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<document id="310723083A8E12781EFB7CAFCAABA139" ID-DOI="10.5281/zenodo.274647" ID-GBIF-Dataset="af5f02d9-635b-453c-8706-1557b7d45435" ID-ISSN="1175-5326" ID-Zenodo-Dep="274647" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459985719703" checkinUser="plazi" docAuthor="Flores, Gustavo E." docDate="2009" docId="03D5ED19FFC8E13B1494BFB1FBD5FF5D" docLanguage="en" docName="zt01985p033.pdf" docOrigin="Zootaxa 1985" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Emmallodera obesa subsp. costata Kulzer" docType="treatment" docVersion="7" lastPageNumber="29" masterDocId="FFEC9561FFC0E1321403BA1CFFE9FF85" masterDocTitle="Revision of some types of the South American tribes Nycteliini, Praocini, and Scotobiini (Coleoptera: Tenebrionidae), with new synonymies" masterLastPageNumber="33" masterPageNumber="21" pageNumber="29" updateTime="1698232258373" updateUser="plazi">
<mods:mods id="6C408292DEA7D8165B5078D92221C498" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AAC8049DC8221115E560419DFA073D4F">
<mods:title id="446E35DA892F2677F2453604E74B104D">Revision of some types of the South American tribes Nycteliini, Praocini, and Scotobiini (Coleoptera: Tenebrionidae), with new synonymies</mods:title>
</mods:titleInfo>
<mods:name id="F0F834F77FBAE34DE6FC9C9C9A9D6AD7" type="personal">
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<mods:roleTerm id="69D9F1244C863E1B6C43C8DDCEE9D71A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="142E8D9F64F71E7252006AFE94012BA2">Flores, Gustavo E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="FE2616ADDA74787B6B286CD7EEA5843A">text</mods:typeOfResource>
<mods:relatedItem id="512DFC8396E0988DD7CC2C4591658BD3" type="host">
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<mods:title id="44BBB565EB76B02111CA32E9593F7589">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="7596CA7D3BD58982C9523DAE8FC71BC4">
<mods:date id="DC345E0F27CDEAC99ACD5AA54680A8F5">2009</mods:date>
<mods:detail id="091445210F02AEE1AF5044A635BFBCB9" type="volume">
<mods:number id="37374DB5B3EBC7778110E6F8049FCB54">1985</mods:number>
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<mods:classification id="EF78B7D0B441ECCDEE4C88DDF5DBB401">journal article</mods:classification>
<mods:identifier id="AC32C5853554B90B4206EB1838E56147" type="DOI">10.5281/zenodo.274647</mods:identifier>
<mods:identifier id="81F189A72587CED83F9835FC540F02F7" type="GBIF-Dataset">af5f02d9-635b-453c-8706-1557b7d45435</mods:identifier>
<mods:identifier id="049DAF39E7D379DB96E5782794A3ED29" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B6E6487B907651025D8010BDBADCCC80" type="Zenodo-Dep">274647</mods:identifier>
</mods:mods>
<treatment id="03D5ED19FFC8E13B1494BFB1FBD5FF5D" ID-DOI="http://doi.org/10.5281/zenodo.6218524" ID-GBIF-Taxon="119379225" ID-Zenodo-Dep="6218524" LSID="urn:lsid:plazi:treatment:03D5ED19FFC8E13B1494BFB1FBD5FF5D" httpUri="http://treatment.plazi.org/id/03D5ED19FFC8E13B1494BFB1FBD5FF5D" lastPageId="9" lastPageNumber="29" pageId="8" pageNumber="29">
<subSubSection id="C3660F84FFC8E13A1494BFB1FEDDFA6F" pageId="8" pageNumber="29" type="nomenclature">
<paragraph id="8BC35C0FFFC8E13A1494BFB1FDD6FA42" blockId="8.[151,575,1453,1514]" box="[151,575,1453,1479]" pageId="8" pageNumber="29">
<heading id="D08BEB63FFC8E13A1494BFB1FDD6FA42" bold="true" box="[151,575,1453,1479]" fontSize="11" level="1" pageId="8" pageNumber="29" reason="1">
<taxonomicName id="4C7C278CFFC8E13A1494BFB1FDD6FA42" authority="Kulzer" authorityName="Kulzer" box="[151,575,1453,1479]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="obesa" subSpecies="costata">
<emphasis id="B908801DFFC8E13A1494BFB1FDD6FA42" bold="true" box="[151,575,1453,1479]" pageId="8" pageNumber="29">
<emphasis id="B908801DFFC8E13A1494BFB1FE0BFA42" bold="true" box="[151,482,1453,1479]" italics="true" pageId="8" pageNumber="29">Emmallodera obesa costata</emphasis>
Kulzer
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC35C0FFFC8E13A1494BFCCFEDDFA6F" blockId="8.[151,575,1453,1514]" box="[151,308,1488,1514]" pageId="8" pageNumber="29">
(
<figureCitation id="1347408AFFC8E13A149CBFCCFEC2FA6F" box="[159,299,1488,1514]" captionStart="FIGURES 16 – 17. 16" captionStartId="9.[151,267,1122,1146]" captionTargetBox="[164,1420,265,1090]" captionTargetId="figure@9.[164,807,250,1090]" captionTargetPageId="9" captionText="FIGURES 16 – 17. 16. Emmallodera obesa costata, paratype; 17. E. ceii, holotype." httpUri="https://zenodo.org/record/274654/files/figure.png" pageId="8" pageNumber="29">Figs. 16–17</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3660F84FFC8E13A1494BC01FCC1F9D1" pageId="8" pageNumber="29" type="reference_group">
<paragraph id="8BC35C0FFFC8E13A1494BC01FD66F9B1" blockId="8.[151,808,1565,1620]" box="[151,655,1565,1588]" pageId="8" pageNumber="29">
<treatmentCitationGroup id="AB6C7B21FFC8E13A1494BC01FD66F9B1" box="[151,655,1565,1588]" pageId="8" pageNumber="29">
<taxonomicName id="4C7C278CFFC8E13A1494BC01FD63F9B1" authority="Kulzer, 1955: 463" authorityName="Kulzer" authorityPageNumber="463" authorityYear="1955" box="[151,650,1565,1588]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="obesa" subSpecies="costata">
<emphasis id="B908801DFFC8E13A1494BC01FE28F9B1" box="[151,449,1565,1588]" italics="true" pageId="8" pageNumber="29">Emmallodera obesa costata</emphasis>
<treatmentCitation id="0ADD7A1EFFC8E13A15C4BC01FD63F9B1" author="Kulzer" box="[455,650,1565,1588]" page="463" pageId="8" pageNumber="29" year="1955">
<bibRefCitation id="EFED21FEFFC8E13A15C4BC01FDBCF9B1" author="Kulzer" box="[455,597,1565,1588]" pageId="8" pageNumber="33" refString="Kulzer, H. (1955) Monographie der Scotobiini (Zehnter Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum Georg Frey, 6, 383 - 478." type="journal article" year="1955">Kulzer, 1955</bibRefCitation>
: 463
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC35C0FFFC8E13A1494BC21FCC1F9D1" blockId="8.[151,808,1565,1620]" box="[151,808,1597,1620]" pageId="8" pageNumber="29">
<treatmentCitationGroup id="AB6C7B21FFC8E13A1494BC21FCC1F9D1" box="[151,808,1597,1620]" pageId="8" pageNumber="29">
<taxonomicName id="4C7C278CFFC8E13A1494BC21FDD5F9D1" authority="Marcuzzi, 1977: 149" authorityName="Marcuzzi" authorityPageNumber="149" authorityYear="1977" box="[151,572,1597,1620]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="ceii">
<emphasis id="B908801DFFC8E13A1494BC21FEBCF9D1" box="[151,341,1597,1620]" italics="true" pageId="8" pageNumber="29">Emmallodera ceii</emphasis>
<treatmentCitation id="0ADD7A1EFFC8E13A155FBC21FDD5F9D1" author="Marcuzzi" box="[348,572,1597,1620]" page="149" pageId="8" pageNumber="29" year="1977">
<bibRefCitation id="EFED21FEFFC8E13A155FBC21FDEFF9D1" author="Marcuzzi" box="[348,518,1597,1620]" pageId="8" pageNumber="33" refString="Marcuzzi, G. (1977) Nuove specie di tenebrionidi dell'Argentina (Coleoptera). Memorie della Societa Entomologica Italiana, 55, 139 - 152 (1976)." type="journal article" year="1977">Marcuzzi, 1977</bibRefCitation>
: 149
</treatmentCitation>
</taxonomicName>
. NEW SYNONYMY.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C3660F84FFC8E13A1494BC9FFE21F8BD" pageId="8" pageNumber="29" type="materials_examined">
<paragraph id="8BC35C0FFFC8E13A1494BC9FFA75F941" blockId="8.[151,1437,1667,2003]" pageId="8" pageNumber="29">
<emphasis id="B908801DFFC8E13A1494BC9FFEA4F918" bold="true" box="[151,333,1667,1693]" pageId="8" pageNumber="29">
<typeStatus id="54C7E2ADFFC8E13A1494BC9FFF3BF918" box="[151,210,1667,1693]" pageId="8" pageNumber="29">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54C7E2ADFFC8E13A1556BC98FE52F91B" box="[341,443,1668,1694]" pageId="8" pageNumber="29" type="paratype">Paratype</typeStatus>
of
<taxonomicName id="4C7C278CFFC8E13A15E0BC98FCC3F918" box="[483,810,1668,1693]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="obesa" subSpecies="costata">
<emphasis id="B908801DFFC8E13A15E0BC98FCC3F918" box="[483,810,1668,1693]" italics="true" pageId="8" pageNumber="29">Emmallodera obesa costata</emphasis>
</taxonomicName>
(
<figureCitation id="1347408AFFC8E13A173FBC98FC78F91B" box="[828,913,1668,1694]" captionStart="FIGURES 16 – 17. 16" captionStartId="9.[151,267,1122,1146]" captionTargetBox="[164,1420,265,1090]" captionTargetId="figure@9.[164,807,250,1090]" captionTargetPageId="9" captionText="FIGURES 16 – 17. 16. Emmallodera obesa costata, paratype; 17. E. ceii, holotype." httpUri="https://zenodo.org/record/274654/files/figure.png" pageId="8" pageNumber="29">Fig. 16</figureCitation>
): [Junín (de los Andes, sic)] [Rep.
<collectingCountry id="F36B1C9FFFC8E13A1120BC98FA7DF91B" box="[1315,1428,1668,1694]" name="Argentina" pageId="8" pageNumber="29">Argentina</collectingCountry>
/ Prov. Neuquén/ H. Richter] [
<typeStatus id="54C7E2ADFFC8E13A15DFBCB6FDA0F941" box="[476,585,1706,1732]" pageId="8" pageNumber="29" type="paratype">Paratype</typeStatus>
] [
<taxonomicName id="4C7C278CFFC8E13A1663BCB7FD16F941" box="[608,767,1707,1732]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B908801DFFC8E13A1663BCB7FD16F941" box="[608,767,1707,1732]" italics="true" pageId="8" pageNumber="29">Emmallodera</emphasis>
</taxonomicName>
/
<taxonomicName id="4C7C278CFFC8E13A170DBCB7FCBBF941" box="[782,850,1707,1732]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="obesa">
<emphasis id="B908801DFFC8E13A170DBCB7FCBBF941" box="[782,850,1707,1732]" italics="true" pageId="8" pageNumber="29">obesa</emphasis>
</taxonomicName>
ssp./
<taxonomicName id="4C7C278CFFC8E13A1795BCB7FBC2F941" authority="Klzr." authorityName="Klzr." box="[918,1067,1706,1732]" class="Insecta" family="Tenebrionidae" genus="Praocis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="costata">
<emphasis id="B908801DFFC8E13A1795BCB7FC03F941" box="[918,1002,1707,1732]" italics="true" pageId="8" pageNumber="29">costata</emphasis>
Klzr.
</taxonomicName>
/ det.
<bibRefCitation id="EFED21FEFFC8E13A106FBCB6FACDF941" author="Kulzer" box="[1132,1316,1706,1732]" pageId="8" pageNumber="33" refString="Kulzer, H. (1955) Monographie der Scotobiini (Zehnter Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum Georg Frey, 6, 383 - 478." type="journal article" year="1955">H. Kulzer 1955</bibRefCitation>
] (
<collectionCode id="ED6DC4CAFFC8E13A1139BCB6FA79F941" box="[1338,1424,1706,1732]" pageId="8" pageNumber="29">MLPA</collectionCode>
).
</paragraph>
<paragraph id="8BC35C0FFFC8E13A14C5BCCDFE21F8BD" blockId="8.[151,1437,1667,2003]" pageId="8" pageNumber="29">
<typeStatus id="54C7E2ADFFC8E13A14C5BCCDFEDAF96E" box="[198,307,1745,1771]" pageId="8" pageNumber="29" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C7C278CFFC8E13A156FBCCDFDAFF96F" box="[364,582,1745,1770]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="ceii">
<emphasis id="B908801DFFC8E13A156FBCCDFDAFF96F" box="[364,582,1745,1770]" italics="true" pageId="8" pageNumber="29">Emmallodera ceii</emphasis>
</taxonomicName>
(
<figureCitation id="1347408AFFC8E13A165CBCCDFD54F96E" box="[607,701,1745,1771]" captionStart="FIGURES 16 – 17. 16" captionStartId="9.[151,267,1122,1146]" captionTargetBox="[164,1420,265,1090]" captionTargetId="figure@9.[164,807,250,1090]" captionTargetPageId="9" captionText="FIGURES 16 – 17. 16. Emmallodera obesa costata, paratype; 17. E. ceii, holotype." httpUri="https://zenodo.org/record/274654/files/figure.png" pageId="8" pageNumber="29">Fig. 17</figureCitation>
): [Neuquén/ Meliquina/
<date id="FFC27ACFFFC8E13A17F2BCCDFB8DF96E" box="[1009,1124,1745,1771]" pageId="8" pageNumber="29" value="1963-12-21">21-XII-63</date>
(pro 1963)] [
<typeStatus id="54C7E2ADFFC8E13A1116BCCDFA7CF96E" box="[1301,1429,1745,1771]" pageId="8" pageNumber="29" type="holotype">Holotypus</typeStatus>
] [
<taxonomicName id="4C7C278CFFC8E13A14A3BCE4FED6F894" box="[160,319,1784,1809]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B908801DFFC8E13A14A3BCE4FED6F894" box="[160,319,1784,1809]" italics="true" pageId="8" pageNumber="29">Emmallodera</emphasis>
</taxonomicName>
/
<taxonomicName id="4C7C278CFFC8E13A1553BCE4FE92F894" box="[336,379,1784,1809]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="ceii">
<emphasis id="B908801DFFC8E13A1553BCE4FE92F894" box="[336,379,1784,1809]" italics="true" pageId="8" pageNumber="29">ceii</emphasis>
</taxonomicName>
mihi] [
<typeStatus id="54C7E2ADFFC8E13A15CEBCE4FDA0F897" box="[461,585,1784,1810]" pageId="8" pageNumber="29" type="holotype">Holotypus</typeStatus>
/
<taxonomicName id="4C7C278CFFC8E13A1658BCE4FD13F894" box="[603,762,1784,1809]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B908801DFFC8E13A1658BCE4FD13F894" box="[603,762,1784,1809]" italics="true" pageId="8" pageNumber="29">Emmallodera</emphasis>
</taxonomicName>
/
<taxonomicName id="4C7C278CFFC8E13A1708BCE4FCDCF894" box="[779,821,1784,1809]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="ceii">
<emphasis id="B908801DFFC8E13A1708BCE4FCDCF894" box="[779,821,1784,1809]" italics="true" pageId="8" pageNumber="29">ceii</emphasis>
</taxonomicName>
/
<bibRefCitation id="EFED21FEFFC8E13A1747BCE4FC12F897" author="Marcuzzi" box="[836,1019,1784,1810]" pageId="8" pageNumber="33" refString="Marcuzzi, G. (1977) Nuove specie di tenebrionidi dell'Argentina (Coleoptera). Memorie della Societa Entomologica Italiana, 55, 139 - 152 (1976)." type="journal article" year="1977">Marcuzzi, 1977</bibRefCitation>
] [Museo Genova/ coll. G. Marcuzzi/ (acquisto 2000)] (
<collectionCode id="ED6DC4CAFFC8E13A1562BD02FE55F8BD" box="[353,444,1822,1848]" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="8" pageNumber="29">MSNG</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C3660F84FFC8E13B14C5BD59FBD5FF5D" lastPageId="9" lastPageNumber="30" pageId="8" pageNumber="29" type="discussion">
<paragraph id="8BC35C0FFFC8E13B14C5BD59FBD5FF5D" blockId="8.[151,1437,1667,2003]" lastBlockId="9.[151,1436,152,216]" lastPageId="9" lastPageNumber="30" pageId="8" pageNumber="29">
<emphasis id="B908801DFFC8E13A14C5BD59FEBBF8DA" bold="true" box="[198,338,1861,1887]" pageId="8" pageNumber="29">Discussion.</emphasis>
The
<typeStatus id="54C7E2ADFFC8E13A1596BD59FE2EF8DA" box="[405,455,1861,1887]" pageId="8" pageNumber="29">type</typeStatus>
specimens of both nominal species present the same characters given by
<bibRefCitation id="EFED21FEFFC8E13A114FBD59FF02F803" author="Kulzer" pageId="8" pageNumber="33" refString="Kulzer, H. (1955) Monographie der Scotobiini (Zehnter Beitrag zur Kenntnis der Tenebrioniden). Entomologische Arbeiten aus dem Museum Georg Frey, 6, 383 - 478." type="journal article" year="1955">Kulzer (1955)</bibRefCitation>
in his key of
<taxonomicName id="4C7C278CFFC8E13A159EBD70FDAEF800" box="[413,583,1900,1925]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="genus">
<emphasis id="B908801DFFC8E13A159EBD70FDAEF800" box="[413,583,1900,1925]" italics="true" pageId="8" pageNumber="29">Emmallodera</emphasis>
</taxonomicName>
and description of
<taxonomicName id="4C7C278CFFC8E13A1743BD70FBFBF800" box="[832,1042,1900,1925]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="obesa" subSpecies="costata">
<emphasis id="B908801DFFC8E13A1743BD70FBFBF800" box="[832,1042,1900,1925]" italics="true" pageId="8" pageNumber="29">E. obesa costata</emphasis>
</taxonomicName>
and by
<bibRefCitation id="EFED21FEFFC8E13A1079BD70FAA4F803" author="Marcuzzi" box="[1146,1357,1900,1926]" pageId="8" pageNumber="33" refString="Marcuzzi, G. (1977) Nuove specie di tenebrionidi dell'Argentina (Coleoptera). Memorie della Societa Entomologica Italiana, 55, 139 - 152 (1976)." type="journal article" year="1977">Marcuzzi (1977)</bibRefCitation>
in his description of
<taxonomicName id="4C7C278CFFC8E13A1542BD8FFE64F829" box="[321,397,1939,1964]" class="Insecta" family="Tenebrionidae" genus="Emmallodera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="29" phylum="Arthropoda" rank="species" species="ceii">
<emphasis id="B908801DFFC8E13A1542BD8FFE64F829" box="[321,397,1939,1964]" italics="true" pageId="8" pageNumber="29">E. ceii</emphasis>
</taxonomicName>
(
<figureCitation id="1347408AFFC8E13A159DBD8EFDC6F829" box="[414,559,1938,1964]" captionStart="FIGURES 16 – 17. 16" captionStartId="9.[151,267,1122,1146]" captionTargetBox="[164,1420,265,1090]" captionTargetId="figure@9.[164,807,250,1090]" captionTargetPageId="9" captionText="FIGURES 16 – 17. 16. Emmallodera obesa costata, paratype; 17. E. ceii, holotype." httpUri="https://zenodo.org/record/274654/files/figure.png" pageId="8" pageNumber="29">Figs. 16–17</figureCitation>
): pronotum with posterior margin same width than anterior margin, lateral margins feebly raised, anterior angles and posterior angles obtuse; each elytron with six carinae on dorsal surface (the seventh carina represents the lateral margin) formed by protuberances in alignment and fused, the three external carinae more evident, intercostal spaces concave, with punctures.
</paragraph>
</subSubSection>
</treatment>
</document>
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