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<mods:title id="AC0CA1393FA678B8785CFBFE0332DFCF">Revision of the spider crab genus Sargassocarcinus Ward, 1936 (Crustacea: Brachyura: Epialtidae)</mods:title>
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Genus
<taxonomicName id="4C4F4D54133BFFDAFF040EA23E42DC89" ID-CoL="7PYSH" authority="Ward, 1936" authorityName="Ward" authorityYear="1936" box="[238,604,470,496]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC5133BFFDAFF040EA23DDADC89" bold="true" box="[238,452,470,496]" italics="true" pageId="1" pageNumber="314">Sargassocarcinus</emphasis>
<bibRefCitation id="EFDE4B26133BFFDAFE210EA23E42DC89" author="Ward" box="[459,604,470,496]" pageId="1" pageNumber="314" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11, 1 - 13." type="journal article" year="1936">Ward, 1936</bibRefCitation>
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<taxonomicName id="4C4F4D54133BFFDAFF7D0D683DFFDF4B" ID-CoL="7PYSH" authority="Ward, 1936: 8" authorityName="Ward" authorityPageNumber="8" authorityYear="1936" box="[151,481,540,562]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC5133BFFDAFF7D0D683D57DF48" box="[151,329,540,561]" italics="true" pageId="1" pageNumber="314">Sargassocarcinus</emphasis>
<treatmentCitation id="0AEE10C6133BFFDAFEA50D683DFFDF4B" author="Ward" box="[335,481,540,562]" page="8" pageId="1" pageNumber="314" year="1936">
<bibRefCitation id="EFDE4B26133BFFDAFEA50D683DD7DF4B" author="Ward" box="[335,457,540,562]" pageId="1" pageNumber="314" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11, 1 - 13." type="journal article" year="1936">Ward, 1936</bibRefCitation>
: 8
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;
<treatmentCitation id="0AEE10C6133BFFDAFE060D683E99DF4B" author="Sakai" box="[492,647,540,562]" page="75" pageId="1" pageNumber="314" year="1965">
<bibRefCitation id="EFDE4B26133BFFDAFE060D683E7FDF4B" author="Sakai" box="[492,609,540,562]" pageId="1" pageNumber="314" refString="Sakai, T. (1965) The Crabs of the Sagami Bay collected by His Majesty the Emperor of Japan. Maruzen Co., Tokyo, xvi + 206 pp. (English section), 27 Figs., 100 pls., 92 pp. (in Japanese) + 26 pp. (Bibliography and index in English) + 6 pp. (pp. 27 - 32, Index in Japanese), 1 map." type="book chapter" year="1965">Sakai 1965</bibRefCitation>
: 75
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;
<treatmentCitation id="0AEE10C6133BFFDAFD7B0D683FBDDF4B" author="Griffin" box="[657,931,540,562]" page="97" pageId="1" pageNumber="314" year="1986">
<bibRefCitation id="EFDE4B26133BFFDAFD7B0D683F63DF4B" author="Griffin" box="[657,893,540,562]" pageId="1" pageNumber="314" refString="Griffin, D. J. G. &amp; Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga- Expeditie, 39 (C 4), 1 - 335." type="journal article" year="1986">Griffin &amp; Tranter 1986</bibRefCitation>
: 97
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;
<treatmentCitation id="0AEE10C6133BFFDAFC440D683844DF4B" author="Davie" box="[942,1114,540,562]" page="267" pageId="1" pageNumber="314" year="2002">
<bibRefCitation id="EFDE4B26133BFFDAFC440D683836DF4B" author="Davie" box="[942,1064,540,562]" pageId="1" pageNumber="314" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca. Eucarida (Part 2). Decapoda - Anomura, Brachyura. Zoological Catalogue of Australia. Vol. 19.3 B. CSIRO Publications, Melbourne, 641 pp. [Wells, A. &amp; Houston, W. W. K. (Eds.)]" type="book" year="2002">Davie 2002</bibRefCitation>
: 267
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.
<typeStatus id="54F48875133BFFDAFF7D0D4F3CD7DF29" box="[151,201,571,592]" pageId="1" pageNumber="314">Type</typeStatus>
species:
<taxonomicName id="4C4F4D54133BFFDAFECC0D4F3EB6DF29" ID-CoL="79MMK" authority="Ward, 1936" authorityName="Ward" authorityYear="1936" box="[294,680,570,592]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="314" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B93BEAC5133BFFDAFECC0D4F3E34DF29" box="[294,554,570,592]" italics="true" pageId="1" pageNumber="314">Sargassocarcinus foliatus</emphasis>
<bibRefCitation id="EFDE4B26133BFFDAFDDB0D4E3EB6DF29" author="Ward" box="[561,680,570,592]" pageId="1" pageNumber="314" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11, 1 - 13." type="journal article" year="1936">Ward, 1936</bibRefCitation>
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, by original designation.
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<paragraph id="8BF036D7133BFFDAFF7D0D0938C1D9AF" blockId="1.[151,1437,637,1994]" pageId="1" pageNumber="314">
<emphasis id="B93BEAC5133BFFDAFF7D0D093D0FDFEF" bold="true" box="[151,273,637,662]" pageId="1" pageNumber="314">Diagnosis.</emphasis>
Surfaces of carapace, chelipeds, ambulatory legs covered by clavate setae, sometimes dense, foliose, spongiform along margins; dorsal surface of carapace with regions not well defined; proto-, metagastric regions elevated, each with median tubercle; cardiac region distinctly raised, with median tubercle; pseudorostrum short, bilobed, not deflexed, with distinct elongated, bifurcated rostral-like structure (pararostral teeth) below pseudorostrum formed by distal rim of antennular fossae; preorbital lobe large, triangular, directed anteriorly; postorbital lobe low to almost undiscernible; hepatic, lateral regions strongly expanded laterally to form wing-like lateral expansion with 2 broadly triangular lateral lobes; posterior carapace margin cristate. Eye dorsoventrally flattened, fused to orbital margin, immobile. Third maxilliped merus subtriangular, anteroexternal angle subauriculiform, carpus inserted on inner subdistal margin of merus, propodus, dactylus short. Chela short, with dorsal, ventral margins of palm distinctly cristate, dorsal margin of dactylus flattened, fingers of large males strongly curved forming prominent median gape when closed, cutting edge of dactylus with distinct subproximal tooth. Ambulatory legs covered with clavate setae, forming dense pubescence, expanded, foliose, along dorsal margins of merus, carpus, propodus forming “pseudo-cristae”; ventral margin of dactylus lined with pair of short subdistal spines, small denticles usually present posteriorly. Thoracic sternum wide, sternites 1
<emphasis id="B93BEAC5133BFFDAFB1D0B25391BD913" bold="true" box="[1271,1285,1105,1130]" pageId="1" pageNumber="314"></emphasis>
4 completely fused, surface of sternites 1
<emphasis id="B93BEAC5133BFFDAFE220B013DC8D9F7" bold="true" box="[456,470,1141,1166]" pageId="1" pageNumber="314"></emphasis>
3 distinctly concave. Male abdomen with somites 3
<emphasis id="B93BEAC5133BFFDAFBF90B01383FD9F7" bold="true" box="[1043,1057,1141,1166]" pageId="1" pageNumber="314"></emphasis>
5 separated by distinct sutures but somites immobile. G1 elongated, gently sinuous, distally bent outwards, distal part dilated but tapering to sharp tip; G2 short. Adult female abdomen with somites 13, telson free, somites 46 fused, domed outline.
</paragraph>
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<paragraph id="8BF036D7133BFFDAFF2D0B953FE4DA6F" blockId="1.[151,1437,637,1994]" pageId="1" pageNumber="314">
<emphasis id="B93BEAC5133BFFDAFF2D0B953D25D983" bold="true" box="[199,315,1249,1274]" pageId="1" pageNumber="314">Remarks.</emphasis>
The entire surface of all three
<taxonomicName id="4C4F4D54133BFFDAFD780B963F49D980" box="[658,855,1250,1273]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC5133BFFDAFD780B963F49D980" box="[658,855,1250,1273]" italics="true" pageId="1" pageNumber="314">Sargassocarcinus</emphasis>
</taxonomicName>
species are covered with unusual clavate setae. The setae are low and appear squamiform or reticulate on the dorsal surface of the carapace and thoracic sternum, especially when dried (
<figureCitation id="13742A52133BFFDAFE740A5D3DFDD83B" box="[414,483,1321,1346]" captionStart="FIGURE 3" captionStartId="5.[151,250,1912,1934]" captionTargetBox="[172,1414,399,1899]" captionTargetId="figure@5.[172,1414,399,1900]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of carapace. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan." httpUri="https://zenodo.org/record/266605/files/figure.png" pageId="1" pageNumber="314">Fig. 3</figureCitation>
). The setae form crest-like, lobiform or dentiform structures on the ambulatory legs and chelipeds (e.g.,
<figureCitation id="13742A52133BFFDAFE6C0A393DD1D81F" box="[390,463,1357,1382]" captionStart="FIGURE 6" captionStartId="9.[151,250,1597,1619]" captionTargetBox="[150,1435,179,1575]" captionTargetId="figure@9.[150,1436,175,1575]" captionTargetPageId="9" captionText="FIGURE 6. Ambulatory legs and cheliped. A H, Sargassocarcinus sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines [all same scale]; I N, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan [all same scale]. A, left first ambulatory leg; B, right first ambulatory leg (denuded); C, J, left second ambulatory leg; D, right second ambulatory leg; E, K, left third ambulatory leg; F, L, right third ambulatory leg; G, left fourth ambulatory leg; H, M, right fourth ambulatory leg; I, right first ambulatory leg; N, left cheliped. Scales = 1.0 mm." httpUri="https://zenodo.org/record/266609/files/figure.png" pageId="1" pageNumber="314">Fig. 6</figureCitation>
). For the setae to form crest-like structures is unusual, even among majoids. The pubescence is particularly low at the sutures of the thoracic sternites and abdomen, exposing the underlying structures (
<figureCitation id="13742A52133BFFDAFEFE0AE13D90D8D7" box="[276,398,1429,1454]" captionStart="FIGURE 5. A E, H" captionStartId="8.[151,250,1565,1587]" captionTargetBox="[151,1435,195,1542]" captionTargetId="figure@8.[151,1437,195,1543]" captionTargetPageId="8" captionText="FIGURE 5. A E, H, Sargassocarcinus sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; G, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan. A, anterior male thoracic sternum and abdomen; B, male thoracic sternum and abdomen; C, left cheliped; D, orbits, antennules, antennae and epistome; E, F, frontal views of cephalothorax; H, posterior view of cephalothorax. Scales = 1.0 mm." httpUri="https://zenodo.org/record/266607/files/figure.png" pageId="1" pageNumber="314">Fig. 5A, B</figureCitation>
). This pubescence is prominently denser and expanded on the dorsal and ventral margins of the ambulatory legs (more so in the former), with the structure appearing distinctly cristate (e.g.,
<figureCitation id="13742A52133BFFDAFB160ACD395ED8AB" box="[1276,1344,1465,1490]" captionStart="FIGURE 6" captionStartId="9.[151,250,1597,1619]" captionTargetBox="[150,1435,179,1575]" captionTargetId="figure@9.[150,1436,175,1575]" captionTargetPageId="9" captionText="FIGURE 6. Ambulatory legs and cheliped. A H, Sargassocarcinus sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines [all same scale]; I N, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan [all same scale]. A, left first ambulatory leg; B, right first ambulatory leg (denuded); C, J, left second ambulatory leg; D, right second ambulatory leg; E, K, left third ambulatory leg; F, L, right third ambulatory leg; G, left fourth ambulatory leg; H, M, right fourth ambulatory leg; I, right first ambulatory leg; N, left cheliped. Scales = 1.0 mm." httpUri="https://zenodo.org/record/266609/files/figure.png" pageId="1" pageNumber="314">Fig. 6</figureCitation>
A) even though the surfaces are actually rounded (
<figureCitation id="13742A52133BFFDAFD9F0AA93EA4D88F" box="[629,698,1501,1526]" captionStart="FIGURE 6" captionStartId="9.[151,250,1597,1619]" captionTargetBox="[150,1435,179,1575]" captionTargetId="figure@9.[150,1436,175,1575]" captionTargetPageId="9" captionText="FIGURE 6. Ambulatory legs and cheliped. A H, Sargassocarcinus sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines [all same scale]; I N, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan [all same scale]. A, left first ambulatory leg; B, right first ambulatory leg (denuded); C, J, left second ambulatory leg; D, right second ambulatory leg; E, K, left third ambulatory leg; F, L, right third ambulatory leg; G, left fourth ambulatory leg; H, M, right fourth ambulatory leg; I, right first ambulatory leg; N, left cheliped. Scales = 1.0 mm." httpUri="https://zenodo.org/record/266609/files/figure.png" pageId="1" pageNumber="314">Fig. 6</figureCitation>
B).
<bibRefCitation id="EFDE4B26133BFFDAFD0F0AA93F8DD88F" author="Ward" box="[741,915,1501,1526]" pageId="1" pageNumber="314" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11, 1 - 13." type="journal article" year="1936">Ward (1936: 9)</bibRefCitation>
remains the only author to have remarked on this pubescence, but only for the surface of the carapace: “covered with fine tomentum that imparts a punctate appearance”.
<bibRefCitation id="EFDE4B26133BFFDAFEDA09513DC2DB47" author="Ward" box="[304,476,1573,1598]" pageId="1" pageNumber="314" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11, 1 - 13." type="journal article" year="1936">Ward (1936: 9)</bibRefCitation>
described the legs of
<taxonomicName id="4C4F4D54133BFFDAFD3F09523F58DB44" box="[725,838,1573,1597]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="314" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B93BEAC5133BFFDAFD3F09523F58DB44" box="[725,838,1573,1597]" italics="true" pageId="1" pageNumber="314">S. foliatus</emphasis>
</taxonomicName>
as “sharply carinated” without mentioning the setae, suggesting he may have thought these were actually part of the exoskeleton.
<bibRefCitation id="EFDE4B26133BFFDAFBE8093D38F0DB1B" author="Rathbun" box="[1026,1262,1609,1634]" pageId="1" pageNumber="314" refString="Rathbun, M. J. (1916) New species of crabs of the families Inachidae and Parthenopidae. Scientific results of the Philippine Cruise of the Fisheries Steamer &quot; Albatross, &quot; 1907 - 1910. - No. 34. Proceedings of the United States National Museum, 50 (2135), 527 - 559. http: // dx. doi. org / 10.5479 / si. 00963801.2135.527" type="journal article" year="1916">Rathbun (1916: 536)</bibRefCitation>
also noted that the ambulatory meri of
<taxonomicName id="4C4F4D54133BFFDAFE76091A3E0BDBFC" box="[412,533,1645,1669]" class="Malacostraca" family="Epialtidae" genus="Peltinia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="314" phylum="Arthropoda" rank="species" species="sublimis">
<emphasis id="B93BEAC5133BFFDAFE76091A3E0BDBFC" box="[412,533,1645,1669]" italics="true" pageId="1" pageNumber="314">S. sublimis</emphasis>
</taxonomicName>
are cristate but this is actually not the case when denuded (
<figureCitation id="13742A52133BFFDAFB4C091938F7DBFF" box="[1190,1257,1645,1670]" captionStart="FIGURE 6" captionStartId="9.[151,250,1597,1619]" captionTargetBox="[150,1435,179,1575]" captionTargetId="figure@9.[150,1436,175,1575]" captionTargetPageId="9" captionText="FIGURE 6. Ambulatory legs and cheliped. A H, Sargassocarcinus sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines [all same scale]; I N, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan [all same scale]. A, left first ambulatory leg; B, right first ambulatory leg (denuded); C, J, left second ambulatory leg; D, right second ambulatory leg; E, K, left third ambulatory leg; F, L, right third ambulatory leg; G, left fourth ambulatory leg; H, M, right fourth ambulatory leg; I, right first ambulatory leg; N, left cheliped. Scales = 1.0 mm." httpUri="https://zenodo.org/record/266609/files/figure.png" pageId="1" pageNumber="314">Fig. 6</figureCitation>
B), although the dense clavate setae make it appear so (
<figureCitation id="13742A52133BFFDAFDAD09E53E97DBD3" box="[583,649,1681,1706]" captionStart="FIGURE 6" captionStartId="9.[151,250,1597,1619]" captionTargetBox="[150,1435,179,1575]" captionTargetId="figure@9.[150,1436,175,1575]" captionTargetPageId="9" captionText="FIGURE 6. Ambulatory legs and cheliped. A H, Sargassocarcinus sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines [all same scale]; I N, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan [all same scale]. A, left first ambulatory leg; B, right first ambulatory leg (denuded); C, J, left second ambulatory leg; D, right second ambulatory leg; E, K, left third ambulatory leg; F, L, right third ambulatory leg; G, left fourth ambulatory leg; H, M, right fourth ambulatory leg; I, right first ambulatory leg; N, left cheliped. Scales = 1.0 mm." httpUri="https://zenodo.org/record/266609/files/figure.png" pageId="1" pageNumber="314">Fig. 6</figureCitation>
A, CH).
<bibRefCitation id="EFDE4B26133BFFDAFD1D09E53FAEDBD3" author="Balss" box="[759,944,1681,1706]" pageId="1" pageNumber="314" refString="Balss, H. (1924) Ostasiatische Decapoden. V. Die Oxyrhynchen und Schlussteil. Archiv fur Naturgeschichte, 90, 20 - 84." type="journal article" year="1924">Balss (1924: 28)</bibRefCitation>
describes the dorsal surface of the carapace of
<taxonomicName id="4C4F4D54133BFFDAFF5E09C23D2FDBB4" box="[180,305,1718,1741]" class="Malacostraca" family="Epialtidae" genus="Mimulus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="314" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="B93BEAC5133BFFDAFF5E09C23D2FDBB4" box="[180,305,1718,1741]" italics="true" pageId="1" pageNumber="314">S. cristatus</emphasis>
</taxonomicName>
as smooth but it is not so. In the smaller male (8.2 ×
<quantity id="4CB79B32133BFFDAFC9109C23FCDDBB4" box="[891,979,1718,1742]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="1" pageNumber="314" unit="mm" value="7.4">7.4 mm</quantity>
, USNM 48248), the setae are present but conspicuously short and sparse, being slightly denser on the lateral parts (
<figureCitation id="13742A52133BFFDAFC3109AD3800DB8B" box="[987,1054,1753,1778]" captionStart="FIGURE 3" captionStartId="5.[151,250,1912,1934]" captionTargetBox="[172,1414,399,1899]" captionTargetId="figure@5.[172,1414,399,1900]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of carapace. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan." httpUri="https://zenodo.org/record/266605/files/figure.png" pageId="1" pageNumber="314">Fig. 3</figureCitation>
E). The setae are more spread out but still visible (
<figureCitation id="13742A52133BFFDAFEA709893D90DA6F" box="[333,398,1789,1814]" captionStart="FIGURE 3" captionStartId="5.[151,250,1912,1934]" captionTargetBox="[172,1414,399,1899]" captionTargetId="figure@5.[172,1414,399,1900]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of carapace. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan." httpUri="https://zenodo.org/record/266605/files/figure.png" pageId="1" pageNumber="314">Fig. 3</figureCitation>
F) in the larger male (14.4 ×
<quantity id="4CB79B32133BFFDAFD38098A3F29DA6C" box="[722,823,1790,1814]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="1" pageNumber="314" unit="mm" value="11.7">11.7 mm</quantity>
, ZRC 2015.288).
</paragraph>
<paragraph id="8BF036D7133BFFDAFF2D08563E3ADAB3" blockId="1.[151,1437,637,1994]" pageId="1" pageNumber="314">
An unusual feature of the species of
<taxonomicName id="4C4F4D54133BFFDAFD7C08563F45DA40" box="[662,859,1826,1849]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC5133BFFDAFD7C08563F45DA40" box="[662,859,1826,1849]" italics="true" pageId="1" pageNumber="314">Sargassocarcinus</emphasis>
</taxonomicName>
is the dorso-laterally flattened eyes, which are completely fused to the orbital margins (
<figureCitation id="13742A52133BFFDAFD8408313ED0DA27" box="[622,718,1861,1886]" captionStart="FIGURE 5. A E, H" captionStartId="8.[151,250,1565,1587]" captionTargetBox="[151,1435,195,1542]" captionTargetId="figure@8.[151,1437,195,1543]" captionTargetPageId="8" captionText="FIGURE 5. A E, H, Sargassocarcinus sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; G, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan. A, anterior male thoracic sternum and abdomen; B, male thoracic sternum and abdomen; C, left cheliped; D, orbits, antennules, antennae and epistome; E, F, frontal views of cephalothorax; H, posterior view of cephalothorax. Scales = 1.0 mm." httpUri="https://zenodo.org/record/266607/files/figure.png" pageId="1" pageNumber="314">Figs 5D</figureCitation>
; 7A, B, K; 10B, C). While the ocular peduncle is still clearly visible, it is completely fused to the carapace and immobile. This is unusual for epialtids. Even the similarly looking members of
<taxonomicName id="4C4F4D54133BFFDAFE6208FA3DC5DADC" box="[392,475,1934,1957]" class="Malacostraca" family="Epialtidae" genus="Huenia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC5133BFFDAFE6208FA3DC5DADC" box="[392,475,1934,1957]" italics="true" pageId="1" pageNumber="314">Huenia</emphasis>
</taxonomicName>
have mobile eyes. The eye itself is peculiar in form, with the lateral edges cornea appearing cristate (
<figureCitation id="13742A52133BFFDAFE8708C53DF8DAB0" box="[365,486,1969,1994]" captionStart="FIGURE 7. A, B, D F" captionStartId="10.[151,250,1648,1670]" captionTargetBox="[171,1418,198,1614]" captionTargetId="figure@10.[150,1436,193,1627]" captionTargetPageId="10" captionText="FIGURE 7. A, B, D F, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; C, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; G, I, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; H, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; J, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; K, S. cristatus, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289). A, I, J, frontal margin and orbits; B, frontal margin, orbits, epistome, antennules and antennae; C, thoracic sternum and sternoabdominal cavity; D, inner view of left third maxilliped; E, outer view of left third maxilliped; F, G, dactylus of first ambulatory leg (denuded); K, ventral view of left eye. Scales: A C, F J = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/266610/files/figure.png" pageId="1" pageNumber="314">Fig. 7A, B</figureCitation>
, I, J).
</paragraph>
<caption id="DF30665F1338FFD9FF7D0ACC3D23DB53" httpUri="https://zenodo.org/record/266603/files/figure.png" pageId="2" pageNumber="315" targetBox="[151,1436,177,1441]" targetPageId="2">
<paragraph id="8BF036D71338FFD9FF7D0ACC3D23DB53" blockId="2.[151,1436,1464,1578]" pageId="2" pageNumber="315">
<emphasis id="B93BEAC51338FFD9FF7D0ACC3D0DD8B4" bold="true" box="[151,275,1464,1486]" pageId="2" pageNumber="315">FIGURE 1.</emphasis>
<taxonomicName id="4C4F4D541338FFD9FEF00ACD3E34D8B7" box="[282,554,1465,1486]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="315" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="B93BEAC51338FFD9FEF00ACD3E34D8B7" box="[282,554,1465,1486]" italics="true" pageId="2" pageNumber="315">Sargassocarcinus cristatus</emphasis>
</taxonomicName>
, colour in life. AD, overall habitus; E, frontal view of cephalothorax; F, ventral view of cephalothorax. A, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; B, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289), Taiwan; C, male (12.5 × 10.1 mm) (NMMBCD 4065), Taiwan; DF, ovigerous female (14.3 × 11.1 mm) (RUMF-ZC-3912), Ryukyus, Japan.
</paragraph>
</caption>
<paragraph id="8BF036D71338FFD9FF2D09253F4ADBAF" blockId="2.[151,1437,1617,2038]" pageId="2" pageNumber="315">
The carpus of the third maxilliped of most majoids is inserted on the distal margin of the merus. In the case of
<taxonomicName id="4C4F4D541338FFD9FF7D09023D42DBF4" box="[151,348,1654,1677]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC51338FFD9FF7D09023D42DBF4" box="[151,348,1654,1677]" italics="true" pageId="2" pageNumber="315">Sargassocarcinus</emphasis>
</taxonomicName>
species, the carpus is attached to the inner subdistal margin of the merus. As a result, when the palp (carpus, propodus, and dactylus) is appressed against the merus, most of it is covered (
<figureCitation id="13742A521338FFD9FB7409ED38EEDBC8" box="[1182,1264,1689,1714]" captionStart="FIGURE 10" captionStartId="13.[151,250,1446,1468]" captionTargetBox="[151,1435,193,1424]" captionTargetId="figure@13.[151,1436,193,1425]" captionTargetPageId="13" captionText="FIGURE 10. Sargassocarcinus cristatus. A, B, female (10.8 × 8.9 mm) (NMMBCD 4065), Taiwan; C, D, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289), Taiwan. A, overall habitus; B, ventral view of cephalothorax showing buccal cavity, epistome, eyes and abdomen; D, sterno-abdominal cavity showing vulvae. Abbreviation: v, vulvae." httpUri="https://zenodo.org/record/266613/files/figure.png" pageId="2" pageNumber="315">Fig. 10</figureCitation>
B, C), and only part of the dactylus is visible from the outer view (
<figureCitation id="13742A521338FFD9FD2409C93F0EDBAC" box="[718,784,1725,1750]" captionStart="FIGURE 7. A, B, D F" captionStartId="10.[151,250,1648,1670]" captionTargetBox="[171,1418,198,1614]" captionTargetId="figure@10.[150,1436,193,1627]" captionTargetPageId="10" captionText="FIGURE 7. A, B, D F, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; C, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; G, I, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; H, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; J, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; K, S. cristatus, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289). A, I, J, frontal margin and orbits; B, frontal margin, orbits, epistome, antennules and antennae; C, thoracic sternum and sternoabdominal cavity; D, inner view of left third maxilliped; E, outer view of left third maxilliped; F, G, dactylus of first ambulatory leg (denuded); K, ventral view of left eye. Scales: A C, F J = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/266610/files/figure.png" pageId="2" pageNumber="315">Fig. 7</figureCitation>
D, E).
</paragraph>
<paragraph id="8BF036D71338FFD9FF2D099538DFDA8F" blockId="2.[151,1437,1617,2038]" pageId="2" pageNumber="315">
The structure of the front in
<taxonomicName id="4C4F4D541338FFD9FDE309953ED0DB81" box="[521,718,1761,1784]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC51338FFD9FDE309953ED0DB81" box="[521,718,1761,1784]" italics="true" pageId="2" pageNumber="315">Sargassocarcinus</emphasis>
</taxonomicName>
is noteworthy and could be unique among epialtids. The actual pseudorostrum is actually short, not deflexed, divided into two relatively low triangular lobes by a median concavity and clearly visible from dorsal view (
<figureCitation id="13742A521338FFD9FD25085D3F33DA38" box="[719,813,1833,1858]" captionStart="FIGURE 7. A, B, D F" captionStartId="10.[151,250,1648,1670]" captionTargetBox="[171,1418,198,1614]" captionTargetId="figure@10.[150,1436,193,1627]" captionTargetPageId="10" captionText="FIGURE 7. A, B, D F, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; C, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; G, I, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; H, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; J, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; K, S. cristatus, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289). A, I, J, frontal margin and orbits; B, frontal margin, orbits, epistome, antennules and antennae; C, thoracic sternum and sternoabdominal cavity; D, inner view of left third maxilliped; E, outer view of left third maxilliped; F, G, dactylus of first ambulatory leg (denuded); K, ventral view of left eye. Scales: A C, F J = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/266610/files/figure.png" pageId="2" pageNumber="315">Fig. 7A</figureCitation>
, I, J). Below this pseudorostrum is a pair of slender triangular lobes that are actually the expanded and projecting distal rims of the antennular fossae (
<figureCitation id="13742A521338FFD9FB010839395BDA1F" box="[1259,1349,1869,1894]" captionStart="FIGURE 7. A, B, D F" captionStartId="10.[151,250,1648,1670]" captionTargetBox="[171,1418,198,1614]" captionTargetId="figure@10.[150,1436,193,1627]" captionTargetPageId="10" captionText="FIGURE 7. A, B, D F, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; C, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; G, I, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; H, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; J, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; K, S. cristatus, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289). A, I, J, frontal margin and orbits; B, frontal margin, orbits, epistome, antennules and antennae; C, thoracic sternum and sternoabdominal cavity; D, inner view of left third maxilliped; E, outer view of left third maxilliped; F, G, dactylus of first ambulatory leg (denuded); K, ventral view of left eye. Scales: A C, F J = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/266610/files/figure.png" pageId="2" pageNumber="315">Fig. 7B</figureCitation>
). These are here referred to as pararostral teeth, and are longer than the actual pseudorostral ones.
<bibRefCitation id="EFDE4B261338FFD9FB4308053CD7DAD7" author="Sakai" pageId="2" pageNumber="315" refString="Sakai, T. (1965) The Crabs of the Sagami Bay collected by His Majesty the Emperor of Japan. Maruzen Co., Tokyo, xvi + 206 pp. (English section), 27 Figs., 100 pls., 92 pp. (in Japanese) + 26 pp. (Bibliography and index in English) + 6 pp. (pp. 27 - 32, Index in Japanese), 1 map." type="book chapter" year="1965" yearSuffix="b">Sakai (1965: text-fig. 11b)</bibRefCitation>
clearly noticed this as he illustrated this unusual structure although he did not discuss it. This is a feature not known for other epialtids, even
<taxonomicName id="4C4F4D541338FFD9FDFC08CE3E76DAA8" box="[534,616,1978,2001]" class="Malacostraca" family="Epialtidae" genus="Huenia" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="315" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC51338FFD9FDFC08CE3E76DAA8" box="[534,616,1978,2001]" italics="true" pageId="2" pageNumber="315">Huenia</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFDE4B261338FFD9FD9108CD3F6EDAAB" author="Rathbun" box="[635,880,1977,2002]" pageId="2" pageNumber="315" refString="Rathbun, M. J. (1916) New species of crabs of the families Inachidae and Parthenopidae. Scientific results of the Philippine Cruise of the Fisheries Steamer &quot; Albatross, &quot; 1907 - 1910. - No. 34. Proceedings of the United States National Museum, 50 (2135), 527 - 559. http: // dx. doi. org / 10.5479 / si. 00963801.2135.527" type="journal article" year="1916">Rathbun (1916: 536)</bibRefCitation>
describes the pseudorostrum (as a rostrum) as “deflexed and cut halfway back into two small triangular teeth separated by a triangular sinus”.
</paragraph>
<paragraph id="8BF036D71339FFD8FF2D0FE33FDCDC65" blockId="3.[151,1436,151,536]" pageId="3" pageNumber="316">
<bibRefCitation id="EFDE4B261339FFD8FF2D0FE33E0FDDC9" author="Griffin" box="[199,529,151,176]" pageId="3" pageNumber="316" refString="Griffin, D. J. G. &amp; Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga- Expeditie, 39 (C 4), 1 - 335." type="journal article" year="1986">Griffin &amp; Tranter (1986: 98)</bibRefCitation>
observed that the male abdomen consists of seven free somites (including the telson). While the sutures between somites 35 are visible (
<figureCitation id="13742A521339FFD8FCC60FC83FB7DDAD" box="[812,937,188,213]" captionStart="FIGURE 5. A E, H" captionStartId="8.[151,250,1565,1587]" captionTargetBox="[151,1435,195,1542]" captionTargetId="figure@8.[151,1437,195,1543]" captionTargetPageId="8" captionText="FIGURE 5. A E, H, Sargassocarcinus sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; G, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan. A, anterior male thoracic sternum and abdomen; B, male thoracic sternum and abdomen; C, left cheliped; D, orbits, antennules, antennae and epistome; E, F, frontal views of cephalothorax; H, posterior view of cephalothorax. Scales = 1.0 mm." httpUri="https://zenodo.org/record/266607/files/figure.png" pageId="3" pageNumber="316">Figs 5A, B</figureCitation>
; 8AD), they are unable to articulate and are effectively fused. In females, only somites 46 are fused, with the sutures between the somites in adults barely visible, forming a domed brood pouch for the eggs (
<figureCitation id="13742A521339FFD8FD340E703F3ADC65" box="[734,804,260,285]" captionStart="FIGURE 8. A" captionStartId="11.[151,250,1364,1385]" captionTargetBox="[162,1417,349,1329]" captionTargetId="figure@11.[150,1436,337,1343]" captionTargetPageId="11" captionText="FIGURE 8. A, Sargassocarcinus foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; B, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; C, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; D, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; E H, S. cristatus, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289), Taiwan. A D, male abdomen (somites 3 6 and telson); E, female fused abdominal somites 4 6 and telson; F, female abdominal somites 2 4; G, female abdominal somite 1; H, female vulvae (arrow points to buccal cavity). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266611/files/figure.png" pageId="3" pageNumber="316">Figs 8</figureCitation>
E, F; 10B, C).
</paragraph>
<paragraph id="8BF036D71339FFD8FF2D0E533CFEDF61" blockId="3.[151,1436,151,536]" pageId="3" pageNumber="316">
The carapace of
<taxonomicName id="4C4F4D541339FFD8FE6B0E5C3E58DC46" box="[385,582,296,319]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC51339FFD8FE6B0E5C3E58DC46" box="[385,582,296,319]" italics="true" pageId="3" pageNumber="316">Sargassocarcinus</emphasis>
</taxonomicName>
superficially resembles genera like
<taxonomicName id="4C4F4D541339FFD8FC330E5C38C1DC39" authority="De Haan, 1837" authorityName="De Haan" authorityYear="1837" box="[985,1247,296,320]" class="Malacostraca" family="Epialtidae" genus="Huenia" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC51339FFD8FC330E5C3835DC46" box="[985,1067,296,319]" italics="true" pageId="3" pageNumber="316">Huenia</emphasis>
<bibRefCitation id="EFDE4B261339FFD8FBD90E5C38C1DC39" author="De" box="[1075,1247,296,320]" pageId="3" pageNumber="316" refString="De Haan, H. M. (1837 [1833 - 1849) Crustacea. In: von Siebold, P. F. (Ed.), Fauna Japonica, sive descriptio animalium, quae in itinere per Japoniam, jussu et auspiciis superiorum, qui summum in India Batavia imperium tenent, suscepto, annis 1823 - 1830 collegit, notis, observationibus a adumbrationibus illustravit. Fasc. 1 - 8. Lugduni Batavorum, Leiden, 21 + 11 + 8 + 243 pp. [pp. i - xxi + vii - xvii + ix - xvi + 243]" type="book chapter" year="1837">De Haan, 1837</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C4F4D541339FFD8FAF30E5C3DADDC1D" authority="H. Milne-Edwards, 1834" authorityName="H. Milne-Edwards" authorityYear="1834" class="Malacostraca" family="Epialtidae" genus="Menaethius" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC51339FFD8FAF30E5C3982DC39" box="[1305,1436,296,320]" italics="true" pageId="3" pageNumber="316">Menaethius</emphasis>
H. Milne-Edwards, 1834
</taxonomicName>
, in having wing-like expansions of the lateral margin. The flattened and fused eyes and the unusual pseudorostral and pararostral structures, however, argue against a close relationship between them (cf.
<bibRefCitation id="EFDE4B261339FFD8FF7D0EE03D80DCD4" author="Griffin" box="[151,414,404,429]" pageId="3" pageNumber="316" refString="Griffin, D. J. G. &amp; Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga- Expeditie, 39 (C 4), 1 - 335." type="journal article" year="1986">Griffin &amp; Tranter 1986</bibRefCitation>
;
<bibRefCitation id="EFDE4B261339FFD8FE430EE03EC7DCD5" author="Takeda" box="[425,729,404,429]" pageId="3" pageNumber="316" refString="Takeda, M. &amp; Marumura, M. (2010) Spider crabs of the genus Huenia De Haan, 1837 (Crustacea, Decapoda, Brachyura) from Japan, with descriptions of two new species. Bulletin of the National Museum of Nature and Science, Tokyo, Series A, 36, 39 - 48." type="journal article" year="2010">Takeda &amp; Marumura 2010</bibRefCitation>
). The wing-like expansions of
<taxonomicName id="4C4F4D541339FFD8FBDF0EE038E4DCD2" box="[1077,1274,404,427]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC51339FFD8FBDF0EE038E4DCD2" box="[1077,1274,404,427]" italics="true" pageId="3" pageNumber="316">Sargassocarcinus</emphasis>
</taxonomicName>
also resemble the recently described
<taxonomicName id="4C4F4D541339FFD8FE790ECC3EFDDCA9" authority="Ng &amp; Anker, 2014" authorityName="Ng &amp; Anker" authorityYear="2014" box="[403,739,439,464]" class="Malacostraca" family="Epialtidae" genus="Pteromaja" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC51339FFD8FE790ECC3E16DCB6" box="[403,520,440,463]" italics="true" pageId="3" pageNumber="316">Pteromaja</emphasis>
Ng &amp;
<bibRefCitation id="EFDE4B261339FFD8FDBF0EC33EFDDCA9" author="Anker" box="[597,739,439,464]" pageId="3" pageNumber="316" refString="Ng, P. K. L. &amp; Anker, A. (2014) Pteromaja maklayi gen. et sp. nov., a remarkable new spider crab (Crustacea: Brachyura: Epialtidae) from Papua New Guinea. Marine Biology Research, 10 (8), 816 - 823. http: // dx. doi. org / 10.1080 / 17451000.2013.853127" type="journal article" year="2014">Anker, 2014</bibRefCitation>
</taxonomicName>
, from
<collectingCountry id="F35876471339FFD8FCC10ECC3818DCA9" box="[811,1030,439,464]" name="Papua New Guinea" pageId="3" pageNumber="316">Papua New Guinea</collectingCountry>
, but in that genus the carapace is far more pyriform, the pseudorostrum, eyes, chelipeds, and ambulatory legs are completely different (cf. Ng &amp;
<bibRefCitation id="EFDE4B261339FFD8FABC0EA83CCADF61" author="Anker" pageId="3" pageNumber="316" refString="Ng, P. K. L. &amp; Anker, A. (2014) Pteromaja maklayi gen. et sp. nov., a remarkable new spider crab (Crustacea: Brachyura: Epialtidae) from Papua New Guinea. Marine Biology Research, 10 (8), 816 - 823. http: // dx. doi. org / 10.1080 / 17451000.2013.853127" type="journal article" year="2014">Anker 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C355655C1339FFD8FF7D0D183D75DE52" pageId="3" pageNumber="316" type="reference_group">
<paragraph id="8BF036D71339FFD8FF7D0D183E54DFFC" blockId="3.[151,676,619,680]" box="[151,586,619,645]" pageId="3" pageNumber="316">
<taxonomicName id="4C4F4D541339FFD8FF7D0D183E54DFFC" ID-CoL="79MMK" authority="Ward, 1936" authorityName="Ward" authorityYear="1936" box="[151,586,619,645]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B93BEAC51339FFD8FF7D0D183DDFDFFD" bold="true" box="[151,449,619,644]" italics="true" pageId="3" pageNumber="316">Sargassocarcinus foliatus</emphasis>
<bibRefCitation id="EFDE4B261339FFD8FE220D183E54DFFC" author="Ward" box="[456,586,620,645]" pageId="3" pageNumber="316" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11, 1 - 13." type="journal article" year="1936">Ward, 1936</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BF036D71339FFD8FF7D0DFB3EBADFD1" blockId="3.[151,676,619,680]" box="[151,676,655,680]" pageId="3" pageNumber="316">
(
<figureCitation id="13742A521339FFD8FF750DFB3CFBDFD1" box="[159,229,655,680]" captionStart="FIGURE 2" captionStartId="4.[151,250,1499,1521]" captionTargetBox="[151,1436,193,1477]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURE 2. Overall habitus. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; F, S. cristatus, male (12.5 × 10.1 mm) (NMMBCD 4065), Taiwan." httpUri="https://zenodo.org/record/266604/files/figure.png" pageId="3" pageNumber="316">Figs 2</figureCitation>
AC; 3AC, 4; 7A, B, DF; 8A; 9AE)
</paragraph>
<paragraph id="8BF036D71339FFD8FF7D0DAD3D75DE52" blockId="3.[151,1436,728,811]" pageId="3" pageNumber="316">
<treatmentCitationGroup id="AB5F11F91339FFD8FF7D0DAD3D75DE52" pageId="3" pageNumber="316">
<taxonomicName id="4C4F4D541339FFD8FF7D0DAD3E2CDF97" ID-CoL="79MMK" authority="Ward, 1936: 9" authorityName="Ward" authorityPageNumber="9" authorityYear="1936" box="[151,562,728,750]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B93BEAC51339FFD8FF7D0DAD3D85DF97" box="[151,411,728,750]" italics="true" pageId="3" pageNumber="316">Sargassocarcinus foliatus</emphasis>
<treatmentCitation id="0AEE10C61339FFD8FE480DAC3E2CDF97" author="Ward" box="[418,562,728,750]" page="9" pageId="3" pageNumber="316" year="1936">
<bibRefCitation id="EFDE4B261339FFD8FE480DAC3E04DF97" author="Ward" box="[418,538,728,750]" pageId="3" pageNumber="316" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11, 1 - 13." type="journal article" year="1936">Ward, 1936</bibRefCitation>
: 9
</treatmentCitation>
</taxonomicName>
, pl. 3 figs 46;
<treatmentCitation id="0AEE10C61339FFD8FD3B0DAC3836DF97" author="Springthorpe" box="[721,1064,728,750]" page="100" pageId="3" pageNumber="316" year="1994">
<bibRefCitation id="EFDE4B261339FFD8FD3B0DAC3FEBDF97" author="Springthorpe" box="[721,1013,728,750]" pageId="3" pageNumber="316" refString="Springthorpe, R. T. &amp; Lowry, J. K. (1994) Catalogue of Crustacean Type Specimens in the Australian Museum: Malacostraca. Technical Reports of the Australian Museum, 11, 1 - 134. http: // dx. doi. org / 10.3853 / j. 1031 - 8062.11.1994.68" type="journal article" year="1994">Springthorpe &amp; Lowry 1994</bibRefCitation>
: 100
</treatmentCitation>
.
<taxonomicName id="4C4F4D541339FFD8FF7D0D833EA6DE75" ID-CoL="6XMMP" authority="Griffin &amp; Tranter 1986: 97" authorityName="Griffin &amp; Tranter" authorityPageNumber="97" authorityYear="1986" box="[151,696,758,780]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="sublimis">
<emphasis id="B93BEAC51339FFD8FF7D0D833DBADE75" box="[151,420,758,780]" italics="true" pageId="3" pageNumber="316">Sargassocarcinus sublimis</emphasis>
<treatmentCitation id="0AEE10C61339FFD8FE400D823EA6DE75" author="Griffin" box="[426,696,758,780]" page="97" pageId="3" pageNumber="316" year="1986">
<bibRefCitation id="EFDE4B261339FFD8FE400D823E8BDE75" author="Griffin" box="[426,661,758,780]" pageId="3" pageNumber="316" refString="Griffin, D. J. G. &amp; Tranter, H. A. (1986) The Decapoda Brachyura of the Siboga Expedition. Part VIII. Majidae. Siboga- Expeditie, 39 (C 4), 1 - 335." type="journal article" year="1986">Griffin &amp; Tranter 1986</bibRefCitation>
: 97
</treatmentCitation>
</taxonomicName>
(part);
<treatmentCitation id="0AEE10C61339FFD8FCEE0D823FAEDE75" author="Davie" box="[772,944,758,780]" page="287" pageId="3" pageNumber="316" year="2002">
<bibRefCitation id="EFDE4B261339FFD8FCEE0D823F60DE75" author="Davie" box="[772,894,758,780]" pageId="3" pageNumber="316" refString="Davie, P. J. F. (2002) Crustacea: Malacostraca. Eucarida (Part 2). Decapoda - Anomura, Brachyura. Zoological Catalogue of Australia. Vol. 19.3 B. CSIRO Publications, Melbourne, 641 pp. [Wells, A. &amp; Houston, W. W. K. (Eds.)]" type="book" year="2002">Davie 2002</bibRefCitation>
: 287
</treatmentCitation>
; Ng
<emphasis id="B93BEAC51339FFD8FC350D8C380CDE75" box="[991,1042,758,781]" italics="true" pageId="3" pageNumber="316">et al.</emphasis>
2008: 101 (part) (not
<taxonomicName id="4C4F4D541339FFD8FB1B0D823D7EDE52" authority="Rathbun, 1916" authorityName="Rathbun" authorityYear="1916" class="Malacostraca" family="Epialtidae" genus="Peltinia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="316" phylum="Arthropoda" rank="species" species="sublimis">
<emphasis id="B93BEAC51339FFD8FB1B0D823982DE75" box="[1265,1436,758,780]" italics="true" pageId="3" pageNumber="316">Peltinia sublimis</emphasis>
<bibRefCitation id="EFDE4B261339FFD8FF2D0C613D7EDE52" author="Rathbun" box="[199,352,789,811]" pageId="3" pageNumber="316" refString="Rathbun, M. J. (1916) New species of crabs of the families Inachidae and Parthenopidae. Scientific results of the Philippine Cruise of the Fisheries Steamer &quot; Albatross, &quot; 1907 - 1910. - No. 34. Proceedings of the United States National Museum, 50 (2135), 527 - 559. http: // dx. doi. org / 10.5479 / si. 00963801.2135.527" type="journal article" year="1916">Rathbun, 1916</bibRefCitation>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C355655C1339FFD8FF7D0C233922DEC1" pageId="3" pageNumber="316" type="materials_examined">
<paragraph id="8BF036D71339FFD8FF7D0C233922DEC1" blockId="3.[151,1437,855,1997]" pageId="3" pageNumber="316">
<materialsCitation id="3B273C8A1339FFD8FF7D0C233D00DEEC" ID-GBIF-Occurrence="1306554014" country="Iceland" location="Whitsunday Passage" pageId="3" pageNumber="316" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<emphasis id="B93BEAC51339FFD8FF7D0C233D98DE09" bold="true" box="[151,390,855,880]" pageId="3" pageNumber="316">Material examined.</emphasis>
<typeStatus id="54F488751339FFD8FE780C233DE1DE09" box="[402,511,855,880]" pageId="3" pageNumber="316" type="holotype">Holotype</typeStatus>
: female (7.8 ×
<quantity id="4CB79B321339FFD8FD570C233F04DE09" box="[701,794,855,880]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="3" pageNumber="316" unit="mm" value="6.8">6.8 mm</quantity>
) (QM W739),
<location id="8E90600C1339FFD8FC380C2338DCDE09" LSID="urn:lsid:plazi:treatment:03E687C1133BFFDCFF7D0EA238DDDFB4:8E90600C1339FFD8FC380C2338DCDE09" box="[978,1218,855,880]" country="Iceland" name="Whitsunday Passage" pageId="3" pageNumber="316" stateProvince="Queensland">Whitsunday Passage</location>
, Lindeman
<collectingCountry id="F35876471339FFD8FABB0C233986DE09" box="[1361,1432,855,880]" name="Iceland" pageId="3" pageNumber="316">Island</collectingCountry>
,
<collectingRegion id="498BF8351339FFD8FF7D0C083D00DEEC" box="[151,286,892,917]" country="Australia" name="Queensland" pageId="3" pageNumber="316">Queensland</collectingRegion>
</materialsCitation>
, Australia, coll. 1930s;
<materialsCitation id="3B273C8A1339FFD8FDC50C083926DEC1" ID-GBIF-Occurrence="1306554006" collectingDate="1987-03-25" collectorName="Home Beach &amp; P. J. F. Davie &amp; J. Short" country="Iceland" location="Queensland" pageId="3" pageNumber="316" specimenCount="2" specimenCount-male="2" stateProvince="Queensland" typeStatus="holotype">
<specimenCount id="9D49FD5E1339FFD8FDC50C083E99DEEC" box="[559,647,892,917]" pageId="3" pageNumber="316" type="male">2 males</specimenCount>
(6.4 × 6.0 mm, 5.7 ×
<quantity id="4CB79B321339FFD8FC600C083FFCDEED" box="[906,994,892,917]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="3" pageNumber="316" unit="mm" value="5.4">5.4 mm</quantity>
) (QM
<accessionNumber id="941CAB341339FFD8FBD90C083889DEED" box="[1075,1175,892,916]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W14482" pageId="3" pageNumber="316">W14482</accessionNumber>
), in subtidal branching coral,
<collectorName id="26BA53011339FFD8FF370CD43D6FDEC1" box="[221,369,927,952]" pageId="3" pageNumber="316">Home Beach</collectorName>
, Lindeman
<collectingCountry id="F35876471339FFD8FE1F0CEB3E22DEC1" box="[501,572,927,952]" name="Iceland" pageId="3" pageNumber="316">Island</collectingCountry>
, northeast
<collectingRegion id="498BF8351339FFD8FD5F0CEB3F22DEC1" box="[693,828,927,952]" country="Australia" name="Queensland" pageId="3" pageNumber="316">Queensland</collectingRegion>
, coll.
<collectorName id="26BA53011339FFD8FC950CD4381DDEC1" box="[895,1027,927,952]" pageId="3" pageNumber="316">P.J.F. Davie</collectorName>
&amp;
<collectorName id="26BA53011339FFD8FBCC0CD43861DEC1" box="[1062,1151,927,952]" pageId="3" pageNumber="316">J. Short</collectorName>
,
<date id="FFF110171339FFD8FB600CEB3926DEC1" box="[1162,1336,927,952]" pageId="3" pageNumber="316" value="1987-03-25">
<collectingDate id="EFB5E9FF1339FFD8FB600CEB3926DEC1" box="[1162,1336,927,952]" pageId="3" pageNumber="316" value="1987-03-25">25 March 1987</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C355655C1339FFD8FF2D0CB03D79D9A1" pageId="3" pageNumber="316" type="diagnosis">
<paragraph id="8BF036D71339FFD8FF2D0CB03D79D9A1" blockId="3.[151,1437,855,1997]" pageId="3" pageNumber="316">
<emphasis id="B93BEAC51339FFD8FF2D0CB03D5FDEA4" bold="true" box="[199,321,964,989]" pageId="3" pageNumber="316">Diagnosis.</emphasis>
Carapace with distinct median tubercle on protogastric region; sub-branchial regions gently convex, obliquely raised (
<figureCitation id="13742A521339FFD8FE500C933E1CD979" box="[442,514,999,1024]" captionStart="FIGURE 2" captionStartId="4.[151,250,1499,1521]" captionTargetBox="[151,1436,193,1477]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURE 2. Overall habitus. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; F, S. cristatus, male (12.5 × 10.1 mm) (NMMBCD 4065), Taiwan." httpUri="https://zenodo.org/record/266604/files/figure.png" pageId="3" pageNumber="316">Figs 2</figureCitation>
AC; 3AC); pararostral teeth long, clearly visible from dorsal view (
<figureCitation id="13742A521339FFD8FAF20C933943D979" box="[1304,1373,999,1024]" captionStart="FIGURE 3" captionStartId="5.[151,250,1912,1934]" captionTargetBox="[172,1414,399,1899]" captionTargetId="figure@5.[172,1414,399,1900]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of carapace. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan." httpUri="https://zenodo.org/record/266605/files/figure.png" pageId="3" pageNumber="316">Figs 3</figureCitation>
AC; 7A, B); eye relatively short (
<figureCitation id="13742A521339FFD8FE370B783E38D95D" box="[477,550,1036,1061]" captionStart="FIGURE 4" captionStartId="6.[151,250,1320,1342]" captionTargetBox="[150,1435,193,1297]" captionTargetId="figure@6.[150,1436,193,1298]" captionTargetPageId="6" captionText="FIGURE 4. Sargassocarcinus foliatus. A, D, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, C, E, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia. A, B, ventral view of cephalothorax; C, left cheliped; D, right fourth ambulatory leg; E, right fourth ambulatory leg (partially denuded). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266606/files/figure.png" pageId="3" pageNumber="316">Figs 4</figureCitation>
A; 7B); postorbital lobe low to almost undiscernible, confluent with wing-like lateral expansion (
<figureCitation id="13742A521339FFD8FE9D0B5B3DDCD931" box="[375,450,1071,1096]" captionStart="FIGURE 3" captionStartId="5.[151,250,1912,1934]" captionTargetBox="[172,1414,399,1899]" captionTargetId="figure@5.[172,1414,399,1900]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of carapace. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan." httpUri="https://zenodo.org/record/266605/files/figure.png" pageId="3" pageNumber="316">Fig. 3</figureCitation>
AC); lateral expansion lamelliform, with relatively deep, symmetrical concavity separating 2 subequal lobes, posterior margin of posterior lobe reaching beyond posterolateral margin of carapace as short cristate ridge, joining sub-branchial region (
<figureCitation id="13742A521339FFD8FCFF0B033F7AD9E9" box="[789,868,1143,1168]" captionStart="FIGURE 2" captionStartId="4.[151,250,1499,1521]" captionTargetBox="[151,1436,193,1477]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURE 2. Overall habitus. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; F, S. cristatus, male (12.5 × 10.1 mm) (NMMBCD 4065), Taiwan." httpUri="https://zenodo.org/record/266604/files/figure.png" pageId="3" pageNumber="316">Figs 2</figureCitation>
AC; 3AC). Ambulatory legs (notably merus) relatively longer (
<figureCitation id="13742A521339FFD8FE880BE83DB5D9CD" box="[354,427,1180,1205]" captionStart="FIGURE 2" captionStartId="4.[151,250,1499,1521]" captionTargetBox="[151,1436,193,1477]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURE 2. Overall habitus. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; F, S. cristatus, male (12.5 × 10.1 mm) (NMMBCD 4065), Taiwan." httpUri="https://zenodo.org/record/266604/files/figure.png" pageId="3" pageNumber="316">Figs 2</figureCitation>
AC; 4A, B, D, E; 7F). G1 relatively shorter, stouter, tapering bent distal part relatively short (
<figureCitation id="13742A521339FFD8FF340BCB3D3ED9A1" box="[222,288,1215,1240]" captionStart="FIGURE 9" captionStartId="12.[151,250,1890,1912]" captionTargetBox="[186,1414,302,1867]" captionTargetId="figure@12.[163,1421,297,1877]" captionTargetPageId="12" captionText="FIGURE 9. Gonopods. A E, Sargassocarcinus foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; F I, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; J N, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; O R, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan. A, F, J, O, left G 1 (ventral view); B, G, K, P, left G 1 (dorsal view); C, H, L, Q, distal view of left G 1 (ventral view); D, distal part of left G 1 (dorsal view); E, I, N, R, left G 2; M, distal half of right G 1 (ventral view). Scales: A, B, F, G, I, J, K, M, N, O, P, R = 0.5 mm; C E, H, L, Q = 0.1 mm." httpUri="https://zenodo.org/record/266612/files/figure.png" pageId="3" pageNumber="316">Fig. 9</figureCitation>
A
<emphasis id="B93BEAC51339FFD8FEDE0BCB3D5CD9A1" bold="true" box="[308,322,1215,1240]" pageId="3" pageNumber="316"></emphasis>
D).
</paragraph>
</subSubSection>
<subSubSection id="C355655C1339FFDCFF2D0B903DCEDF44" lastPageId="7" lastPageNumber="320" pageId="3" pageNumber="316" type="description">
<paragraph id="8BF036D71339FFD8FF2D0B903E52DAB5" blockId="3.[151,1437,855,1997]" pageId="3" pageNumber="316">
<emphasis id="B93BEAC51339FFD8FF2D0B903D47D984" bold="true" box="[199,345,1252,1277]" pageId="3" pageNumber="316">Description.</emphasis>
Carapace subhexagonal; wider than long; surfaces covered by short scale-like setae which appears reticulated; relatively smooth when denuded; regions not well defined, delimited by shallow grooves; proto-, metagastric regions elevated, each with median tubercle; cardiac region distinctly raised, with median tubercle; sub-branchial regions gently convex, forming obliquely raised regions (
<figureCitation id="13742A521339FFD8FB480A3B38E9D811" box="[1186,1271,1359,1384]" captionStart="FIGURE 2" captionStartId="4.[151,250,1499,1521]" captionTargetBox="[151,1436,193,1477]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURE 2. Overall habitus. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; F, S. cristatus, male (12.5 × 10.1 mm) (NMMBCD 4065), Taiwan." httpUri="https://zenodo.org/record/266604/files/figure.png" pageId="3" pageNumber="316">Figs 2</figureCitation>
AC; 3AC). Pseudorostrum short, triangular, visible in dorsal view, forming 2 low rounded lobes with gentle median concavity; distinct pararostral teeth present below pseudorostrum, directed obliquely anteriorly, gently bent downwards, with deep median cleft, clearly visible from dorsal view (
<figureCitation id="13742A521339FFD8FD1A0AC83F27D8AD" box="[752,825,1468,1493]" captionStart="FIGURE 2" captionStartId="4.[151,250,1499,1521]" captionTargetBox="[151,1436,193,1477]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURE 2. Overall habitus. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; F, S. cristatus, male (12.5 × 10.1 mm) (NMMBCD 4065), Taiwan." httpUri="https://zenodo.org/record/266604/files/figure.png" pageId="3" pageNumber="316">Figs 2</figureCitation>
AC; 3AC; 7A, B). Preorbital lobe large, triangular, directed anteriorly, separated from pseudorostrum by prominent cleft (
<figureCitation id="13742A521339FFD8FC3C0AAB383ED881" box="[982,1056,1503,1528]" captionStart="FIGURE 3" captionStartId="5.[151,250,1912,1934]" captionTargetBox="[172,1414,399,1899]" captionTargetId="figure@5.[172,1414,399,1900]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of carapace. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan." httpUri="https://zenodo.org/record/266605/files/figure.png" pageId="3" pageNumber="316">Figs 3</figureCitation>
AC; 7A). Postorbital lobe low, visible, confluent with lateral expansion in males (
<figureCitation id="13742A521339FFD8FD3A09703F0DDB65" box="[720,787,1540,1565]" captionStart="FIGURE 3" captionStartId="5.[151,250,1912,1934]" captionTargetBox="[172,1414,399,1899]" captionTargetId="figure@5.[172,1414,399,1900]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of carapace. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan." httpUri="https://zenodo.org/record/266605/files/figure.png" pageId="3" pageNumber="316">Fig. 3</figureCitation>
B, C); undiscernible in females (
<figureCitation id="13742A521339FFD8FB68097038DBDB65" box="[1154,1221,1540,1565]" captionStart="FIGURE 3" captionStartId="5.[151,250,1912,1934]" captionTargetBox="[172,1414,399,1899]" captionTargetId="figure@5.[172,1414,399,1900]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of carapace. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan." httpUri="https://zenodo.org/record/266605/files/figure.png" pageId="3" pageNumber="316">Fig. 3</figureCitation>
A). Hepatic, lateral regions strongly expanded laterally to form lamelliform, wing-like expansion; each expansion directed obliquely upwards with concave ventral surface, with 2 broadly triangular subequal lateral lobes, separated by relatively deep, broad, symmetrical concavity; posterior margin of posterior lobe reaching beyond posterolateral margin of carapace as short cristate ridge, joining raised oblique convexity on sub-branchial region (
<figureCitation id="13742A521339FFD8FAE309E0394BDBD5" box="[1289,1365,1684,1709]" captionStart="FIGURE 3" captionStartId="5.[151,250,1912,1934]" captionTargetBox="[172,1414,399,1899]" captionTargetId="figure@5.[172,1414,399,1900]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of carapace. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan." httpUri="https://zenodo.org/record/266605/files/figure.png" pageId="3" pageNumber="316">Fig. 3</figureCitation>
AC). Posterolateral margin sinuous, subcristate; posterior carapace margin sinuous, distinctly cristate, partially directed upwards (
<figureCitation id="13742A521339FFD8FEE209A83D4FDB8D" box="[264,337,1756,1781]" captionStart="FIGURE 2" captionStartId="4.[151,250,1499,1521]" captionTargetBox="[151,1436,193,1477]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURE 2. Overall habitus. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; F, S. cristatus, male (12.5 × 10.1 mm) (NMMBCD 4065), Taiwan." httpUri="https://zenodo.org/record/266604/files/figure.png" pageId="3" pageNumber="316">Figs 2</figureCitation>
AC; 3AC). Orbits small; supraorbital margin short, entire, concave; suborbital margin entire, gently crenulated; eye short, dorsoventrally flattened, fused to orbital margin, immobile (
<figureCitation id="13742A521339FFD8FB63098B38D1DA61" box="[1161,1231,1791,1816]" captionStart="FIGURE 3" captionStartId="5.[151,250,1912,1934]" captionTargetBox="[172,1414,399,1899]" captionTargetId="figure@5.[172,1414,399,1900]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of carapace. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; F, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan." httpUri="https://zenodo.org/record/266605/files/figure.png" pageId="3" pageNumber="316">Figs 3</figureCitation>
AC; 4A; 7A, B). Proepistome narrow, not spiniform or dentiform (
<figureCitation id="13742A521339FFD8FD0F08503F5ADA45" box="[741,836,1828,1853]" captionStart="FIGURE 7. A, B, D F" captionStartId="10.[151,250,1648,1670]" captionTargetBox="[171,1418,198,1614]" captionTargetId="figure@10.[150,1436,193,1627]" captionTargetPageId="10" captionText="FIGURE 7. A, B, D F, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; C, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; G, I, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; H, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; J, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; K, S. cristatus, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289). A, I, J, frontal margin and orbits; B, frontal margin, orbits, epistome, antennules and antennae; C, thoracic sternum and sternoabdominal cavity; D, inner view of left third maxilliped; E, outer view of left third maxilliped; F, G, dactylus of first ambulatory leg (denuded); K, ventral view of left eye. Scales: A C, F J = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/266610/files/figure.png" pageId="3" pageNumber="316">Fig. 7B</figureCitation>
). Antennular fossa large, subtriangular; antennules folding obliquely (
<figureCitation id="13742A521339FFD8FE8D08333DA1DA19" box="[359,447,1863,1888]" captionStart="FIGURE 7. A, B, D F" captionStartId="10.[151,250,1648,1670]" captionTargetBox="[171,1418,198,1614]" captionTargetId="figure@10.[150,1436,193,1627]" captionTargetPageId="10" captionText="FIGURE 7. A, B, D F, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; C, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; G, I, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; H, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; J, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; K, S. cristatus, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289). A, I, J, frontal margin and orbits; B, frontal margin, orbits, epistome, antennules and antennae; C, thoracic sternum and sternoabdominal cavity; D, inner view of left third maxilliped; E, outer view of left third maxilliped; F, G, dactylus of first ambulatory leg (denuded); K, ventral view of left eye. Scales: A C, F J = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/266610/files/figure.png" pageId="3" pageNumber="316">Fig. 7B</figureCitation>
). Basal antennal article large, subrectangular, completely fused to carapace without trace of sutures except near orbit; flagellum inserted medially (
<figureCitation id="13742A521339FFD8FCDB08183F93DAFD" box="[817,909,1900,1925]" captionStart="FIGURE 7. A, B, D F" captionStartId="10.[151,250,1648,1670]" captionTargetBox="[171,1418,198,1614]" captionTargetId="figure@10.[150,1436,193,1627]" captionTargetPageId="10" captionText="FIGURE 7. A, B, D F, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; C, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; G, I, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; H, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; J, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; K, S. cristatus, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289). A, I, J, frontal margin and orbits; B, frontal margin, orbits, epistome, antennules and antennae; C, thoracic sternum and sternoabdominal cavity; D, inner view of left third maxilliped; E, outer view of left third maxilliped; F, G, dactylus of first ambulatory leg (denuded); K, ventral view of left eye. Scales: A C, F J = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/266610/files/figure.png" pageId="3" pageNumber="316">Fig. 7B</figureCitation>
). Epistome transversely rectangular; posterior margin forming raised rim, sinuous, with median fissure; lateral edges of posterior margin of epistome, pterygostomial region raised (
<figureCitation id="13742A521339FFD8FE0D08C03E21DAB5" box="[487,575,1972,1997]" captionStart="FIGURE 7. A, B, D F" captionStartId="10.[151,250,1648,1670]" captionTargetBox="[171,1418,198,1614]" captionTargetId="figure@10.[150,1436,193,1627]" captionTargetPageId="10" captionText="FIGURE 7. A, B, D F, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; C, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; G, I, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; H, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; J, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; K, S. cristatus, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289). A, I, J, frontal margin and orbits; B, frontal margin, orbits, epistome, antennules and antennae; C, thoracic sternum and sternoabdominal cavity; D, inner view of left third maxilliped; E, outer view of left third maxilliped; F, G, dactylus of first ambulatory leg (denuded); K, ventral view of left eye. Scales: A C, F J = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/266610/files/figure.png" pageId="3" pageNumber="316">Fig. 7B</figureCitation>
).
</paragraph>
<caption id="DF30665F133EFFDFFF7D0AAF392CDB34" httpUri="https://zenodo.org/record/266604/files/figure.png" pageId="4" pageNumber="317" targetBox="[151,1436,193,1477]" targetPageId="4">
<paragraph id="8BF036D7133EFFDFFF7D0AAF392CDB34" blockId="4.[151,1436,1499,1613]" pageId="4" pageNumber="317">
<emphasis id="B93BEAC5133EFFDFFF7D0AAF3D06D889" bold="true" box="[151,280,1499,1521]" pageId="4" pageNumber="317">FIGURE 2.</emphasis>
Overall habitus. A,
<taxonomicName id="4C4F4D54133EFFDFFE170AA83F16D888" box="[509,776,1499,1521]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B93BEAC5133EFFDFFE170AA83F16D888" box="[509,776,1499,1521]" italics="true" pageId="4" pageNumber="317">Sargassocarcinus foliatus</emphasis>
</taxonomicName>
, holotype female (7.8 × 6.8 mm) (QM W739), Queensland, Australia; B,
<taxonomicName id="4C4F4D54133EFFDFFEF50A8E3D9BDB69" box="[287,389,1530,1552]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B93BEAC5133EFFDFFEF50A8E3D9BDB69" box="[287,389,1530,1552]" italics="true" pageId="4" pageNumber="317">S. foliatus</emphasis>
</taxonomicName>
, male (6.4 × 6.0 mm) (QM
<accessionNumber id="941CAB34133EFFDFFD4E0A8E3EE1DB69" box="[676,767,1530,1552]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W14482" pageId="4" pageNumber="317">W14482</accessionNumber>
), Queensland, Australia; C,
<taxonomicName id="4C4F4D54133EFFDFFBF40A8E389ADB69" box="[1054,1156,1530,1552]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B93BEAC5133EFFDFFBF40A8E389ADB69" box="[1054,1156,1530,1552]" italics="true" pageId="4" pageNumber="317">S. foliatus</emphasis>
</taxonomicName>
, male (5.7 × 5.4 mm) (QM
<accessionNumber id="941CAB34133EFFDFFF7D096C3CEFDB57" box="[151,241,1560,1582]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W14482" pageId="4" pageNumber="317">W14482</accessionNumber>
), Queensland, Australia; D,
<taxonomicName id="4C4F4D54133EFFDFFDFB096D3E61DB57" box="[529,639,1560,1582]" class="Malacostraca" family="Epialtidae" genus="Peltinia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="sublimis">
<emphasis id="B93BEAC5133EFFDFFDFB096D3E61DB57" box="[529,639,1560,1582]" italics="true" pageId="4" pageNumber="317">S. sublimis</emphasis>
</taxonomicName>
, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E,
<taxonomicName id="4C4F4D54133EFFDFFACF096D398BDB57" box="[1317,1429,1561,1582]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="B93BEAC5133EFFDFFACF096D398BDB57" box="[1317,1429,1561,1582]" italics="true" pageId="4" pageNumber="317">S. cristatus</emphasis>
</taxonomicName>
, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; F,
<taxonomicName id="4C4F4D54133EFFDFFD40094C3F04DB34" box="[682,794,1592,1613]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="317" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="B93BEAC5133EFFDFFD40094C3F04DB34" box="[682,794,1592,1613]" italics="true" pageId="4" pageNumber="317">S. cristatus</emphasis>
</taxonomicName>
, male (12.5 × 10.1 mm) (NMMBCD 4065), Taiwan.
</paragraph>
</caption>
<paragraph id="8BF036D7133EFFDFFF2D09003FE5DA1C" blockId="4.[151,1437,1652,2037]" pageId="4" pageNumber="317">
Third maxilliped with outer surfaces covered with short pubescence; relatively short, stout, completely covering buccal chamber when closed; basis separated from ischium by distinct suture; ischium slightly longer than broad, distal, proximal margins subparallel; inner margin with crista dentata, without median sulcus; merus subtriangular, proximal quarter folds under distal part of ischium, anteroexternal angle subauriculiform; carpus broad, inserted on inner subdistal margin of merus, articulating obliquely inwards; propodus, dactylus short; exopod stout, outer margin sinuous, distal third tapering, inner margin with low subdistal triangular tooth, flagellum long which extends almost across width of merus (
<figureCitation id="13742A52133EFFDFFCAB08383F94DA1C" box="[833,906,1868,1893]" captionStart="FIGURE 4" captionStartId="6.[151,250,1320,1342]" captionTargetBox="[150,1435,193,1297]" captionTargetId="figure@6.[150,1436,193,1298]" captionTargetPageId="6" captionText="FIGURE 4. Sargassocarcinus foliatus. A, D, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, C, E, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia. A, B, ventral view of cephalothorax; C, left cheliped; D, right fourth ambulatory leg; E, right fourth ambulatory leg (partially denuded). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266606/files/figure.png" pageId="4" pageNumber="317">Figs 4</figureCitation>
B; 7D, E).
</paragraph>
<paragraph id="8BF036D7133EFFDEFF2D08043957DC1D" blockId="4.[151,1437,1652,2037]" lastBlockId="5.[151,1437,151,357]" lastPageId="5" lastPageNumber="318" pageId="4" pageNumber="317">
Chelipeds relatively short; most of surface (except for fingers) covered with low pubescence, denser, longer on margins, forming “pseudo-cristae” (
<figureCitation id="13742A52133EFFDFFDC708E03E6FDAD4" box="[557,625,1940,1965]" captionStart="FIGURE 4" captionStartId="6.[151,250,1320,1342]" captionTargetBox="[150,1435,193,1297]" captionTargetId="figure@6.[150,1436,193,1298]" captionTargetPageId="6" captionText="FIGURE 4. Sargassocarcinus foliatus. A, D, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, C, E, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia. A, B, ventral view of cephalothorax; C, left cheliped; D, right fourth ambulatory leg; E, right fourth ambulatory leg (partially denuded). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266606/files/figure.png" pageId="4" pageNumber="317">Fig. 4</figureCitation>
AC). Basis, ischium, merus fused; basis-ischium short, suture between ischium and merus visible; merus long, trigonal in cross-section, with subdistal tooth on dorsal margin, rest of margin uneven to weakly spinous. Carpus subovate, short, outer margin with low cristae, dorsal median part with distinct concavity and low rounded tubercle behind it (
<figureCitation id="13742A52133FFFDEFCE40FE33F4CDDC9" box="[782,850,151,176]" captionStart="FIGURE 2" captionStartId="4.[151,250,1499,1521]" captionTargetBox="[151,1436,193,1477]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURE 2. Overall habitus. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; F, S. cristatus, male (12.5 × 10.1 mm) (NMMBCD 4065), Taiwan." httpUri="https://zenodo.org/record/266604/files/figure.png" pageId="5" pageNumber="318">Fig. 2</figureCitation>
AC). Chela with fingers shorter than palm; dorsal margin of palm with strong crista along entire length; ventral margin cristate along entire length to pollex; dactylus curved, dorsal margin distinctly flattened with lateral margins subcristate; pollex curved; cutting edges of fingers evenly lined with small teeth in smaller specimens (
<figureCitation id="13742A52133FFFDEFD010E703F2EDC65" box="[747,816,260,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1320,1342]" captionTargetBox="[150,1435,193,1297]" captionTargetId="figure@6.[150,1436,193,1298]" captionTargetPageId="6" captionText="FIGURE 4. Sargassocarcinus foliatus. A, D, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, C, E, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia. A, B, ventral view of cephalothorax; C, left cheliped; D, right fourth ambulatory leg; E, right fourth ambulatory leg (partially denuded). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266606/files/figure.png" pageId="5" pageNumber="318">Fig. 4</figureCitation>
AC); in large males, fingers arcuate, forming distinct gape when closed, distal part of cutting edges with teeth, median parts unarmed except for subproximal tooth on dactylus (
<figureCitation id="13742A52133FFFDEFEEE0E383D56DC1D" box="[260,328,332,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1320,1342]" captionTargetBox="[150,1435,193,1297]" captionTargetId="figure@6.[150,1436,193,1298]" captionTargetPageId="6" captionText="FIGURE 4. Sargassocarcinus foliatus. A, D, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, C, E, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia. A, B, ventral view of cephalothorax; C, left cheliped; D, right fourth ambulatory leg; E, right fourth ambulatory leg (partially denuded). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266606/files/figure.png" pageId="5" pageNumber="318">Fig. 4</figureCitation>
B, C); in females, chela relatively more slender, fingers not gaping even in adults (
<figureCitation id="13742A52133FFFDEFB080E38393BDC1D" box="[1250,1317,332,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1320,1342]" captionTargetBox="[150,1435,193,1297]" captionTargetId="figure@6.[150,1436,193,1298]" captionTargetPageId="6" captionText="FIGURE 4. Sargassocarcinus foliatus. A, D, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, C, E, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia. A, B, ventral view of cephalothorax; C, left cheliped; D, right fourth ambulatory leg; E, right fourth ambulatory leg (partially denuded). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266606/files/figure.png" pageId="5" pageNumber="318">Fig. 4</figureCitation>
A).
</paragraph>
<caption id="DF30665F133FFFDEFF7D080C38F5DA93" httpUri="https://zenodo.org/record/266605/files/figure.png" pageId="5" pageNumber="318" targetBox="[172,1414,399,1899]" targetPageId="5">
<paragraph id="8BF036D7133FFFDEFF7D080C38F5DA93" blockId="5.[151,1436,1912,2026]" pageId="5" pageNumber="318">
<emphasis id="B93BEAC5133FFFDEFF7D080C3D0CDAF4" bold="true" box="[151,274,1912,1934]" pageId="5" pageNumber="318">FIGURE 3.</emphasis>
Dorsal view of carapace. A,
<taxonomicName id="4C4F4D54133FFFDEFDD0080D3F20DAF7" box="[570,830,1912,1934]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B93BEAC5133FFFDEFDD0080D3F20DAF7" box="[570,830,1912,1934]" italics="true" pageId="5" pageNumber="318">Sargassocarcinus foliatus</emphasis>
</taxonomicName>
, holotype female (7.8 × 6.8 mm) (QM W739), Queensland, Australia; B,
<taxonomicName id="4C4F4D54133FFFDEFEF508E33D9BDAD5" box="[287,389,1942,1964]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B93BEAC5133FFFDEFEF508E33D9BDAD5" box="[287,389,1942,1964]" italics="true" pageId="5" pageNumber="318">S. foliatus</emphasis>
</taxonomicName>
, male (6.4 × 6.0 mm) (QM
<accessionNumber id="941CAB34133FFFDEFD4E08E23EE1DAD5" box="[676,767,1942,1964]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W14482" pageId="5" pageNumber="318">W14482</accessionNumber>
), Queensland, Australia; C,
<taxonomicName id="4C4F4D54133FFFDEFBF408E3389ADAD5" box="[1054,1156,1942,1964]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B93BEAC5133FFFDEFBF408E3389ADAD5" box="[1054,1156,1942,1964]" italics="true" pageId="5" pageNumber="318">S. foliatus</emphasis>
</taxonomicName>
, male (5.7 × 5.4 mm) (QM
<accessionNumber id="941CAB34133FFFDEFF7D08C13CEFDAB2" box="[151,241,1973,1995]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W14482" pageId="5" pageNumber="318">W14482</accessionNumber>
), Queensland, Australia; D,
<taxonomicName id="4C4F4D54133FFFDEFDFB08C23E61DAB2" box="[529,639,1973,1995]" class="Malacostraca" family="Epialtidae" genus="Peltinia" higherTaxonomySource="CoL" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="sublimis">
<emphasis id="B93BEAC5133FFFDEFDFB08C23E61DAB2" box="[529,639,1973,1995]" italics="true" pageId="5" pageNumber="318">S. sublimis</emphasis>
</taxonomicName>
, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E,
<taxonomicName id="4C4F4D54133FFFDEFACF08C2398BDAB2" box="[1317,1429,1974,1995]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="B93BEAC5133FFFDEFACF08C2398BDAB2" box="[1317,1429,1974,1995]" italics="true" pageId="5" pageNumber="318">S. cristatus</emphasis>
</taxonomicName>
, male (8.2 × 7.4 mm) (USNM 48248), Japan; F,
<taxonomicName id="4C4F4D54133FFFDEFD9308A03EF4DA93" box="[633,746,2004,2026]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="318" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="B93BEAC5133FFFDEFD9308A03EF4DA93" box="[633,746,2004,2026]" italics="true" pageId="5" pageNumber="318">S. cristatus</emphasis>
</taxonomicName>
, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan.
</paragraph>
</caption>
<caption id="DF30665F133CFFDDFF7D0A5C382AD802" httpUri="https://zenodo.org/record/266606/files/figure.png" pageId="6" pageNumber="319" targetBox="[150,1435,193,1297]" targetPageId="6">
<paragraph id="8BF036D7133CFFDDFF7D0A5C382AD802" blockId="6.[151,1436,1320,1403]" pageId="6" pageNumber="319">
<emphasis id="B93BEAC5133CFFDDFF7D0A5C3D0AD844" bold="true" box="[151,276,1320,1342]" pageId="6" pageNumber="319">FIGURE 4.</emphasis>
<taxonomicName id="4C4F4D54133CFFDDFEF60A5D3E3DD847" box="[284,547,1320,1342]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="319" phylum="Arthropoda" rank="species" species="foliatus">
<emphasis id="B93BEAC5133CFFDDFEF60A5D3E3DD847" box="[284,547,1320,1342]" italics="true" pageId="6" pageNumber="319">Sargassocarcinus foliatus</emphasis>
</taxonomicName>
. A, D, holotype female (7.8 × 6.8 mm) (QM W739), Queensland, Australia; B, C, E, male (6.4 × 6.0 mm) (QM
<accessionNumber id="941CAB34133CFFDDFE590A323E10D825" box="[435,526,1350,1372]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/W14482" pageId="6" pageNumber="319">W14482</accessionNumber>
), Queensland, Australia. A, B, ventral view of cephalothorax; C, left cheliped; D, right fourth ambulatory leg; E, right fourth ambulatory leg (partially denuded). Scales = 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BF036D7133CFFDDFF2D0AD63988DB86" blockId="6.[151,1437,1442,2007]" pageId="6" pageNumber="319">
Ambulatory legs similar to those in
<taxonomicName id="4C4F4D54133CFFDDFDBD0AD73F02D8C3" box="[599,796,1443,1466]" class="Malacostraca" family="Epialtidae" genus="Sargassocarcinus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="319" phylum="Arthropoda" rank="genus">
<emphasis id="B93BEAC5133CFFDDFDBD0AD73F02D8C3" box="[599,796,1443,1466]" italics="true" pageId="6" pageNumber="319">Sargassocarcinus</emphasis>
</taxonomicName>
species, short, stout; leg decrease progressively in length posteriorly; covered with clavate setae, forming pubescence which is densest, expanded along dorsal margins of merus, carpus, propodus foliose, forming “pseudo-cristae”; surfaces generally smooth when denuded (
<figureCitation id="13742A52133CFFDDFAF20A9E3941DB7A" box="[1304,1375,1514,1539]" captionStart="FIGURE 2" captionStartId="4.[151,250,1499,1521]" captionTargetBox="[151,1436,193,1477]" captionTargetId="figure@4.[151,1436,193,1478]" captionTargetPageId="4" captionText="FIGURE 2. Overall habitus. A, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; C, S. foliatus, male (5.7 × 5.4 mm) (QM W 14482), Queensland, Australia; D, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; E, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; F, S. cristatus, male (12.5 × 10.1 mm) (NMMBCD 4065), Taiwan." httpUri="https://zenodo.org/record/266604/files/figure.png" pageId="6" pageNumber="319">Figs 2</figureCitation>
AC; 4A, B, D, E; 7F). Merus laterally flattened, margins not cristate, appears to be so because of dense pubescence; dorsal margin uneven, may be lined with small scattered granules, with large, rounded distal tooth; ventral margin uneven, may be lined with scattered granules (
<figureCitation id="13742A52133CFFDDFD4B09223EF4DB16" box="[673,746,1622,1647]" captionStart="FIGURE 4" captionStartId="6.[151,250,1320,1342]" captionTargetBox="[150,1435,193,1297]" captionTargetId="figure@6.[150,1436,193,1298]" captionTargetPageId="6" captionText="FIGURE 4. Sargassocarcinus foliatus. A, D, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, C, E, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia. A, B, ventral view of cephalothorax; C, left cheliped; D, right fourth ambulatory leg; E, right fourth ambulatory leg (partially denuded). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266606/files/figure.png" pageId="6" pageNumber="319">Figs 4</figureCitation>
D, E; 7F). Carpus with tooth on dorsal margin, appears larger because of pubescence before denuding (
<figureCitation id="13742A52133CFFDDFDB5090E3EB8DBEA" box="[607,678,1658,1683]" captionStart="FIGURE 4" captionStartId="6.[151,250,1320,1342]" captionTargetBox="[150,1435,193,1297]" captionTargetId="figure@6.[150,1436,193,1298]" captionTargetPageId="6" captionText="FIGURE 4. Sargassocarcinus foliatus. A, D, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, C, E, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia. A, B, ventral view of cephalothorax; C, left cheliped; D, right fourth ambulatory leg; E, right fourth ambulatory leg (partially denuded). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266606/files/figure.png" pageId="6" pageNumber="319">Figs 4</figureCitation>
D, E; 7F). Propodus subrectangular; dorsal margin with 2 low lobes, appears larger because of pubescence before denuding (
<figureCitation id="13742A52133CFFDDFCF809EA3F45DBCE" box="[786,859,1694,1719]" captionStart="FIGURE 4" captionStartId="6.[151,250,1320,1342]" captionTargetBox="[150,1435,193,1297]" captionTargetId="figure@6.[150,1436,193,1298]" captionTargetPageId="6" captionText="FIGURE 4. Sargassocarcinus foliatus. A, D, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, C, E, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia. A, B, ventral view of cephalothorax; C, left cheliped; D, right fourth ambulatory leg; E, right fourth ambulatory leg (partially denuded). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266606/files/figure.png" pageId="6" pageNumber="319">Figs 4</figureCitation>
D, E; 7F). Dactylo-propodal articulation, lock well developed, prominent; dactylus covered with low pubescence except for pectinated tip, gently falcate, ventral margin lined with pair of short subdistal spines, usually with several small denticles behind them (
<figureCitation id="13742A52133CFFDDFB3509923939DB86" box="[1247,1319,1766,1791]" captionStart="FIGURE 4" captionStartId="6.[151,250,1320,1342]" captionTargetBox="[150,1435,193,1297]" captionTargetId="figure@6.[150,1436,193,1298]" captionTargetPageId="6" captionText="FIGURE 4. Sargassocarcinus foliatus. A, D, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, C, E, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia. A, B, ventral view of cephalothorax; C, left cheliped; D, right fourth ambulatory leg; E, right fourth ambulatory leg (partially denuded). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266606/files/figure.png" pageId="6" pageNumber="319">Figs 4</figureCitation>
D, E; 7F).
</paragraph>
<paragraph id="8BF036D7133CFFDCFF2D087E3E32DD81" blockId="6.[151,1437,1442,2007]" lastBlockId="7.[151,1437,151,717]" lastPageId="7" lastPageNumber="320" pageId="6" pageNumber="319">
Male thoracic sternum wide, covered with low pubescence; smooth when denuded; sternites 1
<emphasis id="B93BEAC5133CFFDDFB11087E3917DA5A" bold="true" box="[1275,1289,1802,1827]" pageId="6" pageNumber="319"></emphasis>
4 completely fused except for low, rounded ridge demarcating sternites 3, 4; surface of sternites 1
<emphasis id="B93BEAC5133CFFDDFB86085A3864DA3E" bold="true" box="[1132,1146,1838,1863]" pageId="6" pageNumber="319"></emphasis>
3 distinctly concave; low curved rim demarcating sterno-abdominal cavity; sterno-abdominal cavity reaching to just before demarcation between sternites 3 and 4, with prominent rim on sternite 4; prominent tubercle of male abdominal locking mechanism on distal edge of sternite 5 (
<figureCitation id="13742A52133CFFDDFD8A08EE3EBBDACA" box="[608,677,1946,1971]" captionStart="FIGURE 7. A, B, D F" captionStartId="10.[151,250,1648,1670]" captionTargetBox="[171,1418,198,1614]" captionTargetId="figure@10.[150,1436,193,1627]" captionTargetPageId="10" captionText="FIGURE 7. A, B, D F, Sargassocarcinus foliatus, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; C, S. foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; G, I, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; H, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; J, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; K, S. cristatus, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289). A, I, J, frontal margin and orbits; B, frontal margin, orbits, epistome, antennules and antennae; C, thoracic sternum and sternoabdominal cavity; D, inner view of left third maxilliped; E, outer view of left third maxilliped; F, G, dactylus of first ambulatory leg (denuded); K, ventral view of left eye. Scales: A C, F J = 1.0 mm; D, E = 0.5 mm." httpUri="https://zenodo.org/record/266610/files/figure.png" pageId="6" pageNumber="319">Fig. 7</figureCitation>
C). Penis short, coxal, coming out below condyle. Female thoracic sternum conspicuously wide, sterno-abdominal cavity round, deeply concave, with low anterior rim, completely covered by abdomen (
<figureCitation id="13742A52133DFFDCFE720FE33DF5DDC9" box="[408,491,151,176]" captionStart="FIGURE 10" captionStartId="13.[151,250,1446,1468]" captionTargetBox="[151,1435,193,1424]" captionTargetId="figure@13.[151,1436,193,1425]" captionTargetPageId="13" captionText="FIGURE 10. Sargassocarcinus cristatus. A, B, female (10.8 × 8.9 mm) (NMMBCD 4065), Taiwan; C, D, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289), Taiwan. A, overall habitus; B, ventral view of cephalothorax showing buccal cavity, epistome, eyes and abdomen; D, sterno-abdominal cavity showing vulvae. Abbreviation: v, vulvae." httpUri="https://zenodo.org/record/266613/files/figure.png" pageId="7" pageNumber="320">Fig. 10</figureCitation>
BD). Sternites medially interrupted; sternite 8 separated into 2 halves by median gap. Vulva with operculum, positioned on raised mount on sternite 6, produced obliquely forwards, pushing suture between sternites 5 and 6 anteriorly.
</paragraph>
<paragraph id="8BF036D7133DFFDCFF2D0E703DCEDCA9" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="320">
Surfaces of male, female abdomens covered by low pubescence, smooth when denuded. Male somites 1, 2 transversely narrow, free, reaching bases of coxae of fifth ambulatory leg, completely covering sternite 8 when closed; somites 3
<emphasis id="B93BEAC5133DFFDCFEB30E383D79DC1C" bold="true" box="[345,359,332,357]" pageId="7" pageNumber="320"></emphasis>
5 separated by distinct sutures, immobile; somite 6 subrectangular with convex lateral margins; telson triangular with convex lateral margins (
<figureCitation id="13742A52133DFFDCFD710E1B3EEBDCF1" box="[667,757,367,392]" captionStart="FIGURE 8. A" captionStartId="11.[151,250,1364,1385]" captionTargetBox="[162,1417,349,1329]" captionTargetId="figure@11.[150,1436,337,1343]" captionTargetPageId="11" captionText="FIGURE 8. A, Sargassocarcinus foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; B, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; C, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; D, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan; E H, S. cristatus, ovigerous female (16.2 × 12.1 mm) (ZRC 2015.289), Taiwan. A D, male abdomen (somites 3 6 and telson); E, female fused abdominal somites 4 6 and telson; F, female abdominal somites 2 4; G, female abdominal somite 1; H, female vulvae (arrow points to buccal cavity). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266611/files/figure.png" pageId="7" pageNumber="320">Fig. 8A</figureCitation>
). Female somites 13 free, reaching to bases of coxae of last ambulatory legs; somites 46 fused, with sutures barely or not visible in adults, forming domed structure; telson semicircular, free (
<figureCitation id="13742A52133DFFDCFE830EC33DB2DCA9" box="[361,428,439,464]" captionStart="FIGURE 4" captionStartId="6.[151,250,1320,1342]" captionTargetBox="[150,1435,193,1297]" captionTargetId="figure@6.[150,1436,193,1298]" captionTargetPageId="6" captionText="FIGURE 4. Sargassocarcinus foliatus. A, D, holotype female (7.8 × 6.8 mm) (QM W 739), Queensland, Australia; B, C, E, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia. A, B, ventral view of cephalothorax; C, left cheliped; D, right fourth ambulatory leg; E, right fourth ambulatory leg (partially denuded). Scales = 1.0 mm." httpUri="https://zenodo.org/record/266606/files/figure.png" pageId="7" pageNumber="320">Fig. 4</figureCitation>
A).
</paragraph>
<paragraph id="8BF036D7133DFFDCFF2D0EA838D9DF61" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="320">
G1 elongated, gently sinuous, distally bent outwards, distal part dilated but tapering to sharp tip; groove for G2 ventrally positioned (
<figureCitation id="13742A52133DFFDCFE6F0E8B3DD9DF61" box="[389,455,511,536]" captionStart="FIGURE 9" captionStartId="12.[151,250,1890,1912]" captionTargetBox="[186,1414,302,1867]" captionTargetId="figure@12.[163,1421,297,1877]" captionTargetPageId="12" captionText="FIGURE 9. Gonopods. A E, Sargassocarcinus foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; F I, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; J N, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; O R, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan. A, F, J, O, left G 1 (ventral view); B, G, K, P, left G 1 (dorsal view); C, H, L, Q, distal view of left G 1 (ventral view); D, distal part of left G 1 (dorsal view); E, I, N, R, left G 2; M, distal half of right G 1 (ventral view). Scales: A, B, F, G, I, J, K, M, N, O, P, R = 0.5 mm; C E, H, L, Q = 0.1 mm." httpUri="https://zenodo.org/record/266612/files/figure.png" pageId="7" pageNumber="320">Fig. 9</figureCitation>
A
<emphasis id="B93BEAC5133DFFDCFE360E8B3DF4DF61" bold="true" box="[476,490,511,536]" pageId="7" pageNumber="320"></emphasis>
D). G2 short, distal segment very short, subspatuliform (
<figureCitation id="13742A52133DFFDCFB890E8B38B8DF61" box="[1123,1190,511,536]" captionStart="FIGURE 9" captionStartId="12.[151,250,1890,1912]" captionTargetBox="[186,1414,302,1867]" captionTargetId="figure@12.[163,1421,297,1877]" captionTargetPageId="12" captionText="FIGURE 9. Gonopods. A E, Sargassocarcinus foliatus, male (6.4 × 6.0 mm) (QM W 14482), Queensland, Australia; F I, S. sublimis, holotype male (13.3 × 10.8 mm) (USNM 48247), Philippines; J N, S. cristatus, male (8.2 × 7.4 mm) (USNM 48248), Japan; O R, S. cristatus, male (14.4 × 11.7 mm) (ZRC 2015.288), Taiwan. A, F, J, O, left G 1 (ventral view); B, G, K, P, left G 1 (dorsal view); C, H, L, Q, distal view of left G 1 (ventral view); D, distal part of left G 1 (dorsal view); E, I, N, R, left G 2; M, distal half of right G 1 (ventral view). Scales: A, B, F, G, I, J, K, M, N, O, P, R = 0.5 mm; C E, H, L, Q = 0.1 mm." httpUri="https://zenodo.org/record/266612/files/figure.png" pageId="7" pageNumber="320">Fig. 9</figureCitation>
E).
</paragraph>
<paragraph id="8BF036D7133DFFDCFF2D0D503DCEDF44" blockId="7.[151,1437,151,717]" box="[199,464,548,573]" pageId="7" pageNumber="320">
<emphasis id="B93BEAC5133DFFDCFF2D0D503D4DDF44" bold="true" box="[199,339,548,573]" pageId="7" pageNumber="320">Live colour.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C355655C133DFFDCFF2D0D333EFBDFD1" pageId="7" pageNumber="320" type="discussion">
<paragraph id="8BF036D7133DFFDCFF2D0D333EFBDFD1" blockId="7.[151,1437,151,717]" pageId="7" pageNumber="320">
<emphasis id="B93BEAC5133DFFDCFF2D0D333D25DF19" bold="true" box="[199,315,583,608]" pageId="7" pageNumber="320">Remarks.</emphasis>
<bibRefCitation id="EFDE4B26133DFFDCFEAE0D333DCADF19" author="Ward" box="[324,468,583,608]" pageId="7" pageNumber="320" refString="Ward, M. (1936) Crustacea Brachyura from the coasts of Queensland. Memoirs of Queensland Museum, 11, 1 - 13." type="journal article" year="1936">Ward (1936)</bibRefCitation>
established this species from nine specimens. One female (carapace width 8.0 mm) was designated as the
<typeStatus id="54F48875133DFFDCFE710D183E1EDFFC" box="[411,512,620,645]" pageId="7" pageNumber="320" type="holotype">holotype</typeStatus>
; making the remaining four females (carapace widths
<quantity id="4CB79B32133DFFDCFB9C0D183934DFFD" box="[1142,1322,620,645]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="10.5" metricValueMin="8.5" pageId="7" pageNumber="320" unit="mm" value="9.5" valueMax="10.5" valueMin="8.5">8.5 to 10.5 mm</quantity>
) and four males (carapace widths
<quantity id="4CB79B32133DFFDCFE490DFB3E24DFD1" box="[419,570,655,680]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" metricValueMax="10.0" metricValueMin="7.5" pageId="7" pageNumber="320" unit="mm" value="8.75" valueMax="10.0" valueMin="7.5">7.5 to 10 mm</quantity>
) the
<typeStatus id="54F48875133DFFDCFD980DE43EFFDFD1" box="[626,737,656,680]" pageId="7" pageNumber="320" type="paratype">paratypes</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C355655C133DFFDCFF2D0DC038DDDFB4" box="[199,1219,692,717]" pageId="7" pageNumber="320" type="distribution">
<paragraph id="8BF036D7133DFFDCFF2D0DC038DDDFB4" blockId="7.[151,1437,151,717]" box="[199,1219,692,717]" pageId="7" pageNumber="320">
<emphasis id="B93BEAC5133DFFDCFF2D0DC03D7EDFB4" bold="true" box="[199,352,692,717]" pageId="7" pageNumber="320">Distribution.</emphasis>
Known only from the
<typeStatus id="54F48875133DFFDCFD8A0DC13E8EDFB4" box="[608,656,693,717]" pageId="7" pageNumber="320">type</typeStatus>
locality, Lindeman
<collectingCountry id="F3587647133DFFDCFC9A0DC03FA9DFB4" box="[880,951,692,717]" name="Iceland" pageId="7" pageNumber="320">Island</collectingCountry>
,
<collectingRegion id="498BF835133DFFDCFC280DC03857DFB4" box="[962,1097,692,717]" country="Australia" name="Queensland" pageId="7" pageNumber="320">Queensland</collectingRegion>
,
<collectingCountry id="F3587647133DFFDCFBBF0DC038A1DFB4" box="[1109,1215,692,717]" name="Australia" pageId="7" pageNumber="320">Australia</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>