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<document id="327C9BCEB4A19865B9D1DA527E17CB0A" ID-CLB-Dataset="24900" ID-DOI="10.1093/isd/ixz015" ID-GBIF-Dataset="56172c62-3e7c-4ab7-83d4-1dd9f5c57eb5" ID-Zenodo-Dep="3542130" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573763013583" checkinUser="plazi" docAuthor="Hita-Garcia, Francisco, Lieberman, Ziv, Audisio, Tracy L., Liu, Cong &amp; Economo, Evan P." docDate="2019" docId="03D9AC4AE549FFA5FF64FEE6BE81070A" docLanguage="en" docName="insectSystematicsDiversity.ixz015.pdf" docOrigin="Insect Systematics and Diversity 5" docStyle="DocumentStyle:20C34595BA01015AC07A3B955E96DF74.8:InsectSystDivers.2017-.journal_article" docStyleId="20C34595BA01015AC07A3B955E96DF74" docStyleName="InsectSystDivers.2017-.journal_article" docStyleVersion="8" docTitle="Discothyrea aisnetu Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo 2019, sp. n." docType="treatment" docVersion="6" lastPageNumber="31" masterDocId="FFE0D432E555FFBBFFF6FF95BB120352" masterDocTitle="Revision of the Highly Specialized Ant Genus Discothyrea (Hymenoptera: Formicidae) in the Afrotropics with X-Ray Microtomography and 3 D Cybertaxonomy" masterLastPageNumber="84" masterPageNumber="1" pageNumber="29" updateTime="1698765580234" updateUser="ExternalLinkService">
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<mods:title id="5C42AA58E18D147D0BFBEF87A15DFA0D">Revision of the Highly Specialized Ant Genus Discothyrea (Hymenoptera: Formicidae) in the Afrotropics with X-Ray Microtomography and 3 D Cybertaxonomy</mods:title>
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<mods:namePart id="C87637E718C5B1781C6E59D2F3DAF42E">Hita-Garcia, Francisco</mods:namePart>
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<mods:namePart id="ED99F8F273E086CBE494D8DBF82A07FA">Lieberman, Ziv</mods:namePart>
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<mods:namePart id="D0E46DEA1F883051A096ED1A306ED182">Audisio, Tracy L.</mods:namePart>
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<mods:namePart id="52B159270896DD018E5C1163D9EDB222">Liu, Cong</mods:namePart>
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<mods:namePart id="0199FCFA75B67854984888173767EE04">Economo, Evan P.</mods:namePart>
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<mods:title id="828BCAC0BE152CA2E4F2A958C858B2A2">Insect Systematics and Diversity</mods:title>
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<mods:date id="207ACF8B654A728E2BAA77937632A533">2019</mods:date>
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<mods:number id="0DE802F29D06BE87CBDE3239ED448646">2019-11-14</mods:number>
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<mods:identifier id="8836178841C1A4203A954A0CAF0AABF2" type="DOI">10.1093/isd/ixz015</mods:identifier>
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<treatment id="03D9AC4AE549FFA5FF64FEE6BE81070A" ID-DOI="http://doi.org/10.5281/zenodo.5922590" ID-GBIF-Taxon="160400434" ID-Zenodo-Dep="5922590" LSID="urn:lsid:plazi:treatment:03D9AC4AE549FFA5FF64FEE6BE81070A" httpUri="http://treatment.plazi.org/id/03D9AC4AE549FFA5FF64FEE6BE81070A" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
<subSubSection id="C36A4ED7E549FFA7FF64FEE6B9C002D8" box="[146,722,370,394]" pageId="28" pageNumber="29" type="nomenclature">
<paragraph id="8BCF1D5CE549FFA7FF64FEE6B9C002D8" blockId="28.[146,771,336,726]" box="[146,722,370,394]" pageId="28" pageNumber="29">
<taxonomicName id="4C7066DFE549FFA7FF64FEE6B99C02D8" authority="Hita Garcia &amp; Lieberman" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[146,654,370,394]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="aisnetu" status="sp. nov.">
<emphasis id="B904C14EE549FFA7FF64FEE6BA6002D8" box="[146,370,371,394]" italics="true" pageId="28" pageNumber="29">Discothyrea aisnetu</emphasis>
Hita Garcia &amp; Lieberman
</taxonomicName>
<taxonomicNameLabel id="A2377C35E549FFA7FD63FEE6B9C002D8" box="[661,722,371,394]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C36A4ED7E549FFA7FF64FE00BAC7029E" pageId="28" pageNumber="29" type="description">
<paragraph id="8BCF1D5CE549FFA7FF64FE00BAC7029E" blockId="28.[146,771,336,726]" pageId="28" pageNumber="29">
(
<figureCitation id="134B01D9E549FFA7FF6CFE00BBE102FE" box="[154,243,405,428]" captionStart="Fig" captionStartId="5.[146,173,1283,1300]" captionTargetBox="[153,1435,148,1247]" captionTargetId="figure@5.[152,1435,147,1248]" captionTargetPageId="5" captionText="Fig. 4. Maps showing the known distribution ranges of Afrotropical Discothyrea species." figureDoi="http://doi.org/10.5281/zenodo.3542140" httpUri="https://zenodo.org/record/3542140/files/figure.png" pageId="28" pageNumber="29">Figs. 4C</figureCitation>
,
<figureCitation id="134B01D9E549FFA7FEF6FE00BA0C02FE" box="[256,286,405,428]" captionStart="Fig" captionStartId="10.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1418]" captionTargetId="figure@10.[154,1434,147,1419]" captionTargetPageId="10" captionText="Fig. 6. Diagnostic plate showing still images from surface volume renderings of the head in full-face view (the remainder of the body virtually removed). All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542144" httpUri="https://zenodo.org/record/3542144/files/figure.png" pageId="28" pageNumber="29">6C</figureCitation>
,
<figureCitation id="134B01D9E549FFA7FEDDFE00BA5B02FE" box="[299,329,405,428]" captionStart="Fig" captionStartId="11.[146,173,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@11.[152,1435,147,1422]" captionTargetPageId="11" captionText="Fig. 7. Diagnostic plate showing still images from surface volume renderings of the head in profile (the remainder of the body virtually removed).All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation. Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542148" httpUri="https://zenodo.org/record/3542148/files/figure.png" pageId="28" pageNumber="29">7C</figureCitation>
,
<figureCitation id="134B01D9E549FFA7FEA0FE00BA6102FE" box="[342,371,405,428]" captionStart="Fig" captionStartId="12.[146,172,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@12.[152,1435,147,1422]" captionTargetPageId="12" captionText="Fig. 8. Diagnostic plate showing still images from surface volume renderings of the right mandible in dorsal view (the remainder of the body virtually removed).All Afrotropical species are shown,species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542150" httpUri="https://zenodo.org/record/3542150/files/figure.png" pageId="28" pageNumber="29">8C</figureCitation>
,
<figureCitation id="134B01D9E549FFA7FE77FE00BA8C02FE" box="[385,414,405,428]" captionStart="Fig" captionStartId="13.[146,173,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@13.[152,1435,147,1422]" captionTargetPageId="13" captionText="Fig. 9. Diagnostic plate showing still images from surface volume renderings of the mesosoma in profile (the remainder of the body virtually removed). All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542152" httpUri="https://zenodo.org/record/3542152/files/figure.png" pageId="28" pageNumber="29">9C</figureCitation>
,
<figureCitation id="134B01D9E549FFA7FE5DFE00BAC102FE" box="[427,467,405,428]" captionStart="Fig" captionStartId="14.[146,173,1457,1474]" captionTargetBox="[153,1435,149,1421]" captionTargetId="figure@14.[152,1435,147,1422]" captionTargetPageId="14" captionText="Fig. 10. Diagnostic plate showing still images from surface volume renderings of the body in dorsal view with focus on mesosoma.All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation. Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542154" httpUri="https://zenodo.org/record/3542154/files/figure.png" pageId="28" pageNumber="29">10C</figureCitation>
,
<figureCitation id="134B01D9E549FFA7FE14FE00B91402FE" box="[482,518,405,428]" captionStart="Fig" captionStartId="15.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1418]" captionTargetId="figure@15.[154,1434,147,1419]" captionTargetPageId="15" captionText="Fig. 11. Diagnostic plate showing still images from surface volume renderings of the posterior propodeum in posterior view (the remainder of the body virtually removed). All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation. Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542156" httpUri="https://zenodo.org/record/3542156/files/figure.png" pageId="28" pageNumber="29">11C</figureCitation>
,
<figureCitation id="134B01D9E549FFA7FDE1FE00B95302FE" box="[535,577,405,428]" captionStart="Fig" captionStartId="16.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1418]" captionTargetId="figure@16.[154,1434,147,1419]" captionTargetPageId="16" captionText="Fig. 12. Diagnostic plate showing still images from surface volume renderings of the petiole and gaster in profile (the remainder of the body virtually removed).All Afrotropical species are shown, species names are embedded in each particular image for better comparison and orientation.Specimen codes are given inTable 1." figureDoi="http://doi.org/10.5281/zenodo.3542158" httpUri="https://zenodo.org/record/3542158/files/figure.png" pageId="28" pageNumber="29">12C</figureCitation>
,
<figureCitation id="134B01D9E549FFA7FDB8FE00B96502FE" box="[590,631,405,428]" captionStart="Fig" captionStartId="18.[146,173,1454,1471]" captionTargetBox="[155,1434,149,1419]" captionTargetId="figure@18.[154,1434,147,1419]" captionTargetPageId="18" captionText="Fig. 14. Diagnostic plate showing differences in body pilosity. (A) D. mixta (CASENT0235473),(B) D. oculata (CASENT0235467),(C) D. aisnetu (CASENT0235475), (D) D.athene (CASENT0235476),(E) D.chimera (CASENT0235471),(F) D.damato (CASENT0247362),(G) D.dryad (CASENT0247371),(H) D.gaia (CASENT0247040), (I) D. gryphon (CASENT0247367), (J) D. hawkesi (CASENT0235470), (K) D. kalypso (CASENT0235468), (L) D. maia (CASENT0790541), (M) D. michelae (CASENT0235469), (N) D. patrizii (CASENT0235472), (O) D. penthos (CASENT0247383), (P) D. poweri (SAM-ENT-0011509), (Q) D. schulzei (CASENT0247370), (R) D. traegaordhi (CASENT0790122), (S) D. venus (CASENT0247017), (T) D. wakanda (CASENT0790326).With the exception of F, L, R,T, all other images are from https://www.antweb.org—photographers Michele Esposito and Will Ericson." figureDoi="http://doi.org/10.5281/zenodo.3542162" httpUri="https://zenodo.org/record/3542162/files/figure.png" pageId="28" pageNumber="29">14C</figureCitation>
,
<figureCitation id="134B01D9E549FFA7FD70FE00B9B302FE" box="[646,673,405,428]" captionStart="Fig" captionStartId="28.[146,173,1892,1909]" captionTargetBox="[147,768,978,1857]" captionTargetId="figure@28.[147,768,978,1857]" captionTargetPageId="28" captionText="Fig. 23. Stacked digital color images of D. aisnetu sp. n. holotype (CASENT0235475 - from https://www.antweb.org, photographerWill Ericson). (A) body in profile, (B) body in dorsal view, (C) head in full-face view." figureDoi="http://doi.org/10.5281/zenodo.3542180" httpUri="https://zenodo.org/record/3542180/files/figure.png" pageId="28" pageNumber="29">23</figureCitation>
,
<figureCitation id="134B01D9E549FFA7FD58FE00B9DB02FE" box="[686,713,405,428]" captionStart="Fig" captionStartId="29.[146,173,1782,1799]" captionTargetBox="[202,1386,147,1747]" captionTargetId="figure@29.[202,1386,147,1747]" captionTargetPageId="29" captionText="Fig. 24. Still images from shaded surface display volume renderings of D. aisnetu sp. n. holotype (CASENT0235475) showing virtually segmented body parts. (A) Body in dorsal view, (B) body in profile, (C) head in full-face view, (D) head in dorsal view, (E) head in anterodorsal view, (F) head in anterior view, (G) head in profile, (H) head in ventral view, (I) posterior propodeum in posterior view, (J) petiole in profile, (K) petiole and gaster in profile, (L) petiole in dorsal view, (M) petiole in anterior view, (N) petiole oblique anterior view, (O) gaster in dorsal view, (P) petiole in ventral view, (Q) abdominal sternite 3 in ventral view, (R) mesotibia and mesotarsus in anterior view." figureDoi="http://doi.org/10.5281/zenodo.3542182" httpUri="https://zenodo.org/record/3542182/files/figure.png" pageId="28" pageNumber="29">24</figureCitation>
; Supp Video S3 [online only])
</paragraph>
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<subSubSection id="C36A4ED7E549FFA7FF64FE4DBEB103B1" pageId="28" pageNumber="29" type="materials_examined">
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<emphasis id="B904C14EE549FFA7FF64FE4DBA0602B9" bold="true" box="[146,276,472,491]" pageId="28" pageNumber="29">Type Material</emphasis>
</paragraph>
<paragraph id="8BCF1D5CE549FFA7FF64FE60BADD0184" blockId="28.[146,771,336,726]" pageId="28" pageNumber="29">
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<typeStatus id="54CBA3FEE549FFA7FF64FE60BA19015A" box="[146,267,501,520]" pageId="28" pageNumber="29" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
, pinned worker,
<collectingCountry id="F3675DCCE549FFA7FE4CFE63B923015B" box="[442,561,502,521]" name="Tanzania" pageId="28" pageNumber="29">TANZANIA</collectingCountry>
,
<collectingRegion id="49B4D3BEE549FFA7FDB6FE60B9EC015A" box="[576,766,501,520]" country="Tanzania" name="Kilimanjaro" pageId="28" pageNumber="29">Kilimanjaro Region</collectingRegion>
,
<collectingMunicipality id="6BAB8726E549FFA7FF64FD86BBF70174" box="[146,229,531,550]" pageId="28" pageNumber="29">Mwanga</collectingMunicipality>
,
<location id="8EAF4B87E549FFA7FF07FD86BAFA0174" LSID="urn:lsid:plazi:treatment:03D9AC4AE549FFA5FF64FEE6BE81070A:8EAF4B87E549FFA7FF07FD86BAFA0174" box="[241,488,531,550]" country="Tanzania" latitude="-3.7452" longLatPrecision="5" longitude="37.64267" municipality="Mwanga" name="Kindoroko Forest Reserve" pageId="28" pageNumber="29" stateProvince="Kilimanjaro">Kindoroko Forest Reserve</location>
,
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<geoCoordinate id="EE447B9BE549FFA7FDBBFD86B9BA0174" box="[589,680,531,550]" degrees="37.64267" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="37.64267">37.64267</geoCoordinate>
,
<quantity id="4C88B0B9E549FFA7FD45FD86B9EC0174" box="[691,766,531,550]" metricMagnitude="3" metricUnit="m" metricValue="1.739" pageId="28" pageNumber="29" unit="m" value="1739.0">
<elevation id="005DFA6FE549FFA7FD45FD86B9EC0174" box="[691,766,531,550]" metricMagnitude="3" metricUnit="m" metricValue="1.739" pageId="28" pageNumber="29" unit="m" value="1739.0">1739 m</elevation>
</quantity>
, primary forest, leaf litter,
<collectorName id="2685788AE549FFA7FE78FDA5BACA0111" box="[398,472,560,579]" pageId="28" pageNumber="29">Winkler</collectorName>
, collection code CEPF-TZ-5.2,
<date id="FFCE3B9CE549FFA7FF64FDD8BA1F0132" box="[146,269,589,608]" pageId="28" pageNumber="29" value="2005-09-05" valueMax="2005-09-08" valueMin="2005-09-05">
<collectingDate id="EF8AC274E549FFA7FF64FDD8BA1F0132" box="[146,269,589,608]" pageId="28" pageNumber="29" value="2005-09-05" valueMax="2005-09-08" valueMin="2005-09-05">5.–8.IX.2005</collectingDate>
</date>
(
<emphasis id="B904C14EE549FFA7FED5FDDBB9BA0133" box="[291,680,589,609]" italics="true" pageId="28" pageNumber="29">
<collectorName id="2685788AE549FFA7FED5FDDBBA9F0132" box="[291,397,589,608]" pageId="28" pageNumber="29">P. Hawkes</collectorName>
,
<collectorName id="2685788AE549FFA7FE69FDDBB91C0132" box="[415,526,589,608]" pageId="28" pageNumber="29">J. Makwati</collectorName>
&amp;
<collectorName id="2685788AE549FFA7FDB6FDDBB9B30133" box="[576,673,590,609]" pageId="28" pageNumber="29">R. Mtana</collectorName>
)
</emphasis>
(
<collectionCode id="ED618599E549FFA7FD4BFDDBB9ED0133" box="[701,767,590,609]" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="28" pageNumber="29" type="Museum">SAMC</collectionCode>
:
<specimenCode id="DBD6B527E549FFA7FF64FDFEBA29012C" box="[146,315,619,638]" collectionCode="SAMC" country="South Africa" httpUri="http://grbio.org/cool/94p5-pepz" name="Iziko Museums of Cape Town" pageId="28" pageNumber="29" type="Museum">CASENT0235475</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B181701E549FFA7FEA7FDFEBAD90184" ID-GBIF-Occurrence="2451560215" collectingDate="2005-09-05" collectingDateMax="2005-09-08" collectingDateMin="2005-09-05" collectionCode="BMNH, CASC" collectorName="Winkler &amp; P. Hawkes &amp; J. Makwati &amp; R. Mtana" country="Tanzania" elevation="1739" latitude="-3.7452" location="Kindoroko Forest Reserve" longLatPrecision="5" longitude="37.64267" municipality="Mwanga" pageId="28" pageNumber="29" specimenCode="CASENT0250399, CASENT0250394" specimenCount="2" stateProvince="Kilimanjaro" typeStatus="paratype">
<emphasis id="B904C14EE549FFA7FEA7FDFEBAD9012C" bold="true" box="[337,459,619,638]" pageId="28" pageNumber="29">
<typeStatus id="54CBA3FEE549FFA7FEA7FDFEBAD9012C" box="[337,459,619,638]" pageId="28" pageNumber="29" type="paratype">PARATYPES</typeStatus>
</emphasis>
, one worker with same data as holotype (
<collectionCode id="ED618599E549FFA7FF07FD1DBA2901C9" box="[241,315,648,667]" pageId="28" pageNumber="29">BMNH</collectionCode>
:
<specimenCode id="DBD6B527E549FFA7FEB0FD1DBAE001C9" box="[326,498,648,667]" collectionCode="BMNH" pageId="28" pageNumber="29">CASENT0250399</specimenCode>
); and one worker with same data as
<typeStatus id="54CBA3FEE549FFA7FF2FFD30BA3801EA" box="[217,298,677,696]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
except collection code CEPF-TZ-5.4 and hand collected (
<collectionCode id="ED618599E549FFA7FF25FD56BA1D0184" box="[211,271,707,726]" pageId="28" pageNumber="29">CASC</collectionCode>
:
<specimenCode id="DBD6B527E549FFA7FEEEFD56BAD70184" box="[280,453,707,726]" collectionCode="CASC" pageId="28" pageNumber="29">CASENT0250394</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BCF1D5CE549FFA7FF64FC95BEB103B1" blockId="28.[146,770,768,934]" lastBlockId="28.[818,1443,149,228]" pageId="28" pageNumber="29">
<emphasis id="B904C14EE549FFA7FF64FC95BBE50041" box="[146,247,768,787]" italics="true" pageId="28" pageNumber="29">Cybertype.</emphasis>
Volumetric raw data (in DICOM format), 3D rotation video, still images of surface volume rendering, and 3D surface (in PLY format) of the physical
<typeStatus id="54CBA3FEE549FFA7FE52FCAEBAE7001C" box="[420,501,827,846]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
(CASENT0235475) in addition to stacked digital color images illustrating head in full-face view, profile and dorsal views of the body.The data are deposited at Dryad (Hita Garcia et al. 2019, http://doi.org/10.5061/dryad.3qm4183) and can be freely accessed as virtual representation of the type. In addition to the cybertype data at Dryad, we also provide a freely accessible 3D surface model of the
<typeStatus id="54CBA3FEE549FFA7FB85FF45BFD603B1" box="[1139,1220,208,227]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
at Sketchfab (Model 3).
</paragraph>
</subSubSection>
<subSubSection id="C36A4ED7E549FFA7FCC4FE85BE310281" pageId="28" pageNumber="29" type="diagnosis">
<paragraph id="8BCF1D5CE549FFA7FCC4FE85B89E0271" blockId="28.[818,1442,272,468]" box="[818,908,272,291]" pageId="28" pageNumber="29">
<heading id="D087AA30E549FFA7FCC4FE85B89E0271" bold="true" box="[818,908,272,291]" fontSize="8" level="3" pageId="28" pageNumber="29" reason="6">
<emphasis id="B904C14EE549FFA7FCC4FE85B89E0271" bold="true" box="[818,908,272,291]" pageId="28" pageNumber="29">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE549FFA7FCC4FEB8BE310281" blockId="28.[818,1442,272,468]" pageId="28" pageNumber="29">
The following character combination separates
<taxonomicName id="4C7066DFE549FFA7FB1AFEBBBE590212" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[1260,1355,302,320]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="aisnetu">
<emphasis id="B904C14EE549FFA7FB1AFEBBBE590212" box="[1260,1355,302,320]" italics="true" pageId="28" pageNumber="29">D. aisnetu</emphasis>
</taxonomicName>
from the other species of the complex: mesosomal profile high-rounded and extremely convex; relatively large species (HW 0.57–0.61; WL 0.80– 0.90); in dorsal view mesosoma relatively thin and slender (DMI 53–58; DMI2 85–88); mesotibia without apicoventral spur; AT3 around 1.1 to 1.2 times longer than AT4 (ASI 85–90).
</paragraph>
</subSubSection>
<subSubSection id="C36A4ED7E549FFA7FCC4FD9DBF370439" pageId="28" pageNumber="29" type="description">
<paragraph id="8BCF1D5CE549FFA7FCC4FD9DBFA70149" blockId="28.[818,1442,520,716]" box="[818,1205,520,539]" pageId="28" pageNumber="29">
<heading id="D087AA30E549FFA7FCC4FD9DBFA70149" bold="true" box="[818,1205,520,539]" fontSize="8" level="3" pageId="28" pageNumber="29" reason="6">
<emphasis id="B904C14EE549FFA7FCC4FD9DBFA70149" bold="true" box="[818,1205,520,539]" pageId="28" pageNumber="29">
Worker Measurements and Indices (
<emphasis id="B904C14EE549FFA7FB8BFD9DBF9B0149" bold="true" box="[1149,1161,520,539]" italics="true" pageId="28" pageNumber="29">n</emphasis>
= 3)
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE549FFA7FCC4FDB3BF830199" blockId="28.[818,1442,520,716]" pageId="28" pageNumber="29">
EL 0.03; HL 0.68–0.70; HW 0.57–0.61; SL 0.43–0.45;
<collectingCountry id="F3675DCCE549FFA7FA72FDB3BEB0016B" box="[1412,1442,550,569]" name="Philippines" pageId="28" pageNumber="29">PH</collectingCountry>
0.40–0.43; PW 0.46–0.48; DML 0.52–0.57; PrH 0.49–0.53; WL 0.80–0.90; HFL 0.55–0.58; PeL 0.15–0.17; PeW 0.33–0.37; PeH 0.30–0.31; LT3 0.50–0.56; LT4 0.45–0.50; OI 4; CI 84–87; SI 63–67; LMI 44–51; DMI 53–58; DMI2 85–88; ASI 85–90; HFI 64–69; DPeI 218–220; LPeI 184–200.
</paragraph>
<paragraph id="8BCF1D5CE549FFA7FCC4FD6BB8F50042" blockId="28.[818,1443,765,1958]" box="[818,999,765,785]" pageId="28" pageNumber="29">
<heading id="D087AA30E549FFA7FCC4FD6BB8F50042" bold="true" box="[818,999,765,785]" fontSize="8" level="3" pageId="28" pageNumber="29" reason="6">
<emphasis id="B904C14EE549FFA7FCC4FD6BB8F50042" bold="true" box="[818,999,765,785]" pageId="28" pageNumber="29">Worker Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE549FFA7FCC4FC8EBFF20682" blockId="28.[818,1443,765,1958]" pageId="28" pageNumber="29">
<emphasis id="B904C14EE549FFA7FCC4FC8EB877007C" box="[818,869,795,814]" italics="true" pageId="28" pageNumber="29">Head</emphasis>
slightly longer than broad (CI 84–87), posterior head margin nearly straight; posterodorsal corners of head rounded; sides of head slightly constricted laterally between eyes and anterolateral corner of gena, this region appearing concave in frontal view; eyes present, small (OI 4), round, comprising several ommatidia, situated around one-third of way between corner of gena and posterior head margin; eyes just visible in frontal view; frontal lamella triangular in profile, apex rounded; lamella more or less evenly translucent across its disc, without fenestra; medial clypeus broad, gently convex, lateral clypeus curving gently between antennal sockets and anterolateral corners of head, bearing short, dense, curved setae.
<emphasis id="B904C14EE549FFA7FC46FBCBBF120723" box="[944,1024,1118,1137]" italics="true" pageId="28" pageNumber="29">Antenna</emphasis>
with long scape (SI 63–67), scape somewhat expanded apically, gently bent; pedicel campaniform, longer than broad; true antennomere count nine; apparent antennomere count nine to eleven, flagellomeres basad apical club relatively long, taken together longer than apical club; club relatively narrow.
<emphasis id="B904C14EE549FFA7FCC4FB65B8BB0651" box="[818,937,1264,1283]" italics="true" pageId="28" pageNumber="29">Ventral head</emphasis>
with postoccipital ridge well-defined, with very short anteromedian carina; hypostoma very wide overall, median region bluntly triangular, arms only slightly narrowed, similar in width across their entire length; palpal formula not examined.
<emphasis id="B904C14EE549FFA7FCC4FAF0B899062A" box="[818,907,1381,1400]" italics="true" pageId="28" pageNumber="29">Mandible</emphasis>
edentate; basal angle rounded; ectal face with low carina originating near basal angle and becoming confluent with masticatory margin at apical one-fourth, leaving narrow, arcuate, depressed medial region on masticatory margin.
</paragraph>
<paragraph id="8BCF1D5CE549FFA7FCA4FA4EBFD705A4" blockId="28.[818,1443,765,1958]" pageId="28" pageNumber="29">
<emphasis id="B904C14EE549FFA7FCA4FA4EB8A606BF" box="[850,948,1499,1517]" italics="true" pageId="28" pageNumber="29">Mesosoma</emphasis>
in profile robust, high-rounded and extremely convex; in dorsal view mesosoma relatively thin and slender (DMI 53–58; DMI2 85–88) and distinctly narrowed posteriorly, pronotum significantly wider than propodeum; pronotal humeri broadly rounded; posterior propodeal margin indistinct or appearing straight; in profile, propodeum without distinct juncture between dorsal and declivitous faces, lacking angles or denticles, not marginate; propodeal declivity sloping, at most barely concave in oblique posterior view; propodeal spiracle distinct, aperture round, directed posteriorly; propodeal lobes well-developed, flange-like.
</paragraph>
<paragraph id="8BCF1D5CE549FFA7FCA4F894BF370439" blockId="28.[818,1443,765,1958]" pageId="28" pageNumber="29">
<emphasis id="B904C14EE549FFA7FCA4F894B86F0441" box="[850,893,1793,1811]" italics="true" pageId="28" pageNumber="29">Legs</emphasis>
moderately long to long and narrow (HFI 64–69); mesotibia without apicoventral spur, with small but distinct seta inserted in apical pit; mesobasitarsus very elongate, almost as long as remaining tarsomeres taken together.
</paragraph>
</subSubSection>
<caption id="DF0F4DD4E549FFA7FF64F8F1B9D504FA" ID-DOI="http://doi.org/10.5281/zenodo.3542180" ID-Zenodo-Dep="3542180" httpUri="https://zenodo.org/record/3542180/files/figure.png" pageId="28" pageNumber="29" startId="28.[146,173,1892,1909]" targetBox="[147,768,978,1857]" targetPageId="28">
<paragraph id="8BCF1D5CE549FFA7FF64F8F1B9D504FA" blockId="28.[146,772,1892,1960]" pageId="28" pageNumber="29">
<emphasis id="B904C14EE549FFA7FF64F8F1BBC50427" bold="true" box="[146,215,1892,1909]" pageId="28" pageNumber="29">Fig. 23.</emphasis>
Stacked digital color images of
<taxonomicName id="4C7066DFE549FFA7FDFBF8F1B97A0427" authority="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo, 2019" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[525,616,1892,1909]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="aisnetu" status="sp. nov.">
<emphasis id="B904C14EE549FFA7FDFBF8F1B97A0427" box="[525,616,1892,1909]" italics="true" pageId="28" pageNumber="29">D. aisnetu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377C35E549FFA7FD80F8F1B9BE0427" box="[630,684,1892,1909]" pageId="28" pageNumber="29" rank="species">sp. n.</taxonomicNameLabel>
holotype (CASENT0235475 - from https://www.antweb.org, photographerWill Ericson). (A) body in profile, (B) body in dorsal view, (C) head in full-face view.
</paragraph>
</caption>
<subSubSection id="C36A4ED7E549FFA5FCA4F8E0BA0701AC" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BCF1D5CE549FFA5FCA4F8E0BA0701AC" blockId="28.[818,1443,765,1958]" lastBlockId="30.[146,772,673,766]" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29">
<emphasis id="B904C14EE549FFA7FCA4F8E0B89D04DA" box="[850,911,1909,1928]" italics="true" pageId="28" pageNumber="29">Petiole</emphasis>
node quite thick, about 1.8 to 2.0 times higher than long (LPeI 184–200), rounded-cuneate in profile; in profile anterior
<emphasis id="B904C14EE54BFFA5FF64FD34BBCB01E0" bold="true" box="[146,217,673,690]" pageId="30" pageNumber="31">Model 3.</emphasis>
3D surface model of
<taxonomicName id="4C7066DFE54BFFA5FE79FD34BAF001E0" authority="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo, 2019" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[399,482,673,690]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="aisnetu" status="sp. nov.">
<emphasis id="B904C14EE54BFFA5FE79FD34BAF001E0" box="[399,482,673,690]" italics="true" pageId="30" pageNumber="31">D. aisnetu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377C35E54BFFA5FE1EFD34B90401E0" box="[488,534,673,690]" pageId="30" pageNumber="31" rank="species">sp. n.</taxonomicNameLabel>
<typeStatus id="54CBA3FEE54BFFA5FDECFD34B97001E0" box="[538,610,673,690]" pageId="30" pageNumber="31" type="holotype">holotype</typeStatus>
(CASENT0235475). An interactive version of this model is available in the HTML version of this article online and at https://sketchfab.com/3d-models/1bf459295c6941d6b11 51aa006e3382a.
</paragraph>
</subSubSection>
<caption id="DF0F4DD4E548FFA6FF64F963BAE4043E" ID-DOI="http://doi.org/10.5281/zenodo.3542182" ID-Zenodo-Dep="3542182" httpUri="https://zenodo.org/record/3542182/files/figure.png" pageId="29" pageNumber="30" startId="29.[146,173,1782,1799]" targetBox="[202,1386,147,1747]" targetPageId="29">
<paragraph id="8BCF1D5CE548FFA6FF64F963BAE4043E" blockId="29.[146,1444,1782,1900]" pageId="29" pageNumber="30">
<emphasis id="B904C14EE548FFA6FF64F963BBDC0455" bold="true" box="[146,206,1782,1799]" pageId="29" pageNumber="30">Fig. 24.</emphasis>
Still images from shaded surface display volume renderings of
<taxonomicName id="4C7066DFE548FFA6FD16F963B8210455" authority="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo, 2019" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[736,819,1782,1799]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="aisnetu" status="sp. nov.">
<emphasis id="B904C14EE548FFA6FD16F963B8210455" box="[736,819,1782,1799]" italics="true" pageId="29" pageNumber="30">D. aisnetu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2377C35E548FFA6FCCEF963B8740455" box="[824,870,1782,1799]" pageId="29" pageNumber="30" rank="species">sp. n.</taxonomicNameLabel>
holotype (CASENT0235475) showing virtually segmented body parts. (A) Body in dorsal view, (B) body in profile, (C) head in full-face view, (D) head in dorsal view, (E) head in anterodorsal view, (F) head in anterior view, (G) head in profile, (H) head in ventral view, (I) posterior propodeum in posterior view, (J) petiole in profile, (K) petiole and gaster in profile, (L) petiole in dorsal view, (M) petiole in anterior view, (N) petiole oblique anterior view, (O) gaster in dorsal view, (P) petiole in ventral view, (Q) abdominal sternite 3 in ventral view, (R) mesotibia and mesotarsus in anterior view.
</paragraph>
</caption>
<subSubSection id="C36A4ED7E54BFFA5FF64FCB1BE7F0269" pageId="30" pageNumber="31" type="description">
<paragraph id="8BCF1D5CE54BFFA5FF64FCB1BA870770" blockId="30.[146,770,804,1909]" pageId="30" pageNumber="31">face of node sloping posterodorsally, gently curved, apex rounded, hence without distinct posterior face; in dorsal view, node roundedrectangular to nearly elliptical, around 2.2 times broader than long (DPeI 218–220), sides convex; in anterior view, petiolar outline broadly disciform, without distinct angles, dorsally rounded; in ventral view, sides distinctly curved, outline campaniform; subpetiolar process elongate-triangular, weakly spiniform to digitate, anteriorly projecting, apex approximately rounded; petiolar spiracles very large, round in ventral view.</paragraph>
<paragraph id="8BCF1D5CE54BFFA5FF44FBB9BBE1065E" blockId="30.[146,770,804,1909]" pageId="30" pageNumber="31">
<emphasis id="B904C14EE54BFFA5FF44FBB9BA6C076D" box="[178,382,1068,1087]" italics="true" pageId="30" pageNumber="31">Abdominal segment 3</emphasis>
campaniform, widest posteriorly, sternite rounded and distinctly bulging in profile;
<collectingCountry id="F3675DCCE54BFFA5FDD1FBDCB956070E" box="[551,580,1097,1116]" name="American Samoa" pageId="30" pageNumber="31">AS</collectingCountry>
3 with weak medial carina; prora present as a fine carina, weakly concave in ventral view; AT3 around 1.1 to 1.2 times longer than AT4 (ASI 85–90); AT4 gently recurved hemidemispherical;
<collectingCountry id="F3675DCCE54BFFA5FDFCFB34B93707E6" box="[522,549,1185,1204]" name="American Samoa" pageId="30" pageNumber="31">AS</collectingCountry>
4 with anterior lip short and narrow, scarcely overlapping
<collectingCountry id="F3675DCCE54BFFA5FE24FB2ABAE20780" box="[466,496,1215,1234]" name="American Samoa" pageId="30" pageNumber="31">AS</collectingCountry>
3, anterior margin straight in ventral view; successive abdominal segments short, telescopic, often concealed.
</paragraph>
<paragraph id="8BCF1D5CE54BFFA5FF44FA82BA99051D" blockId="30.[146,770,804,1909]" pageId="30" pageNumber="31">
<emphasis id="B904C14EE54BFFA5FF44FA82BA1B0678" box="[178,265,1303,1322]" italics="true" pageId="30" pageNumber="31">Sculpture</emphasis>
on head evenly punctate-reticulate, sculpture equally developed over head except medial clypeus distinctly smoother; mandibles fairly smooth but with fine piligerous punctulae; mesosoma similarly punctate-reticulate on dorsum and sides, punctae sometimes arranged in concentric lines just anterad pronotomesepisternal junction; declivitious face of propodeum finely reticulate to confused-rugulose; petiole and abdominal segment 3 (tergite and sternite) similarly punctate-reticulate, punctures on AT3 piligerous; abdominal tergite 4 distinctively shinier and less coarsely sculptured than AT3 with fine piligerous punctulae and posteriorly with a few scattered, shallow punctae.
</paragraph>
<paragraph id="8BCF1D5CE54BFFA5FF44F9CFB8D5024C" blockId="30.[146,770,804,1909]" lastBlockId="30.[818,1442,149,316]" pageId="30" pageNumber="31">
<emphasis id="B904C14EE54BFFA5FF44F9CFBBEF053F" box="[178,253,1626,1645]" italics="true" pageId="30" pageNumber="31">Setation</emphasis>
on head and scape fairly short, dense, appressed and white pubescence; ectal face of mandible with fairly long, thin, curved, spaced setae; masticatory margin with row of straight, stout setae; mesosoma with well-developed appressed pubescence, more abundant on dorsum and lateral pronotum than remainder of mesosoma, shortest and most diffuse on sides of propodeum; petiole heavily setose, setae appressed to decumbent, quite long and thick, especially on dorsum of node; setae obscuring petiolar sculpture; ventral margin of petiolar sternite with row of straight, erect hairs; abdominal segment 3 with fairly long, mostly appressed setae, similar on tergite and sternite; AT4 more heavily setose; dorsum most strongly pubescent, becoming more diffuse ventrolaterad; successive abdominal segments with long, flexuous, curved pilosity; legs with appressed, fine pubescence, on tibiae and tarsi generally longer than on femora.
</paragraph>
<paragraph id="8BCF1D5CE54BFFA5FCA4FEBDBE7F0269" blockId="30.[818,1442,149,316]" box="[850,1389,296,316]" pageId="30" pageNumber="31">
<emphasis id="B904C14EE54BFFA5FCA4FEBDB8950269" box="[850,903,296,315]" italics="true" pageId="30" pageNumber="31">Color</emphasis>
luteous-orange to deep iron-red, appendages yellow.
</paragraph>
</subSubSection>
<subSubSection id="C36A4ED7E54BFFA5FCC4FEEEBFC002B4" pageId="30" pageNumber="31" type="etymology">
<paragraph id="8BCF1D5CE54BFFA5FCC4FEEEB88702DC" blockId="30.[818,1442,379,486]" box="[818,917,379,398]" pageId="30" pageNumber="31">
<heading id="D087AA30E54BFFA5FCC4FEEEB88702DC" bold="true" box="[818,917,379,398]" fontSize="8" level="3" pageId="30" pageNumber="31" reason="6">
<emphasis id="B904C14EE54BFFA5FCC4FEEEB88702DC" bold="true" box="[818,917,379,398]" pageId="30" pageNumber="31">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE54BFFA5FCC4FE0DBFC002B4" blockId="30.[818,1442,379,486]" pageId="30" pageNumber="31">The specific epithet is the Latin interjection ‘Aisne tu?’ meaning roughly ‘Could it be?’ (lit. ‘Do you say?’), recalling the authors’ reactions upon first viewing this aberrant species.</paragraph>
</subSubSection>
<subSubSection id="C36A4ED7E54BFFA5FCC4FD86BF1C01C9" pageId="30" pageNumber="31" type="distribution">
<paragraph id="8BCF1D5CE54BFFA5FCC4FD86BF0A0174" blockId="30.[818,1442,531,667]" box="[818,1048,531,550]" pageId="30" pageNumber="31">
<heading id="D087AA30E54BFFA5FCC4FD86BF0A0174" bold="true" box="[818,1048,531,550]" fontSize="8" level="3" pageId="30" pageNumber="31" reason="6">
<emphasis id="B904C14EE54BFFA5FCC4FD86BF0A0174" bold="true" box="[818,1048,531,550]" pageId="30" pageNumber="31">Distribution and Biology</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE54BFFA5FCC4FDA5BF1C01C9" blockId="30.[818,1442,531,667]" pageId="30" pageNumber="31">
At present the species is known only from the
<typeStatus id="54CBA3FEE54BFFA5FAF5FDA4BE380116" box="[1283,1322,561,580]" pageId="30" pageNumber="31">type</typeStatus>
locality, the Kindoroko Forest Reserve in
<collectingCountry id="F3675DCCE54BFFA5FBCAFDD8BF9D0132" box="[1084,1167,589,608]" name="Tanzania" pageId="30" pageNumber="31">Tanzania</collectingCountry>
(
<figureCitation id="134B01D9E54BFFA5FB6AFDD8BFCD0133" box="[1180,1247,589,609]" captionStart="Fig" captionStartId="5.[146,173,1283,1300]" captionTargetBox="[153,1435,148,1247]" captionTargetId="figure@5.[152,1435,147,1248]" captionTargetPageId="5" captionText="Fig. 4. Maps showing the known distribution ranges of Afrotropical Discothyrea species." figureDoi="http://doi.org/10.5281/zenodo.3542140" httpUri="https://zenodo.org/record/3542140/files/figure.png" pageId="30" pageNumber="31">Fig. 4C</figureCitation>
), which is a montane rainforest located at an altitude of
<quantity id="4C88B0B9E54BFFA5FB86FDFEBFA8012C" box="[1136,1210,619,638]" metricMagnitude="3" metricUnit="m" metricValue="1.739" pageId="30" pageNumber="31" unit="m" value="1739.0">1739 m</quantity>
.
<taxonomicName id="4C7066DFE54BFFA5FB35FDFEBE6A012F" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[1219,1400,619,638]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="aisnetu">
<emphasis id="B904C14EE54BFFA5FB35FDFEBE6A012F" box="[1219,1400,619,638]" italics="true" pageId="30" pageNumber="31">Discothyrea aisnetu</emphasis>
</taxonomicName>
was sampled from leaf litter.
</paragraph>
</subSubSection>
<subSubSection id="C36A4ED7E54BFFA5FCC4FD5DBE81070A" pageId="30" pageNumber="31" type="discussion">
<paragraph id="8BCF1D5CE54BFFA5FCC4FD5DB8840189" blockId="30.[818,1443,712,966]" box="[818,918,712,731]" pageId="30" pageNumber="31">
<heading id="D087AA30E54BFFA5FCC4FD5DB8840189" bold="true" box="[818,918,712,731]" fontSize="8" level="3" pageId="30" pageNumber="31" reason="6">
<emphasis id="B904C14EE54BFFA5FCC4FD5DB8840189" bold="true" box="[818,918,712,731]" pageId="30" pageNumber="31">Comments</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE54BFFA5FCC4FD70BEB20094" blockId="30.[818,1443,712,966]" pageId="30" pageNumber="31">
This highly characteristic species displays a unique mesosomal shape not seen in the remainder of the African
<taxonomicName id="4C7066DFE54BFFA5FB47FC96BE0E0044" authorityName="Roger" authorityYear="1863" box="[1201,1308,771,790]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis id="B904C14EE54BFFA5FB47FC96BE0E0044" box="[1201,1308,771,790]" italics="true" pageId="30" pageNumber="31">Discothyrea</emphasis>
</taxonomicName>
fauna. Within the
<taxonomicName id="4C7066DFE54BFFA5FCA0FCB5B8AE0061" authorityName="Santschi" authorityYear="1913" box="[854,956,800,819]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="traegaordhi">
<emphasis id="B904C14EE54BFFA5FCA0FCB5B8AE0061" box="[854,956,800,819]" italics="true" pageId="30" pageNumber="31">traegaordhi</emphasis>
</taxonomicName>
-complex it is a rather large species with long limbs, comparatively distinct flagellomeres and relatively narrow apical club. This is a character set shared only with
<taxonomicName id="4C7066DFE54BFFA5FB7BFCCEBFF5003F" baseAuthorityName="Arnold" baseAuthorityYear="1916" box="[1165,1255,859,877]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="poweri">
<emphasis id="B904C14EE54BFFA5FB7BFCCEBFF5003F" box="[1165,1255,859,877]" italics="true" pageId="30" pageNumber="31">D. poweri</emphasis>
</taxonomicName>
. However, it remains unclear if these two species are closely related since apart from the differently shaped mesosomal outline,
<taxonomicName id="4C7066DFE54BFFA5FB9AFC03BFD500FA" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[1132,1223,918,936]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="aisnetu">
<emphasis id="B904C14EE54BFFA5FB9AFC03BFD500FA" box="[1132,1223,918,936]" italics="true" pageId="30" pageNumber="31">D. aisnetu</emphasis>
</taxonomicName>
also lacks the mesotibial spur present in
<taxonomicName id="4C7066DFE54BFFA5FC4DFC26BF070097" baseAuthorityName="Arnold" baseAuthorityYear="1916" box="[955,1045,947,965]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="poweri">
<emphasis id="B904C14EE54BFFA5FC4DFC26BF070097" box="[955,1045,947,965]" italics="true" pageId="30" pageNumber="31">D. poweri</emphasis>
</taxonomicName>
, which is an important diagnostic character.
</paragraph>
<paragraph id="8BCF1D5CE54BFFA5FCC4FC78B89A0752" blockId="30.[818,1442,1005,1113]" box="[818,904,1005,1024]" pageId="30" pageNumber="31">
<heading id="D087AA30E54BFFA5FCC4FC78B89A0752" bold="true" box="[818,904,1005,1024]" fontSize="8" level="3" pageId="30" pageNumber="31" reason="6">
<emphasis id="B904C14EE54BFFA5FCC4FC78B89A0752" bold="true" box="[818,904,1005,1024]" pageId="30" pageNumber="31">Variation</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF1D5CE54BFFA5FCC4FB9EBE81070A" blockId="30.[818,1442,1005,1113]" pageId="30" pageNumber="31">
<taxonomicName id="4C7066DFE54BFFA5FCC4FB9EB8F8074F" authorityName="Hita-Garcia &amp; Lieberman &amp; Audisio &amp; Liu &amp; Economo" authorityYear="2019" box="[818,1002,1035,1054]" class="Insecta" family="Formicidae" genus="Discothyrea" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="aisnetu">
<emphasis id="B904C14EE54BFFA5FCC4FB9EB8F8074F" box="[818,1002,1035,1054]" italics="true" pageId="30" pageNumber="31">Discothyrea aisnetu</emphasis>
</taxonomicName>
varies slightly in color and in the pattern of punctae on the lateral pronotum just anterad the pronotomesepisternal juncture: they are more or less arranged in concentric broken lines.
</paragraph>
</subSubSection>
</treatment>
</document>
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