id="B93BEAC1BF618318FF4C1473FE7E1ACF" box="[245,474,872,896]" italics="true" pageId="22" pageNumber="23">Tryonia chuviscarae</emphasis>
|
||
</taxonomicName>
|
||
was delineated as a well supported lineage in the Bayesian tree (
|
||
<figureCitation id="13742A56BF618318FB08147DFB521A30" box="[1201,1270,871,895]" captionStart="FIGURE 2" captionStartId="5.[151,244,1938,1959]" captionTargetBox="[244,1345,828,1915]" captionTargetId="figure@5.[244,1346,828,1920]" captionTargetPageId="5" captionText="FIGURE 2. Bayesian tree based on the COI dataset. Posterior probabilities are provided when> 95 %. The names of the species treated herein are in bold face. Terminals and specimen codes are labeled as in Table 1. The two well supported clades composed of new species described herein are labeled A – B." httpUri="https://zenodo.org/record/278444/files/figure.png" pageId="22" pageNumber="23">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |