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<document id="D1947051F1467E0C7604736F34717FD3" ID-CLB-Dataset="24139" ID-DOI="10.11646/zootaxa.4731.3.6" ID-GBIF-Dataset="daf28c32-4733-4f63-a3a1-85ce4c73ef7e" ID-ISSN="1175-5326" ID-Zenodo-Dep="3653632" ID-ZooBank="A6944DC9-38EE-4688-B431-3918BD6091E6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581059480110" checkinUser="plazi" docAuthor="Luo, Yongting &amp; Segers, Hendrik" docDate="2020" docId="03E687B48A5FFFAA7AB6D0E9B6CBBF9B" docLanguage="en" docName="zootaxa.4731.3.6.pdf" docOrigin="Zootaxa 4731 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Squatinella Bory de St. Vincent 1924" docType="treatment" docVersion="9" lastPageNumber="380" masterDocId="FFDFFFCC8A58FFA37A21D64CB638BA5D" masterDocTitle="Eight new Lepadellidae (Rotifera, Monogononta) from the Congo bring to level endemism in Africa’s rotifers" masterLastPageNumber="387" masterPageNumber="371" pageNumber="378" updateTime="1698780921035" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B0E5CB82D399DD68241E5C7BCA0E1C34">Eight new Lepadellidae (Rotifera, Monogononta) from the Congo bring to level endemism in Africa’s rotifers</mods:title>
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<mods:namePart id="E7A02499CE35A6A87375BC989DA6F747">Luo, Yongting</mods:namePart>
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<treatment id="03E687B48A5FFFAA7AB6D0E9B6CBBF9B" ID-DOI="http://doi.org/10.5281/zenodo.3665201" ID-GBIF-Taxon="162054228" ID-Zenodo-Dep="3665201" LSID="urn:lsid:plazi:treatment:03E687B48A5FFFAA7AB6D0E9B6CBBF9B" httpUri="http://treatment.plazi.org/id/03E687B48A5FFFAA7AB6D0E9B6CBBF9B" lastPageId="9" lastPageNumber="380" pageId="7" pageNumber="378">
<subSubSection id="C35565298A5FFFA47AB6D0E9B7AABDE1" pageId="7" pageNumber="378" type="nomenclature">
<paragraph id="8BF036A28A5FFFA47AB6D0E9B485BCE2" blockId="7.[151,701,1701,1728]" box="[151,701,1701,1728]" pageId="7" pageNumber="378">
<heading id="D0B881CE8A5FFFA47AB6D0E9B485BCE2" bold="true" box="[151,701,1701,1728]" fontSize="11" level="1" pageId="7" pageNumber="378" reason="1">
<emphasis id="B93BEAB08A5FFFA47AB6D0E9B485BCE2" bold="true" box="[151,701,1701,1728]" pageId="7" pageNumber="378">
Genus
<taxonomicName id="4C4F4D218A5FFFA47ACFD0E9B485BCE2" ID-CoL="7LRT" ID-ENA="525910" authority="Bory de St. Vincent, 1924" authorityName="Bory de St. Vincent" authorityYear="1924" box="[238,701,1701,1728]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="7" pageNumber="378" phylum="Rotifera" rank="genus">
<emphasis id="B93BEAB08A5FFFA47ACFD0E9B741BC9D" bold="true" box="[238,377,1701,1728]" italics="true" pageId="7" pageNumber="378">Squatinella</emphasis>
Bory de St. Vincent, 1924
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF036A28A5FFFA47AB6D0A1B7AABDE1" blockId="7.[151,1437,1773,1980]" pageId="7" pageNumber="378">
Our material contains two distinct taxa of
<taxonomicName id="4C4F4D218A5FFFA47855D0A1B4CCBD5A" authorityName="Bory de St. Vincent" authorityYear="1924" box="[628,756,1773,1799]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="7" pageNumber="378" phylum="Rotifera" rank="genus">
<emphasis id="B93BEAB08A5FFFA47855D0A1B4CCBD5A" box="[628,756,1773,1799]" italics="true" pageId="7" pageNumber="378">Squatinella</emphasis>
</taxonomicName>
, both belonging to the
<taxonomicName id="4C4F4D218A5FFFA479DAD0A2B339BD5A" box="[1019,1281,1773,1800]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="7" pageNumber="378" phylum="Rotifera" rank="species" species="longispinata-leydigii">
<emphasis id="B93BEAB08A5FFFA479DAD0A2B339BD5A" box="[1019,1281,1773,1800]" italics="true" pageId="7" pageNumber="378">S. longispinata-leydigii</emphasis>
</taxonomicName>
group as recognized by
<bibRefCitation id="EFDE4B538A5FFFA47B3BD15EB789BD76" author="Koste, W." box="[282,433,1809,1836]" pageId="7" pageNumber="378" refId="ref10488" refString="Koste, W. (1978) Rotatoria. Die Radertiere Mitteleuropas. Gebr. Borntraeger, Berlin and Stuttgart, 673 pp., 234 pls." type="book" year="1978">Koste (1978)</bibRefCitation>
. The taxonomy of the members of this group is complex and confused due to the rarity of the species and the sometimes contradictory reports. Based on an examination of original sources and the present and other material seen by us, we follow Myers’ (1942),
<bibRefCitation id="EFDE4B538A5FFFA4793DD115B5E5BD2E" author="Pourriot, R." box="[796,989,1881,1907]" pageId="7" pageNumber="378" pagination="97 - 101" refId="ref11135" refString="Pourriot, R. (1971) Deux formes nouvelles du genre Squatinella (Rotiferes) observees en Sologne. Cahiers des naturalistes, Bulletin des Naturalistes Parisiens, 27, 97 - 101." type="journal article" year="1971">Pourriot’s (1971)</bibRefCitation>
and Koste &amp; Shiel’s (1989) conclusion that the group contains at least two well-definable species-level taxa, diagnosed as follows (see
<tableCitation id="C6CD03198A5FFFA47E9CD131B328BDCA" box="[1213,1296,1917,1943]" captionStart="TABLE 3" captionStartId="9.[151,239,154,178]" captionTargetPageId="9" captionText="TABLE 3. Comparison of species in the S. longispinata-leydigii group" httpUri="http://table.plazi.org/id/DF30662A8A51FFAA7AB6D6D6B5B3BAEF" pageId="7" pageNumber="378" tableUuid="DF30662A8A51FFAA7AB6D6D6B5B3BAEF">Table 3</tableCitation>
), albeit with changes in synonymy:
</paragraph>
</subSubSection>
<caption id="DF30662A8A50FFAB7AB6D13CB32FBDA8" ID-DOI="http://doi.org/10.5281/zenodo.3653636" ID-Zenodo-Dep="3653636" httpUri="https://zenodo.org/record/3653636/files/figure.png" pageId="8" pageNumber="379" startId="8.[151,250,1904,1929]" targetBox="[189,1399,187,1872]" targetPageId="8">
<paragraph id="8BF036A28A50FFAB7AB6D13CB32FBDA8" blockId="8.[151,1436,1904,2037]" pageId="8" pageNumber="379">
<emphasis id="B93BEAB08A50FFAB7AB6D13CB708BDD4" bold="true" box="[151,304,1904,1929]" pageId="8" pageNumber="379">FIGURE 5–9.</emphasis>
<taxonomicName id="4C4F4D218A50FFAB7B18D13DB7A6BDD4" authorityName="Bory de St Vincent" authorityYear="1826" box="[313,414,1905,1929]" class="Eurotatoria" family="Lepadellidae" genus="Lepadella" kingdom="Animalia" order="Ploima" pageId="8" pageNumber="379" phylum="Rotifera" rank="genus">
<emphasis id="B93BEAB08A50FFAB7B18D13DB7A6BDD4" box="[313,414,1905,1929]" italics="true" pageId="8" pageNumber="379">Lepadella</emphasis>
</taxonomicName>
spp., line drawings. Figure 5a–c:
<taxonomicName id="4C4F4D218A50FFAB792BD13DB5CFBDD4" authority="Luo &amp; Segers, 2020" authorityName="Luo &amp; Segers" authorityYear="2020" box="[778,1015,1905,1929]" class="Eurotatoria" family="Lepadellidae" genus="Lepadella" kingdom="Animalia" order="Ploima" pageId="8" pageNumber="379" phylum="Rotifera" rank="species" species="hanneloreae" status="sp. nov.">
<emphasis id="B93BEAB08A50FFAB792BD13DB5CFBDD4" box="[778,1015,1905,1929]" italics="true" pageId="8" pageNumber="379">Lepadella hanneloreae</emphasis>
</taxonomicName>
<emphasis id="B93BEAB08A50FFAB7E21D13DB204BDD4" bold="true" box="[1024,1084,1905,1929]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="A20857CB8A50FFAB7E21D13DB204BDD4" box="[1024,1084,1905,1929]" pageId="8" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus. a: ventral; b: dorsal; c: frontal. Figure 6a–c:
<taxonomicName id="4C4F4D218A50FFAB7B55D1D9B40DBDF0" authority="Luo &amp; Segers, 2020" authorityName="Luo &amp; Segers" authorityYear="2020" box="[372,565,1941,1965]" class="Eurotatoria" family="Lepadellidae" genus="Lepadella" kingdom="Animalia" order="Ploima" pageId="8" pageNumber="379" phylum="Rotifera" rank="species" species="jingruae" status="sp. nov.">
<emphasis id="B93BEAB08A50FFAB7B55D1D9B40DBDF0" box="[372,565,1941,1965]" italics="true" pageId="8" pageNumber="379">Lepadella jingruae</emphasis>
</taxonomicName>
<emphasis id="B93BEAB08A50FFAB781AD1D9B44DBDF0" bold="true" box="[571,629,1941,1965]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="A20857CB8A50FFAB781AD1D9B44DBDF0" box="[571,629,1941,1965]" pageId="8" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus. a: ventral; b: lateral; c: frontal. Figure 7a–c:
<taxonomicName id="4C4F4D218A50FFAB7E87D1D9B36EBDF0" authority="Luo &amp; Segers, 2020" authorityName="Luo &amp; Segers" authorityYear="2020" box="[1190,1366,1941,1965]" class="Eurotatoria" family="Lepadellidae" genus="Lepadella" kingdom="Animalia" order="Ploima" pageId="8" pageNumber="379" phylum="Rotifera" rank="species" species="weijiai" status="sp. nov.">
<emphasis id="B93BEAB08A50FFAB7E87D1D9B36EBDF0" box="[1190,1366,1941,1965]" italics="true" pageId="8" pageNumber="379">Lepadella weijiai</emphasis>
</taxonomicName>
<emphasis id="B93BEAB08A50FFAB7F7CD1D9B3AEBDF0" bold="true" box="[1373,1430,1941,1965]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="A20857CB8A50FFAB7F7CD1D9B3AEBDF0" box="[1373,1430,1941,1965]" pageId="8" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus. a: ventral; b: dorsal; c: frontal. Figure 8a–c:
<taxonomicName id="4C4F4D218A50FFAB78E2D1F5B5ABBD8C" authority="Luo &amp; Segers, 2020" authorityName="Luo &amp; Segers" authorityYear="2020" box="[707,915,1977,2001]" class="Eurotatoria" family="Lepadellidae" genus="Lepadella" kingdom="Animalia" order="Ploima" pageId="8" pageNumber="379" phylum="Rotifera" rank="species" species="yangambi" status="sp. nov.">
<emphasis id="B93BEAB08A50FFAB78E2D1F5B5ABBD8C" box="[707,915,1977,2001]" italics="true" pageId="8" pageNumber="379">Lepadella yangambi</emphasis>
</taxonomicName>
<emphasis id="B93BEAB08A50FFAB79BBD1F5B5EDBD8C" bold="true" box="[922,981,1977,2001]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="A20857CB8A50FFAB79BBD1F5B5EDBD8C" box="[922,981,1977,2001]" pageId="8" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus. a: ventral; b: dorsal; c: lateral; d: caudal. Figure 9a–c:
<taxonomicName id="4C4F4D218A50FFAB7B53D191B472BDA8" authority="Luo &amp; Segers, 2020" authorityName="Luo &amp; Segers" authorityYear="2020" box="[370,586,2013,2037]" class="Eurotatoria" family="Lepadellidae" genus="Lepadella" kingdom="Animalia" order="Ploima" pageId="8" pageNumber="379" phylum="Rotifera" rank="species" species="wilungulai" status="sp. nov.">
<emphasis id="B93BEAB08A50FFAB7B53D191B472BDA8" box="[370,586,2013,2037]" italics="true" pageId="8" pageNumber="379">Lepadella wilungulai</emphasis>
</taxonomicName>
<emphasis id="B93BEAB08A50FFAB7871D191B4B2BDA8" bold="true" box="[592,650,2013,2037]" pageId="8" pageNumber="379">
<taxonomicNameLabel id="A20857CB8A50FFAB7871D191B4B2BDA8" box="[592,650,2013,2037]" pageId="8" pageNumber="379" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, habitus. a: ventral; b: dorsal; c: frontal. (all scale bars: 50µm).
</paragraph>
</caption>
<caption id="DF30662A8A51FFAA7AB6D6D6B5B3BAEF" ID-Table-UUID="DF30662A8A51FFAA7AB6D6D6B5B3BAEF" box="[151,907,154,178]" httpUri="http://table.plazi.org/id/DF30662A8A51FFAA7AB6D6D6B5B3BAEF" pageId="9" pageNumber="380" targetBox="[166,1406,190,688]" targetIsTable="true" targetPageId="9">
<paragraph id="8BF036A28A51FFAA7AB6D6D6B5B3BAEF" blockId="9.[151,907,154,178]" box="[151,907,154,178]" pageId="9" pageNumber="380">
<emphasis id="B93BEAB08A51FFAA7AB6D6D6B732BAEF" bold="true" box="[151,266,154,178]" pageId="9" pageNumber="380">TABLE 3.</emphasis>
Comparison of species in the
<taxonomicName id="4C4F4D218A51FFAA786CD6D6B57EBAEF" box="[589,838,154,178]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="longispinata-leydigii">
<emphasis id="B93BEAB08A51FFAA786CD6D6B57EBAEF" box="[589,838,154,178]" italics="true" pageId="9" pageNumber="380">S. longispinata-leydigii</emphasis>
</taxonomicName>
group
</paragraph>
</caption>
<paragraph id="8BF036A28A51FFAA7BA9D6F2B24BB8ED" pageId="9" pageNumber="380">
<table id="F94FC4028A51005C7A87D6F2B346B8ED" box="[166,1406,190,688]" gridcols="5" gridrows="12" pageId="9" pageNumber="380">
<tr id="357F34E08A51005C7A87D6F2B346BA8B" box="[166,1406,190,214]" gridrow="0" pageId="9" pageNumber="380" rowspan-0="1">
<th id="76AE5D9C8A51005C7BA9D6F2B45FBA8B" box="[392,615,190,214]" gridcol="1" gridrow="0" pageId="9" pageNumber="380">
<taxonomicName id="4C4F4D218A51FFAA7BA9D6F2B418BA8B" baseAuthorityName="Tatem" baseAuthorityYear="1867" box="[392,544,190,214]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="longispinata">
<emphasis id="B93BEAB08A51FFAA7BA9D6F2B418BA8B" box="[392,544,190,214]" italics="true" pageId="9" pageNumber="380">S. longispinata</emphasis>
</taxonomicName>
</th>
<th id="76AE5D9C8A51005C78A6D6F2B55EBA8B" box="[647,870,190,214]" gridcol="2" gridrow="0" pageId="9" pageNumber="380">
<taxonomicName id="4C4F4D218A51FFAA78A6D6F2B4C4BA8B" authority="Luo &amp; Segers, 2020" authorityName="Luo &amp; Segers" authorityYear="2020" box="[647,764,190,214]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="longipila" status="sp. nov.">
<emphasis id="B93BEAB08A51FFAA78A6D6F2B4C4BA8B" box="[647,764,190,214]" italics="true" pageId="9" pageNumber="380">S. longipila</emphasis>
</taxonomicName>
<emphasis id="B93BEAB08A51FFAA7923D6F2B503BA8B" bold="true" box="[770,827,190,214]" pageId="9" pageNumber="380">
<taxonomicNameLabel id="A20857CB8A51FFAA7923D6F2B503BA8B" box="[770,827,190,214]" pageId="9" pageNumber="380" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76AE5D9C8A51005C79A7D6F2B241BA8B" box="[902,1145,190,214]" gridcol="3" gridrow="0" pageId="9" pageNumber="380">
<taxonomicName id="4C4F4D218A51FFAA79A7D6F2B238BA8B" baseAuthorityName="Levander" baseAuthorityYear="1894" box="[902,1024,190,214]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="variegata">
<emphasis id="B93BEAB08A51FFAA79A7D6F2B238BA8B" box="[902,1024,190,214]" italics="true" pageId="9" pageNumber="380">S. variegata</emphasis>
</taxonomicName>
</th>
<th id="76AE5D9C8A51005C7EB7D6F2B346BA8B" box="[1174,1406,190,214]" gridcol="4" gridrow="0" pageId="9" pageNumber="380">
<taxonomicName id="4C4F4D218A51FFAA7EB7D6F2B323BA8B" authority="Luo &amp; Segers, 2020" authorityName="Luo &amp; Segers" authorityYear="2020" box="[1174,1307,190,214]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="curvispina" status="sp. nov.">
<emphasis id="B93BEAB08A51FFAA7EB7D6F2B323BA8B" box="[1174,1307,190,214]" italics="true" pageId="9" pageNumber="380">S. curvispina</emphasis>
</taxonomicName>
<emphasis id="B93BEAB08A51FFAA7F00D6F2B362BA8B" bold="true" box="[1313,1370,190,214]" pageId="9" pageNumber="380">
<taxonomicNameLabel id="A20857CB8A51FFAA7F00D6F2B362BA8B" box="[1313,1370,190,214]" pageId="9" pageNumber="380" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="357F34E08A51005C7A87D6AFB346BAA1" box="[166,1406,227,252]" gridrow="1" pageId="9" pageNumber="380">
<th id="76AE5D9C8A51005C7A87D6AFB752BAA1" box="[166,362,227,252]" gridcol="0" gridrow="1" pageId="9" pageNumber="380">Dorsal spine shape</th>
<td id="76AE5D9C8A51005C7BA9D6AFB45FBAA1" box="[392,615,227,252]" gridcol="1" gridrow="1" pageId="9" pageNumber="380">straight</td>
<td id="76AE5D9C8A51005C78A6D6AFB55EBAA1" box="[647,870,227,252]" gridcol="2" gridrow="1" pageId="9" pageNumber="380">weakly bent basally,</td>
<td id="76AE5D9C8A51005C79A7D6AFB241BAA1" box="[902,1145,227,252]" gridcol="3" gridrow="1" pageId="9" pageNumber="380">straight</td>
<td id="76AE5D9C8A51005C7EB7D6AFB346BAA1" box="[1174,1406,227,252]" gridcol="4" gridrow="1" pageId="9" pageNumber="380">S-curved</td>
</tr>
<tr id="357F34E08A51005C7A87D74BB346BB1C" box="[166,1406,263,321]" gridrow="2" pageId="9" pageNumber="380">
<th id="76AE5D9C8A51005C7A87D74BB752BB1C" box="[166,362,263,321]" gridcol="0" gridrow="2" pageId="9" pageNumber="380">Dorsal spine tip</th>
<td id="76AE5D9C8A51005C7BA9D74BB45FBB1C" box="[392,615,263,321]" gridcol="1" gridrow="2" pageId="9" pageNumber="380">sharp</td>
<td id="76AE5D9C8A51005C78A6D74BB55EBB1C" box="[647,870,263,321]" gridcol="2" gridrow="2" pageId="9" pageNumber="380">curved terminally sharp</td>
<td id="76AE5D9C8A51005C79A7D74BB241BB1C" box="[902,1145,263,321]" gridcol="3" gridrow="2" pageId="9" pageNumber="380">blunt</td>
<td id="76AE5D9C8A51005C7EB7D74BB346BB1C" box="[1174,1406,263,321]" gridcol="4" gridrow="2" pageId="9" pageNumber="380">blunt</td>
</tr>
<tr id="357F34E08A51005C7A87D702B346BB3A" box="[166,1406,334,359]" gridrow="3" pageId="9" pageNumber="380">
<th id="76AE5D9C8A51005C7A87D702B752BB3A" box="[166,362,334,359]" gridcol="0" gridrow="3" pageId="9" pageNumber="380">Dorsal spine basis</th>
<td id="76AE5D9C8A51005C7BA9D702B45FBB3A" box="[392,615,334,359]" gridcol="1" gridrow="3" pageId="9" pageNumber="380">reports contradictory</td>
<td id="76AE5D9C8A51005C78A6D702B55EBB3A" box="[647,870,334,359]" gridcol="2" gridrow="3" pageId="9" pageNumber="380">elongate triangular</td>
<td id="76AE5D9C8A51005C79A7D702B241BB3A" box="[902,1145,334,359]" gridcol="3" gridrow="3" pageId="9" pageNumber="380">reports contradictory</td>
<td id="76AE5D9C8A51005C7EB7D702B346BB3A" box="[1174,1406,334,359]" gridcol="4" gridrow="3" pageId="9" pageNumber="380">Not broadened</td>
</tr>
<tr id="357F34E08A51005C7A87D73FB346BBD1" box="[166,1406,371,396]" gridrow="4" pageId="9" pageNumber="380">
<th id="76AE5D9C8A51005C7A87D73FB752BBD1" box="[166,362,371,396]" gridcol="0" gridrow="4" pageId="9" pageNumber="380">Foot</th>
<td id="76AE5D9C8A51005C7BA9D73FB45FBBD1" box="[392,615,371,396]" gridcol="1" gridrow="4" pageId="9" pageNumber="380">three pseudosegments</td>
<td id="76AE5D9C8A51005C78A6D73FB55EBBD1" box="[647,870,371,396]" gridcol="2" gridrow="4" pageId="9" pageNumber="380">three pseudosegments</td>
<td id="76AE5D9C8A51005C79A7D73FB241BBD1" box="[902,1145,371,396]" gridcol="3" gridrow="4" pageId="9" pageNumber="380">two pseudosegments,</td>
<td id="76AE5D9C8A51005C7EB7D73FB346BBD1" box="[1174,1406,371,396]" gridcol="4" gridrow="4" pageId="9" pageNumber="380">two pseudosegments,</td>
</tr>
<tr id="357F34E08A51005C7A87D7DBB346BB8C" box="[166,1406,407,465]" gridrow="5" pageId="9" pageNumber="380">
<th id="76AE5D9C8A51005C7A87D7DBB752BB8C" box="[166,362,407,465]" gridcol="0" gridrow="5" pageId="9" pageNumber="380">Toes</th>
<td id="76AE5D9C8A51005C7BA9D7DBB45FBB8C" box="[392,615,407,465]" gridcol="1" gridrow="5" pageId="9" pageNumber="380">two separate</td>
<td id="76AE5D9C8A51005C78A6D7DBB55EBB8C" box="[647,870,407,465]" gridcol="2" gridrow="5" pageId="9" pageNumber="380">two separate</td>
<td id="76AE5D9C8A51005C79A7D7DBB241BB8C" box="[902,1145,407,465]" gridcol="3" gridrow="5" pageId="9" pageNumber="380">terminal fused element fused, terminal fissure</td>
<td id="76AE5D9C8A51005C7EB7D7DBB346BB8C" box="[1174,1406,407,465]" gridcol="4" gridrow="5" pageId="9" pageNumber="380">terminal fused element fused, terminal fissure</td>
</tr>
<tr id="357F34E08A51005C7A87D792B346BBAA" box="[166,1406,478,503]" gridrow="6" pageId="9" pageNumber="380">
<th id="76AE5D9C8A51005C7A87D792B752BBAA" box="[166,362,478,503]" gridcol="0" gridrow="6" pageId="9" pageNumber="380">Add. foot spine</th>
<td id="76AE5D9C8A51005C7BA9D792B45FBBAA" box="[392,615,478,503]" gridcol="1" gridrow="6" pageId="9" pageNumber="380">present</td>
<td id="76AE5D9C8A51005C78A6D792B55EBBAA" box="[647,870,478,503]" gridcol="2" gridrow="6" pageId="9" pageNumber="380">absent</td>
<td id="76AE5D9C8A51005C79A7D792B241BBAA" box="[902,1145,478,503]" gridcol="3" gridrow="6" pageId="9" pageNumber="380">absent</td>
<td id="76AE5D9C8A51005C7EB7D792B346BBAA" box="[1174,1406,478,503]" gridcol="4" gridrow="6" pageId="9" pageNumber="380">absent</td>
</tr>
<tr id="357F34E08A51005C7A87D448B346B840" box="[166,1406,516,541]" gridrow="7" pageId="9" pageNumber="380">
<th id="76AE5D9C8A51005C7A87D448B752B840" box="[166,362,516,541]" gridcol="0" gridrow="7" pageId="9" pageNumber="380">Total length</th>
<td id="76AE5D9C8A51005C7BA9D448B45FB840" box="[392,615,516,541]" gridcol="1" gridrow="7" pageId="9" pageNumber="380">82–140</td>
<td id="76AE5D9C8A51005C78A6D448B55EB840" box="[647,870,516,541]" gridcol="2" gridrow="7" pageId="9" pageNumber="380">117–130</td>
<td id="76AE5D9C8A51005C79A7D448B241B840" box="[902,1145,516,541]" gridcol="3" gridrow="7" pageId="9" pageNumber="380">180–240</td>
<td id="76AE5D9C8A51005C7EB7D448B346B840" box="[1174,1406,516,541]" gridcol="4" gridrow="7" pageId="9" pageNumber="380">219–257</td>
</tr>
<tr id="357F34E08A51005C7A87D466B346B81E" box="[166,1406,554,579]" gridrow="8" pageId="9" pageNumber="380">
<th id="76AE5D9C8A51005C7A87D466B752B81E" box="[166,362,554,579]" gridcol="0" gridrow="8" pageId="9" pageNumber="380">Dorsal spine length</th>
<td id="76AE5D9C8A51005C7BA9D466B45FB81E" box="[392,615,554,579]" gridcol="1" gridrow="8" pageId="9" pageNumber="380">96–180</td>
<td id="76AE5D9C8A51005C78A6D466B55EB81E" box="[647,870,554,579]" gridcol="2" gridrow="8" pageId="9" pageNumber="380">251–303</td>
<td id="76AE5D9C8A51005C79A7D466B241B81E" box="[902,1145,554,579]" gridcol="3" gridrow="8" pageId="9" pageNumber="380">190–378</td>
<td id="76AE5D9C8A51005C7EB7D466B346B81E" box="[1174,1406,554,579]" gridcol="4" gridrow="8" pageId="9" pageNumber="380">146–190</td>
</tr>
<tr id="357F34E08A51005C7A87D403B346B835" box="[166,1406,591,616]" gridrow="9" pageId="9" pageNumber="380">
<th id="76AE5D9C8A51005C7A87D403B752B835" box="[166,362,591,616]" gridcol="0" gridrow="9" pageId="9" pageNumber="380">Ratio total length:</th>
<td id="76AE5D9C8A51005C7BA9D403B45FB835" box="[392,615,591,616]" gridcol="1" gridrow="9" pageId="9" pageNumber="380">~ 0.8</td>
<td id="76AE5D9C8A51005C78A6D403B55EB835" box="[647,870,591,616]" gridcol="2" gridrow="9" pageId="9" pageNumber="380">~ 2.0–2.6</td>
<td id="76AE5D9C8A51005C79A7D403B241B835" box="[902,1145,591,616]" gridcol="3" gridrow="9" pageId="9" pageNumber="380">
~ 0.8 (f.
<emphasis id="B93BEAB08A51FFAA79FFD41DB224B835" box="[990,1052,593,616]" italics="true" pageId="9" pageNumber="380">typica</emphasis>
)
</td>
<td id="76AE5D9C8A51005C7EB7D403B346B835" box="[1174,1406,591,616]" gridcol="4" gridrow="9" pageId="9" pageNumber="380">~ 0.83</td>
</tr>
<tr id="357F34E08A51005C7A87D43FB346B8D1" box="[166,1406,627,652]" gridrow="10" pageId="9" pageNumber="380" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<th id="76AE5D9C8A51005C7A87D43FB752B8D1" box="[166,362,627,652]" gridcol="0" gridrow="10" pageId="9" pageNumber="380">dorsal spine length</th>
<td id="76AE5D9C8A51005C79A7D43FB241B8D1" box="[902,1145,627,652]" gridcol="3" gridrow="10" pageId="9" pageNumber="380">
~ 1.1–1.2 (f.
<taxonomicName id="4C4F4D218A51FFAA7E31D439B249B8D1" baseAuthorityName="Levander" baseAuthorityYear="1894" box="[1040,1137,629,652]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="variegata">
<emphasis id="B93BEAB08A51FFAA7E31D439B249B8D1" box="[1040,1137,629,652]" italics="true" pageId="9" pageNumber="380">variegata</emphasis>
</taxonomicName>
)
</td>
</tr>
<tr id="357F34E08A51005C7A87D4DBB346B8ED" box="[166,1406,663,688]" gridrow="11" pageId="9" pageNumber="380" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1">
<td id="76AE5D9C8A51005C79A7D4DBB241B8ED" box="[902,1145,663,688]" gridcol="3" gridrow="11" pageId="9" pageNumber="380">
&gt; or = 1.6 (f.
<emphasis id="B93BEAB08A51FFAA7E2FD4D4B253B8ED" box="[1038,1131,664,688]" italics="true" pageId="9" pageNumber="380">longiseta</emphasis>
)
</td>
</tr>
</table>
</paragraph>
<subSubSection id="C35565298A51FFAA7AB6D491B6CBBF9B" pageId="9" pageNumber="380" type="reference_group">
<paragraph id="8BF036A28A51FFAA7AB6D491B520B993" blockId="9.[151,1437,732,1478]" pageId="9" pageNumber="380">
-
<taxonomicName id="4C4F4D218A51FFAA7AE6D490B4BBB8AA" ID-CoL="6ZFH2" authority="(Tatem, 1867)" baseAuthorityName="Tatem" baseAuthorityYear="1867" box="[199,643,732,759]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="longispinata">
<emphasis id="B93BEAB08A51FFAA7AE6D490B7E3B8AB" box="[199,475,732,758]" italics="true" pageId="9" pageNumber="380">Squatinella longispinata</emphasis>
(
<bibRefCitation id="EFDE4B538A51FFAA7BCAD490B443B8AA" author="Tatem, T. G." box="[491,635,732,759]" pageId="9" pageNumber="380" pagination="251 - 253" refId="ref11925" refString="Tatem, T. G. (1867) New Species of Microscopic Animals. Quarterly Journal of Microscopical Science, 7, 251 - 253." type="journal article" year="1867">Tatem, 1867</bibRefCitation>
)
</taxonomicName>
(synonyms:
<taxonomicName id="4C4F4D218A51FFAA7936D490B262B8AA" ID-CoL="6ZFH5" authority="(Zacharias, 1886)" baseAuthorityName="Zacharias" baseAuthorityYear="1886" box="[791,1114,732,759]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="leydigii">
<emphasis id="B93BEAB08A51FFAA7936D490B5B1B8AB" box="[791,905,732,758]" italics="true" pageId="9" pageNumber="380">S. leydigii</emphasis>
(
<bibRefCitation id="EFDE4B538A51FFAA79B8D490B26BB8AA" author="Zacharias, O." box="[921,1107,732,759]" pageId="9" pageNumber="380" pagination="252 - 276" refId="ref12162" refString="Zacharias, O. (1886) Ergebnisse einer zoologischen Exkursion in das Glatzer-, Iser- und Riesengebirge. Zeitschrift fur wissenschaftliche Zoologie, 43, 252 - 276." type="journal article" year="1886">Zacharias, 1886</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C4F4D218A51FFAA7E49D490B37DB8AA" ID-CoL="52CM2" authority="(Collins)" baseAuthorityName="Collins" box="[1128,1349,732,759]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="uniseta">
<emphasis id="B93BEAB08A51FFAA7E49D490B2EEB8AB" box="[1128,1238,732,758]" italics="true" pageId="9" pageNumber="380">S. uniseta</emphasis>
(Collins)
</taxonomicName>
,
<taxonomicName id="4C4F4D218A51FFAA7F73D490B7FCB947" ID-CoL="52CM3" authority="(Hudson &amp; Gosse)" baseAuthorityName="Hudson &amp; Gosse" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="unisetatus">
<emphasis id="B93BEAB08A51FFAA7F73D490B6DDB947" italics="true" pageId="9" pageNumber="380">S. unisetatus</emphasis>
(Hudson &amp; Gosse)
</taxonomicName>
,
<taxonomicName id="4C4F4D218A51FFAA7BF0D54CB494B946" ID-CoL="52CLZ" authority="(Hudson))" baseAuthorityName="Hudson" box="[465,684,768,795]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="tripus">
<emphasis id="B93BEAB08A51FFAA7BF0D54CB417B947" box="[465,559,768,794]" italics="true" pageId="9" pageNumber="380">S. tripus</emphasis>
(Hudson))
</taxonomicName>
is here considered to be a relatively small species, with three foot pseudosegments and bearing a minute spine dorsally and terminally on the last foot pseudosegment. Its dorsal lorica spine narrows gradually into a sharp terminal point. Note that neither the (rather superficial) original description of
<taxonomicName id="4C4F4D218A51FFAA7AB6D520B778B9DB" ID-CoL="6ZFH2" baseAuthorityName="Tatem" baseAuthorityYear="1867" box="[151,320,876,902]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="longispinata">
<emphasis id="B93BEAB08A51FFAA7AB6D520B778B9DB" box="[151,320,876,902]" italics="true" pageId="9" pageNumber="380">S. longispinata</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFDE4B538A51FFAA7B4AD521B43BB9DA" author="Tatem, T. G." box="[363,515,876,903]" pageId="9" pageNumber="380" pagination="251 - 253" refId="ref11925" refString="Tatem, T. G. (1867) New Species of Microscopic Animals. Quarterly Journal of Microscopical Science, 7, 251 - 253." type="journal article" year="1867">Tatem (1867)</bibRefCitation>
nor that of
<taxonomicName id="4C4F4D218A51FFAA78A8D520B4C2B9DB" ID-CoL="6ZFH5" baseAuthorityName="Zacharias" baseAuthorityYear="1886" box="[649,762,876,902]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="leydigii">
<emphasis id="B93BEAB08A51FFAA78A8D520B4C2B9DB" box="[649,762,876,902]" italics="true" pageId="9" pageNumber="380">S. leydigii</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFDE4B538A51FFAA7904D520B5DDB9DA" author="Zacharias, O." box="[805,997,876,903]" pageId="9" pageNumber="380" pagination="252 - 276" refId="ref12162" refString="Zacharias, O. (1886) Ergebnisse einer zoologischen Exkursion in das Glatzer-, Iser- und Riesengebirge. Zeitschrift fur wissenschaftliche Zoologie, 43, 252 - 276." type="journal article" year="1886">Zacharias (1886)</bibRefCitation>
mention a foot spine, which may have been overlooked in these early reports. The relevance of the short spines flanking the dorsal spine, as reported for
<taxonomicName id="4C4F4D218A51FFAA7FA6D5DCB71BB993" ID-CoL="6ZFH2" baseAuthorityName="Tatem" baseAuthorityYear="1867" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="longispinata">
<emphasis id="B93BEAB08A51FFAA7FA6D5DCB71BB993" italics="true" pageId="9" pageNumber="380">S. longispinata</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFDE4B538A51FFAA7B6CD5F8B7D8B992" author="Pejler, B." box="[333,480,948,975]" pageId="9" pageNumber="380" pagination="327 - 422" refId="ref11070" refString="Pejler, B. (1962) On the taxonomy and ecology of benthic and periphytic Rotatoria. Investigation in northern Swedish Lappland. Zoologiska Bidrag fran Uppsala, 33, 327 - 422." type="journal article" year="1962">Pejler (1962)</bibRefCitation>
requires further evaluation.
</paragraph>
<paragraph id="8BF036A28A51FFAA7AB6D595B448BF4F" blockId="9.[151,1437,732,1478]" pageId="9" pageNumber="380">
-
<taxonomicName id="4C4F4D218A51FFAA7AE6D594B4BEB9AF" authority="(Levander, 1894)" baseAuthorityName="Levander" baseAuthorityYear="1894" box="[199,646,984,1010]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="variegata">
<emphasis id="B93BEAB08A51FFAA7AE6D594B782B9AF" box="[199,442,984,1010]" italics="true" pageId="9" pageNumber="380">Squatinella variegata</emphasis>
(
<bibRefCitation id="EFDE4B538A51FFAA7BEAD594B446B9AF" author="Levander, K. M." box="[459,638,984,1010]" pageId="9" pageNumber="380" pagination="1 - 72" refId="ref10716" refString="Levander, K. M. (1894) Materialien zur Kenntniss der Wasserfauna in der Umgebung von Helsingfors, mit besonderer Berucksichtigung der Meeresfauna. II. Rotatoria. Acta Societatis pro Fauna et Flora Fennica, 12, 1 - 72." type="journal article" year="1894">Levander, 1894</bibRefCitation>
)
</taxonomicName>
(note that
<bibRefCitation id="EFDE4B538A51FFAA7922D594B59DB9AF" author="Koste, W." box="[771,933,984,1010]" pageId="9" pageNumber="380" refId="ref10488" refString="Koste, W. (1978) Rotatoria. Die Radertiere Mitteleuropas. Gebr. Borntraeger, Berlin and Stuttgart, 673 pp., 234 pls." type="book" year="1978">Koste’s (1978</bibRefCitation>
: T. 58 fig. 2a) reproduction of the foot of
<taxonomicName id="4C4F4D218A51FFAA7FA6D594B73ABE4B" baseAuthorityName="Levander" baseAuthorityYear="1894" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="variegata">
<emphasis id="B93BEAB08A51FFAA7FA6D594B73ABE4B" italics="true" pageId="9" pageNumber="380">S. variegata</emphasis>
</taxonomicName>
differs in some minute but important aspects from
<bibRefCitation id="EFDE4B538A51FFAA7963D5B0B22ABE4B" author="Levander, K. M." box="[834,1042,1020,1046]" pageId="9" pageNumber="380" pagination="1 - 72" refId="ref10716" refString="Levander, K. M. (1894) Materialien zur Kenntniss der Wasserfauna in der Umgebung von Helsingfors, mit besonderer Berucksichtigung der Meeresfauna. II. Rotatoria. Acta Societatis pro Fauna et Flora Fennica, 12, 1 - 72." type="journal article" year="1894">Levander’s (1894)</bibRefCitation>
original figure), a relatively robust species with two short foot pseudosegments and a third terminal element. Based on the peculiar morphology of the terminal foot pseudosegment, we believe that, in
<taxonomicName id="4C4F4D218A51FFAA789FD208B57DBE03" baseAuthorityName="Levander" baseAuthorityYear="1894" box="[702,837,1092,1118]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="variegata">
<emphasis id="B93BEAB08A51FFAA789FD208B57DBE03" box="[702,837,1092,1118]" italics="true" pageId="9" pageNumber="380">S. variegata</emphasis>
</taxonomicName>
, the third foot pseudosegment and toes are fused into a single element, while the vestiges of the toes are indicated by a short terminal fissure. This we conclude from the position of the dorsal sensory groove, which is situated basally and dorsally on the third foot pseudosegment in
<taxonomicName id="4C4F4D218A51FFAA7A95D2FCB770BE97" ID-CoL="BVX" authorityName="H.K.Harring" authorityYear="1913" box="[180,328,1200,1226]" class="Eurotatoria" family="Lepadellidae" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="family">Lepadellidae</taxonomicName>
, and is situated in the same position on the terminal element of the foot in
<taxonomicName id="4C4F4D218A51FFAA7EB0D2FCB320BE97" baseAuthorityName="Levander" baseAuthorityYear="1894" box="[1169,1304,1200,1226]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="variegata">
<emphasis id="B93BEAB08A51FFAA7EB0D2FCB320BE97" box="[1169,1304,1200,1226]" italics="true" pageId="9" pageNumber="380">S. variegata</emphasis>
</taxonomicName>
and related taxa (see below,
<figureCitation id="13742A278A51FFAA7B71D298B785BEB2" box="[336,445,1236,1263]" captionStart="FIGURES 1–4" captionStartId="4.[151,264,1860,1885]" captionTargetBox="[163,1421,497,1826]" captionTargetId="figure@4.[151,1436,481,1840]" captionTargetPageId="4" captionText="FIGURES 1–4. Lepadellidae, line drawings. Figure 1a–b: Squatinella curviseta n. sp., a: habitus, lateral (scale bar: 100µm); b: detail of foot, lateral (scale bar: 50µm). Figure 2a–c: Squatinella longipila n. sp., habitus. a: detail, body (scale bar: 50µm); b–c: complete specimens (scale bar: 50µm). Figure 3a–c: Colurella asymmetrica n. sp. a: caudal region and foot, left; b: habitus, right; c: habitus, ventral view. Figure 4: Colurella uncinata bicuspidata: habitus, right (3, 4: scale bar: 50µm)." figureDoi="http://doi.org/10.5281/zenodo.3653634" httpUri="https://zenodo.org/record/3653634/files/figure.png" pageId="9" pageNumber="380">Figure 1b</figureCitation>
). There is no additional toe spine and the dorsal spine is more or less parallel sided along much of its length, and ends in a blunt tip.
</paragraph>
<paragraph id="8BF036A28A51FFAA7AE6D350B6CBBF9B" blockId="9.[151,1437,732,1478]" pageId="9" pageNumber="380">
Relatively large specimens reported by
<bibRefCitation id="EFDE4B538A51FFAA785FD350B512BF6B" author="Pourriot, R." box="[638,810,1308,1334]" pageId="9" pageNumber="380" pagination="97 - 101" refId="ref11135" refString="Pourriot, R. (1971) Deux formes nouvelles du genre Squatinella (Rotiferes) observees en Sologne. Cahiers des naturalistes, Bulletin des Naturalistes Parisiens, 27, 97 - 101." type="journal article" year="1971">Pourriot (1971)</bibRefCitation>
as having a dorsal spine of more than 1.5 times the body length (“f.
<emphasis id="B93BEAB08A51FFAA7B33D30CB741BF07" box="[274,377,1344,1370]" italics="true" pageId="9" pageNumber="380">longiseta</emphasis>
” after
<bibRefCitation id="EFDE4B538A51FFAA7BE6D30CB45DBF07" author="Pourriot, R." box="[455,613,1344,1370]" pageId="9" pageNumber="380" pagination="97 - 101" refId="ref11135" refString="Pourriot, R. (1971) Deux formes nouvelles du genre Squatinella (Rotiferes) observees en Sologne. Cahiers des naturalistes, Bulletin des Naturalistes Parisiens, 27, 97 - 101." type="journal article" year="1971">Pourriot 1971</bibRefCitation>
), may yet be another taxon, in view of the report of them having a different foot structure with two clearly separated toes (
<bibRefCitation id="EFDE4B538A51FFAA78EBD328B56FBF23" author="Koste, W." box="[714,855,1380,1406]" pageId="9" pageNumber="380" refId="ref10488" refString="Koste, W. (1978) Rotatoria. Die Radertiere Mitteleuropas. Gebr. Borntraeger, Berlin and Stuttgart, 673 pp., 234 pls." type="book" year="1978">Koste, 1978</bibRefCitation>
). The two taxa in our material differ in a number of significant aspects from either
<taxonomicName id="4C4F4D218A51FFAA7BD8D3C4B49BBFFF" ID-CoL="6ZFH2" baseAuthorityName="Tatem" baseAuthorityYear="1867" box="[505,675,1416,1442]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="longispinata">
<emphasis id="B93BEAB08A51FFAA7BD8D3C4B49BBFFF" box="[505,675,1416,1442]" italics="true" pageId="9" pageNumber="380">S. longispinata</emphasis>
</taxonomicName>
or
<taxonomicName id="4C4F4D218A51FFAA78EDD3C4B56DBFFF" baseAuthorityName="Levander" baseAuthorityYear="1894" box="[716,853,1416,1442]" class="Eurotatoria" family="Lepadellidae" genus="Squatinella" kingdom="Animalia" order="Ploima" pageId="9" pageNumber="380" phylum="Rotifera" rank="species" species="variegata">
<emphasis id="B93BEAB08A51FFAA78EDD3C4B56DBFFF" box="[716,853,1416,1442]" italics="true" pageId="9" pageNumber="380">S. variegata</emphasis>
</taxonomicName>
, prompting us to describe them as new species, as follows.
</paragraph>
</subSubSection>
</treatment>
</document>
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