266 lines
29 KiB
XML
266 lines
29 KiB
XML
<document id="538A32E4A40CE7592616AB28A0DF85D3" ID-DOI="10.11646/palaeoentomology.4.5.11" ID-ISSN="2624-2834" ID-Zenodo-Dep="5530149" ID-ZooBank="50F1EB3E-450C-4B2C-A4F8-11E113C0FD02" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632739847826" checkinUser="plazi" docAuthor="BASHKUEV, ALEXEY S. & SUKATSHEVA, IRINA D." docDate="2021" docId="03D8B42BF236FF9BFF65F926CC06F7D0" docLanguage="en" docName="palaeoentomology.4.5.11.pdf" docOrigin="Palaeoentomology 4 (5)" docSource="http://dx.doi.org/10.11646/palaeoentomology.4.5.11" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Kamopanorpa rotunda BASHKUEV & SUKATSHEVA 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="458" masterDocId="FFE1CC53F232FF9EFFEDFFF8C913FFD7" masterDocTitle="New species of Kamopanorpa Martynov from the Permian of South Siberia with comments on the systematic position of Microptysmatidae (Protomeropina = Permotrichoptera)" masterLastPageNumber="461" masterPageNumber="453" pageNumber="457" updateTime="1699415050900" updateUser="plazi">
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<mods:titleInfo id="FFB9D7C043604C09F621169D8D7FAC6E">
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<mods:title id="5BD5D79A3A9395D23AA24A01BF276A04">New species of Kamopanorpa Martynov from the Permian of South Siberia with comments on the systematic position of Microptysmatidae (Protomeropina = Permotrichoptera)</mods:title>
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<mods:name id="0493A02D2B3B90CF4F476AED502C70C0" type="personal">
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<mods:roleTerm id="13C04A0DCF174925ADC1E5E92148DCB4">Author</mods:roleTerm>
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<mods:namePart id="740CE387E459F98AB7D2D13D340BACB2">BASHKUEV, ALEXEY S.</mods:namePart>
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<mods:name id="FEEF067667CAECAD23303617699B4140" type="personal">
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<mods:roleTerm id="BD8F07757CA06E98C5C6738FF4C9DEB0">Author</mods:roleTerm>
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<mods:namePart id="050A7CD489FEFD1C1A3E6CADDF16649B">SUKATSHEVA, IRINA D.</mods:namePart>
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<mods:typeOfResource id="90AA3B213144B47C7BD24768F0D26FA5">text</mods:typeOfResource>
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<mods:titleInfo id="15E631001161B19E04B60EC1038BABC1">
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<mods:title id="1354FFD89A73E003756C508E9A69CEBB">Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part id="5AA966C353112BB360340652FF2A170E">
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<mods:date id="85EFAC70074F02BF21B6D2C01E97C22E">2021</mods:date>
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<mods:detail id="40617953C9086E10F52ECB9DFEC3B314" type="pubDate">
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<mods:number id="521EA0B7A3BBE26CD9D560A00EE4B699">2021-09-24</mods:number>
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<mods:detail id="C2EFF739FD89705818BFC0ACF3DF66A8" type="volume">
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<mods:number id="C37521C11B70C4808C93FFFF56AA4E78">4</mods:number>
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<mods:detail id="F237D14E2F1A358FD6BF1F65EE01316B" type="issue">
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<mods:number id="0053513C0CC00F9FC00A599DF91A3D4B">5</mods:number>
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<mods:location id="6E949BB12020856AACD43FB8CF3FED84">
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<mods:url id="0C60183763EECDE958417C7C12F04244">http://dx.doi.org/10.11646/palaeoentomology.4.5.11</mods:url>
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<mods:classification id="BB2DA981A648637DA398D15AC34624C9">journal article</mods:classification>
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<mods:identifier id="00678967B43F9F1CC88EF2DE81D0D3EA" type="DOI">10.11646/palaeoentomology.4.5.11</mods:identifier>
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<mods:identifier id="AE39F2BCE805B8858E579210481E9C9E" type="ISSN">2624-2834</mods:identifier>
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</mods:mods>
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<treatment id="03D8B42BF236FF9BFF65F926CC06F7D0" LSID="urn:lsid:plazi:treatment:03D8B42BF236FF9BFF65F926CC06F7D0" httpUri="http://treatment.plazi.org/id/03D8B42BF236FF9BFF65F926CC06F7D0" lastPageId="5" lastPageNumber="458" pageId="4" pageNumber="457">
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<subSubSection id="C36B56B6F236FF9AFF65F926C8E3F92F" box="[136,496,1757,1784]" pageId="4" pageNumber="457" type="nomenclature">
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<paragraph id="8BCE053DF236FF9AFF65F926C8E3F92F" blockId="4.[136,496,1757,1821]" box="[136,496,1757,1784]" pageId="4" pageNumber="457">
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<heading id="D086B251F236FF9AFF65F926C8E3F92F" bold="true" box="[136,496,1757,1784]" fontSize="10" level="2" pageId="4" pageNumber="457" reason="3">
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<emphasis id="B905D92FF236FF9AFF65F926C8E3F92F" bold="true" box="[136,496,1757,1784]" pageId="4" pageNumber="457">
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<taxonomicName id="4C717EBEF236FF9AFF65F926C883F92F" authority="BASHKUEV & SUKATSHEVA, 2021" authorityName="BASHKUEV & SUKATSHEVA" authorityYear="2021" box="[136,400,1757,1784]" class="Insecta" family="Cladochoristidae" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="457" phylum="Arthropoda" rank="species" species="rotunda" status="sp. nov.">
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<emphasis id="B905D92FF236FF9AFF65F926C883F92F" bold="true" box="[136,400,1757,1784]" italics="true" pageId="4" pageNumber="457">Kamopanorpa rotunda</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2366454F236FF9AFE7AF926C8E3F92F" box="[407,496,1758,1784]" pageId="4" pageNumber="457" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B56B6F236FF9AFF65F8FAC832F8CA" box="[136,289,1794,1821]" pageId="4" pageNumber="457" type="description">
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<paragraph id="8BCE053DF236FF9AFF65F8FAC832F8CA" blockId="4.[136,496,1757,1821]" box="[136,289,1794,1821]" pageId="4" pageNumber="457">
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(
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<figureCitation id="134A19B8F236FF9AFF7DF8FAC9F8F8CA" box="[144,235,1794,1821]" captionStart="FIGURE 1" captionStartId="3.[136,229,1027,1049]" captionTargetBox="[222,1353,246,987]" captionTargetId="figure-428@3.[215,1371,223,1003]" captionTargetPageId="3" captionText="FIGURE 1. A, B, Kamopanorpa rasnitsyni sp. nov. A, Holotype PIN 5386/9. B, Paratype PIN 5386/3. C, K. rotunda sp. nov., holotype PIN 5386/5. D, K. sivchikovi sp. nov., holotype PIN 5386/1. E, F, Kamopanorpa spp. E, 5386/8. F, 5386/11.All to the same scale." figureDoi="http://doi.org/10.5281/zenodo.5530151" httpUri="https://zenodo.org/record/5530151/files/figure.png" pageId="4" pageNumber="457">Figs 1C</figureCitation>
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,
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<figureCitation id="134A19B8F236FF9AFF1AF8FBC80AF8CA" box="[247,281,1795,1821]" captionStart="FIGURE 2" captionStartId="4.[136,229,1083,1105]" captionTargetBox="[137,776,190,1059]" captionTargetId="figure-479@4.[137,776,190,1059]" captionTargetPageId="4" captionText="FIGURE 2. A, Kamopanorpa rasnitsyni sp. nov. Holotype PIN 5386/9. B, K. sivchikovi sp. nov., holotype PIN 5386/1. C, K. rotunda sp. nov., holotype PIN 5386/5. SEM images." figureDoi="http://doi.org/10.5281/zenodo.5530153" httpUri="https://zenodo.org/record/5530153/files/figure.png" pageId="4" pageNumber="457">2C</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B56B6F236FF9AFF65F8B5C8E4F85B" pageId="4" pageNumber="457" type="materials_examined">
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<paragraph id="8BCE053DF236FF9AFF65F8B5C8E4F85B" blockId="4.[136,778,1868,2086]" pageId="4" pageNumber="457">
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<emphasis id="B905D92FF236FF9AFF65F8B5C82AF8B1" bold="true" box="[136,313,1868,1895]" pageId="4" pageNumber="457">Type material.</emphasis>
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<materialsCitation id="3B190F60F236FF9AFEA1F8B5C8E4F85B" collectionCode="PIN" pageId="4" pageNumber="457" specimenCode="PIN 5386" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54CABB9FF236FF9AFEA1F8B5C8A7F8B0" box="[332,436,1869,1895]" pageId="4" pageNumber="457" type="holotype">Holotype</typeStatus>
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<specimenCode id="DBD7AD46F236FF9AFE53F8B5CB23F8B0" box="[446,560,1868,1895]" collectionCode="PIN" country="Russia" httpUri="http://grbio.org/cool/kjjc-ziy5" name="Paleontological Institute, Russian Academy of Sciences" pageId="4" pageNumber="457">PIN 5386</specimenCode>
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/5±, well-preserved forewing with anal area missing.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B56B6F236FF9AFF50F86FC8A1F801" pageId="4" pageNumber="457" type="etymology">
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<paragraph id="8BCE053DF236FF9AFF50F86FC8A1F801" blockId="4.[136,778,1868,2086]" pageId="4" pageNumber="457">
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<emphasis id="B905D92FF236FF9AFF50F86FC857F866" bold="true" box="[189,324,1943,1969]" pageId="4" pageNumber="457">Etymology.</emphasis>
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From Latin
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<emphasis id="B905D92FF236FF9AFE35F86FCB28F866" box="[472,571,1943,1969]" italics="true" pageId="4" pageNumber="457">rotundus</emphasis>
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(round), referring to roundish shape of wing.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36B56B6F236FF9AFF50F818CCBFFE62" pageId="4" pageNumber="457" type="diagnosis">
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<paragraph id="8BCE053DF236FF9AFF50F818CA19F7F7" blockId="4.[136,778,1868,2086]" pageId="4" pageNumber="457">
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<emphasis id="B905D92FF236FF9AFF50F818C82BF82D" bold="true" box="[189,312,2016,2042]" pageId="4" pageNumber="457">Diagnosis.</emphasis>
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Wing short and rounded; SC ending at R before level of RS
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<subScript id="17F50778F236FF9AFEB2F7E0C874F7F1" attach="left" box="[351,359,2072,2086]" fontSize="6" pageId="4" pageNumber="457">1</subScript>
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fork, with four subcostal crossveins
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</paragraph>
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<paragraph id="8BCE053DF236FF9AFCC7FF47CD02FE90" blockId="4.[810,1452,191,1136]" pageId="4" pageNumber="457">
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(besides apical one); subcostal space broad with long sigmoid crossvein. Clearly differs from all congeners by simplified venation of M with only four branches and very short M
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<subScript id="17F50778F236FF9AFC57FEB8CAC7FE99" attach="left" box="[954,980,320,334]" fontSize="6" pageId="4" pageNumber="457">3+4</subScript>
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fork.
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</paragraph>
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<paragraph id="8BCE053DF236FF9AFCB2FEAACCB8FE46" blockId="4.[810,1452,191,1136]" pageId="4" pageNumber="457">
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<emphasis id="B905D92FF236FF9AFCB2FEAACD7FFEBB" bold="true" box="[863,1132,338,364]" pageId="4" pageNumber="457">Locality and horizon.</emphasis>
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<materialsCitation id="3B190F60F236FF9AFB94FEAACCB8FE46" location="Belyi Yar" pageId="4" pageNumber="457" specimenCount="1" stateProvince="Khakasiya">
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Izykh Coal Mines, quarry no. 1, south of
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<location id="8EAE53E6F236FF9AFC07FE8FCD48FE46" LSID="urn:lsid:plazi:treatment:03D8B42BF236FF9BFF65F926CC06F7D0:8EAE53E6F236FF9AFC07FE8FCD48FE46" box="[1002,1115,375,401]" name="Belyi Yar" pageId="4" pageNumber="457" stateProvince="Khakasiya">Belyi Yar</location>
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,
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<collectingRegion id="49B5CBDFF236FF9AFB81FE8FCDF1FE46" box="[1132,1250,375,401]" country="Russia" name="Khakasiya" pageId="4" pageNumber="457">Khakassia</collectingRegion>
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, Minusinsk Coal
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</materialsCitation>
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</paragraph>
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<paragraph id="8BCE053DF236FF9AFCC7FE63CCBFFE62" blockId="4.[810,1452,191,1136]" box="[810,1452,411,437]" pageId="4" pageNumber="457">Basin, South Siberia; Izykh Formation, Middle Permian.</paragraph>
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</subSubSection>
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<subSubSection id="C36B56B6F236FF9BFCB2FE38CC06F7D0" lastPageId="5" lastPageNumber="458" pageId="4" pageNumber="457" type="description">
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<paragraph id="8BCE053DF236FF9AFCB2FE38CCB8FE0D" blockId="4.[810,1452,191,1136]" box="[863,1451,448,474]" pageId="4" pageNumber="457">
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<emphasis id="B905D92FF236FF9AFCB2FE38CAE2FE0D" bold="true" box="[863,1009,448,474]" pageId="4" pageNumber="457">Description.</emphasis>
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Wing short, broadly ovoid (L:W ratio
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</paragraph>
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<paragraph id="8BCE053DF236FF9AFCC7FE1DCCBFFD0C" blockId="4.[810,1452,191,1136]" pageId="4" pageNumber="457">
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1:2.2) with rounded apex at RS
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<subScript id="17F50778F236FF9AFB49FE0FCDBFFDD2" attach="left" box="[1188,1196,503,517]" fontSize="6" pageId="4" pageNumber="457">3</subScript>
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–RS
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<subScript id="17F50778F236FF9AFB31FE0FCDF7FDD2" attach="left" box="[1244,1252,503,517]" fontSize="6" pageId="4" pageNumber="457">4</subScript>
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; anterior margin convex at 1/3 of wing length. SC reaching 2/3 of wing length, visually ending at R with short and weak veinlet to costal margin. Besides that apical vein, there are four short subcostal crossveins at equidistance from one another. Subcostal space broad, crossed by strong sigmoid crossvein sc–r at level of R–RS forking point.
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</paragraph>
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<paragraph id="8BCE053DF236FF9AFCC7FD1ECCB8FC6F" blockId="4.[810,1452,191,1136]" pageId="4" pageNumber="457">
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Pterostigma not preserved. R curved basally, without pterostigmal branches. RS forking well before middle of wing, with five branches. Cells DC and MC open. M with four branches; M
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<subScript id="17F50778F236FF9AFC01FC9FCD15FCA2" attach="left" box="[1004,1030,871,885]" fontSize="6" pageId="4" pageNumber="457">3+4</subScript>
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forking very late, distal to M
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<subScript id="17F50778F236FF9AFAB9FC9FCC7DFCA2" attach="left" box="[1364,1390,871,885]" fontSize="6" pageId="4" pageNumber="457">1+2</subScript>
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fork. Base of CuA long, strongly oblique, M
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<subScript id="17F50778F236FF9AFB3EFC73CDC8FC4E" attach="left" box="[1235,1243,907,921]" fontSize="6" pageId="4" pageNumber="457">5</subScript>
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much shorter. CuP preserved only by its distal half after cua-cup crossvein,
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</paragraph>
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<paragraph id="8BCE053DF236FF9AFCC7FC3BCACFFBF0" blockId="4.[810,1452,191,1136]" pageId="4" pageNumber="457">
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anal area missing. Crossveins sc–r, r–rs
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<subScript id="17F50778F236FF9AFB19FC2DCDEFFC34" attach="right" box="[1268,1276,981,995]" fontSize="6" pageId="4" pageNumber="457">1</subScript>
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, m
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<subScript id="17F50778F236FF9AFACEFC2DCC2EFC34" attach="left" box="[1315,1341,981,995]" fontSize="6" pageId="4" pageNumber="457">3+4</subScript>
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–cua, and cua–cup distinct, others completely invisible, except for very faint rs–m.
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</paragraph>
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<paragraph id="8BCE053DF236FF9AFCB2FBC9CCB8FB9C" blockId="4.[810,1452,191,1136]" box="[863,1451,1073,1099]" pageId="4" pageNumber="457">
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<emphasis id="B905D92FF236FF9AFCB2FBC9CD06FB9C" bold="true" box="[863,1045,1073,1099]" pageId="4" pageNumber="457">Measurements.</emphasis>
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Forewing length,
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<quantity id="4C89A8D8F236FF9AFB18FBC9CC40FB9C" box="[1269,1363,1073,1099]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="4" pageNumber="457" unit="mm" value="3.6">3.6 mm</quantity>
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, width,
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</paragraph>
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<paragraph id="8BCE053DF236FF9AFCC7FBAECA94FBA7" blockId="4.[810,1452,191,1136]" box="[810,903,1110,1136]" pageId="4" pageNumber="457">
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<quantity id="4C89A8D8F236FF9AFCC7FBAECA90FBA7" box="[810,899,1110,1136]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="4" pageNumber="457" unit="mm" value="1.6">1.6 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BCE053DF236FF9AFCC7FB67CD50FB6E" blockId="4.[810,1452,1183,1430]" box="[810,1091,1183,1209]" pageId="4" pageNumber="457">
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<emphasis id="B905D92FF236FF9AFCC7FB67CD50FB6E" box="[810,1091,1183,1209]" italics="true" pageId="4" pageNumber="457">Other material examined</emphasis>
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</paragraph>
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<paragraph id="8BCE053DF236FF9AFCC7FB3CCCB8FA41" blockId="4.[810,1452,1183,1430]" pageId="4" pageNumber="457">
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In addition to the described species, there are two wellpreserved hindwings, PIN 5386/2± and 10± (
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<figureCitation id="134A19B8F236FF9AFAC5FB11CC7FFAD4" box="[1320,1388,1257,1283]" captionStart="FIGURE 3" captionStartId="4.[810,903,2024,2046]" captionTargetBox="[830,1438,1474,1989]" captionTargetId="figure-517@4.[809,1452,1463,2000]" captionTargetPageId="4" captionText="FIGURE 3. Kamopanorpa spp., hindwing venation. A, PIN 5386/2. B, PIN 5386/10." figureDoi="http://doi.org/10.5281/zenodo.5530155" httpUri="https://zenodo.org/record/5530155/files/figure.png" pageId="4" pageNumber="457">Fig. 3</figureCitation>
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), and also several forewings (PIN 5386/4, 6±, 7±, 8±, 11±, 12, 13±) that are too fragmentary or poorly preserved to be referred to any particular species. Some of them show additional forkings on RS and M branches, notably
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</paragraph>
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<caption id="DF0E55B5F236FF9AFCC7F810CD04F7F5" ID-DOI="http://doi.org/10.5281/zenodo.5530155" ID-Zenodo-Dep="5530155" httpUri="https://zenodo.org/record/5530155/files/figure.png" pageId="4" pageNumber="457" startId="4.[810,903,2024,2046]" targetBox="[830,1438,1474,1989]" targetPageId="4">
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<paragraph id="8BCE053DF236FF9AFCC7F810CD04F7F5" blockId="4.[810,1452,2024,2082]" pageId="4" pageNumber="457">
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<emphasis id="B905D92FF236FF9AFCC7F810CABBF82A" bold="true" box="[810,936,2024,2046]" pageId="4" pageNumber="457">FIGURE 3.</emphasis>
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<taxonomicName id="4C717EBEF236FF9AFC5AF811CD68F82A" box="[951,1147,2024,2046]" class="Insecta" family="Cladochoristidae" genus="Kamopanorpa" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="457" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B905D92FF236FF9AFC5AF811CD51F829" box="[951,1090,2025,2046]" italics="true" pageId="4" pageNumber="457">Kamopanorpa</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, hindwing venation.
|
||
<emphasis id="B905D92FF236FF9AFAB3F810CC7CF82A" bold="true" box="[1374,1391,2024,2045]" pageId="4" pageNumber="457">A</emphasis>
|
||
, PIN 5386/2.
|
||
<emphasis id="B905D92FF236FF9AFC95F7F4CA9BF7F6" bold="true" box="[888,904,2060,2081]" pageId="4" pageNumber="457">B</emphasis>
|
||
, PIN 5386/10.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0E55B5F237FF9BFF65FA2FCBE3F9E2" ID-DOI="http://doi.org/10.5281/zenodo.5530157" ID-Zenodo-Dep="5530157" httpUri="https://zenodo.org/record/5530157/files/figure.png" pageId="5" pageNumber="458" startId="5.[136,229,1495,1517]" targetBox="[215,1372,190,1471]" targetPageId="5">
|
||
<paragraph id="8BCE053DF237FF9BFF65FA2FCBBDF9E2" blockId="5.[136,1452,1495,1589]" pageId="5" pageNumber="458">
|
||
<emphasis id="B905D92FF237FF9BFF65FA2FC9EDFA3B" bold="true" box="[136,254,1495,1517]" pageId="5" pageNumber="458">FIGURE 4.</emphasis>
|
||
Details of the hindwings.
|
||
<emphasis id="B905D92FF237FF9BFE16FA2FCB3AFA3A" bold="true" box="[507,553,1495,1517]" pageId="5" pageNumber="458">A–C</emphasis>
|
||
,
|
||
<taxonomicName id="4C717EBEF237FF9BFDD8FA20CBF7FA3A" box="[565,740,1496,1517]" class="Insecta" family="Cladochoristidae" genus="Kamopanorpa" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B905D92FF237FF9BFDD8FA20CBD3FA3A" box="[565,704,1496,1517]" italics="true" pageId="5" pageNumber="458">Kamopanorpa</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, PIN 5386/10 (mirror images).
|
||
<emphasis id="B905D92FF237FF9BFBFFFA20CD31FA3A" bold="true" box="[1042,1058,1496,1517]" pageId="5" pageNumber="458">B</emphasis>
|
||
, Bases of setae on SC.
|
||
<emphasis id="B905D92FF237FF9BFAEEFA20CC07FA3A" bold="true" box="[1283,1300,1496,1517]" pageId="5" pageNumber="458">D</emphasis>
|
||
,
|
||
<taxonomicName id="4C717EBEF237FF9BFACCFA20C8BCF9C6" authority="(Sukatsheva, 1976)" baseAuthorityName="Sukatsheva" baseAuthorityYear="1976" class="Insecta" family="Cladochoristidae" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="pritykinae">
|
||
<emphasis id="B905D92FF237FF9BFACCFA20C9FAF9C7" italics="true" pageId="5" pageNumber="458">Kamopanorpa pritykinae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE078CCF237FF9BFF15FA03C8BBF9C6" author="Sukatsheva, I. D." box="[248,424,1531,1553]" pageId="5" pageNumber="458" pagination="198 - 209" refId="ref6016" refString="Sukatsheva, I. D. (1976) Caddisflies of the suborder Permotrichoptera. Paleontological Journal, 10 (2), 198 - 209." type="journal article" year="1976">Sukatsheva, 1976</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, paratype PIN 2781/265.
|
||
<emphasis id="B905D92FF237FF9BFD43FA04CBADF9C6" bold="true" box="[686,702,1532,1553]" pageId="5" pageNumber="458">E</emphasis>
|
||
and
|
||
<emphasis id="B905D92FF237FF9BFD1CFA04CA13F9C6" bold="true" box="[753,768,1532,1553]" pageId="5" pageNumber="458">F</emphasis>
|
||
,
|
||
<taxonomicName id="4C717EBEF237FF9BFCE0FA04CAADF9C6" box="[781,958,1532,1553]" class="Insecta" family="Cladochoristidae" genus="Kamopanorpa" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B905D92FF237FF9BFCE0FA04CA8BF9C6" box="[781,920,1532,1553]" italics="true" pageId="5" pageNumber="458">Kamopanorpa</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
, PIN 5386/2. Arrows indicate putative position of crossveins on
|
||
<emphasis id="B905D92FF237FF9BFEFDF9E7C832F9E2" bold="true" box="[272,289,1567,1589]" pageId="5" pageNumber="458">C</emphasis>
|
||
and
|
||
<emphasis id="B905D92FF237FF9BFEBDF9D8C84CF9E2" bold="true" box="[336,351,1568,1589]" pageId="5" pageNumber="458">F</emphasis>
|
||
, and unsclerotized apical part of A
|
||
</paragraph>
|
||
<paragraph id="8BCE053DF237FF9BFD56F9D8CBE3F9E2" blockId="5.[136,1452,1495,1589]" box="[699,752,1568,1589]" pageId="5" pageNumber="458">
|
||
on
|
||
<emphasis id="B905D92FF237FF9BFD34F9D8CBF9F9E2" bold="true" box="[729,746,1568,1589]" pageId="5" pageNumber="458">D</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCE053DF237FF9BFD43F9D7CBA6F9EC" box="[686,693,1583,1595]" pageId="5" pageNumber="458">2</paragraph>
|
||
<paragraph id="8BCE053DF237FF9BFF65F981CB6CF90B" blockId="5.[136,778,1657,2080]" pageId="5" pageNumber="458">
|
||
5386/8± (
|
||
<figureCitation id="134A19B8F237FF9BFF18F981C859F943" box="[245,330,1657,1684]" captionStart="FIGURE 1" captionStartId="3.[136,229,1027,1049]" captionTargetBox="[222,1353,246,987]" captionTargetId="figure-428@3.[215,1371,223,1003]" captionTargetPageId="3" captionText="FIGURE 1. A, B, Kamopanorpa rasnitsyni sp. nov. A, Holotype PIN 5386/9. B, Paratype PIN 5386/3. C, K. rotunda sp. nov., holotype PIN 5386/5. D, K. sivchikovi sp. nov., holotype PIN 5386/1. E, F, Kamopanorpa spp. E, 5386/8. F, 5386/11.All to the same scale." figureDoi="http://doi.org/10.5281/zenodo.5530151" httpUri="https://zenodo.org/record/5530151/files/figure.png" pageId="5" pageNumber="458">Fig. 1E</figureCitation>
|
||
), which has RS with 8 branches in total with deep fork on RS
|
||
<subScript id="17F50778F237FF9BFE6DF948C89BF969" attach="left" box="[384,392,1712,1726]" fontSize="6" pageId="5" pageNumber="458">4</subScript>
|
||
(the latter is only rarely found in
|
||
<taxonomicName id="4C717EBEF237FF9BFF65F93ACB61F90B" authority=": Novokshonov, 1992" authorityName=": Novokshonov" authorityYear="1992" box="[136,626,1729,1756]" class="Insecta" family="Cladochoristidae" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B905D92FF237FF9BFF65F93AC839F90C" box="[136,298,1730,1755]" italics="true" pageId="5" pageNumber="458">Kamopanorpa</emphasis>
|
||
:
|
||
<bibRefCitation id="EFE078CCF237FF9BFED4F939CB0EF90B" author="Novokshonov, V. G." box="[313,541,1729,1756]" pageId="5" pageNumber="458" pagination="136 - 141" refId="ref5823" refString="Novokshonov, V. G. (1992) Caddisflies of the genus Kamopanorpa (Trichoptera, Microptysmatidae) from the Kungurian of Chekarda (Perm District). Paleontological Journal, 26 (3), 136 - 141." type="journal article" year="1992">Novokshonov, 1992</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFE078CCF237FF9BFDC5F939CB61F90B" author="Novokshonov, V. G." box="[552,626,1729,1756]" pageId="5" pageNumber="458" pagination="90 - 101" refId="ref5895" refString="Novokshonov, V. G. (1993 b) Caddis flies (Insecta: Trichoptera: Microptysmatidae). Paleontological Journal, 27 (1 A), 90 - 101." type="journal article" year="1993">1993b</bibRefCitation>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCE053DF237FF9BFF50F91ECDD6F8D3" blockId="5.[136,778,1657,2080]" lastBlockId="5.[810,1452,1657,2056]" pageId="5" pageNumber="458">
|
||
In microptysmatid hindwings, as was shown by
|
||
<bibRefCitation id="EFE078CCF237FF9BFF65F8F1C8B6F8F3" author="Novokshonov, V. G." box="[136,421,1801,1828]" pageId="5" pageNumber="458" refId="ref5928" refString="Novokshonov, V. G. (1997) Rannyaya evolyutsiya skorpionnits (Insecta: Panorpida) [Early evolution of the scorpion flies (Insecta: Panorpida)]. Moscow, Nauka Press, 140 pp." type="book" year="1997">Novokshonov (1997: 94)</bibRefCitation>
|
||
, the anal veins have a tendency to looping, changing from freely reaching wing margin in Early Permian forms to coalesced apically and forming distinct anal loop in the Late Permian ones.Re-examination of the hindwing of
|
||
<taxonomicName id="4C717EBEF237FF9BFE88F862CBCFF863" authority="(Sukatsheva, 1976)" baseAuthorityName="Sukatsheva" baseAuthorityYear="1976" box="[357,732,1945,1972]" class="Insecta" family="Cladochoristidae" genus="Kamopanorpa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="458" phylum="Arthropoda" rank="species" species="pritykinae">
|
||
<emphasis id="B905D92FF237FF9BFE88F862C8E9F864" box="[357,506,1945,1971]" italics="true" pageId="5" pageNumber="458">K. pritykinae</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE078CCF237FF9BFDE4F861CBC0F863" author="Sukatsheva, I. D." box="[521,723,1945,1972]" pageId="5" pageNumber="458" pagination="198 - 209" refId="ref6016" refString="Sukatsheva, I. D. (1976) Caddisflies of the suborder Permotrichoptera. Paleontological Journal, 10 (2), 198 - 209." type="journal article" year="1976">Sukatsheva, 1976</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
has shown that A
|
||
<subScript id="17F50778F237FF9BFEF7F828C820F809" attach="left" box="[282,307,2000,2014]" fontSize="6" pageId="5" pageNumber="458">2+3</subScript>
|
||
does not just end blindly in membrane but its unsclerotized apical portion terminates on A
|
||
<subScript id="17F50778F237FF9BFD62F80CCB84F7D5" attach="left" box="[655,663,2036,2050]" fontSize="6" pageId="5" pageNumber="458">1</subScript>
|
||
(
|
||
<figureCitation id="134A19B8F237FF9BFD49F819CBEDF82B" box="[676,766,2017,2044]" captionStart="FIGURE 4" captionStartId="5.[136,229,1495,1517]" captionTargetBox="[215,1372,190,1471]" captionTargetId="figure-232@5.[215,1372,190,1471]" captionTargetPageId="5" captionText="FIGURE 4. Details of the hindwings. A–C, Kamopanorpa sp., PIN 5386/10 (mirror images). B, Bases of setae on SC. D, Kamopanorpa pritykinae (Sukatsheva, 1976), paratype PIN 2781/265. E and F, Kamopanorpa sp., PIN 5386/2. Arrows indicate putative position of crossveins on C and F, and unsclerotized apical part of A on D." figureDoi="http://doi.org/10.5281/zenodo.5530157" httpUri="https://zenodo.org/record/5530157/files/figure.png" pageId="5" pageNumber="458">Fig. 4D</figureCitation>
|
||
). The new specimens show an arrangement of anal veins of transitional nature: the A
|
||
<subScript id="17F50778F237FF9BFBD2F973CD54F94E" attach="left" box="[1087,1095,1675,1689]" fontSize="6" pageId="5" pageNumber="458">2</subScript>
|
||
and A
|
||
<subScript id="17F50778F237FF9BFB7DF973CD8BF94E" attach="left" box="[1168,1176,1675,1689]" fontSize="6" pageId="5" pageNumber="458">3</subScript>
|
||
meet inner wing margin and each is distally connected to the next anal by a very weak curved crossvein. Apical part of A
|
||
<subScript id="17F50778F237FF9BFB00F92DCDE6F934" attach="left" box="[1261,1269,1749,1763]" fontSize="6" pageId="5" pageNumber="458">3</subScript>
|
||
is unsclerotized in both specimens (
|
||
<figureCitation id="134A19B8F237FF9BFBEEF911CD59F8D3" box="[1027,1098,1769,1796]" captionStart="FIGURE 3" captionStartId="4.[810,903,2024,2046]" captionTargetBox="[830,1438,1474,1989]" captionTargetId="figure-517@4.[809,1452,1463,2000]" captionTargetPageId="4" captionText="FIGURE 3. Kamopanorpa spp., hindwing venation. A, PIN 5386/2. B, PIN 5386/10." figureDoi="http://doi.org/10.5281/zenodo.5530155" httpUri="https://zenodo.org/record/5530155/files/figure.png" pageId="5" pageNumber="458">Figs 3</figureCitation>
|
||
,
|
||
<figureCitation id="134A19B8F237FF9BFBBAF912CDABF8D3" box="[1111,1208,1770,1796]" captionStart="FIGURE 4" captionStartId="5.[136,229,1495,1517]" captionTargetBox="[215,1372,190,1471]" captionTargetId="figure-232@5.[215,1372,190,1471]" captionTargetPageId="5" captionText="FIGURE 4. Details of the hindwings. A–C, Kamopanorpa sp., PIN 5386/10 (mirror images). B, Bases of setae on SC. D, Kamopanorpa pritykinae (Sukatsheva, 1976), paratype PIN 2781/265. E and F, Kamopanorpa sp., PIN 5386/2. Arrows indicate putative position of crossveins on C and F, and unsclerotized apical part of A on D." figureDoi="http://doi.org/10.5281/zenodo.5530157" httpUri="https://zenodo.org/record/5530157/files/figure.png" pageId="5" pageNumber="458">4B, E, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BCE053DF237FF9BFCB2F8F7CC06F7D0" blockId="5.[810,1452,1657,2056]" pageId="5" pageNumber="458">
|
||
An interesting detail was observed in specimen 5386/10±. On the posterior side of subcosta along its basal part up to the first crossvein, there is a dense row of sockets (bases of setae), directed basad and upwards, which in all probability was a part of a wing coupling apparatus (the first direct evidence for wing coupling in Palaeozoic amphiesmenopterans) (
|
||
<figureCitation id="134A19B8F237FF9BFB42F815CC1BF7D0" box="[1199,1288,2029,2055]" captionStart="FIGURE 4" captionStartId="5.[136,229,1495,1517]" captionTargetBox="[215,1372,190,1471]" captionTargetId="figure-232@5.[215,1372,190,1471]" captionTargetPageId="5" captionText="FIGURE 4. Details of the hindwings. A–C, Kamopanorpa sp., PIN 5386/10 (mirror images). B, Bases of setae on SC. D, Kamopanorpa pritykinae (Sukatsheva, 1976), paratype PIN 2781/265. E and F, Kamopanorpa sp., PIN 5386/2. Arrows indicate putative position of crossveins on C and F, and unsclerotized apical part of A on D." figureDoi="http://doi.org/10.5281/zenodo.5530157" httpUri="https://zenodo.org/record/5530157/files/figure.png" pageId="5" pageNumber="458">Fig. 4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |