184 lines
25 KiB
XML
184 lines
25 KiB
XML
<document id="6362244BFED2A2DE56A156711C31868A" ID-DOI="10.11646/zootaxa.3619.5.1" ID-GBIF-Dataset="dcfb62d8-4460-4e6a-8cd5-70039bb48512" ID-ISSN="1175-5326" ID-Zenodo-Dep="223148" ID-ZooBank="17D18CD6-3C12-4CD3-B334-E3BCDA4D22F0" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460578587007" checkinUser="plazi" docAuthor="Krikken, J. & Huijbregts, J." docDate="2013" docId="03E687A4DD2DFFDBFF2807EA1347FDE1" docLanguage="en" docName="zt03619p525.pdf" docOrigin="Zootaxa 3619 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus bituberoculus Krikken & Huijbregts, new species" docType="treatment" docVersion="9" lastPageNumber="513" masterDocId="FFDFFFDCDD27FFD7FFBF012D177EFFDC" masterDocTitle="New Guinea Onthophagus: taxonomy of ten small, unicolored new species (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="525" masterPageNumber="501" pageNumber="511" updateTime="1698323726565" updateUser="plazi">
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<mods:mods id="7D973E06EABF29FF17ABFC6933A0BB99" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="80CB8115F9B900BF31ED644E10FB9AC2">
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<mods:title id="D803908FE85C41EF715A3FAC3B7F9305">New Guinea Onthophagus: taxonomy of ten small, unicolored new species (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="B63CED675746B86F5A3146F1DCE89A3C" type="personal">
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<mods:namePart id="42CA9FFED727A71D93C4D8999C378906">Krikken, J.</mods:namePart>
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<mods:name id="F73D2640C7511E50123D6EBA9E92E1C4" type="personal">
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<mods:roleTerm id="E9E41CD03C7FD5C3A8D5F0374586B1AD">Author</mods:roleTerm>
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<mods:namePart id="E87E4829EF77693208E674EB21B04662">Huijbregts, J.</mods:namePart>
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<mods:typeOfResource id="36B8044D01AA44B4F3B238ADCB529C38">text</mods:typeOfResource>
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<mods:title id="407E59B332E08335EE44E351E9193569">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="EE43E8E6F4B4047E36C6474C29A29C0E">
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<mods:date id="1842C81D46CEBB341A97358D4BFB2185">2013</mods:date>
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<mods:detail id="462199095A69400F01452CF1AB81918E" type="volume">
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<mods:number id="5AB4C948379C973F4D2E8F1B2CE7A6C6">3619</mods:number>
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<mods:detail id="C50FD79CD7695ADC46EEBE2C8DD8DAE4" type="issue">
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<mods:number id="72ED509346F4B706FED09A22C7809543">5</mods:number>
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<mods:classification id="65315C872CFB1BCB38ADCA0C74BA573C">journal article</mods:classification>
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<mods:identifier id="731A8033E10FDF10ADDCCE7199467599" type="DOI">10.11646/zootaxa.3619.5.1</mods:identifier>
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<mods:identifier id="DE564995AB36AA773F71980182EDF554" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E687A4DD2DFFDBFF2807EA1347FDE1" ID-DOI="http://doi.org/10.5281/zenodo.6161071" ID-GBIF-Taxon="119560718" ID-Zenodo-Dep="6161071" LSID="urn:lsid:plazi:treatment:03E687A4DD2DFFDBFF2807EA1347FDE1" httpUri="http://treatment.plazi.org/id/03E687A4DD2DFFDBFF2807EA1347FDE1" lastPageId="12" lastPageNumber="513" pageId="10" pageNumber="511">
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<subSubSection id="C3556539DD2DFFDDFF2807EA160AF8DE" pageId="10" pageNumber="511" type="nomenclature">
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<paragraph id="8BF036B2DD2DFFDDFF2807EA14D8F93D" blockId="10.[151,934,1735,1794]" box="[151,934,1735,1761]" pageId="10" pageNumber="511">
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<heading id="D0B881DEDD2DFFDDFF2807EA14D8F93D" bold="true" box="[151,934,1735,1761]" fontSize="11" level="1" pageId="10" pageNumber="511" reason="1">
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<emphasis id="B93BEAA0DD2DFFDDFF2807EA14D8F93D" bold="true" box="[151,934,1735,1761]" pageId="10" pageNumber="511">
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<taxonomicName id="4C4F4D31DD2DFFDDFF2807EA1477F93D" ID-CoL="7XMH6" authority="Krikken & Huijbregts" authorityName="Krikken & Huijbregts" box="[151,777,1735,1761]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="511" phylum="Arthropoda" rank="species" species="bituberoculus" status="sp. nov.">
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<emphasis id="B93BEAA0DD2DFFDDFF2807EA1697F93D" bold="true" box="[151,489,1735,1761]" italics="true" pageId="10" pageNumber="511">Onthophagus bituberoculus</emphasis>
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Krikken & Huijbregts
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</taxonomicName>
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,
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<taxonomicNameLabel id="A20857DBDD2DFFDDFCA907EA14D8F93D" box="[790,934,1735,1761]" pageId="10" pageNumber="511" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF036B2DD2DFFDDFF2807C7160AF8DE" blockId="10.[151,934,1735,1794]" box="[151,372,1770,1794]" pageId="10" pageNumber="511">
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||
(
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<figureCitation id="13742A37DD2DFFDDFF2007C71790F8DE" box="[159,238,1770,1794]" captionStart="FIGURES 1 – 8" captionStartId="18.[151,264,1895,1918]" captionTargetBox="[200,1375,205,1851]" captionTargetId="figure@18.[198,1392,195,1860]" captionTargetPageId="18" captionText="FIGURES 1 – 8. Habitus (oblique view) of Onthophagus species, with indication of body length. 1, O. acerus, holotype ,, 6 mm; 2, O. aceroides, holotype, 5 mm; 3, O. baiyericus, holotype, 5 mm; 4, O. mimikanus, female holotype, 6 mm; 5, O. kokopygus, holotype, 3 mm; 6, O. daymanus, holotype, 4.5 mm; 7, O. kokocellosus, holotype, 4.5 mm; 8, O. bituberoculus, female paratype, Timika, 7 mm." httpUri="https://zenodo.org/record/223149/files/figure.png" pageId="10" pageNumber="511">Figs. 8</figureCitation>
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,
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<figureCitation id="13742A37DD2DFFDDFF4507C71666F8DE" box="[250,280,1770,1794]" captionStart="FIGURES 13 – 24" captionStartId="19.[151,264,1917,1940]" captionTargetBox="[166,1406,183,1813]" captionTargetId="figure@19.[151,1436,175,1851]" captionTargetPageId="19" captionText="FIGURES 13 – 24. Parameres (lateral view) of Onthophagus species. 13, O. acerus, holotype; 14, O. aceroides, holotype; 15, O. baiyericus, holotype; 16, O. daymanus, holotype; 17, O. kokopygus, holotype; 18, O. kokocellosus, holotype; 19, O. bituberoculus, holotype; 20, O. kukali, holotype; 21, O. kokodentatus, holotype; 22 – 23, O. dissidentatus, paratype, Wau (22), paratype. N Port Moresby (23); 24, O. ofianus, holotype. Scale lines (on top of relevant parameres) equal 0.5 mm." httpUri="https://zenodo.org/record/223150/files/figure.png" pageId="10" pageNumber="511">19</figureCitation>
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,
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<figureCitation id="13742A37DD2DFFDDFE9A07C71615F8DE" box="[293,363,1770,1794]" captionStart="FIGURES 58 – 73" captionStartId="22.[151,264,1913,1936]" captionTargetBox="[203,1368,200,1863]" captionTargetId="figure@22.[203,1390,193,1870]" captionTargetPageId="22" captionText="FIGURES 58 – 73. Onthophagus species, body parts. 58 – 62, O. bituberoculus, holotype; 63, O. bituberoculus (or near), male; 64 – 68, O. kokocellosus, holotype; 69 – 73, O. kukali, holotype. — Head (full-face) 58, 63, 64, 69; pronotum (dorsal) 59, 65, 70; left elytron (dorsal) 60, 66, 71; protibia (upper side) 61, 67, 72; metatibia (upper side) 62, (underside) 68, 73. Scale lines equal 1 mm." httpUri="https://zenodo.org/record/223153/files/figure.png" pageId="10" pageNumber="511">58–62</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556539DD2DFFDDFF28061E16FCF84E" pageId="10" pageNumber="511" type="materials_examined">
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<paragraph id="8BF036B2DD2DFFDDFF28061E16FCF84E" blockId="10.[151,1436,1842,2010]" pageId="10" pageNumber="511">
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<emphasis id="B93BEAA0DD2DFFDDFF28061E16B5F897" bold="true" box="[151,459,1842,1867]" pageId="10" pageNumber="511">
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<typeStatus id="54F48810DD2DFFDDFF28061E17ACF897" box="[151,210,1843,1867]" pageId="10" pageNumber="511">Type</typeStatus>
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material.
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<typeStatus id="54F48810DD2DFFDDFEE1061E16B5F897" box="[350,459,1843,1867]" pageId="10" pageNumber="511" type="holotype">Holotype</typeStatus>
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</emphasis>
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male (MBBJ) from
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<collectingCountry id="F3587622DD2DFFDDFD69061F1439F896" box="[726,839,1842,1866]" name="Indonesia" pageId="10" pageNumber="511">Indonesia</collectingCountry>
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: Irian Jaya: Freeport Concession,
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<date id="FFF11072DD2DFFDDFB4C061F1213F896" box="[1267,1389,1842,1866]" pageId="10" pageNumber="511" value="1997-03-11">11.iii.1997</date>
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, R. Ubaidillah, Dpt 5-5 97206, dry lowland forest, 4˚ 26’ 21” S–136˚ 52’ 59” E,
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<quantity id="4CB79B57DD2DFFDDFBAD067B1327F8B2" box="[1042,1113,1878,1902]" metricMagnitude="1" metricUnit="m" metricValue="9.144" pageId="10" pageNumber="511" unit="ft" value="300.0">300 ft</quantity>
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, dung pitfall trap.
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<typeStatus id="54F48810DD2DFFDDFA90067A12E2F8B3" box="[1327,1436,1879,1903]" pageId="10" pageNumber="511" type="paratype">Paratypes</typeStatus>
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listed in Appendix 1.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556539DD2DFFDCFF7606B31475FEB8" lastPageId="11" lastPageNumber="512" pageId="10" pageNumber="511" type="diagnosis">
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<paragraph id="8BF036B2DD2DFFDCFF7606B31475FEB8" blockId="10.[151,1436,1842,2010]" lastBlockId="11.[151,1437,151,1976]" lastPageId="11" lastPageNumber="512" pageId="10" pageNumber="511">
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<emphasis id="B93BEAA0DD2DFFDDFF7606B3163DF86B" bold="true" box="[201,323,1950,1975]" pageId="10" pageNumber="511">Diagnosis.</emphasis>
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A quasi outsider in the group with large eye foramina by having a fine but distinct clypeofrontal ridge and an interocular pair of (albeit low) conical protrusions. Genal border fully, evenly rounded, seemingly short, but posteriorly connected to occipital side by fine, pigmented line. Color varying from yellow to mediumbrown (one black male may represent a different species, see Comments, below). Dorsum shiny, punctation generally superficial, poorly defined, somewhat more distinct and denser on pronotum; interstrial punctation very vague, minute. Eyes very broadly elliptic, separated over frons by about two eye-widths. Clypeal apex shallowly bisinuate, margin slightly reflexed. Pygidium scattered with punctation varying in size, several punctures with long seta. Comparatively large, body length usually
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<quantity id="4CB79B57DD2CFFDCFD1800611479FEB8" box="[679,775,332,356]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="11" pageNumber="512" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556539DD2CFFDBFF7600421636FF2B" lastPageId="12" lastPageNumber="513" pageId="11" pageNumber="512" type="description">
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<paragraph id="8BF036B2DD2CFFDCFF76004215D6FE28" blockId="11.[151,1437,151,1976]" pageId="11" pageNumber="512">
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<emphasis id="B93BEAA0DD2CFFDCFF760042162AFE54" bold="true" box="[201,340,367,392]" pageId="11" pageNumber="512">Description</emphasis>
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(
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<typeStatus id="54F48810DD2CFFDCFEDC004216B7FE5B" box="[355,457,367,391]" pageId="11" pageNumber="512" type="holotype">holotype</typeStatus>
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, male). Body length approximately
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<quantity id="4CB79B57DD2CFFDCFCDE004214C5FE5B" box="[865,955,367,391]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="11" pageNumber="512" unit="mm" value="6.5">6.5 mm</quantity>
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. Habitus convex, dorsum deplanate. Color largely medium-brown, shiny (vaguely shagreened), underside and legs lighter brown. Dorsum with fine, superficial punctation, most distinct on pronotum, lacking distinct setae or shorter bristles, underside with numerous pale-brown bristles and longer setae.
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</paragraph>
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<paragraph id="8BF036B2DD2CFFDCFF7600D216A0FCC8" blockId="11.[151,1437,151,1976]" pageId="11" pageNumber="512">
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Head brown, frons lacking distinct punctation, virtually smooth, genae with some indistinct punctures. Clypeogenal border widely, evenly rounded from genal side to slightly reflexed, bisinuate clypeal apex; clypeogenal transition continuous at border, suture vague; clypeofrontal transition with distinct, low, slightly arcuate transverse ridge reaching clypeogenal suture; clypeal surface superficially, transversely rugulate. Genal tip fully rounded, posterior extension tapering to fine dark line, reaching occipital side (
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<emphasis id="B93BEAA0DD2CFFDCFBC103BC13DFFD74" box="[1150,1185,657,680]" italics="true" pageId="11" pageNumber="512">i.e.</emphasis>
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, eyes laterally quasiundivided). Frontal surface halfway eyes with pair of small conical tubercles, close to eye border. Eye foramina very broadly elliptic, with approximately 21 facet rows across widest point. Ratio interocular distance / maximum eye width approximately 2.1.
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</paragraph>
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<paragraph id="8BF036B2DD2CFFDCFF76020C14F2FC30" blockId="11.[151,1437,151,1976]" pageId="11" pageNumber="512">Pronotum brown, shiny with scattered, with sparse to abundant, fine, shallow, poorly-defined punctation (qualification depending on angle of view); surface evenly convex (lacking midline impression); anterior declivity gradual, unmodified; all borders finely marginate; anterior section of lateral border almost straight (full-face view), anterolateral angle shortly rounded, subrectangular; posterior section of lateral border slightly sinuate (full-face view), almost straight in dorsal view, posterolateral angle rounded off, continuing to basal border; basal border of pronotum widely rounded in dorsal view, with vague median angle.</paragraph>
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<paragraph id="8BF036B2DD2CFFDCFF7602DA1538FB7C" blockId="11.[151,1437,151,1976]" pageId="11" pageNumber="512">Elytra brown, shiny; scutellum indistinct, shape of base and apex unmodified; epipleuron with seta-bearing punctures, light brown, shiny. Elytra broad, with 8 fine, distinct, straight to slightly curved, striae; stria 7 evenly, slightly sinuate in front, ending on humeral umbone; strial punctures fine, on disc mostly separated by 6–10 puncture diameters, slightly crenulating interstriae. Interstrial surfaces slightly convex, with scattered, sparse, minute, simple, superficial punctation.</paragraph>
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<paragraph id="8BF036B2DD2CFFDCFF760581135CFA40" blockId="11.[151,1437,151,1976]" pageId="11" pageNumber="512">Antennal club yellow, scapus unmodified. Mentum shallowly emarginate in front. Postocular area of propectus slightly concave, matt, posteriorly delimited by simple coxal-marginal ridge. Propectoral sides with several fine seta-bearing hemi-punctures, setae subappressed. Metasternum moderately shiny, anterior lobe slightly, evenly convex in front; metasternal disc damaged by needle; metasternal flanks matt, with numerous fine, seta-bearing ocellate punctures. Abdominal ventrites matt, each laterally with row of small, superficial seta-bearing punctures. Pygidium with general surface evenly convex, light brown, matt, with numerous punctures of different sizes, several with seta; base with transverse ridge, sides and apex distinctly marginate.</paragraph>
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<paragraph id="8BF036B2DD2CFFDCFF7604841308F8F4" blockId="11.[151,1437,151,1976]" pageId="11" pageNumber="512">
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Legs slender, generally shiny brown. Protibia robust, with 3+1 larger external denticles (distal 3 long, slender, acuminate, proximal one small but distinct), distinctly separated by serration; proximal serration consisting of 6 small denticles; apico-external denticle oblique to tibial axis; apico-internal angle unmodified, terminal spur fine, elongate-acuminate, curved; protibial underside with low longitudinal crest; protarsus slender. Profemur robust, underside shiny, with numerous (partly seta-bearing) punctures. Mesofemoral and metafemoral undersides with fewer punctures, very few with seta. Mesotibiae and metatibiae slender, dilated distad to transversely subelliptic crest, which is fringed with short fossorial spines below, and some longer spines above; externally with approximately 5 sets of very slight, spine-bearing fossorial protrusions; mesotibia slightly sinuate. Mesotarsi and metatarsi generally very slender, with distinct sickle-shaped claws; mesotarsomeres and metatarsomeres 1 long, virtually straight, unmodified. Terminal spurs on mesotibiae and metatibiae elongate-acuminate. Length proportions terminal spur // metatarsomeres
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<quantity id="4CB79B57DD2CFFDCFD37063D15B1F8F4" box="[648,719,1808,1832]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="12.7" metricValueMin="2.54" pageId="11" pageNumber="512" unit="in" value="3.0" valueMax="5.0" valueMin="1.0">1–5 in</quantity>
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approximate integers: 15//
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<date id="FFF11072DD2CFFDCFC42063D133BF8F4" box="[1021,1093,1808,1832]" pageId="11" pageNumber="512" value="1918-04-07">18/7/4</date>
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/3/6.
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</paragraph>
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<paragraph id="8BF036B2DD2CFFDCFF7606191570F8AC" blockId="11.[151,1437,151,1976]" pageId="11" pageNumber="512">
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Aedeagus distorted; parameres broad at base, short, each with strongly tapering stalk (full-face view), curved downward (lateral view),
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<figureCitation id="13742A37DD2CFFDCFE0706751574F8AC" box="[440,522,1880,1904]" captionStart="FIGURES 13 – 24" captionStartId="19.[151,264,1917,1940]" captionTargetBox="[166,1406,183,1813]" captionTargetId="figure@19.[151,1436,175,1851]" captionTargetPageId="19" captionText="FIGURES 13 – 24. Parameres (lateral view) of Onthophagus species. 13, O. acerus, holotype; 14, O. aceroides, holotype; 15, O. baiyericus, holotype; 16, O. daymanus, holotype; 17, O. kokopygus, holotype; 18, O. kokocellosus, holotype; 19, O. bituberoculus, holotype; 20, O. kukali, holotype; 21, O. kokodentatus, holotype; 22 – 23, O. dissidentatus, paratype, Wau (22), paratype. N Port Moresby (23); 24, O. ofianus, holotype. Scale lines (on top of relevant parameres) equal 0.5 mm." httpUri="https://zenodo.org/record/223150/files/figure.png" pageId="11" pageNumber="512">Fig. 19</figureCitation>
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.
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</paragraph>
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<paragraph id="8BF036B2DD2CFFDCFF76065014D6F864" blockId="11.[151,1437,151,1976]" pageId="11" pageNumber="512">Measurements in mm (dorsal view). Maximum width of head 2.0. Median length of pronotum 2.1, maximum width 3.2. Sutural length of elytra 3.0, maximum width combined 3.8.</paragraph>
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<paragraph id="8BF036B2DD2BFFDBFF7601BA1636FF2B" blockId="12.[151,1436,151,573]" pageId="12" pageNumber="513">
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<emphasis id="B93BEAA0DD2BFFDBFF7601BA1511FF6C" bold="true" box="[201,623,151,176]" pageId="12" pageNumber="513">Variation and sexual dimorphism.</emphasis>
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Females very similar to male
|
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<typeStatus id="54F48810DD2BFFDBFC5E01BA1338FF73" box="[993,1094,151,175]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
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, any differences gradual and minimal. General color varies from yellow through medium brown to black (see also Comments, hereafter). Body length
|
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<quantity id="4CB79B57DD2BFFDBFF5B01F2163AFF2B" box="[228,324,223,247]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="12" pageNumber="513" unit="mm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 mm</quantity>
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||
.
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||
</paragraph>
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</subSubSection>
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||
<subSubSection id="C3556539DD2BFFDBFF76002E1434FDCB" pageId="12" pageNumber="513" type="discussion">
|
||
<paragraph id="8BF036B2DD2BFFDBFF76002E16D1FE5B" blockId="12.[151,1436,151,573]" pageId="12" pageNumber="513">
|
||
<emphasis id="B93BEAA0DD2BFFDBFF76002E162FFEC0" bold="true" box="[201,337,259,284]" pageId="12" pageNumber="513">Comments.</emphasis>
|
||
Three light brown to medium brown females and a single male from two regions of West New
|
||
<collectingCountry id="F3587622DD2BFFDBFF28000A1796FEE3" box="[151,232,295,319]" name="Guinea-Bissau" pageId="12" pageNumber="513">Guinea</collectingCountry>
|
||
are referred to
|
||
<taxonomicName id="4C4F4D31DD2BFFDBFE280005152DFE9C" box="[407,595,296,320]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="bituberoculus">
|
||
<emphasis id="B93BEAA0DD2BFFDBFE280005152DFE9C" box="[407,595,296,320]" italics="true" pageId="12" pageNumber="513">O. bituberoculus</emphasis>
|
||
</taxonomicName>
|
||
. In spite of the male specimen being poorly cohesive, the parts being a bit loose, it is designated
|
||
<typeStatus id="54F48810DD2BFFDBFE310061168AFEB8" box="[398,500,332,356]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
|
||
; caution: aedeagus of this
|
||
<typeStatus id="54F48810DD2BFFDBFCA500611402FEB8" box="[794,892,332,356]" pageId="12" pageNumber="513" type="holotype">holotype</typeStatus>
|
||
very weakly sclerotized, fragile, and contours in
|
||
<figureCitation id="13742A37DD2BFFDBFF2800421798FE5B" box="[151,230,367,391]" captionStart="FIGURES 13 – 24" captionStartId="19.[151,264,1917,1940]" captionTargetBox="[166,1406,183,1813]" captionTargetId="figure@19.[151,1436,175,1851]" captionTargetPageId="19" captionText="FIGURES 13 – 24. Parameres (lateral view) of Onthophagus species. 13, O. acerus, holotype; 14, O. aceroides, holotype; 15, O. baiyericus, holotype; 16, O. daymanus, holotype; 17, O. kokopygus, holotype; 18, O. kokocellosus, holotype; 19, O. bituberoculus, holotype; 20, O. kukali, holotype; 21, O. kokodentatus, holotype; 22 – 23, O. dissidentatus, paratype, Wau (22), paratype. N Port Moresby (23); 24, O. ofianus, holotype. Scale lines (on top of relevant parameres) equal 0.5 mm." httpUri="https://zenodo.org/record/223150/files/figure.png" pageId="12" pageNumber="513">
|
||
Fig.
|
||
<date id="FFF11072DD2BFFDBFF7500421798FE5B" box="[202,230,367,391]" pageId="12" pageNumber="513" value="1919-05">19</date>
|
||
</figureCitation>
|
||
may be distorted.
|
||
</paragraph>
|
||
<paragraph id="8BF036B2DD2BFFDBFF7600B81434FDCB" blockId="12.[151,1436,151,573]" pageId="12" pageNumber="513">
|
||
In Papuasia and elsewhere, in the
|
||
<collectingCountry id="F3587622DD2BFFDBFDEC00B915BEFE70" box="[595,704,404,428]" name="Indonesia" pageId="12" pageNumber="513">Moluccas</collectingCountry>
|
||
and Sundaland, more relatives of
|
||
<taxonomicName id="4C4F4D31DD2BFFDBFBF300B81274FE70" box="[1100,1290,404,428]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="bituberoculus">
|
||
<emphasis id="B93BEAA0DD2BFFDBFBF300B81274FE70" box="[1100,1290,404,428]" italics="true" pageId="12" pageNumber="513">O. bituberoculus</emphasis>
|
||
</taxonomicName>
|
||
occur, some unnamed (and not listed here) because only inadequate singletons are available. An example is a small, black male from
|
||
<collectingCountry id="F3587622DD2BFFDBFF6800F016CAFE28" box="[215,436,476,500]" name="Papua New Guinea" pageId="12" pageNumber="513">Papua New Guinea</collectingCountry>
|
||
(vicinity of Wau), basically similar to
|
||
<taxonomicName id="4C4F4D31DD2BFFDBFCCA00F0134DFE28" box="[885,1075,476,500]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="513" phylum="Arthropoda" rank="species" species="bituberoculus">
|
||
<emphasis id="B93BEAA0DD2BFFDBFCCA00F014F7FE28" box="[885,905,477,500]" italics="true" pageId="12" pageNumber="513">O</emphasis>
|
||
.
|
||
<emphasis id="B93BEAA0DD2BFFDBFC2500F1134DFE28" box="[922,1075,476,500]" italics="true" pageId="12" pageNumber="513">bituberoculus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="B93BEAA0DD2BFFDBFBFA00F01316FE28" box="[1093,1128,477,500]" italics="true" pageId="12" pageNumber="513">i.e.</emphasis>
|
||
, with clypeofrontal ridge), here illustrated (head,
|
||
<figureCitation id="13742A37DD2BFFDBFE2E00D2169AFDCB" box="[401,484,511,535]" captionStart="FIGURES 58 – 73" captionStartId="22.[151,264,1913,1936]" captionTargetBox="[203,1368,200,1863]" captionTargetId="figure@22.[203,1390,193,1870]" captionTargetPageId="22" captionText="FIGURES 58 – 73. Onthophagus species, body parts. 58 – 62, O. bituberoculus, holotype; 63, O. bituberoculus (or near), male; 64 – 68, O. kokocellosus, holotype; 69 – 73, O. kukali, holotype. — Head (full-face) 58, 63, 64, 69; pronotum (dorsal) 59, 65, 70; left elytron (dorsal) 60, 66, 71; protibia (upper side) 61, 67, 72; metatibia (upper side) 62, (underside) 68, 73. Scale lines equal 1 mm." httpUri="https://zenodo.org/record/223153/files/figure.png" pageId="12" pageNumber="513">Fig. 62</figureCitation>
|
||
); excluded from the
|
||
<typeStatus id="54F48810DD2BFFDBFD74032D1585FDC4" box="[715,763,512,536]" pageId="12" pageNumber="513">type</typeStatus>
|
||
series.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556539DD2BFFDBFF7603091347FDE1" box="[201,1081,548,573]" pageId="12" pageNumber="513" type="etymology">
|
||
<paragraph id="8BF036B2DD2BFFDBFF7603091347FDE1" blockId="12.[151,1436,151,573]" box="[201,1081,548,573]" pageId="12" pageNumber="513">
|
||
<emphasis id="B93BEAA0DD2BFFDBFF760309162EFDE0" bold="true" box="[201,336,548,572]" pageId="12" pageNumber="513">Etymology.</emphasis>
|
||
Name is a contraction, referring to pair of tubercles between eyes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |