227 lines
19 KiB
XML
227 lines
19 KiB
XML
<document ID-DOI="10.11646/zootaxa.3219.1.1" ID-GBIF-Dataset="3fa772dc-c5b2-40c9-b28e-cd067ca9a3f6" ID-ISSN="1175-5326" ID-Zenodo-Dep="280223" checkinTime="1460447300036" checkinUser="plazi" docAuthor="Fikáček, Martin, Jia, Fenglong & Prokin, Alexander" docDate="2012" docId="03E687BAFFE3F337FF5EFD560DC5C131" docLanguage="en" docName="zt03219p053.pdf" docOrigin="Zootaxa 3219" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.3:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="3" docTitle="Australocyon keralensis Hebauer 2002, comb. nov." docType="treatment" docVersion="13" lastPageNumber="48" masterDocId="FFDFFFC2FFCCF318FFC9FFDB0C07C424" masterDocTitle="A review of the Asian species of the genus Pachysternum (Coleoptera: Hydrophilidae: Sphaeridiinae)" masterLastPageNumber="53" masterPageNumber="1" pageNumber="48" updateTime="1678199402054" updateUser="tatiana">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the Asian species of the genus Pachysternum (Coleoptera: Hydrophilidae: Sphaeridiinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fikáček, Martin</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jia, Fenglong</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Prokin, Alexander</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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||
<mods:date>2012</mods:date>
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||
<mods:detail type="pubDate">
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||
<mods:number>2012-03-02</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>3219</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>53</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.3219.1.1</mods:identifier>
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<mods:identifier type="GBIF-Dataset">3fa772dc-c5b2-40c9-b28e-cd067ca9a3f6</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">280223</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6180450" ID-GBIF-Taxon="119509777" ID-Zenodo-Dep="6180450" LSID="urn:lsid:plazi:treatment:03E687BAFFE3F337FF5EFD560DC5C131" httpUri="http://treatment.plazi.org/id/03E687BAFFE3F337FF5EFD560DC5C131" lastPageNumber="48" pageId="47" pageNumber="48">
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<subSubSection box="[151,797,653,679]" pageId="47" pageNumber="48" type="nomenclature">
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<paragraph blockId="47.[151,797,653,679]" box="[151,797,653,679]" pageId="47" pageNumber="48">
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<heading bold="true" box="[151,797,653,679]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
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<emphasis bold="true" box="[151,797,653,679]" pageId="47" pageNumber="48">
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<taxonomicName authority="Hebauer, 2002" authorityName="Hebauer" authorityYear="2002" box="[151,649,653,679]" class="Insecta" family="Hydrophilidae" genus="Australocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="keralensis" status="comb. nov.">
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<emphasis bold="true" box="[151,441,653,679]" italics="true" pageId="47" pageNumber="48">Australocyon keralensis</emphasis>
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(
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<bibRefCitation author="Hebauer, F." box="[456,641,653,679]" pageId="47" pageNumber="48" pagination="3 - 24" refId="ref42136" refString="Hebauer, F. (2002 a) New Hydrophilidae of the Old World (Coleoptera, Hydrophiloidea). Acta Coleopterologica 18, 3, 3 - 24." type="journal article" year="2002">
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Hebauer,
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<number box="[579,641,653,679]" pageId="47" pageNumber="48" value="2002.0">2002</number>
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</bibRefCitation>
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||
)
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</taxonomicName>
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,
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<taxonomicNameLabel box="[662,797,653,679]" pageId="47" pageNumber="48">comb. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection pageId="47" pageNumber="48" type="reference_group">
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<paragraph blockId="47.[151,827,722,779]" box="[151,610,722,744]" pageId="47" pageNumber="48">
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<treatmentCitationGroup box="[151,610,722,744]" pageId="47" pageNumber="48">
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<taxonomicName authority="Hebauer, 2002a: 16" authorityName="Hebauer" authorityPageNumber="16" authorityYear="2002" box="[151,606,722,744]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="keralense">
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<emphasis box="[151,399,722,744]" italics="true" pageId="47" pageNumber="48">Pachysternum keralense</emphasis>
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<treatmentCitation author="Hebauer" box="[405,606,722,744]" page="16" pageId="47" pageNumber="48" year="2002">
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<bibRefCitation author="Hebauer" box="[405,606,722,744]" firstAuthor="Hebauer" page="16" pageId="47" pageNumber="48" pagination="3 - 24" refId="ref42136" refString="Hebauer, F. (2002 a) New Hydrophilidae of the Old World (Coleoptera, Hydrophiloidea). Acta Coleopterologica 18, 3, 3 - 24." type="journal article" year="2002" yearSuffix="a">
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Hebauer,
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<number box="[503,557,723,744]" pageId="47" pageNumber="48" value="2002.0">2002</number>
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a:
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<number box="[578,606,722,744]" pageId="47" pageNumber="48" value="16.0">16</number>
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</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="47.[151,827,722,779]" box="[151,827,757,779]" pageId="47" pageNumber="48">
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<treatmentCitationGroup box="[151,827,757,779]" pageId="47" pageNumber="48">
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<taxonomicName authority="Short & Hebauer" authorityName="Short & Hebauer" authorityYear="2006" box="[151,823,757,779]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="keralense">
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<emphasis box="[151,399,757,779]" italics="true" pageId="47" pageNumber="48">Pachysternum keralense</emphasis>
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:
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<bibRefCitation author="Short" box="[412,823,757,779]" pageId="47" pageNumber="48" pagination="315" refId="ref43527" refString="Short, A. E. Z. & Hebauer, F. (2006) World catalogue of Hydrophiloidea - additions and corrections, 1 (1999 - 2005) (Coleoptera). Koleopterologische Rundschau 76, 315 - 359." type="journal article" year="2006">
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<treatmentCitation box="[412,704,757,779]" pageId="47" pageNumber="48">
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Short & Hebauer (
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<number box="[598,654,757,779]" pageId="47" pageNumber="48" value="2006.0">2006</number>
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:
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<number box="[664,704,757,778]" pageId="47" pageNumber="48" value="349.0">349</number>
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</treatmentCitation>
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, catalogue)
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</bibRefCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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||
</subSubSection>
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<subSubSection pageId="47" pageNumber="48" type="materials_examined">
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<paragraph blockId="47.[151,1437,826,1301]" pageId="47" pageNumber="48">
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||
<emphasis bold="true" box="[151,447,826,851]" pageId="47" pageNumber="48">
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<typeStatus box="[151,210,827,851]" pageId="47" pageNumber="48">Type</typeStatus>
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material examined.
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="3499776309" collectingDate="1989-09-02" collectionCode="NMPC" collectorName="A. Riedel" country="India" location="Peryar W. L. S." municipality="Thekkady" pageId="47" pageNumber="48" specimenCount="1" stateProvince="Kerala" typeStatus="paratype">
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<typeStatus box="[455,553,827,851]" pageId="47" pageNumber="48">Paratype</typeStatus>
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||
:
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<specimenCount box="[568,647,826,850]" pageId="47" pageNumber="48" type="generic">
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<number box="[568,582,826,850]" pageId="47" pageNumber="48" value="1.0">1</number>
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spec.
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</specimenCount>
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||
(
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<collectionCode box="[661,747,826,850]" collectionName="Czech Republic, Prague, National Museum (Natural History)" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="47" pageNumber="48">NMPC</collectionCode>
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): “
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<collectingCountry box="[777,880,826,850]" name="India" pageId="47" pageNumber="48">INDIEN</collectingCountry>
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:
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<collectingRegion box="[891,966,826,850]" country="India" name="Kerala" pageId="47" pageNumber="48">Kerala</collectingRegion>
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/
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<collectingMunicipality box="[988,1101,826,850]" pageId="47" pageNumber="48">Thekkady</collectingMunicipality>
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,
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<location LSID="urn:lsid:plazi:treatment:03E687BAFFE3F337FF5EFD560DC5C131:8E906077FFE3F337FB91FCE00902C776" box="[1112,1285,826,851]" country="India" municipality="Thekkady" name="Peryar W. L. S." pageId="47" pageNumber="48" stateProvince="Kerala">Peryar W. L. S.</location>
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/
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<date box="[1308,1421,826,850]" pageId="47" pageNumber="48" value="1989-09-02">
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<collectingDate box="[1308,1421,826,850]" pageId="47" pageNumber="48" value="1989-09-02">
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<number box="[1308,1353,826,850]" pageId="47" pageNumber="48" value="2.9">2. 9</number>
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.
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<number box="[1365,1421,826,850]" pageId="47" pageNumber="48" value="1989.0">1989</number>
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</collectingDate>
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</date>
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/ leg.
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<collectorName box="[199,308,861,886]" pageId="47" pageNumber="48">A. Riedel</collectorName>
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”
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="47" pageNumber="48" type="discussion">
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||
<paragraph blockId="47.[151,1437,826,1301]" pageId="47" pageNumber="48">
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<emphasis bold="true" box="[201,397,895,920]" pageId="47" pageNumber="48">Taxonomic note.</emphasis>
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The species bears all diagnostic characters of the
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<taxonomicName box="[958,1276,896,920]" class="Insecta" family="Hydrophilidae" genus="Australocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pilocnemoides">
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<emphasis box="[958,1276,896,920]" italics="true" pageId="47" pageNumber="48">Australocyon pilocnemoides</emphasis>
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</taxonomicName>
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species group mentioned by
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<bibRefCitation author="Hansen" box="[311,479,930,954]" firstAuthor="Hansen" pageId="47" pageNumber="48" pagination="53 - 84" refId="ref41916" refString="Hansen, M. (2003) Discovery of Australocyon Hansen and Pilocnema Hansen (Coleoptera, Hydrophilidae) outside the Australian region. In: Cuccudoro, G. & Leschen, R. A. B. (eds.) Systematics of Coleoptera: papers celebrating the retirement of Ivan Lobl. Memoirs on Entomology International 17, 53 - 84." type="journal article" year="2003">
|
||
Hansen (
|
||
<number box="[410,471,930,954]" pageId="47" pageNumber="48" value="2003.0">2003</number>
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||
)
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||
</bibRefCitation>
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||
, and moreover differs from
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<taxonomicName box="[794,954,931,955]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
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<emphasis box="[794,954,931,955]" italics="true" pageId="47" pageNumber="48">Pachysternum</emphasis>
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</taxonomicName>
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species by the antennal grooves not reaching the lateral margin of hypomeron, and metaventrite lacking any trace of the femoral lines. Morphology of its genitalia (see
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<bibRefCitation author="Hebauer" box="[311,487,999,1023]" firstAuthor="Hebauer" pageId="47" pageNumber="48" pagination="3 - 24" refId="ref42136" refString="Hebauer, F. (2002 a) New Hydrophilidae of the Old World (Coleoptera, Hydrophiloidea). Acta Coleopterologica 18, 3, 3 - 24." type="journal article" year="2002" yearSuffix="a">
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Hebauer
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<number box="[416,475,999,1023]" pageId="47" pageNumber="48" value="2002.0">2002</number>
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||
a
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||
</bibRefCitation>
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||
,
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||
<figureCitation box="[501,584,999,1023]" captionStart="FIGURES 12 – 25" captionStartId="5.[151,264,1863,1886]" captionTargetBox="[151,1436,193,1841]" captionTargetId="figure@5.[151,1436,193,1842]" captionTargetPageId="5" captionText="FIGURES 12 – 25. General morphology of the Asian Pachysternum. 12 – 14: pronotal punctation (12: P. nigrovittatum, 13: P. haemorrhoum, 14: P. r u g o s u m); 15: meso- and metathorax of P. nigrovittatum, ventral view; 16 – 19: preepisternal elevation of mesothorax (16: P. haemorrhoum, 17: P. s u l a w e s i c u m, 18: P. apicatum, 19: P. rugosum); 20 – 21: surface sculpture of metaventrite mesally and laterally of femoral line (20: P. nigrovittatum, 21: P. haemorrhoum); 22: elytral punctation of P. nigrovittatum; 23: scutellum of P. nigrovittatum; 24: anterior leg of P. nigrovittatum; 25: abdomen of P. nigrovitattum." httpUri="https://zenodo.org/record/280225/files/figure.png" pageId="47" pageNumber="48">
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||
Fig.
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<number box="[555,584,999,1023]" pageId="47" pageNumber="48" value="22.0">22</number>
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||
</figureCitation>
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||
) also do not correspond with the
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<taxonomicName box="[976,1136,1000,1024]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
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||
<emphasis box="[976,1136,1000,1024]" italics="true" pageId="47" pageNumber="48">Pachysternum</emphasis>
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</taxonomicName>
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species, but largely agree with those illustrated by
|
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<bibRefCitation author="Hansen" box="[430,742,1034,1058]" pageId="47" pageNumber="48" pagination="53 - 84" refId="ref41916" refString="Hansen, M. (2003) Discovery of Australocyon Hansen and Pilocnema Hansen (Coleoptera, Hydrophilidae) outside the Australian region. In: Cuccudoro, G. & Leschen, R. A. B. (eds.) Systematics of Coleoptera: papers celebrating the retirement of Ivan Lobl. Memoirs on Entomology International 17, 53 - 84." type="journal article" year="2003">
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||
Hansen (
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||
<number box="[529,589,1034,1058]" pageId="47" pageNumber="48" value="2003.0">2003</number>
|
||
,
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<figureCitation box="[600,734,1034,1058]" captionStart="FIGURES 12 – 25" captionStartId="5.[151,264,1863,1886]" captionTargetBox="[151,1436,193,1841]" captionTargetId="figure@5.[151,1436,193,1842]" captionTargetPageId="5" captionText="FIGURES 12 – 25. General morphology of the Asian Pachysternum. 12 – 14: pronotal punctation (12: P. nigrovittatum, 13: P. haemorrhoum, 14: P. r u g o s u m); 15: meso- and metathorax of P. nigrovittatum, ventral view; 16 – 19: preepisternal elevation of mesothorax (16: P. haemorrhoum, 17: P. s u l a w e s i c u m, 18: P. apicatum, 19: P. rugosum); 20 – 21: surface sculpture of metaventrite mesally and laterally of femoral line (20: P. nigrovittatum, 21: P. haemorrhoum); 22: elytral punctation of P. nigrovittatum; 23: scutellum of P. nigrovittatum; 24: anterior leg of P. nigrovittatum; 25: abdomen of P. nigrovitattum." httpUri="https://zenodo.org/record/280225/files/figure.png" pageId="47" pageNumber="48">
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||
Figs.
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||
<number box="[663,734,1034,1058]" pageId="47" pageNumber="48" value="19.5" valueMax="21.0" valueMin="18.0">18–21</number>
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||
</figureCitation>
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||
)
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||
</bibRefCitation>
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||
for the species of the
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<taxonomicName box="[991,1310,1035,1059]" class="Insecta" family="Hydrophilidae" genus="Australocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pilocnemoides">
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<emphasis box="[991,1310,1035,1059]" italics="true" pageId="47" pageNumber="48">Australocyon pilocnemoides</emphasis>
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</taxonomicName>
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group. For that reason, the species is transferred here to the genus
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<taxonomicName box="[767,916,1069,1093]" class="Insecta" family="Hydrophilidae" genus="Australocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
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||
<emphasis box="[767,916,1069,1093]" italics="true" pageId="47" pageNumber="48">Australocyon</emphasis>
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||
</taxonomicName>
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||
.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="47" pageNumber="48" type="reference_group">
|
||
<paragraph blockId="47.[151,1437,826,1301]" pageId="47" pageNumber="48">
|
||
<taxonomicName authority="(Hebauer, 2002)" authorityName="Hebauer" box="[201,470,1104,1128]" class="Insecta" family="Hydrophilidae" genus="Australocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="keralensis">
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||
<emphasis box="[201,470,1104,1128]" italics="true" pageId="47" pageNumber="48">Australocyon keralensis</emphasis>
|
||
</taxonomicName>
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||
may be diagnosed as follows: dorsal surface uniformly dark brown; antennal club unicolored; pronotum without sublateral depressions; posterolateral corners of pronotum bluntly rounded; pronotal punctation moderately coarse but sparse, consisting of punctures of two slightly different sizes; pronotal interstices bearing strong mesh-like microsculpture; elytra evenly convex, not humpy in humeral areas. Based on the key provided by
|
||
<bibRefCitation author="Hansen" box="[256,425,1242,1266]" firstAuthor="Hansen" pageId="47" pageNumber="48" pagination="53 - 84" refId="ref41916" refString="Hansen, M. (2003) Discovery of Australocyon Hansen and Pilocnema Hansen (Coleoptera, Hydrophilidae) outside the Australian region. In: Cuccudoro, G. & Leschen, R. A. B. (eds.) Systematics of Coleoptera: papers celebrating the retirement of Ivan Lobl. Memoirs on Entomology International 17, 53 - 84." type="journal article" year="2003">
|
||
Hansen (
|
||
<number box="[356,417,1242,1266]" pageId="47" pageNumber="48" value="2003.0">2003</number>
|
||
)
|
||
</bibRefCitation>
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||
, the species seems to be very similar especially to
|
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<taxonomicName authority="Hansen, 2003" authorityName="Hansen" authorityYear="2003" box="[1000,1356,1242,1267]" class="Insecta" family="Hydrophilidae" genus="Australocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pilocnemoides">
|
||
<emphasis box="[1000,1194,1243,1267]" italics="true" pageId="47" pageNumber="48">A. pilocnemoides</emphasis>
|
||
<bibRefCitation author="Hansen" box="[1201,1356,1242,1266]" firstAuthor="Hansen" pageId="47" pageNumber="48" pagination="53 - 84" refId="ref41916" refString="Hansen, M. (2003) Discovery of Australocyon Hansen and Pilocnema Hansen (Coleoptera, Hydrophilidae) outside the Australian region. In: Cuccudoro, G. & Leschen, R. A. B. (eds.) Systematics of Coleoptera: papers celebrating the retirement of Ivan Lobl. Memoirs on Entomology International 17, 53 - 84." type="journal article" year="2003">
|
||
Hansen,
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||
<number box="[1300,1356,1242,1266]" pageId="47" pageNumber="48" value="2003.0">2003</number>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
and
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<taxonomicName authority="Hansen, 2003" authorityName="Hansen" authorityYear="2003" class="Insecta" family="Hydrophilidae" genus="Australocyon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="puncticollis">
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<emphasis italics="true" pageId="47" pageNumber="48">A. puncticollis</emphasis>
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<bibRefCitation author="Hansen" box="[288,446,1277,1301]" firstAuthor="Hansen" pageId="47" pageNumber="48" pagination="53 - 84" refId="ref41916" refString="Hansen, M. (2003) Discovery of Australocyon Hansen and Pilocnema Hansen (Coleoptera, Hydrophilidae) outside the Australian region. In: Cuccudoro, G. & Leschen, R. A. B. (eds.) Systematics of Coleoptera: papers celebrating the retirement of Ivan Lobl. Memoirs on Entomology International 17, 53 - 84." type="journal article" year="2003">
|
||
Hansen,
|
||
<number box="[387,446,1277,1301]" pageId="47" pageNumber="48" value="2003.0">2003</number>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |