564 lines
75 KiB
XML
564 lines
75 KiB
XML
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<mods:title id="B12248BC344A825A2C5CF0CBB1457521">Systematics of the Podarcis hispanicus complex (Sauria, Lacertidae) III: valid nomina of the western and central Iberian forms</mods:title>
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<mods:namePart id="1E1FCFD38756873669A8C9FDBFC467D0">Sá-Sousa, Paulo</mods:namePart>
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<mods:namePart id="F3D32ECB6C79711C5C5AB700E20A69CA">Guillaume, Claude P.</mods:namePart>
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<mods:namePart id="8C18303E9C5987F60F9446D482635447">Crochet, Pierre-André</mods:namePart>
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<mods:number id="369DCD25522CE0DAC4145EBC1BBBA6C1">3794</mods:number>
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<treatment id="03E68784FFAFC20C85C2F869FA14FAEA" ID-DOI="http://doi.org/10.5281/zenodo.5694217" ID-GBIF-Taxon="127667737" ID-Zenodo-Dep="5694217" LSID="urn:lsid:plazi:treatment:03E68784FFAFC20C85C2F869FA14FAEA" httpUri="http://treatment.plazi.org/id/03E68784FFAFC20C85C2F869FA14FAEA" lastPageId="13" lastPageNumber="14" pageId="7" pageNumber="8">
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<subSubSection id="C3556519FFAFC20685C2F869FC8EF829" box="[151,770,1953,1979]" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="8BF03692FFAFC20685C2F869FC8EF829" blockId="7.[151,770,1953,1979]" box="[151,770,1953,1979]" pageId="7" pageNumber="8">
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<heading id="D0B881FEFFAFC20685C2F869FC8EF829" bold="true" box="[151,770,1953,1979]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
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<taxonomicName id="4C4F4D11FFAFC20685C2F869FC8EF829" authority="Bosca, 1916" authorityName="Bosca" authorityYear="1916" box="[151,770,1953,1979]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
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<emphasis id="B93BEA80FFAFC20685C2F869FC8EF829" bold="true" box="[151,770,1953,1979]" pageId="7" pageNumber="8">
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<emphasis id="B93BEA80FFAFC20685C2F869FDD9F829" bold="true" box="[151,597,1953,1979]" italics="true" pageId="7" pageNumber="8">Podarcis guadarramae guadarramae</emphasis>
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(
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<bibRefCitation id="EFDE4B63FFAFC2068731F869FD76F829" author="Bosca" box="[612,762,1953,1979]" pageId="7" pageNumber="8" refString="Bosca, E. (1916) Dos observaciones a proposito de Lacerta muralis en Espana. Boletin de la Real Sociedad Espanola de Historia Natural, 16, 327 - 330." type="journal article" year="1916">Boscá, 1916</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556519FFAFC20885C2F815FE40F8A3" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="reference_group">
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<paragraph id="8BF03692FFAFC20685C2F815FB74F860" blockId="7.[151,1272,2012,2034]" box="[151,1272,2012,2034]" pageId="7" pageNumber="8">
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<taxonomicName id="4C4F4D11FFAFC20685C2F815FDC3F860" authority="Bosca, 1916" authorityName="Bosca" authorityYear="1916" box="[151,591,2012,2034]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="8" phylum="Chordata" rank="subSpecies" species="muralis" subSpecies="guadarramae">
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<emphasis id="B93BEA80FFAFC20685C2F815FE46F860" box="[151,458,2012,2034]" italics="true" pageId="7" pageNumber="8">Lacerta muralis guadarramae</emphasis>
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<bibRefCitation id="EFDE4B63FFAFC2068485F814FDC3F860" author="Bosca" box="[464,591,2012,2034]" pageId="7" pageNumber="8" refString="Bosca, E. (1916) Dos observaciones a proposito de Lacerta muralis en Espana. Boletin de la Real Sociedad Espanola de Historia Natural, 16, 327 - 330." type="journal article" year="1916">Boscá, 1916</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C4F4D11FFAFC206870CF814FD28F860" box="[601,676,2012,2034]" class="Reptilia" family="Lacertidae" genus="Boletin" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="26" phylum="Chordata" rank="genus">Boletín</taxonomicName>
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de la Real Sociedad Española de Historia Natural 16: 330.
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</paragraph>
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<caption id="DF30661AFFA0C20985C2F97BFEC5F8B6" httpUri="https://zenodo.org/record/285984/files/figure.png" pageId="8" pageNumber="9" targetBox="[303,1284,193,1692]" targetPageId="8">
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<paragraph id="8BF03692FFA0C20985C2F97BFEC5F8B6" blockId="8.[151,1437,1715,1828]" pageId="8" pageNumber="9">
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<emphasis id="B93BEA80FFA0C20985C2F97BFE98F95B" bold="true" box="[151,276,1715,1737]" pageId="8" pageNumber="9">FIGURE 3.</emphasis>
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Neotype of
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<taxonomicName id="4C4F4D11FFA0C20984C3F97CFCC0F95B" authority="Bosca 1916" authorityName="Bosca" authorityYear="1916" box="[406,844,1715,1737]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="8" pageNumber="9" phylum="Chordata" rank="subSpecies" species="muralis" subSpecies="guadarramae">
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<emphasis id="B93BEA80FFA0C20984C3F97CFD40F95B" box="[406,716,1715,1737]" italics="true" pageId="8" pageNumber="9">Lacerta muralis guadarramae</emphasis>
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Boscá 1916
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</taxonomicName>
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MNHN 2012.0262 from the village of Lozoya, Sierra de Guadarrama (province of Madrid, Spain). Note the well flattened head, the strongly black-spotted pileus, the lack of vertebral stripe, the wide light dorsolateral and black supradorsolateral stripes on the dorsum, continuing on tail, and the light orange iris. Photos P. Geniez.
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</paragraph>
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</caption>
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<caption id="DF30661AFFA1C20885C2FAA7FB8AF9CE" httpUri="https://zenodo.org/record/285985/files/figure.png" pageId="9" pageNumber="10" targetBox="[151,1436,193,1369]" targetPageId="9">
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<paragraph id="8BF03692FFA1C20885C2FAA7FB8AF9CE" blockId="9.[151,1436,1391,1628]" pageId="9" pageNumber="10">
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<emphasis id="B93BEA80FFA1C20885C2FAA7FE98FA17" bold="true" box="[151,276,1391,1413]" pageId="9" pageNumber="10">FIGURE 4.</emphasis>
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Variation in males of
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<taxonomicName id="4C4F4D11FFA1C20884ABFAA7FCF1FA17" box="[510,893,1391,1413]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
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<emphasis id="B93BEA80FFA1C20884ABFAA7FCF1FA17" box="[510,893,1391,1413]" italics="true" pageId="9" pageNumber="10">Podarcis guadarramae guadarramae</emphasis>
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</taxonomicName>
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. For GPS coordinates (WGS84 in decimal degrees), see Appendix 1.
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<emphasis id="B93BEA80FFA1C2088416FA46FED1FA36" bold="true" box="[323,349,1422,1444]" pageId="9" pageNumber="10">A)</emphasis>
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Spain, Sierra de Gredos, 1.5 km N. Laguna Grande, 1,897 m (province of Ávila), 17.08.2007 (PGe.1107). Photo O. Buisson.
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<emphasis id="B93BEA80FFA1C208840CFA64FEE6FA50" bold="true" box="[345,362,1452,1474]" pageId="9" pageNumber="10">B</emphasis>
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) Spain, Sierra de Guadarrama, Boca del Asno, 7 km past La Granja de San Ildefonso towards Puerto de Navacerrada (province of Segovia), 19.08.1990 (PGe.1028). Photo P. Geniez.
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<emphasis id="B93BEA80FFA1C20886EFFA03FC59FA73" bold="true" box="[954,981,1483,1505]" pageId="9" pageNumber="10">C)</emphasis>
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Spain, Hiendelaencina, in the village, Plaza Major, 1,080 m (province of Guadalajara), 17.07.2010 (BEV.10983). Photo P. Geniez & P.-A. Crochet.
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<emphasis id="B93BEA80FFA1C20881F2FA22FB4DF992" bold="true" box="[1191,1217,1514,1536]" pageId="9" pageNumber="10">D)</emphasis>
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Spain, San Martín de Valdeiglesias (province of Madrid), 27.06.2001 (BEV.2021). Photo P. Geniez.
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<emphasis id="B93BEA80FFA1C2088682F9C1FC63F98D" bold="true" box="[983,1007,1545,1567]" pageId="9" pageNumber="10">E)</emphasis>
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Spain, El Tiemblo (province of Ávila), 27.06.2001 (BEV.2009). Photo P. Geniez.
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<emphasis id="B93BEA80FFA1C208871AF9EFFDEAF9AF" bold="true" box="[591,614,1575,1597]" pageId="9" pageNumber="10">F)</emphasis>
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Specimen with greenish tint on the back. Spain, Sierra de Gredos, 4 km NE. Laguna Grande, 2000 m (province of Ávila), 16.07.1992 (PGe.1035). Photo P. Geniez.
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</paragraph>
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</caption>
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<paragraph id="8BF03692FFA1C20885C2F940FE40F8A3" blockId="9.[151,1437,1672,2022]" pageId="9" pageNumber="10">
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<emphasis id="B93BEA80FFA1C20885C2F940FE0BF933" bold="true" box="[151,391,1672,1697]" pageId="9" pageNumber="10">
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Name-bearing
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<typeStatus id="54F48830FFA1C208841EF940FE0EF933" box="[331,386,1672,1697]" pageId="9" pageNumber="10">type</typeStatus>
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:
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</emphasis>
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Muséum National d’Histoire Naturelle MNHN 2012.0262,
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<typeStatus id="54F48830FFA1C208816CF941FB1EF933" box="[1081,1170,1673,1697]" pageId="9" pageNumber="10" type="neotype">neotype</typeStatus>
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by present designation (
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<figureCitation id="13742A17FFA1C20885CAF964FF6AF957" box="[159,230,1708,1733]" captionStart="FIGURE 3" captionStartId="8.[151,250,1715,1737]" captionTargetBox="[303,1284,193,1692]" captionTargetId="figure@8.[303,1284,193,1693]" captionTargetPageId="8" captionText="FIGURE 3. Neotype of Lacerta muralis guadarramae Boscá 1916 MNHN 2012.0262 from the village of Lozoya, Sierra de Guadarrama (province of Madrid, Spain). Note the well flattened head, the strongly black-spotted pileus, the lack of vertebral stripe, the wide light dorsolateral and black supradorsolateral stripes on the dorsum, continuing on tail, and the light orange iris. Photos P. Geniez." httpUri="https://zenodo.org/record/285984/files/figure.png" pageId="9" pageNumber="10">Fig. 3</figureCitation>
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). Originally an adult male from “San Ildefonso” (= La Granja de San Ildefonso, Sierra de Guadarrama, province of Segovia,
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<collectingCountry id="F3587602FFA1C20884C4F918FE59F97B" box="[401,469,1744,1769]" name="Spain" pageId="9" pageNumber="10">Spain</collectingCountry>
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||
),
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<typeStatus id="54F48830FFA1C20884B9F918FDC2F97B" box="[492,590,1744,1769]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
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||
by monotypy, currently lost.
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<typeStatus id="54F48830FFA1C20886FCF919FC6CF97B" box="[937,992,1745,1769]" pageId="9" pageNumber="10">Type</typeStatus>
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locality: Village of Lozoya, Sierra de Guadarrama, province of Madrid (
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<collectingCountry id="F3587602FFA1C208877EF93CFDE2F89F" box="[555,622,1780,1805]" name="Spain" pageId="9" pageNumber="10">Spain</collectingCountry>
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) [WGS84 =
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<geoCoordinate id="EE7B5055FFA1C2088658F93CFCF6F89F" box="[781,890,1780,1805]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="55" value="40.951">40.951°N</geoCoordinate>
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/
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<geoCoordinate id="EE7B5055FFA1C20886CCF93DFB8DF89F" box="[921,1025,1781,1805]" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="55" value="-3.792">3.792°W</geoCoordinate>
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] after present
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<typeStatus id="54F48830FFA1C20881FAF93EFA84F89C" box="[1199,1288,1782,1806]" pageId="9" pageNumber="10" type="neotype">neotype</typeStatus>
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designation. Originally “San Ildefonso”.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3556519FFA1C20C8592F8F5FE40FC7F" lastPageId="13" lastPageNumber="14" pageId="9" pageNumber="10" type="diagnosis">
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<paragraph id="8BF03692FFA1C20B8592F8F5FA90FCCF" blockId="9.[151,1437,1672,2022]" lastBlockId="10.[151,1437,151,2013]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
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||
<emphasis id="B93BEA80FFA1C2088592F8F5FECDF8C4" bold="true" box="[199,321,1853,1878]" pageId="9" pageNumber="10">Diagnosis.</emphasis>
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This is the lineage referred to as
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<taxonomicName id="4C4F4D11FFA1C2088780F8F5FC30F8C4" box="[725,956,1853,1878]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hispanicus">
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<emphasis id="B93BEA80FFA1C2088780F8F5FC30F8C4" box="[725,956,1853,1878]" italics="true" pageId="9" pageNumber="10">Podarcis hispanicus</emphasis>
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||
</taxonomicName>
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“
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<typeStatus id="54F48830FFA1C2088684F8F5FB8FF8C7" box="[977,1027,1853,1877]" pageId="9" pageNumber="10">type</typeStatus>
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1B” by
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<bibRefCitation id="EFDE4B63FFA1C2088138F8F5FAC8F8C4" author="Pinho" box="[1133,1348,1853,1878]" pageId="9" pageNumber="10" refString="Pinho, C., Ferrand, N. & Harris, D. J. (2006) Reexamination of the Iberian and North African Podarcis (Squamata: Lacertidae) phylogeny based on increased mitochondrial DNA sequencing. Molecular Phylogenetics and Evolution, 38, 266 - 273. http: // dx. doi. org / 10.1016 / j. ympev. 2005.06.012" type="journal article" year="2006">
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Pinho
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<emphasis id="B93BEA80FFA1C20881EFF8F6FB7AF8C4" box="[1210,1270,1853,1878]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
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(2006
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</bibRefCitation>
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||
, 2007),
|
||
<bibRefCitation id="EFDE4B63FFA1C20885C2F8A9FEDFF8EB" author="Carretero" box="[151,339,1889,1913]" pageId="9" pageNumber="10" refString="Carretero, M. A. (2008) An integrated assessment of a group with complex systematics: the Iberomaghrebian lizards genus Podarcis (Squamata, Lacertidae). Integrative Zoology, 4, 247 - 266. http: // dx. doi. org / 10.1111 / j. 1749 - 4877.2008.00102. x" type="journal article" year="2008">Carretero (2008)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFDE4B63FFA1C20884DFF8A8FD5CF8EB" author="Kaliontzopoulou" box="[394,720,1888,1913]" pageId="9" pageNumber="10" refString="Kaliontzopoulou, A., Pinho, C., Harris, D. J. & Carretero, M. A. (2011) When cryptic diversity blurs the picture: a cautionary tale from Iberian and North African Podarcis wall lizards. Biological Journal of the Linnean Society, 103, 779 - 800. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2011.01703. x" type="journal article" year="2011">
|
||
Kaliontzopoulou
|
||
<emphasis id="B93BEA80FFA1C2088718F8AAFD0AF8EB" box="[589,646,1888,1913]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
|
||
(2011
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDE4B63FFA1C2088789F8A9FC95F8EB" author="Kaliontzopoulou" box="[732,793,1889,1913]" pageId="9" pageNumber="10" refString="Kaliontzopoulou, A., Carretero, M. A. & Llorente, G. A. (2012) Morphology of the Podarcis wall lizards (Squamata: Lacertidae) from the Iberian Peninsula and North Africa: patterns of variation in a putative cryptic species complex. Zoological Journal of the Linnean Society, 164, 173 - 193. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00760. x" type="journal article" year="2012">2012</bibRefCitation>
|
||
). A typical Iberian wall lizard of the
|
||
<taxonomicName id="4C4F4D11FFA1C20881ECF8A8FA10F8EB" box="[1209,1436,1888,1913]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="9" pageNumber="10" phylum="Chordata" rank="species" species="hispanicus">
|
||
<emphasis id="B93BEA80FFA1C20881ECF8A8FA10F8EB" box="[1209,1436,1888,1913]" italics="true" pageId="9" pageNumber="10">Podarcis hispanicus</emphasis>
|
||
</taxonomicName>
|
||
complex characterized by the following features (see also
|
||
<figureCitation id="13742A17FFA1C2088670F84DFCFEF80F" box="[805,882,1925,1950]" captionStart="FIGURE 4" captionStartId="9.[151,250,1391,1413]" captionTargetBox="[151,1436,193,1369]" captionTargetId="figure@9.[151,1436,193,1370]" captionTargetPageId="9" captionText="FIGURE 4. Variation in males of Podarcis guadarramae guadarramae. For GPS coordinates (WGS 84 in decimal degrees), see Appendix 1. A) Spain, Sierra de Gredos, 1.5 km N. Laguna Grande, 1,897 m (province of Ávila), 17.08. 2007 (PGe. 1107). Photo O. Buisson. B) Spain, Sierra de Guadarrama, Boca del Asno, 7 km past La Granja de San Ildefonso towards Puerto de Navacerrada (province of Segovia), 19.08. 1990 (PGe. 1028). Photo P. Geniez. C) Spain, Hiendelaencina, in the village, Plaza Major, 1,080 m (province of Guadalajara), 17.07. 2010 (BEV. 10983). Photo P. Geniez & P. - A. Crochet. D) Spain, San Martín de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2021). Photo P. Geniez. E) Spain, El Tiemblo (province of Ávila), 27.06. 2001 (BEV. 2009). Photo P. Geniez. F) Specimen with greenish tint on the back. Spain, Sierra de Gredos, 4 km NE. Laguna Grande, 2000 m (province of Ávila), 16.07. 1992 (PGe. 1035). Photo P. Geniez." httpUri="https://zenodo.org/record/285985/files/figure.png" pageId="9" pageNumber="10">Figs. 4</figureCitation>
|
||
&
|
||
<figureCitation id="13742A17FFA1C20886C2F84DFC2BF80C" box="[919,935,1925,1950]" captionStart="FIGURE 5" captionStartId="11.[151,250,1782,1804]" captionTargetBox="[151,1436,517,1760]" captionTargetId="figure@11.[151,1436,517,1761]" captionTargetPageId="11" captionText="FIGURE 5. Variation in females, undersides and juveniles of Podarcis guadarramae guadarramae. A) Female. Spain, El Tiemblo (province of Ávila). 27.06. 2001 (BEV. 2007). Photo P. Geniez. B) Female with the “ guadarramae striped pattern ”. Spain, Navalmoral (province of Ávila), 27.6. 2001 (BEV. 1999). Photo P. Geniez. C) Typical underside of a male, Spain, Navalilla, in the village (province of Segovia), 13.05. 1996 (BEV. 4487). Photo P. Geniez. D) Male with a white underside. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2018). Photo P. Geniez. E) Juvenile male. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2025). Photo P. Geniez. F) Juvenile female. Spain, La Alberca, Iglesia de Nuestra Señora de la Asunción (province of Salamanca), 18.09. 2007 (PGe. 790). Photo P. - A. Crochet." httpUri="https://zenodo.org/record/285986/files/figure.png" pageId="9" pageNumber="10">5</figureCitation>
|
||
): large proportion of black elements in body coloration; medium body size (average of adult male SVL:
|
||
<quantity id="4CB79B77FFA1C2088619F860FC39F852" box="[844,949,1960,1985]" metricMagnitude="-2" metricUnit="m" metricValue="5.38" pageId="9" pageNumber="10" unit="mm" value="53.8">53.8 mm</quantity>
|
||
, range 41 to 64, with 83% of adult males above
|
||
<quantity id="4CB79B77FFA1C20885B7F805FEBFF877" box="[226,307,1997,2022]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="10" unit="mm" value="50.0">50 mm</quantity>
|
||
SVL, average of adult female SVL:
|
||
<quantity id="4CB79B77FFA1C208878EF805FCCEF877" box="[731,834,1997,2022]" metricMagnitude="-2" metricUnit="m" metricValue="5.16" pageId="9" pageNumber="10" unit="mm" value="51.6">51.6 mm</quantity>
|
||
, range
|
||
<quantity id="4CB79B77FFA1C20886C2F805FC6BF877" box="[919,999,1997,2021]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="9" pageNumber="10" unit="mm" value="43.0">43 mm</quantity>
|
||
to
|
||
<quantity id="4CB79B77FFA1C208815BF805FBEDF877" box="[1038,1121,1997,2022]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="9" pageNumber="10" unit="mm" value="59.0">59 mm</quantity>
|
||
, with 67% of adult females above
|
||
<quantity id="4CB79B77FFA2C20B85B0FF5FFEB5FF3D" box="[229,313,151,176]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="11" unit="mm" value="50.0">50 mm</quantity>
|
||
SVL); head distinctly flattened, body slender and flattened; vertebral stripe generally absent or vestigial, and then usually limited to the anterior part of the dorsum; pale dorsolateral stripes in males bright and contrasting, narrower than the dark supradorsolateral stripes, and reaching the anterior part of the tail, usually made of series of elongated whitish spots on a very pale background; many females with pale dorsolateral stripes straight and uninterrupted, without irregular pattern (called here “
|
||
<taxonomicName id="4C4F4D11FFA2C20B8649FEEFFC23FED2" box="[796,943,295,320]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="guadarramae">guadarramae</taxonomicName>
|
||
striped pattern”, cf.
|
||
<figureCitation id="13742A17FFA2C20B81CDFEEFFB55FED2" box="[1176,1241,295,320]" captionStart="FIGURE 5" captionStartId="11.[151,250,1782,1804]" captionTargetBox="[151,1436,517,1760]" captionTargetId="figure@11.[151,1436,517,1761]" captionTargetPageId="11" captionText="FIGURE 5. Variation in females, undersides and juveniles of Podarcis guadarramae guadarramae. A) Female. Spain, El Tiemblo (province of Ávila). 27.06. 2001 (BEV. 2007). Photo P. Geniez. B) Female with the “ guadarramae striped pattern ”. Spain, Navalmoral (province of Ávila), 27.6. 2001 (BEV. 1999). Photo P. Geniez. C) Typical underside of a male, Spain, Navalilla, in the village (province of Segovia), 13.05. 1996 (BEV. 4487). Photo P. Geniez. D) Male with a white underside. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2018). Photo P. Geniez. E) Juvenile male. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2025). Photo P. Geniez. F) Juvenile female. Spain, La Alberca, Iglesia de Nuestra Señora de la Asunción (province of Salamanca), 18.09. 2007 (PGe. 790). Photo P. - A. Crochet." httpUri="https://zenodo.org/record/285986/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
|
||
B), other females with pale dorsolateral stripes fragmented in series of elongated spots like males (
|
||
<figureCitation id="13742A17FFA2C20B8161FE84FBF4FEF7" box="[1076,1144,332,357]" captionStart="FIGURE 5" captionStartId="11.[151,250,1782,1804]" captionTargetBox="[151,1436,517,1760]" captionTargetId="figure@11.[151,1436,517,1761]" captionTargetPageId="11" captionText="FIGURE 5. Variation in females, undersides and juveniles of Podarcis guadarramae guadarramae. A) Female. Spain, El Tiemblo (province of Ávila). 27.06. 2001 (BEV. 2007). Photo P. Geniez. B) Female with the “ guadarramae striped pattern ”. Spain, Navalmoral (province of Ávila), 27.6. 2001 (BEV. 1999). Photo P. Geniez. C) Typical underside of a male, Spain, Navalilla, in the village (province of Segovia), 13.05. 1996 (BEV. 4487). Photo P. Geniez. D) Male with a white underside. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2018). Photo P. Geniez. E) Juvenile male. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2025). Photo P. Geniez. F) Juvenile female. Spain, La Alberca, Iglesia de Nuestra Señora de la Asunción (province of Salamanca), 18.09. 2007 (PGe. 790). Photo P. - A. Crochet." httpUri="https://zenodo.org/record/285986/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
|
||
A); juveniles with similar pattern (
|
||
<figureCitation id="13742A17FFA2C20B85A3FEA7FEC9FE1A" box="[246,325,367,392]" captionStart="FIGURE 5" captionStartId="11.[151,250,1782,1804]" captionTargetBox="[151,1436,517,1760]" captionTargetId="figure@11.[151,1436,517,1761]" captionTargetPageId="11" captionText="FIGURE 5. Variation in females, undersides and juveniles of Podarcis guadarramae guadarramae. A) Female. Spain, El Tiemblo (province of Ávila). 27.06. 2001 (BEV. 2007). Photo P. Geniez. B) Female with the “ guadarramae striped pattern ”. Spain, Navalmoral (province of Ávila), 27.6. 2001 (BEV. 1999). Photo P. Geniez. C) Typical underside of a male, Spain, Navalilla, in the village (province of Segovia), 13.05. 1996 (BEV. 4487). Photo P. Geniez. D) Male with a white underside. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2018). Photo P. Geniez. E) Juvenile male. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2025). Photo P. Geniez. F) Juvenile female. Spain, La Alberca, Iglesia de Nuestra Señora de la Asunción (province of Salamanca), 18.09. 2007 (PGe. 790). Photo P. - A. Crochet." httpUri="https://zenodo.org/record/285986/files/figure.png" pageId="10" pageNumber="11">Figs. 5</figureCitation>
|
||
E & 5F); pileus strongly spotted with black; ground color of the dorsum generally brown, rarely with green hue except in mountains (
|
||
<figureCitation id="13742A17FFA2C20B871FFE5CFD1DFE3E" box="[586,657,404,429]" captionStart="FIGURE 4" captionStartId="9.[151,250,1391,1413]" captionTargetBox="[151,1436,193,1369]" captionTargetId="figure@9.[151,1436,193,1370]" captionTargetPageId="9" captionText="FIGURE 4. Variation in males of Podarcis guadarramae guadarramae. For GPS coordinates (WGS 84 in decimal degrees), see Appendix 1. A) Spain, Sierra de Gredos, 1.5 km N. Laguna Grande, 1,897 m (province of Ávila), 17.08. 2007 (PGe. 1107). Photo O. Buisson. B) Spain, Sierra de Guadarrama, Boca del Asno, 7 km past La Granja de San Ildefonso towards Puerto de Navacerrada (province of Segovia), 19.08. 1990 (PGe. 1028). Photo P. Geniez. C) Spain, Hiendelaencina, in the village, Plaza Major, 1,080 m (province of Guadalajara), 17.07. 2010 (BEV. 10983). Photo P. Geniez & P. - A. Crochet. D) Spain, San Martín de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2021). Photo P. Geniez. E) Spain, El Tiemblo (province of Ávila), 27.06. 2001 (BEV. 2009). Photo P. Geniez. F) Specimen with greenish tint on the back. Spain, Sierra de Gredos, 4 km NE. Laguna Grande, 2000 m (province of Ávila), 16.07. 1992 (PGe. 1035). Photo P. Geniez." httpUri="https://zenodo.org/record/285985/files/figure.png" pageId="10" pageNumber="11">Fig. 4</figureCitation>
|
||
F), with a pale area between the two dark supra-dorsolateral bands usually moderately wide (more than 5 scales in 77% of males and 84% of females); throat normally whitish, exceptionally pink, with numerous black points especially in males; ventral face whitish, pink or salmon, sometimes brick red (never yellow or yellowish), frequently with the anterior part of the underside whitish becoming progressively reddish toward the belly (
|
||
<figureCitation id="13742A17FFA2C20B878CFDECFC97FDAF" box="[729,795,548,573]" captionStart="FIGURE 5" captionStartId="11.[151,250,1782,1804]" captionTargetBox="[151,1436,517,1760]" captionTargetId="figure@11.[151,1436,517,1761]" captionTargetPageId="11" captionText="FIGURE 5. Variation in females, undersides and juveniles of Podarcis guadarramae guadarramae. A) Female. Spain, El Tiemblo (province of Ávila). 27.06. 2001 (BEV. 2007). Photo P. Geniez. B) Female with the “ guadarramae striped pattern ”. Spain, Navalmoral (province of Ávila), 27.6. 2001 (BEV. 1999). Photo P. Geniez. C) Typical underside of a male, Spain, Navalilla, in the village (province of Segovia), 13.05. 1996 (BEV. 4487). Photo P. Geniez. D) Male with a white underside. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2018). Photo P. Geniez. E) Juvenile male. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2025). Photo P. Geniez. F) Juvenile female. Spain, La Alberca, Iglesia de Nuestra Señora de la Asunción (province of Salamanca), 18.09. 2007 (PGe. 790). Photo P. - A. Crochet." httpUri="https://zenodo.org/record/285986/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
|
||
C), underside often white in females, infrequently so in males (
|
||
<figureCitation id="13742A17FFA2C20B85B9FD8FFEBFFDF2" box="[236,307,583,608]" captionStart="FIGURE 5" captionStartId="11.[151,250,1782,1804]" captionTargetBox="[151,1436,517,1760]" captionTargetId="figure@11.[151,1436,517,1761]" captionTargetPageId="11" captionText="FIGURE 5. Variation in females, undersides and juveniles of Podarcis guadarramae guadarramae. A) Female. Spain, El Tiemblo (province of Ávila). 27.06. 2001 (BEV. 2007). Photo P. Geniez. B) Female with the “ guadarramae striped pattern ”. Spain, Navalmoral (province of Ávila), 27.6. 2001 (BEV. 1999). Photo P. Geniez. C) Typical underside of a male, Spain, Navalilla, in the village (province of Segovia), 13.05. 1996 (BEV. 4487). Photo P. Geniez. D) Male with a white underside. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2018). Photo P. Geniez. E) Juvenile male. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06. 2001 (BEV. 2025). Photo P. Geniez. F) Juvenile female. Spain, La Alberca, Iglesia de Nuestra Señora de la Asunción (province of Salamanca), 18.09. 2007 (PGe. 790). Photo P. - A. Crochet." httpUri="https://zenodo.org/record/285986/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
|
||
D); marginal ventral plates, and more rarely medium and central plates, with a quadrangular or roundish, more rarely triangular, black mark; iris whitish to pale orange; masseteric shield generally small, absent in 34% of males and 43% of females; numerous longitudinal rows of dorsal scales at midbody (52 to 72 for males, average: 59.9 and 48 to 68 for females, average 57.9); large number of femoral pores (15 to 23 for males, average 18.5, 15 to 20 for females, average 17.7). Diagnostic positions in the DNA sequence of the mitochondrial NADH dehydrogenase subunit 4 (ND4) gene relative to other lineages of the
|
||
<taxonomicName id="4C4F4D11FFA2C20B86F5FD35FBBFFC87" box="[928,1075,764,789]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="hispanicus">
|
||
<emphasis id="B93BEA80FFA2C20B86F5FD35FBBFFC87" box="[928,1075,764,789]" italics="true" pageId="10" pageNumber="11">P. hispanicus</emphasis>
|
||
</taxonomicName>
|
||
complex include a C at position 10888, T at position 10929, A at position 11097, A at position 11365 and A at position 11418 (positions numbered according to the
|
||
<taxonomicName id="4C4F4D11FFA2C20B8407FC8DFE4FFCCF" box="[338,451,836,861]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="muralis">
|
||
<emphasis id="B93BEA80FFA2C20B8407FC8DFE4FFCCF" box="[338,451,836,861]" italics="true" pageId="10" pageNumber="11">P. muralis</emphasis>
|
||
</taxonomicName>
|
||
mitochondrion complete genome GenBank accession number NC_
|
||
<date id="FFF11052FFA2C20B81EFFC8CFA9DFCCF" box="[1210,1297,836,861]" pageId="10" pageNumber="11" value="07-01-16">011607</date>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF03692FFA2C20B8592FCAFFE57F972" blockId="10.[151,1437,151,2013]" pageId="10" pageNumber="11">
|
||
<emphasis id="B93BEA80FFA2C20B8592FCAFFE27FC12" bold="true" box="[199,427,871,896]" pageId="10" pageNumber="11">Range and ecology.</emphasis>
|
||
<taxonomicName id="4C4F4D11FFA2C20B84E7FCAFFCD8FC12" box="[434,852,871,896]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
|
||
<emphasis id="B93BEA80FFA2C20B84E7FCAFFCD8FC12" box="[434,852,871,896]" italics="true" pageId="10" pageNumber="11">Podarcis guadarramae guadarramae</emphasis>
|
||
</taxonomicName>
|
||
occurs only in
|
||
<collectingCountry id="F3587602FFA2C20B86AAFCAFFBCEFC12" box="[1023,1090,871,896]" name="Spain" pageId="10" pageNumber="11">Spain</collectingCountry>
|
||
, in the Spanish Central System mountains, including Sierras de Guadarrama, Béjar, Gredos, Peña de
|
||
<collectingCountry id="F3587602FFA2C20B8696FC44FB9BFC37" box="[963,1047,908,933]" name="France" pageId="10" pageNumber="11">Francia</collectingCountry>
|
||
and Gata and the smaller sierras around these main massifs, and in siliceous plains with
|
||
<taxonomicName id="4C4F4D11FFA2C20B866BFC79FC64FC5A" box="[830,1000,945,968]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="pinaster">
|
||
<emphasis id="B93BEA80FFA2C20B866BFC79FC64FC5A" box="[830,1000,945,968]" italics="true" pageId="10" pageNumber="11">Pinus pinaster</emphasis>
|
||
</taxonomicName>
|
||
forests north of the Central System mountains. Range limits to the north-west are still poorly known due to difficulty of morphological separation from
|
||
<emphasis id="B93BEA80FFA2C20B85C2FC31FEC4FB82" box="[151,328,1015,1040]" italics="true" pageId="10" pageNumber="11">P. g. l u s i t an i c u s</emphasis>
|
||
, north-westernmost localities confirmed with genetic analysis (mtDNA sequencing) are
|
||
<collectingCountry id="F3587602FFA2C20B8017FC3FFAF4FB82" box="[1346,1400,1015,1040]" name="United Kingdom" pageId="10" pageNumber="11">Alba</collectingCountry>
|
||
de Tormes and Ciudad Rodrigo in province of Salamanca (
|
||
<bibRefCitation id="EFDE4B63FFA2C20B8663FBD4FB08FBA6" author="Kaliontzopoulou" box="[822,1156,1052,1077]" pageId="10" pageNumber="11" refString="Kaliontzopoulou, A., Pinho, C., Harris, D. J. & Carretero, M. A. (2011) When cryptic diversity blurs the picture: a cautionary tale from Iberian and North African Podarcis wall lizards. Biological Journal of the Linnean Society, 103, 779 - 800. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2011.01703. x" type="journal article" year="2011">
|
||
Kaliontzopoulou
|
||
<emphasis id="B93BEA80FFA2C20B8154FBD5FBB3FBA7" box="[1025,1087,1052,1077]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
) while western limit is currently La Alberca, Salamanca (Pinho
|
||
<emphasis id="B93BEA80FFA2C20B873DFB89FD2EFBCA" box="[616,674,1087,1112]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||
2007). Further east, populations assigned to
|
||
<taxonomicName id="4C4F4D11FFA2C20B81F1FB89FAF4FBCA" box="[1188,1400,1087,1112]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
|
||
<emphasis id="B93BEA80FFA2C20B81F1FB89FAF4FBCA" box="[1188,1400,1087,1112]" italics="true" pageId="10" pageNumber="11">P. g. guadarramae</emphasis>
|
||
</taxonomicName>
|
||
by their typical morphology and their location reach the north of the province of Segovia where they abruptly meet
|
||
<taxonomicName id="4C4F4D11FFA2C20B80D2FBADFF66FB32" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="liolepis">
|
||
<emphasis id="B93BEA80FFA2C20B80D2FBADFF66FB32" italics="true" pageId="10" pageNumber="11">P. liolepis</emphasis>
|
||
</taxonomicName>
|
||
(see contact zones below). Eastern and southern limits better known as the adjacent
|
||
<taxonomicName id="4C4F4D11FFA2C20B8190FB41FAC7FB32" box="[1221,1355,1161,1184]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="virescens">
|
||
<emphasis id="B93BEA80FFA2C20B8190FB41FAC7FB32" box="[1221,1355,1161,1184]" italics="true" pageId="10" pageNumber="11">P. virescens</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4F4D11FFA2C20B80D2FB41FF66FB57" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="liolepis">
|
||
<emphasis id="B93BEA80FFA2C20B80D2FB41FF66FB57" italics="true" pageId="10" pageNumber="11">P. liolepis</emphasis>
|
||
</taxonomicName>
|
||
are reasonably easy to separate morphologically (see below). The limit of the distribution south of the Spanish Central System seems to follow closely the transition between the dry, sparsely vegetated plateau of Central
|
||
<collectingCountry id="F3587602FFA2C20B85A4FB3CFEBDFA9F" box="[241,305,1268,1293]" name="Spain" pageId="10" pageNumber="11">Spain</collectingCountry>
|
||
(inhabited by
|
||
<taxonomicName id="4C4F4D11FFA2C20B8481FB3DFDB7FA9E" box="[468,571,1269,1292]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="virescens">
|
||
<emphasis id="B93BEA80FFA2C20B8481FB3DFDB7FA9E" box="[468,571,1269,1292]" italics="true" pageId="10" pageNumber="11">virescens</emphasis>
|
||
</taxonomicName>
|
||
) and the more vegetated foothills of the Central System massifs (pers. obs.).
|
||
<taxonomicName id="4C4F4D11FFA2C20B85C2FADFFDB6FAA2" box="[151,570,1303,1328]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
|
||
<emphasis id="B93BEA80FFA2C20B85C2FADFFDB6FAA2" box="[151,570,1303,1328]" italics="true" pageId="10" pageNumber="11">Podarcis guadarramae guadarramae</emphasis>
|
||
</taxonomicName>
|
||
is mostly a rock lizard adapted to rupicolous habitats including granite rocks, artificial stone walls, rubbles and detritus etc., mostly on siliceous substratum. It reaches at least
|
||
<quantity id="4CB79B77FFA2C20B81B0FAF4FACEFAC6" box="[1253,1346,1340,1364]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="11" unit="m" value="2000.0">2,000 m</quantity>
|
||
a.s.l. in the Sierra de Gredos (pers. obs.). In parts of its range where there is no competitors (e.g.
|
||
<taxonomicName id="4C4F4D11FFA2C20B818AFAA9FADAFAEA" box="[1247,1366,1375,1400]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="muralis">
|
||
<emphasis id="B93BEA80FFA2C20B818AFAA9FADAFAEA" box="[1247,1366,1375,1400]" italics="true" pageId="10" pageNumber="11">P. muralis</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="4C4F4D11FFA2C20B80D2FAA9FE86FA0F" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="carbonelli">
|
||
<emphasis id="B93BEA80FFA2C20B80D2FAA9FE86FA0F" italics="true" pageId="10" pageNumber="11">P. carbonelli</emphasis>
|
||
</taxonomicName>
|
||
), it can also live on the ground, especially in pine
|
||
<taxonomicName id="4C4F4D11FFA2C20B860CFA4DFC72FA0E" box="[857,1022,1413,1436]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="pinaster">
|
||
<emphasis id="B93BEA80FFA2C20B860CFA4DFC72FA0E" box="[857,1022,1413,1436]" italics="true" pageId="10" pageNumber="11">Pinus pinaster</emphasis>
|
||
</taxonomicName>
|
||
forests on siliceous sandy soils. An (apparently) isolated population inhabits the Castillo de Trujillo (province of Cáceres,
|
||
<collectingCountry id="F3587602FFA2C20B81CFFA6FFB52FA52" box="[1178,1246,1447,1472]" name="Spain" pageId="10" pageNumber="11">Spain</collectingCountry>
|
||
), surrounded by localities where only
|
||
<typeStatus id="54F48830FFA2C20B84D9FA05FE30FA77" box="[396,444,1485,1509]" pageId="10" pageNumber="11">type</typeStatus>
|
||
2 has been documented; first photographed by V. Joubert in 1995 (photos in PG photo collection, see Appendix 1), this occurrence was later confirmed by genetic data (
|
||
<bibRefCitation id="EFDE4B63FFA2C20B8173FA27FB66F99A" author="Pinho" box="[1062,1258,1519,1544]" pageId="10" pageNumber="11" refString="Pinho, C., Ferrand, N. & Harris, D. J. (2006) Reexamination of the Iberian and North African Podarcis (Squamata: Lacertidae) phylogeny based on increased mitochondrial DNA sequencing. Molecular Phylogenetics and Evolution, 38, 266 - 273. http: // dx. doi. org / 10.1016 / j. ympev. 2005.06.012" type="journal article" year="2006">
|
||
Pinho
|
||
<emphasis id="B93BEA80FFA2C20B8125FA39FB25F99A" box="[1136,1193,1519,1544]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||
2006
|
||
</bibRefCitation>
|
||
, specimen Trj1, collected exactly at the same location as confirmed by Pinho pers. com.). This occurrence seems to be linked with the ecological situation of Trujillo which is built on a rocky outcrop emerging from the surrounding plains. In addition,
|
||
<bibRefCitation id="EFDE4B63FFA2C20B8450F994FDECF9E6" box="[261,608,1628,1653]" pageId="10" pageNumber="11" refString="Kaliontzopoulou, A., Pinho, C., Harris, D. J. & Carretero, M. A. (2011) When cryptic diversity blurs the picture: a cautionary tale from Iberian and North African Podarcis wall lizards. Biological Journal of the Linnean Society, 103, 779 - 800. http: // dx. doi. org / 10.1111 / j. 1095 - 8312.2011.01703. x" type="journal article">
|
||
Kaliontzopoulou
|
||
<emphasis id="B93BEA80FFA2C20B8498F995FD87F9E7" box="[461,523,1628,1653]" italics="true" pageId="10" pageNumber="11">et al.</emphasis>
|
||
(2011)
|
||
</bibRefCitation>
|
||
report an isolated occurrence of
|
||
<taxonomicName id="4C4F4D11FFA2C20B86BBF995FB4BF9E7" box="[1006,1223,1628,1653]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
|
||
<emphasis id="B93BEA80FFA2C20B86BBF995FB4BF9E7" box="[1006,1223,1628,1653]" italics="true" pageId="10" pageNumber="11">P. g. guadarramae</emphasis>
|
||
</taxonomicName>
|
||
in Torrejón de la Calzada (province of Madrid), a locality surrounded by
|
||
<taxonomicName id="4C4F4D11FFA2C20B866CF949FC4DF90A" box="[825,961,1665,1688]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="virescens">
|
||
<emphasis id="B93BEA80FFA2C20B866CF949FC4DF90A" box="[825,961,1665,1688]" italics="true" pageId="10" pageNumber="11">P. virescens</emphasis>
|
||
</taxonomicName>
|
||
populations and in a typical ecological situation for that species. Further sampling would be needed to fully exclude a mistake in sample labeling or sample processing in the lab.
|
||
</paragraph>
|
||
<paragraph id="8BF03692FFA2C20A8592F924FEA1FE42" blockId="10.[151,1437,151,2013]" lastBlockId="11.[151,1436,151,464]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
<emphasis id="B93BEA80FFA2C20B8592F924FD4FF897" bold="true" box="[199,707,1772,1797]" pageId="10" pageNumber="11">Situation in contact zones with other taxa.</emphasis>
|
||
<taxonomicName id="4C4F4D11FFA2C20B879FF924FBE3F897" box="[714,1135,1772,1797]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
|
||
<emphasis id="B93BEA80FFA2C20B879FF924FBE3F897" box="[714,1135,1772,1797]" italics="true" pageId="10" pageNumber="11">Podarcis guadarramae guadarramae</emphasis>
|
||
</taxonomicName>
|
||
can coexist (sometimes in syntopy) with two other
|
||
<taxonomicName id="4C4F4D11FFA2C20B84F8F8C7FD9DF8BA" box="[429,529,1807,1832]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
|
||
<emphasis id="B93BEA80FFA2C20B84F8F8C7FD9DF8BA" box="[429,529,1807,1832]" italics="true" pageId="10" pageNumber="11">Podarcis</emphasis>
|
||
</taxonomicName>
|
||
species:
|
||
<taxonomicName id="4C4F4D11FFA2C20B872FF8D9FD60F8BA" box="[634,748,1807,1832]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="muralis">
|
||
<emphasis id="B93BEA80FFA2C20B872FF8D9FD60F8BA" box="[634,748,1807,1832]" italics="true" pageId="10" pageNumber="11">P. muralis</emphasis>
|
||
</taxonomicName>
|
||
(e.g. Sierra de Guadarrama) and
|
||
<taxonomicName id="4C4F4D11FFA2C20B8133F8D9FB79F8BA" box="[1126,1269,1807,1832]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="carbonelli">
|
||
<emphasis id="B93BEA80FFA2C20B8133F8D9FB79F8BA" box="[1126,1269,1807,1832]" italics="true" pageId="10" pageNumber="11">P. carbonelli</emphasis>
|
||
</taxonomicName>
|
||
(e.g. Sierra de la Peña de
|
||
<collectingCountry id="F3587602FFA2C20B8440F8FCFEE2F8DF" box="[277,366,1844,1869]" name="France" pageId="10" pageNumber="11">Francia</collectingCountry>
|
||
). It is parapatric with
|
||
<taxonomicName id="4C4F4D11FFA2C20B8732F8FDFD67F8DE" box="[615,747,1845,1868]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="virescens">
|
||
<emphasis id="B93BEA80FFA2C20B8732F8FDFD67F8DE" box="[615,747,1845,1868]" italics="true" pageId="10" pageNumber="11">P. virescens</emphasis>
|
||
</taxonomicName>
|
||
south of the Central System (cf. below), and with
|
||
<emphasis id="B93BEA80FFA2C20B8078F8FDFA14F8DE" box="[1325,1432,1844,1869]" italics="true" pageId="10" pageNumber="11">P. l i o l e p i s</emphasis>
|
||
along a contact zone that runs from the north of the province of Segovia (Villaverde de Iscar, Remondo, Chañe, Arroyo de Cuéllar, Campo de Cuéllar, Valleruela de Sepúlveda et Duruelo) to the easternmost foothills of the Central Spanish System south of Atienza (province of Guadalajara). The contact zone in the province of Segovia runs precisely along the border between cultivated calcareous plateau in the north (home of
|
||
<emphasis id="B93BEA80FFA2C20B81FFF80DFA99F84E" box="[1194,1301,1988,2013]" italics="true" pageId="10" pageNumber="11">P. l i ol e pi s</emphasis>
|
||
) and sandy siliceous plains with
|
||
<taxonomicName id="4C4F4D11FFA3C20A84DCFF51FDA2FF22" box="[393,558,153,176]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="pinaster">
|
||
<emphasis id="B93BEA80FFA3C20A84DCFF51FDA2FF22" box="[393,558,153,176]" italics="true" pageId="11" pageNumber="12">Pinus pinaster</emphasis>
|
||
</taxonomicName>
|
||
woods (home of
|
||
<taxonomicName id="4C4F4D11FFA3C20A87AEFF51FC42FF22" box="[763,974,151,176]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
|
||
<emphasis id="B93BEA80FFA3C20A87AEFF51FC42FF22" box="[763,974,151,176]" italics="true" pageId="11" pageNumber="12">P. g. guadarramae</emphasis>
|
||
</taxonomicName>
|
||
). We have identified very few localities where
|
||
<taxonomicName id="4C4F4D11FFA3C20A85B7FF75FE3EFF47" box="[226,434,188,213]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
|
||
<emphasis id="B93BEA80FFA3C20A85B7FF75FE3EFF47" box="[226,434,188,213]" italics="true" pageId="11" pageNumber="12">P. g. guadarramae</emphasis>
|
||
</taxonomicName>
|
||
coexists with these species, as populations are usually a few kilometres apart. Within the abrupt contact zone with
|
||
<taxonomicName id="4C4F4D11FFA3C20A84E6FF29FDADFF6A" box="[435,545,223,248]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="liolepis">
|
||
<emphasis id="B93BEA80FFA3C20A84E6FF29FDADFF6A" box="[435,545,223,248]" italics="true" pageId="11" pageNumber="12">P. liolepis</emphasis>
|
||
</taxonomicName>
|
||
in the north of the province of Segovia pure populations of either species were found only tens to hundreds of meters apart and local syntopy has even been observed in one spot between cultivated lands on calcareous substratum and pinewoods on sand (
|
||
<bibRefCitation id="EFDE4B63FFA3C20A8699FEEFFBE9FED2" author="Geniez" box="[972,1125,295,320]" pageId="11" pageNumber="12" refString="Geniez, P. (2001) Variation geographique des lezards du genre Podarcis (Reptilia, Sauria, Lacertidae) dans la peninsule Iberique, l'Afrique du Nord et le sud de la France. Unpublished E. P. H. E. diploma thesis. Universite Montpellier II and Ecole Pratique des Hautes Etudes, Montpellier, 342 pp." type="book" year="2001">Geniez 2001</bibRefCitation>
|
||
) (see
|
||
<figureCitation id="13742A17FFA3C20A81EDFEEFFA8FFED2" box="[1208,1283,295,320]" captionStart="FIGURE 6" captionStartId="12.[151,250,1817,1839]" captionTargetBox="[151,1435,193,1795]" captionTargetId="figure@12.[151,1436,193,1796]" captionTargetPageId="12" captionText="FIGURE 6. Podarcis liolepis BEV. 4122 (above) and Podarcis guadarramae guadarramae PGe. 1006, two males found in strict syntopy. Spain, Road C. 603, 1.4 km past San Miguel de Bernuy towards Cantalejo (province of Segovia), 7.05. 1986. Photos P. Geniez. Note in P. liolepis the complete vertebral stripe, the unbroken pale dorsolateral stripes, the less flattened head, the big masseteric shield and the less minute temporal and dorsal scales." httpUri="https://zenodo.org/record/285987/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
|
||
). No visible morphological introgression has been observed there or in any other population close to the contact zones with
|
||
<taxonomicName id="4C4F4D11FFA3C20A80D2FE85FF72FE1A" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="virescens">
|
||
<emphasis id="B93BEA80FFA3C20A80D2FE85FF72FE1A" italics="true" pageId="11" pageNumber="12">P. virescens</emphasis>
|
||
</taxonomicName>
|
||
or with
|
||
<taxonomicName id="4C4F4D11FFA3C20A8435FEB9FE42FE1A" box="[352,462,367,392]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="liolepis">
|
||
<emphasis id="B93BEA80FFA3C20A8435FEB9FE42FE1A" box="[352,462,367,392]" italics="true" pageId="11" pageNumber="12">P. liolepis</emphasis>
|
||
</taxonomicName>
|
||
, althoug detailed analysis of morphological variation accross contact zones remain to be done. We have no personal information on contact zones with
|
||
<emphasis id="B93BEA80FFA3C20A8622FE5CFC7CFE3F" box="[887,1008,404,429]" italics="true" pageId="11" pageNumber="12">lusitanicus</emphasis>
|
||
(but see “systematic ranks of the new taxa” above).
|
||
</paragraph>
|
||
<caption id="DF30661AFFA3C20A85C2F93EFAECF856" httpUri="https://zenodo.org/record/285986/files/figure.png" pageId="11" pageNumber="12" targetBox="[151,1436,517,1760]" targetPageId="11">
|
||
<paragraph id="8BF03692FFA3C20A85C2F93EFAECF856" blockId="11.[151,1437,1782,1988]" pageId="11" pageNumber="12">
|
||
<emphasis id="B93BEA80FFA3C20A85C2F93EFE9AF89E" bold="true" box="[151,278,1782,1804]" pageId="11" pageNumber="12">FIGURE 5.</emphasis>
|
||
Variation in females, undersides and juveniles of
|
||
<taxonomicName id="4C4F4D11FFA3C20A867EF93EFB21F89E" box="[811,1197,1782,1804]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
|
||
<emphasis id="B93BEA80FFA3C20A867EF93EFB21F89E" box="[811,1197,1782,1804]" italics="true" pageId="11" pageNumber="12">Podarcis guadarramae guadarramae</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="B93BEA80FFA3C20A81E9F93EFB5BF89E" bold="true" box="[1212,1239,1782,1804]" pageId="11" pageNumber="12">A)</emphasis>
|
||
Female. Spain, El Tiemblo (province of Ávila). 27.06.2001 (BEV.2007). Photo P. Geniez.
|
||
<emphasis id="B93BEA80FFA3C20A86DDF8DCFC2CF8B8" bold="true" box="[904,928,1812,1834]" pageId="11" pageNumber="12">B)</emphasis>
|
||
Female with the “
|
||
<taxonomicName id="4C4F4D11FFA3C20A8132F8DCFB67F8B8" box="[1127,1259,1812,1834]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="guadarramae">guadarramae</taxonomicName>
|
||
striped pattern”. Spain, Navalmoral (province of Ávila), 27.6.2001 (BEV.1999). Photo P. Geniez.
|
||
<emphasis id="B93BEA80FFA3C20A86A2F8FBFB9EF8DB" bold="true" box="[1015,1042,1843,1865]" pageId="11" pageNumber="12">C)</emphasis>
|
||
Typical underside of a male, Spain, Navalilla, in the village (province of Segovia), 13.05.1996 (BEV.4487). Photo P. Geniez.
|
||
<emphasis id="B93BEA80FFA3C20A8111F89AFBD2F8FA" bold="true" box="[1092,1118,1874,1896]" pageId="11" pageNumber="12">D)</emphasis>
|
||
Male with a white underside. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06.2001 (BEV.2018). Photo P. Geniez.
|
||
<emphasis id="B93BEA80FFA3C20A81CAF8B9FB34F815" bold="true" box="[1183,1208,1905,1927]" pageId="11" pageNumber="12">E)</emphasis>
|
||
Juvenile male. Spain, San Martin de Valdeiglesias (province of Madrid), 27.06.2001 (BEV.2025). Photo P. Geniez.
|
||
<emphasis id="B93BEA80FFA3C20A8130F847FBF2F837" bold="true" box="[1125,1150,1935,1957]" pageId="11" pageNumber="12">F)</emphasis>
|
||
Juvenile female. Spain, La Alberca, Iglesia de Nuestra Señora de la Asunción (province of Salamanca), 18.09.2007 (PGe.790). Photo P.-A. Crochet.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF30661AFFA4C20D85C2F8D1FCFFF819" httpUri="https://zenodo.org/record/285987/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1435,193,1795]" targetPageId="12">
|
||
<paragraph id="8BF03692FFA4C20D85C2F8D1FCFFF819" blockId="12.[151,1436,1817,1931]" pageId="12" pageNumber="13">
|
||
<emphasis id="B93BEA80FFA4C20D85C2F8D1FE9AF8BD" bold="true" box="[151,278,1817,1839]" pageId="12" pageNumber="13">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C4F4D11FFA4C20D8475F8D1FE43F8BD" box="[288,463,1817,1839]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="liolepis">
|
||
<emphasis id="B93BEA80FFA4C20D8475F8D1FE43F8BD" box="[288,463,1817,1839]" italics="true" pageId="12" pageNumber="13">Podarcis liolepis</emphasis>
|
||
</taxonomicName>
|
||
BEV.4122 (above) and
|
||
<taxonomicName id="4C4F4D11FFA4C20D879AF8D1FB4DF8BD" box="[719,1217,1817,1839]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
|
||
<emphasis id="B93BEA80FFA4C20D879AF8D1FBDDF8BD" box="[719,1105,1817,1839]" italics="true" pageId="12" pageNumber="13">Podarcis guadarramae guadarramae</emphasis>
|
||
PGe.1006
|
||
</taxonomicName>
|
||
, two males found in strict syntopy. Spain, Road C.603, 1.4 km past San Miguel de Bernuy towards Cantalejo (province of Segovia), 7.05.1986. Photos P. Geniez. Note in
|
||
<taxonomicName id="4C4F4D11FFA4C20D84C8F89FFD8CF8FE" box="[413,512,1878,1900]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="liolepis">
|
||
<emphasis id="B93BEA80FFA4C20D84C8F89FFD8CF8FE" box="[413,512,1878,1900]" italics="true" pageId="12" pageNumber="13">P. liolepis</emphasis>
|
||
</taxonomicName>
|
||
the complete vertebral stripe, the unbroken pale dorsolateral stripes, the less flattened head, the big masseteric shield and the less minute temporal and dorsal scales.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF03692FFA5C20C8592FF5FFEE2FF6A" blockId="13.[151,1437,151,1400]" pageId="13" pageNumber="14">
|
||
<emphasis id="B93BEA80FFA5C20C8592FF5FFE67FF22" bold="true" box="[199,491,151,176]" pageId="13" pageNumber="14">Geographical variation.</emphasis>
|
||
none evident to us but mountain populations tend to have a more contrasted coloration with larger proportion of dark elements, more contrasted pale dorsolateral lines and sometimes a green hue on the dorsum.
|
||
</paragraph>
|
||
<paragraph id="8BF03692FFA5C20C8592FECCFE40FC7F" blockId="13.[151,1437,151,1400]" pageId="13" pageNumber="14">
|
||
<emphasis id="B93BEA80FFA5C20C8592FECCFBA6FE8F" bold="true" box="[199,1066,259,285]" pageId="13" pageNumber="14">
|
||
Comparison with other species within the
|
||
<taxonomicName id="4C4F4D11FFA5C20C8799FECBFC3AFE8E" box="[716,950,259,284]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hispanicus">
|
||
<emphasis id="B93BEA80FFA5C20C8799FECBFC3AFE8E" bold="true" box="[716,950,259,284]" italics="true" pageId="13" pageNumber="14">Podarcis hispanicus</emphasis>
|
||
</taxonomicName>
|
||
complex.
|
||
</emphasis>
|
||
Separation of
|
||
<taxonomicName id="4C4F4D11FFA5C20C8183FECCFA10FE8F" box="[1238,1436,260,285]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="bocagei">
|
||
<emphasis id="B93BEA80FFA5C20C8183FECCFA10FE8F" box="[1238,1436,260,285]" italics="true" pageId="13" pageNumber="14">Podarcis bocagei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4F4D11FFA5C20C8592FEE1FEDAFED2" box="[199,342,295,320]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="carbonelli">
|
||
<emphasis id="B93BEA80FFA5C20C8592FEE1FEDAFED2" box="[199,342,295,320]" italics="true" pageId="13" pageNumber="14">P. carbonelli</emphasis>
|
||
</taxonomicName>
|
||
from the other forms of the
|
||
<taxonomicName id="4C4F4D11FFA5C20C87C2FEE1FCA7FED2" box="[663,811,295,320]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hispanicus">
|
||
<emphasis id="B93BEA80FFA5C20C87C2FEE1FCA7FED2" box="[663,811,295,320]" italics="true" pageId="13" pageNumber="14">P. hispanicus</emphasis>
|
||
</taxonomicName>
|
||
complex has been well treated in e.g.
|
||
<bibRefCitation id="EFDE4B63FFA5C20C8188FEEFFF6FFEF6" pageId="13" pageNumber="14" refString="Engelmann, W. - E., Fritzche, J., Gunther, R. & Obst, F. J. (1993) Lurche und Kriechtiere Europas. Neumann Verlag, Radebeul, 440 pp." type="book">
|
||
Engelmann
|
||
<emphasis id="B93BEA80FFA5C20C8036FEE1FA10FED2" box="[1379,1436,295,320]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
|
||
(1993)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDE4B63FFA5C20C85BAFE84FDD3FEF6" author="Salvador" box="[239,607,332,357]" pageId="13" pageNumber="14" refString="Salvador, A. & Pleguezuelos, J. M. (2002) Reptiles Espanoles. Identificacion, historia natural y distribucion. Canseco Editores, Talavera de la Reina, 493 pp." type="book" year="2002">
|
||
<collectingCountry id="F3587602FFA5C20C85BAFE84FEDFFEF7" box="[239,339,332,357]" name="El Salvador" pageId="13" pageNumber="14">Salvador</collectingCountry>
|
||
& Pleguezuelos (2002)
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDE4B63FFA5C20C873FFE85FD77FEF6" author="Kwet" box="[618,763,332,356]" pageId="13" pageNumber="14" refString="Kwet, A. (2009) Guide photographique des reptiles et amphibiens d'Europe. 130 especes et 60 sous-especes. Delachaux et Niestle, Paris, 252 pp." type="book" year="2009">Kwet (2009)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDE4B63FFA5C20C8653FE84FBBFFEF6" author="Arnold" box="[774,1075,332,357]" pageId="13" pageNumber="14" refString="Arnold, N. & Ovenden, D. (2010) Le guide herpeto. 228 amphibiens et reptiles d'Europe (troisieme edition). Delachaux et Niestle, Paris, 290 pp." type="book" year="2010">Arnold & Ovenden (2010)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFDE4B63FFA5C20C816AFE84FB6CFEF6" author="Glandt" box="[1087,1248,332,357]" pageId="13" pageNumber="14" refString="Glandt, D. (2010) Taschenlexikon der Amphibien und Reptilien Europas. Quelle & Meyer, Wiebelscheim, 633 pp." type="book" year="2010">Glandt (2010)</bibRefCitation>
|
||
. Separation of
|
||
<taxonomicName id="4C4F4D11FFA5C20C80D2FE85FE83FE1A" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="hispanicus">
|
||
<emphasis id="B93BEA80FFA5C20C80D2FE85FE83FE1A" italics="true" pageId="13" pageNumber="14">P. hispanicus</emphasis>
|
||
</taxonomicName>
|
||
has been covered in
|
||
<bibRefCitation id="EFDE4B63FFA5C20C84AFFEA7FD56FE1A" box="[506,730,367,392]" pageId="13" pageNumber="14" refString="Geniez, P., Cluchier, A., Sa-Sousa, P., Guillaume, C. P. & Crochet, P. - A. (2007) Systematics of the Podarcis hispanicus - complex (Sauria, Lacertidae) I: Redefinition, morphology and distribution of the nominotypical taxon. Herpetological Journal, 17, 69 - 80." type="journal article">
|
||
Geniez
|
||
<emphasis id="B93BEA80FFA5C20C8705FEB9FD04FE1A" box="[592,648,367,392]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
|
||
(2007)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFDE4B63FFA5C20C8645FEA7FC3CFE1A" author="Glandt" box="[784,944,367,392]" pageId="13" pageNumber="14" refString="Glandt, D. (2010) Taschenlexikon der Amphibien und Reptilien Europas. Quelle & Meyer, Wiebelscheim, 633 pp." type="book" year="2010">Glandt (2010)</bibRefCitation>
|
||
, separation from
|
||
<taxonomicName id="4C4F4D11FFA5C20C8126FEB9FB6CFE1A" box="[1139,1248,367,392]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="liolepis">
|
||
<emphasis id="B93BEA80FFA5C20C8126FEB9FB6CFE1A" box="[1139,1248,367,392]" italics="true" pageId="13" pageNumber="14">P. liolepis</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<bibRefCitation id="EFDE4B63FFA5C20C8051FEA7FF7DFE3E" pageId="13" pageNumber="14" refString="Renoult, J. P., Geniez, P., Bacquet, P., Guillaume, C. P. & Crochet, P. - A. (2010 a) Systematics of the Podarcis hispanicus - complex (Sauria, Lacertidae) II: the valid name of the north-eastern Spanish form. Zootaxa, 2500, 58 - 68." type="journal article">
|
||
Renoult
|
||
<emphasis id="B93BEA80FFA5C20C8031FEB9FA10FE1A" box="[1380,1436,367,392]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
|
||
(2010a)
|
||
</bibRefCitation>
|
||
. See
|
||
<taxonomicName id="4C4F4D11FFA5C20C847BFE5DFE3EFE3E" box="[302,434,405,428]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="virescens">
|
||
<emphasis id="B93BEA80FFA5C20C847BFE5DFE3EFE3E" box="[302,434,405,428]" italics="true" pageId="13" pageNumber="14">P. virescens</emphasis>
|
||
</taxonomicName>
|
||
account below for separation from that species.
|
||
<taxonomicName id="4C4F4D11FFA5C20C8689FE5CFA0DFE3F" box="[988,1409,404,429]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
|
||
<emphasis id="B93BEA80FFA5C20C8689FE5CFA0DFE3F" box="[988,1409,404,429]" italics="true" pageId="13" pageNumber="14">Podarcis guadarramae guadarramae</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="4C4F4D11FFA5C20C8407FE71FD86FE42" box="[338,522,439,464]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="lusitanicus">
|
||
<emphasis id="B93BEA80FFA5C20C8407FE71FD86FE42" box="[338,522,439,464]" italics="true" pageId="13" pageNumber="14">P. g. lusitanicus</emphasis>
|
||
</taxonomicName>
|
||
but many populations can be separated by coloration patterns of adult males. Average differences in pholidosis and morphometry (see
|
||
<bibRefCitation id="EFDE4B63FFA5C20C866DFE14FB0CFE66" author="Kaliontzopoulou" box="[824,1152,476,501]" pageId="13" pageNumber="14" refString="Kaliontzopoulou, A., Carretero, M. A. & Llorente, G. A. (2012) Morphology of the Podarcis wall lizards (Squamata: Lacertidae) from the Iberian Peninsula and North Africa: patterns of variation in a putative cryptic species complex. Zoological Journal of the Linnean Society, 164, 173 - 193. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.2011.00760. x" type="journal article" year="2012">
|
||
Kaliontzopoulou
|
||
<emphasis id="B93BEA80FFA5C20C8155FE15FBB1FE67" box="[1024,1085,476,501]" italics="true" pageId="13" pageNumber="14">et al.</emphasis>
|
||
2012
|
||
</bibRefCitation>
|
||
and
|
||
<emphasis id="B93BEA80FFA5C20C8195FE14FAB6FE67" box="[1216,1338,476,501]" italics="true" pageId="13" pageNumber="14">lusitanicus</emphasis>
|
||
account below) are of little use for identification, even if
|
||
<emphasis id="B93BEA80FFA5C20C87B1FE37FCD1FD8A" box="[740,861,511,536]" italics="true" pageId="13" pageNumber="14">lusitanicus</emphasis>
|
||
is often a visibly more flattened lizard. In
|
||
<taxonomicName id="4C4F4D11FFA5C20C8033FDC9FE9CFDAF" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="lusitanicus">
|
||
<emphasis id="B93BEA80FFA5C20C8033FDC9FE9CFDAF" italics="true" pageId="13" pageNumber="14">P. g. lusitanicus</emphasis>
|
||
</taxonomicName>
|
||
, pale spots in the light dorsolateral stripes are usually more obvious and better separated from each other because they stand out from a very dark background; the dark supradorsolateral stripes are broader and more fragmented, often reducing the pale background coloration to a narrow mid-dorsal band (dark supradorsolateral bands are narrower with more regular inner edges in
|
||
<taxonomicName id="4C4F4D11FFA5C20C87BFFD47FC0EFD3A" box="[746,898,655,680]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="guadarramae">
|
||
<emphasis id="B93BEA80FFA5C20C87BFFD47FC0EFD3A" box="[746,898,655,680]" italics="true" pageId="13" pageNumber="14">guadarramae</emphasis>
|
||
</taxonomicName>
|
||
), in some males the dark coloration invades the whole dorsum, pale coloration being limited to isolated round spots or ocelli, green-backed males more frequent than in
|
||
<taxonomicName id="4C4F4D11FFA5C20C85B9FD1FFE08FD62" box="[236,388,727,752]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="guadarramae">
|
||
<emphasis id="B93BEA80FFA5C20C85B9FD1FFE08FD62" box="[236,388,727,752]" italics="true" pageId="13" pageNumber="14">guadarramae</emphasis>
|
||
</taxonomicName>
|
||
; north-eastern populations of
|
||
<emphasis id="B93BEA80FFA5C20C8782FD1FFCDCFD62" box="[727,848,727,752]" italics="true" pageId="13" pageNumber="14">lusitanicus</emphasis>
|
||
(Cantabria, Asturias, north of the province of León, see “geographical variation” in
|
||
<emphasis id="B93BEA80FFA5C20C84A9FD34FDF9FC87" box="[508,629,764,789]" italics="true" pageId="13" pageNumber="14">lusitanicus</emphasis>
|
||
account) are better differentiated as they are paler with less developed, sometimes strongly reduced dark supradorsolateral stripes. Many specimens of the wholly allopatric
|
||
<emphasis id="B93BEA80FFA5C20C8049FCE9FA14FCAA" box="[1308,1432,799,824]" italics="true" pageId="13" pageNumber="14">P. v a uc h e r i</emphasis>
|
||
can be separated from both
|
||
<taxonomicName id="4C4F4D11FFA5C20C8480FC8DFD24FCCF" box="[469,680,836,861]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="guadarramae">
|
||
<emphasis id="B93BEA80FFA5C20C8480FC8DFD24FCCF" box="[469,680,836,861]" italics="true" pageId="13" pageNumber="14">P. g. guadarramae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C4F4D11FFA5C20C87B7FC8DFC19FCCF" box="[738,917,836,861]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="guadarramae" subSpecies="lusitanicus">
|
||
<emphasis id="B93BEA80FFA5C20C87B7FC8DFC19FCCF" box="[738,917,836,861]" italics="true" pageId="13" pageNumber="14">P. g. lusitanicus</emphasis>
|
||
</taxonomicName>
|
||
by their greener back (especially males from mountain populations), less flattened head and body, narrower dark supradorsolateal stripes, and rounder pale dorsolateral spots (more elongated in
|
||
<taxonomicName id="4C4F4D11FFA5C20C8705FC45FC8BFC37" box="[592,775,908,933]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="guadarramae">
|
||
<emphasis id="B93BEA80FFA5C20C8705FC45FC8BFC37" box="[592,775,908,933]" italics="true" pageId="13" pageNumber="14">P. guadarramae</emphasis>
|
||
</taxonomicName>
|
||
), although some specimens can bear an almost identical dorsal pattern;
|
||
<taxonomicName id="4C4F4D11FFA5C20C8405FC79FE5AFC5A" box="[336,470,943,968]" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="vaucheri">
|
||
<emphasis id="B93BEA80FFA5C20C8405FC79FE5AFC5A" box="[336,470,943,968]" italics="true" pageId="13" pageNumber="14">P. vaucheri</emphasis>
|
||
</taxonomicName>
|
||
generally exhibits some obvious yellow coloration on the underside which
|
||
<taxonomicName id="4C4F4D11FFA5C20C80D2FC79FEBCFC7F" class="Reptilia" family="Lacertidae" genus="Podarcis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="guadarramae">
|
||
<emphasis id="B93BEA80FFA5C20C80D2FC79FEBCFC7F" italics="true" pageId="13" pageNumber="14">P. guadarramae</emphasis>
|
||
</taxonomicName>
|
||
always lacks.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556519FFA5C20C8592FC3FFA14FAEA" pageId="13" pageNumber="14" type="description">
|
||
<paragraph id="8BF03692FFA5C20C8592FC3FFA14FAEA" blockId="13.[151,1437,151,1400]" pageId="13" pageNumber="14">
|
||
<emphasis id="B93BEA80FFA5C20C8592FC3FFDA8FB82" bold="true" box="[199,548,1015,1040]" pageId="13" pageNumber="14">
|
||
Description of the
|
||
<typeStatus id="54F48830FFA5C20C8495FC3FFD93FB82" box="[448,543,1015,1040]" pageId="13" pageNumber="14" type="neotype">neotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult male showing the following morphological features (
|
||
<figureCitation id="13742A17FFA5C20C8047FC3FFAECFB82" box="[1298,1376,1015,1040]" captionStart="FIGURE 3" captionStartId="8.[151,250,1715,1737]" captionTargetBox="[303,1284,193,1692]" captionTargetId="figure@8.[303,1284,193,1693]" captionTargetPageId="8" captionText="FIGURE 3. Neotype of Lacerta muralis guadarramae Boscá 1916 MNHN 2012.0262 from the village of Lozoya, Sierra de Guadarrama (province of Madrid, Spain). Note the well flattened head, the strongly black-spotted pileus, the lack of vertebral stripe, the wide light dorsolateral and black supradorsolateral stripes on the dorsum, continuing on tail, and the light orange iris. Photos P. Geniez." httpUri="https://zenodo.org/record/285984/files/figure.png" pageId="13" pageNumber="14">Fig. 3</figureCitation>
|
||
): 59 longitudinal rows of dorsal scales at midbody, 28 gular scales counted along a longitudinal fictive line from the contact between the fourth pair of maxillary scales to the collar, 27 transversal rows of ventral plates from the collar to the anal plates, 16 and 17 femoral pores (left and right), 29 subdigital lamellae beneath the fourth toe, 4 supralabials in front of the subocular but a small further plate inserted between the third and the fourth supralabials, 87 scales on the temporal area, one small masseteric shield on each side, snout-vent length
|
||
<quantity id="4CB79B77FFA5C20C81C9FB64FB61FB56" box="[1180,1261,1196,1221]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="13" pageNumber="14" unit="mm" value="58.0">58 mm</quantity>
|
||
, tail length
|
||
<quantity id="4CB79B77FFA5C20C8027FB64FF49FB7A" metricMagnitude="-1" metricUnit="m" metricValue="1.0699999999999998" pageId="13" pageNumber="14" unit="mm" value="107.0">107 mm</quantity>
|
||
, pileus length 15.0 mm, pileus width
|
||
<quantity id="4CB79B77FFA5C20C8720FB07FD43FB7A" box="[629,719,1231,1256]" metricMagnitude="-3" metricUnit="m" metricValue="6.8" pageId="13" pageNumber="14" unit="mm" value="6.8">6.8 mm</quantity>
|
||
, pileus high
|
||
<quantity id="4CB79B77FFA5C20C8637FB07FC30FB7A" box="[866,956,1231,1256]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="13" pageNumber="14" unit="mm" value="6.4">6.4 mm</quantity>
|
||
, pileus strongly marked by 12 large black blotches, dorsum grey brown with two continuous paler dorsolateral stripes, each bordered on their inner edge by a wide supra-dorsolateral black stripe with light ocelli, reaching the base of the tail; no vertebral stripe, throat whitish with a few dark dots on the sides; belly uniform pale pinkish in life (with no dark markings, a rare character in males of this form but matching the description of the
|
||
<typeStatus id="54F48830FFA5C20C87ADFA97FCD6FAEA" box="[760,858,1375,1400]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4F4D11FFA5C20C86D5FAA9FB5FFAEA" box="[896,1235,1375,1400]" class="Reptilia" family="Lacertidae" genus="Lacerta" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="14" phylum="Chordata" rank="subSpecies" species="muralis" subSpecies="guadarramae">
|
||
<emphasis id="B93BEA80FFA5C20C86D5FAA9FB5FFAEA" box="[896,1235,1375,1400]" italics="true" pageId="13" pageNumber="14">Lacerta muralis guadarramae</emphasis>
|
||
</taxonomicName>
|
||
given by Boscá).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |