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<document id="3D42E109142A3701038985426C641671" ID-CLB-Dataset="23381" ID-DOI="10.5252/zoosystema2019v41a26" ID-GBIF-Dataset="cef5076b-424b-4cdc-8d13-6e472d861e09" ID-ISSN="1638-9387" ID-Zenodo-Dep="3726028" ID-ZooBank="urn:lsid:zoobank.org:pub:CF16A992-0401-44C8-BEEE-842CE7F1D27E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1585071141162" checkinUser="felipe" docAuthor="Ortega, José Rafael &amp; Gofas, Serge" docDate="2019" docId="03E687A0B242FF9C9B75F8E2FCC5FA14" docLanguage="en" docName="zoosystema.41.26.513-551.pdf.imf" docOrigin="Zoosystema 41 (26)" docStyle="DocumentStyle:AFAF64E709DD7C8340B70A24A5804402.4:Zoosystema.2018-2019.journal_article" docStyleId="AFAF64E709DD7C8340B70A24A5804402" docStyleName="Zoosystema.2018-2019.journal_article" docStyleVersion="4" docTitle="Gymnobela multilirata Ortega &amp; Gofas 2019, n. sp." docType="treatment" docUuid="3B5B70FF-0A68-4463-9076-1306DEBC2C7E" docUuidSource="ZooBank" docVersion="8" lastPageNumber="538" masterDocId="FFDFFFD8B258FF879A52FFD1FFCDFFBB" masterDocTitle="The unknown bathyal of the Canaries: new species and new records of deep-sea Mollusca" masterLastPageNumber="551" masterPageNumber="513" pageNumber="537" updateTime="1698828230493" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="52101EDA08CC7248CED24C578E1DEA2B">The unknown bathyal of the Canaries: new species and new records of deep-sea Mollusca</mods:title>
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<mods:name id="1F070299996C27CCDBC6A4DADB1DF556" type="personal">
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<mods:namePart id="5B71FA1D444CBC8F60C51733E4C5EB9D">Ortega, José Rafael</mods:namePart>
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<mods:namePart id="FC2A1321AA45E65305F1A95342417E2E">Gofas, Serge</mods:namePart>
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<mods:title id="CEBF38714E62C3F0587AC8F82A68E52B">Zoosystema</mods:title>
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<mods:date id="99B418C7D945C1C237CC1EC8A4E55E3B">2019</mods:date>
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<mods:number id="A6ED0DCD5D7C2EA7700FC16695B17E0B">2019-11-28</mods:number>
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<mods:identifier id="2EE9FAC5B0369A2144F751FC1A8DD8E5" type="DOI">10.5252/zoosystema2019v41a26</mods:identifier>
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<treatment id="03E687A0B242FF9C9B75F8E2FCC5FA14" ID-DOI="http://doi.org/10.5281/zenodo.3729374" ID-GBIF-Taxon="163283944" ID-Zenodo-Dep="3729374" LSID="urn:lsid:zoobank.org:act:3B5B70FF-0A68-4463-9076-1306DEBC2C7E" httpUri="http://treatment.plazi.org/id/03E687A0B242FF9C9B75F8E2FCC5FA14" lastPageId="27" lastPageNumber="538" pageId="26" pageNumber="537">
<subSubSection id="C355653DB242FF9D9B75F8E2FDA9F8F5" box="[295,612,1843,1870]" pageId="26" pageNumber="537" type="nomenclature">
<paragraph id="8BF036B6B242FF9D9B75F8E2FDA9F8F5" blockId="26.[295,612,1843,1902]" box="[295,612,1843,1870]" pageId="26" pageNumber="537">
<heading id="D0B881DAB242FF9D9B75F8E2FDA9F8F5" box="[295,612,1843,1870]" centered="true" fontSize="11" level="2" pageId="26" pageNumber="537" reason="2">
<taxonomicName id="4C4F4D35B242FF9D9B75F8E2FDEFF8F5" ID-CoL="7RGSW" authority="Ortega &amp; Gofas, 2019" authorityName="Ortega &amp; Gofas" authorityYear="2019" box="[295,546,1843,1870]" class="Gastropoda" family="Raphitomidae" genus="Gymnobela" kingdom="Animalia" order="Neogastropoda" pageId="26" pageNumber="537" phylum="Mollusca" rank="species" species="multilirata" status="sp. nov.">
<emphasis id="B93BEAA4B242FF9D9B75F8E2FDEFF8F5" bold="true" box="[295,546,1843,1870]" italics="true" pageId="26" pageNumber="537">Gymnobela multilirata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857DFB242FF9D987BF8E5FDA9F8F5" box="[553,612,1844,1870]" pageId="26" pageNumber="537" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C355653DB242FF9D9BD3F882FD21F811" pageId="26" pageNumber="537" type="description">
<paragraph id="8BF036B6B242FF9D9BD3F882FDC4F8D5" blockId="26.[295,612,1843,1902]" box="[385,521,1875,1902]" pageId="26" pageNumber="537">
(
<figureCitation id="13742A33B242FF9D9BDBF882FE1EF8D5" box="[393,467,1875,1902]" captionStart="FIG" captionStartId="27.[132,143,707,725]" captionTargetBox="[132,1454,217,662]" captionTargetId="figure@27.[132,1454,205,675]" captionTargetPageId="27" captionText="FIG. 21. — A, B, Gymnobela multilirata n. sp., holotype from DW130 (height 2.2 mm); C, D, paratype, same locality (height 2.7 mm); E, protoconch of another paratype, same locality. Scale bar: 200 µm." figureDoi="http://doi.org/10.5281/zenodo.3726071" httpUri="https://zenodo.org/record/3726071/files/figure.png" pageId="26" pageNumber="537">Fig. 21</figureCitation>
A-E)
</paragraph>
<paragraph id="8BF036B6B242FF9D9ACDF849FD21F811" blockId="26.[159,748,1944,1962]" box="[159,748,1944,1962]" pageId="26" pageNumber="537">
<uri id="FFDE3AB4B242FF9D9ACDF849FD21F811" box="[159,748,1944,1962]" pageId="26" pageNumber="537">
urn:lsid:zoobank.org:act:
<uuid id="FFE90C63B242FF9D9B35F849FD21F811" box="[359,748,1944,1962]" pageId="26" pageNumber="537">3B5B70FF-0A68-4463-9076-1306DEBC2C7E</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C355653DB242FF9D9AD6F802FC48FBD8" pageId="26" pageNumber="537" type="materials_examined">
<paragraph id="8BF036B6B242FF9D9AD6F802FC48FBD8" blockId="26.[132,738,2003,2027]" lastBlockId="26.[813,1456,1072,1123]" pageId="26" pageNumber="537">
<typeStatus id="54F48814B242FF9D9AD6F802FF78F853" box="[132,181,2003,2027]" pageId="26" pageNumber="537">TYPE</typeStatus>
MATERIAL. —
<materialsCitation id="3B273CEBB242FF9D9B16F802FD18F850" ID-GBIF-Occurrence="2580042109" box="[324,725,2003,2027]" country="Djibouti" pageId="26" pageNumber="537" specimenCount="1" typeStatus="holotype">
<emphasis id="B93BEAA4B242FF9D9B16F802FE64F850" bold="true" box="[324,425,2003,2027]" pageId="26" pageNumber="537">
<typeStatus id="54F48814B242FF9D9B16F802FE68F850" box="[324,421,2003,2027]" pageId="26" pageNumber="537" type="holotype">Holotype</typeStatus>
.
</emphasis>
sh., MNHN-IM-2000-34274
</materialsCitation>
.
<materialsCitation id="3B273CEBB242FF9D997FFBE1FC4FFBD8" ID-GBIF-Occurrence="2580042103" country="Djibouti" pageId="26" pageNumber="537" specimenCount="1" typeStatus="paratype">
<emphasis id="B93BEAA4B242FF9D997FFBE1FC41FBF3" bold="true" box="[813,908,1072,1096]" pageId="26" pageNumber="537">
<typeStatus id="54F48814B242FF9D997FFBE1FC41FBF3" box="[813,908,1072,1096]" pageId="26" pageNumber="537" type="paratype">Paratypes</typeStatus>
</emphasis>
. 10 sh., MNHN-IM-2000-34275 from SEAMOUNT 2 DW130
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C355653DB242FF9D997FFBABFAE8FB17" pageId="26" pageNumber="537" type="distribution">
<paragraph id="8BF036B6B242FF9D997FFBABFAE8FB17" blockId="26.[813,1454,1146,1197]" pageId="26" pageNumber="537">
<typeStatus id="54F48814B242FF9D997FFBABFCAEFB34" box="[813,867,1146,1170]" pageId="26" pageNumber="537">TYPE</typeStatus>
LOCALITY. — Off W Gran Canaria,
<geoCoordinate id="EE7B5071B242FF9D9F76FBABFA67FB29" box="[1316,1450,1146,1170]" degrees="28" direction="north" minutes="08.95" orientation="latitude" pageId="26" pageNumber="537" precision="9" value="28.149166">28°08.95N</geoCoordinate>
,
<geoCoordinate id="EE7B5071B242FF9D997FFB44FC68FB16" box="[813,933,1173,1197]" degrees="15" direction="west" minutes="53.11" orientation="longitude" pageId="26" pageNumber="537" precision="9" value="-15.885167">15°53.11W</geoCoordinate>
/
<geoCoordinate id="EE7B5071B242FF9D99FFFB44FBEEFB16" box="[941,1059,1173,1197]" degrees="28" direction="north" minutes="09.06" orientation="latitude" pageId="26" pageNumber="537" precision="9" value="28.151">28°09.06N</geoCoordinate>
,
<geoCoordinate id="EE7B5071B242FF9D9E7CFB44FB68FB16" box="[1070,1189,1173,1197]" degrees="15" direction="west" minutes="52.92" orientation="longitude" pageId="26" pageNumber="537" precision="9" value="-15.882">15°52.92W</geoCoordinate>
,
<quantity id="4CB79B53B242FF9D9EFDFB44FAECFB17" box="[1199,1313,1173,1197]" metricMagnitude="2" metricUnit="m" metricValue="6.575" metricValueMax="6.6" metricValueMin="6.55" pageId="26" pageNumber="537" unit="m" value="657.5" valueMax="660.0" valueMin="655.0">
655-
<specimenCount id="9D49FD3FB242FF9D9E8EFB44FAECFB17" box="[1244,1313,1173,1197]" pageId="26" pageNumber="537" type="generic">660 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C355653DB242FF9D997FFB15FAF8FB4C" pageId="26" pageNumber="537" type="etymology">
<paragraph id="8BF036B6B242FF9D997FFB15FAF8FB4C" blockId="26.[813,1456,1220,1271]" pageId="26" pageNumber="537">
ETYMOLOGY. — An adjective formed from the Latin
<emphasis id="B93BEAA4B242FF9D9F7FFB15FA80FB67" box="[1325,1357,1220,1244]" italics="true" pageId="26" pageNumber="537">lira</emphasis>
, a furrow, based on the sculpture with numerous spiral furrows.
</paragraph>
</subSubSection>
<subSubSection id="C355653DB242FF9D997FFAF7FA7DFA84" box="[813,1456,1318,1343]" pageId="26" pageNumber="537" type="materials_examined">
<paragraph id="8BF036B6B242FF9D997FFAF7FA7DFA84" blockId="26.[813,1456,1318,1343]" box="[813,1456,1318,1343]" pageId="26" pageNumber="537">OTHER MATERIAL EXAMINED. — 66 sh., SEAMOUNT 2 DW130.</paragraph>
</subSubSection>
<subSubSection id="C355653DB242FF9D997FFAA5FBBBF855" pageId="26" pageNumber="537" type="description">
<paragraph id="8BF036B6B242FF9D997FFAA5FC70FA36" blockId="26.[813,1456,1396,2030]" box="[813,957,1396,1422]" pageId="26" pageNumber="537">DESCRIPTION</paragraph>
<paragraph id="8BF036B6B242FF9D997FFA44FBBBF855" blockId="26.[813,1456,1396,2030]" pageId="26" pageNumber="537">
Shell small, whitish, consisting of a little more than three convex, definitely shouldered whorls, with a well marked suture; protoconch (visible part
<quantity id="4CB79B53B242FF9D9ED2FA04FB1EFA54" box="[1152,1235,1493,1520]" metricMagnitude="-1" metricUnit="m" metricValue="5.5" pageId="26" pageNumber="537" unit="mm" value="550.0">550 µm</quantity>
high,
<quantity id="4CB79B53B242FF9D9F44FA04FAA7FA54" box="[1302,1386,1493,1519]" metricMagnitude="-1" metricUnit="m" metricValue="4.0" pageId="26" pageNumber="537" unit="mm" value="400.0">400 µm</quantity>
diameter) protruding and somewhat pointed apically, formed of a little more than one whorl, nearly smooth on the initial part but mostly covered by minute, irregular granules which are loosely aligned spirally. Protoconch/teleoconch limit very definite along an orthocline line. Teleoconch with elevated spiral cords, of which 4-5 on the spire whorls and 19-21 on the last whorl; cords narrower than the interspaces, one along the suture, the strongest one on the shoulder and the 2-3 abapical ones decreasing slightly in size; on the last half-whorl, an additional cord between the suture and the shoulder. The entire teleoconch covered by axial wrinkles which are much finer than the cords and override them. Aperture oval-elongate, somewhat truncated adapically at the shoulder of last whorl; outer lip simple, siphonal canal short and very broad. Dimensions of the
<typeStatus id="54F48814B242FF9D99E3F862FBDEF876" box="[945,1043,1971,1997]" pageId="26" pageNumber="537" type="holotype">holotype</typeStatus>
:
<quantity id="4CB79B53B242FF9D9E4CF862FBB8F875" box="[1054,1141,1971,1998]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="26" pageNumber="537" unit="mm" value="2.2">2.2 mm</quantity>
height ×
<quantity id="4CB79B53B242FF9D9E8DF862FA8EF875" box="[1247,1347,1971,1998]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="26" pageNumber="537" unit="mm" value="1.15">1.15 mm</quantity>
diameter,
<typeStatus id="54F48814B242FF9D997FF805FC5FF855" box="[813,914,2004,2030]" pageId="26" pageNumber="537" type="paratype">paratypes</typeStatus>
up to 2.7 ×
<quantity id="4CB79B53B242FF9D9E4BF802FBBEF855" box="[1049,1139,2003,2030]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="537" unit="mm" value="1.5">1.5 mm</quantity>
.
</paragraph>
</subSubSection>
<caption id="DF30663EB243FF9C9AD6FD12FE20FD57" ID-DOI="http://doi.org/10.5281/zenodo.3726071" ID-Zenodo-Dep="3726071" httpUri="https://zenodo.org/record/3726071/files/figure.png" pageId="27" pageNumber="538" startId="27.[132,143,707,725]" targetBox="[132,1454,217,662]" targetPageId="27">
<paragraph id="8BF036B6B243FF9C9AD6FD12FE20FD57" pageId="27" pageNumber="538">
FIG. 21. —
<emphasis id="B93BEAA4B243FF9C9AB3FD12FF23FD6E" bold="true" box="[225,238,707,725]" pageId="27" pageNumber="538">A</emphasis>
,
<emphasis id="B93BEAA4B243FF9C9AABFD12FECBFD6E" bold="true" box="[249,262,707,725]" pageId="27" pageNumber="538">B</emphasis>
,
<emphasis id="B93BEAA4B243FF9C9B43FD12FE0BFD6E" box="[273,454,707,725]" italics="true" pageId="27" pageNumber="538">Gymnobela multilirata</emphasis>
n. sp., holotype from DW130 (height 2.2 mm);
<emphasis id="B93BEAA4B243FF9C9901FD12FCACFD6E" bold="true" box="[851,865,707,725]" pageId="27" pageNumber="538">C</emphasis>
,
<emphasis id="B93BEAA4B243FF9C993EFD12FCB7FD6E" bold="true" box="[876,890,707,725]" pageId="27" pageNumber="538">D</emphasis>
, paratype, same locality (height 2.7 mm);
<emphasis id="B93BEAA4B243FF9C9E8EFD12FB25FD6E" bold="true" box="[1244,1256,707,725]" pageId="27" pageNumber="538">E</emphasis>
, protoconch of another paratype, same locality. Scale bar: 200 µm.
</paragraph>
</caption>
<subSubSection id="C355653DB243FF9C9AD6FCC6FCC5FA14" pageId="27" pageNumber="538" type="discussion">
<paragraph id="8BF036B6B243FF9C9AD6FCC6FF25FC94" blockId="27.[132,776,791,1456]" box="[132,232,791,817]" pageId="27" pageNumber="538">REMARKS</paragraph>
<paragraph id="8BF036B6B243FF9C9AD6FCE6FCC5FA14" blockId="27.[132,776,791,1456]" pageId="27" pageNumber="538">
<taxonomicName id="4C4F4D35B243FF9C9ADDFCE6FECDFCEA" authorityName="Verrill" authorityYear="1884" box="[143,256,823,849]" class="Gastropoda" family="Raphitomidae" genus="Gymnobela" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="538" phylum="Mollusca" rank="genus">
<emphasis id="B93BEAA4B243FF9C9ADDFCE6FECDFCEA" box="[143,256,823,849]" italics="true" pageId="27" pageNumber="538">Gymnobela</emphasis>
</taxonomicName>
<taxonomicName id="4C4F4D35B243FF9C9B41FCE6FE4DFCEA" authority="Ortega &amp; Gofas, 2019" authorityName="Ortega &amp; Gofas" authorityYear="2019" box="[275,384,823,849]" class="Gastropoda" family="Raphitomidae" genus="Gymnobela" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="538" phylum="Mollusca" rank="species" species="multilirata" status="sp. nov.">
<emphasis id="B93BEAA4B243FF9C9B41FCE6FE4DFCEA" box="[275,384,823,849]" italics="true" pageId="27" pageNumber="538">multilirata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857DFB243FF9C9BDAFCE9FE09FCE9" box="[392,452,824,850]" pageId="27" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
, does not resemble closely any Atlantic species known to us, and even the generic and familial placement is tentative. It is guided by the overall resemblance with species like e.g.
<taxonomicName id="4C4F4D35B243FF9C9B31FC46FEF0FC69" authority="(Dautzenberg &amp; H. Fischer, 1896)" baseAuthorityName="Dautzenberg &amp; H. Fischer" baseAuthorityYear="1896" class="Gastropoda" family="Raphitomidae" genus="Gymnobela" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="538" phylum="Mollusca" rank="species" species="subaraneosa">
<emphasis id="B93BEAA4B243FF9C9B31FC46FD9FFC0A" box="[355,594,919,945]" italics="true" pageId="27" pageNumber="538">Gymnobela subaraneosa</emphasis>
(Dautzenberg &amp; H. Fischer, 1896)
</taxonomicName>
or
<taxonomicName id="4C4F4D35B243FF9C9B30FC66FD53FC6A" authority="(Dautzenberg, 1925)" baseAuthorityName="Dautzenberg" baseAuthorityYear="1925" box="[354,670,950,977]" class="Gastropoda" family="Raphitomidae" genus="Gymnobela" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="538" phylum="Mollusca" rank="species" species="lamyi">
<emphasis id="B93BEAA4B243FF9C9B30FC66FE77FC6A" box="[354,442,951,977]" italics="true" pageId="27" pageNumber="538">G. lamyi</emphasis>
(Dautzenberg, 1925)
</taxonomicName>
(see
<bibRefCitation id="EFDE4B47B243FF9C9881FC66FEBEFC4A" author="BOUCHET P. &amp; WAREN A." pageId="27" pageNumber="538" pagination="1 - 119" refId="ref21933" refString="BOUCHET P. &amp; WAREN A. 1980. - Revision of the North-East Atlantic bathyal and abyssal Turridae (Mollusca: Gastropoda). Journal of Molluscan Studies supplement 8: 1 - 119." type="journal article" year="1980">Bouchet &amp; Warén 1980: 52</bibRefCitation>
) which also have a definite shoulder and a sculpture of widely separated spiral cords. Another superficially similar species is
<taxonomicName id="4C4F4D35B243FF9C9B62FBC7FD6FFB8B" authority="(G. O. Sars, 1878)" baseAuthorityName="G. O. Sars" baseAuthorityYear="1878" box="[304,674,1046,1072]" class="Gastropoda" family="Raphitomidae" genus="Nepotilla" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="538" phylum="Mollusca" rank="species" species="amoena">
<emphasis id="B93BEAA4B243FF9C9B62FBC7FE13FB8B" box="[304,478,1046,1072]" italics="true" pageId="27" pageNumber="538">Nepotilla amoena</emphasis>
(G. O. Sars, 1878)
</taxonomicName>
(see
<bibRefCitation id="EFDE4B47B243FF9C9881FBC6FEB7FBEA" author="BOUCHET P. &amp; WAREN A." pageId="27" pageNumber="538" pagination="1 - 119" refId="ref21933" refString="BOUCHET P. &amp; WAREN A. 1980. - Revision of the North-East Atlantic bathyal and abyssal Turridae (Mollusca: Gastropoda). Journal of Molluscan Studies supplement 8: 1 - 119." type="journal article" year="1980">Bouchet &amp; Warén 1980: 75</bibRefCitation>
, 76) but the latter has fewer and much stronger spirals and a much narrower canal and abapical part. The paucispiral protoconch of
<taxonomicName id="4C4F4D35B243FF9C9B92FBA7FD80FB2B" authority="Ortega &amp; Gofas, 2019" authorityName="Ortega &amp; Gofas" authorityYear="2019" box="[448,589,1142,1168]" class="Gastropoda" family="Raphitomidae" genus="Gymnobela" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="538" phylum="Mollusca" rank="species" species="multilirata" status="sp. nov.">
<emphasis id="B93BEAA4B243FF9C9B92FBA7FD80FB2B" box="[448,589,1142,1168]" italics="true" pageId="27" pageNumber="538">G. multilirata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20857DFB243FF9C9801FBA6FD46FB2A" box="[595,651,1143,1169]" pageId="27" pageNumber="538" rank="species">n. sp.</taxonomicNameLabel>
is typical of a species with non-planktotrophic larval development but does not give a definite clue, although the profile of the protoconch and its definite, orthocline edge resembles the cancellariid
<taxonomicName id="4C4F4D35B243FF9C9881FB07FE22FAAB" authority="(Reeve, 1856)" baseAuthorityName="Reeve" baseAuthorityYear="1856" class="Gastropoda" family="Cancellariidae" genus="Pseudobabylonella" kingdom="Animalia" order="Neogastropoda" pageId="27" pageNumber="538" phylum="Mollusca" rank="species" species="minima">
<emphasis id="B93BEAA4B243FF9C9881FB07FE93FAB4" italics="true" pageId="27" pageNumber="538">Pseudobabylonella minima</emphasis>
(Reeve, 1856)
</taxonomicName>
(see
<bibRefCitation id="EFDE4B47B243FF9C984DFB24FD36FAAB" author="VERHECKEN A." box="[543,763,1269,1296]" pageId="27" pageNumber="538" pagination="281 - 364" refId="ref25184" refString="VERHECKEN A. 2007. - Revision of the Cancellariidae (Mollusca, Neogastropoda, Cancellarioidea) of the eastern Atlantic (40 ° N- 40 ° S) and the Mediterranean. Zoosystema 29 (2): 281 - 364." type="journal article" year="2007">Verhecken 2007: 351</bibRefCitation>
). However, teleoconch characters including the lack of folds on the columella and the elongate, tapering last whorl (usually blunt and short in cancellariids) do not support a placement in the latter family. Despite the uncertainty on its position, we consider this species as distinctive and abundant so as to be named.
</paragraph>
</subSubSection>
</treatment>
</document>