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<document id="E6B0376E1D22057482194ABBF8335641" ID-DOI="10.5281/zenodo.156909" ID-GBIF-Dataset="96f2b8e1-0b79-4d49-bce9-662141a937ca" ID-ISSN="1175­5326" ID-Zenodo-Dep="156909" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459607264213" checkinUser="plazi" docAuthor="Brown, John W. &amp; Adamski, David" docDate="2003" docId="03C7AC266A20460CFE92753EFDFA0F36" docLanguage="en" docName="zt00200.pdf" docOrigin="Zootaxa 200" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Anopinella styraxivora Brown and Adamski, new species" docType="treatment" docVersion="11" lastPageNumber="48" masterDocId="FFFED45E6A0E4623FF9A7070FFF7093E" masterDocTitle="Systematic revision of Anopinella Powell (Lepidoptera: Tortricidae: Euliini) and phylogenetic analysis of the Apolychrosis group of genera" masterLastPageNumber="94" masterPageNumber="1" pageNumber="47" updateTime="1698203016486" updateUser="plazi">
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<mods:title id="AA875B2A9DDBC41BB5492CCD3D99AB4B">Systematic revision of Anopinella Powell (Lepidoptera: Tortricidae: Euliini) and phylogenetic analysis of the Apolychrosis group of genera</mods:title>
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<mods:namePart id="0A6F12E6D1DD0AECD6934D23047CC283">Brown, John W.</mods:namePart>
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<mods:namePart id="CAAF59EBE94073EE6B4C2354314799F2">Adamski, David</mods:namePart>
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<paragraph id="8BD11D306A20460DFE92753EFC330C56" blockId="46.[264,964,1358,1419]" box="[264,964,1358,1384]" pageId="46" pageNumber="47">
<heading id="D099AA5C6A20460DFE92753EFC330C56" bold="true" box="[264,964,1358,1384]" fontSize="11" level="1" pageId="46" pageNumber="47" reason="1">
<emphasis id="B91AC1226A20460DFE92753EFC330C56" bold="true" box="[264,964,1358,1384]" pageId="46" pageNumber="47">
<taxonomicName id="4C6E66B36A20460DFE92753EFCD00C56" ID-CoL="95WDX" authority="Brown and Adamski" authorityName="Brown and Adamski" box="[264,807,1358,1384]" class="Insecta" family="Tortricidae" genus="Anopinella" kingdom="Animalia" order="Lepidoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="styraxivora" status="sp. nov.">
<emphasis id="B91AC1226A20460DFE92753EFDEB0C56" bold="true" box="[264,540,1358,1384]" italics="true" pageId="46" pageNumber="47">Anopinella styraxivora</emphasis>
Brown and Adamski
</taxonomicName>
,
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</emphasis>
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<paragraph id="8BD11D306A20460DFE927501FE490CB5" blockId="46.[264,964,1358,1419]" box="[264,446,1393,1419]" pageId="46" pageNumber="47">
<figureCitation id="135501B56A20460DFE927501FE9E0CB5" box="[264,361,1393,1419]" captionStart="FIGURES 33" captionStartId="74.[264,382,1856,1880]" captionText="FIGURES 33. Male genitalia of Anopinella, valvae spread, aedeagus removed. 33, A. styraxivora." pageId="46" pageNumber="47">Figs. 33</figureCitation>
,
<figureCitation id="135501B56A20460DFEEF7501FE620CB5" box="[373,405,1393,1419]" captionStart="FIGURES 54 ­ 55" captionStartId="85.[264,382,1860,1884]" captionTargetBox="[308,1271,432,1742]" captionTargetId="figure@85.[265,1322,361,1802]" captionTargetPageId="85" captionText="FIGURES 54 ­ 55. Female genitalia of Anopinella, posterior end up. 54, A. rica; 55, A. styraxivora." httpUri="https://zenodo.org/record/156935/files/figure.png" pageId="46" pageNumber="47">55</figureCitation>
,
<figureCitation id="135501B56A20460DFE3B7501FE490CB5" box="[417,446,1393,1419]" captionStart="FIGURES 68 ­ 73" captionStartId="88.[264,382,1784,1808]" captionTargetBox="[270,1316,310,1741]" captionTargetId="figure@88.[234,1345,299,1751]" captionTargetPageId="88" captionText="FIGURES 68 ­ 73. Adults of Anopinella. 68, A. mariana; 69, A. carabayana; 70, A. cafrosana; 71, A. porrasa; 72, A. styraxivora, 73, A. fana." httpUri="https://zenodo.org/record/168725/files/figure.png" pageId="46" pageNumber="47">72</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3744EBB6A20460CFE9275B1FDFA0F36" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="diagnosis">
<paragraph id="8BD11D306A20460DFE9275B1FB460FF5" blockId="46.[264,1323,1473,1899]" pageId="46" pageNumber="47">
Diagnosis. The male genitalia of
<taxonomicName id="4C6E66B36A20460DFD0575B1FCBE0CE4" box="[671,841,1473,1498]" class="Insecta" family="Tortricidae" genus="Anopinella" kingdom="Animalia" order="Lepidoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="styraxivora">
<emphasis id="B91AC1226A20460DFD0575B1FCBE0CE4" box="[671,841,1473,1498]" italics="true" pageId="46" pageNumber="47">A. styraxivora</emphasis>
</taxonomicName>
are unlike those of all other species of
<taxonomicName id="4C6E66B36A20460DFE927599FE7E0F3C" box="[264,393,1513,1538]" class="Insecta" family="Tortricidae" genus="Anopinella" kingdom="Animalia" order="Lepidoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
<emphasis id="B91AC1226A20460DFE927599FE7E0F3C" box="[264,393,1513,1538]" italics="true" pageId="46" pageNumber="47">Anopinella</emphasis>
</taxonomicName>
, with the distal lobes of the gnathos medially fused, forming a dorsoventrally flattened, spinose plate, and the valva narrow and lanceolate, with a large postsaccular lobe bearing a cluster of large spinelike setae. Autapomorphies for the species include these unique features of the male genitalia, plus the shiny forewing scaling. The highly divergent morphology of this species suggests that separate generic status may be justified if and when additional species are discovered to validate these synapomorphies.
</paragraph>
<paragraph id="8BD11D306A20460DFEA276A9FE410E55" blockId="46.[264,1323,1473,1899]" pageId="46" pageNumber="47">Description. Head: Frontoclypeus and vertex white; labial palpus with outer surface white intermixed with few brown scales, inner surface white. Antenna with scape white intermixed with few brown scales; basal 6­8 flagellomeres brown; distal flagellomeres brownish gray.</paragraph>
<paragraph id="8BD11D306A21460CFEA2716CFADC0B70" blockId="47.[264,1325,284,1550]" pageId="47" pageNumber="48">
Thorax: Tegula and mesonotum white intermixed with pale yellowish brown and some brown scales. Forewing (
<figureCitation id="135501B56A21460CFDA97134FD7C0860" box="[563,651,324,350]" captionStart="FIGURES 68 ­ 73" captionStartId="88.[264,382,1784,1808]" captionTargetBox="[270,1316,310,1741]" captionTargetId="figure@88.[234,1345,299,1751]" captionTargetPageId="88" captionText="FIGURES 68 ­ 73. Adults of Anopinella. 68, A. mariana; 69, A. carabayana; 70, A. cafrosana; 71, A. porrasa; 72, A. styraxivora, 73, A. fana." httpUri="https://zenodo.org/record/168725/files/figure.png" pageId="47" pageNumber="48">Fig. 72</figureCitation>
) length 6.0­
<quantity id="4C96B0D56A21460CFC877134FC810860" box="[797,886,324,350]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" pageId="47" pageNumber="48" unit="mm" value="8.1">8.1 mm</quantity>
(n = 2); incomplete basal fascia and costal blotch grayish brown intermixed with brown, reddish brown, and white; separated by a pale narrow white band demarcating costal blotch; costal blotch subtriangular, extending to distal costa, with typical small white spot near posterior end absent; area between CuP and posterior margin pale gray intermixed with few reddish brown scales; area beyond costal blotch pale gray intermixed with white. Fringe with inner portion reddish brown, outer portion gray. Hindwing pale brown basally, gradually darkening to apex.
</paragraph>
<paragraph id="8BD11D306A21460CFEA2722CFB270D10" blockId="47.[264,1325,284,1550]" pageId="47" pageNumber="48">
Abdomen: Male genitalia (
<figureCitation id="135501B56A21460CFD1A722CFD2B0B48" box="[640,732,604,630]" captionStart="FIGURES 33" captionStartId="74.[264,382,1856,1880]" captionText="FIGURES 33. Male genitalia of Anopinella, valvae spread, aedeagus removed. 33, A. styraxivora." pageId="47" pageNumber="48">Fig. 33</figureCitation>
; drawn from JWB slide 557; n = 1) with uncus curved at ca. mid­length. Socius digitate, short, ca. 0.16 length of basal portion of gnathos arms. Gnathos arms fused distally, forming a multispinose, horizontal plate. Valva widest basally, gradually narrowed toward apex; costa straight throughout length, densely setose apically; postsaccular margin abruptly rounded distally forming an subtriangular lobe bearing a dense cluster of stout setae; cucullus elongate, knife­like. Phallus slightly curved medially; vesica densely microtrichiate, with two elongate cornuti. Female genitalia (
<figureCitation id="135501B56A21460CFB61733CFEDF0AB0" captionStart="FIGURES 54 ­ 55" captionStartId="85.[264,382,1860,1884]" captionTargetBox="[308,1271,432,1742]" captionTargetId="figure@85.[265,1322,361,1802]" captionTargetPageId="85" captionText="FIGURES 54 ­ 55. Female genitalia of Anopinella, posterior end up. 54, A. rica; 55, A. styraxivora." httpUri="https://zenodo.org/record/156935/files/figure.png" pageId="47" pageNumber="48">Fig. 55</figureCitation>
; drawn from JWB slide 558; n =1) with broadened sclerotized mesal patch at lamella postvaginalis; eighth tergum with a median longitudinal sinus; ostium wide, ca. 0.33 width of seventh sternum at widest point; ductus bursae long, slender, gradually widened anteriorly; inception of accessory bursae ca. halfway between ostium and corpus bursae. Corpus bursae large, subspherical, weakly spiculate on 0.65 side bearing ductus seminalis.
</paragraph>
<paragraph id="8BD11D306A21460CFEA2744CFB580D98" blockId="47.[264,1325,284,1550]" pageId="47" pageNumber="48">
Pupa (
<figureCitation id="135501B56A21460CFE19744CFDE10D68" box="[387,534,1084,1110]" captionStart="FIGURES 90 ­ 92" captionStartId="92.[264,382,1825,1849]" captionTargetBox="[278,1306,382,1795]" captionTargetId="figure@92.[265,1321,361,1801]" captionTargetPageId="92" captionText="FIGURES 90 ­ 92. Wing venation of Anopinella triquetra (90); pupa of A. styraxivora, dorsum (91) and venter (92)." httpUri="https://zenodo.org/record/156937/files/figure.png" pageId="47" pageNumber="48">Figs. 91, 92</figureCitation>
). Typically tortricinae, without modification of head; no conspicuous sculpturing. Abdominal segments 3­8 with two rows of spines dorsally. Cremaster short, stout, with 6 long hooked setae. Indistinguishable from that of
<taxonomicName id="4C6E66B36A21460CFBAB74FCFB5F0D9B" box="[1073,1192,1164,1189]" class="Insecta" family="Tortricidae" genus="Anopinella" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="powelli">
<emphasis id="B91AC1226A21460CFBAB74FCFB5F0D9B" box="[1073,1192,1164,1189]" italics="true" pageId="47" pageNumber="48">A. powelli</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD11D306A21460CFEA274C4FEBB0C20" blockId="47.[264,1325,284,1550]" pageId="47" pageNumber="48">
<typeStatus id="54D5A3926A21460CFEA274C4FE5F0DF0" box="[312,424,1204,1230]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
, ɗ,
<collectingCountry id="F3795DA06A21460CFE4474C4FD920DF0" box="[478,613,1204,1230]" name="Costa Rica" pageId="47" pageNumber="48">
Costa
<taxonomicName id="4C6E66B36A21460CFDB674C4FD920DF0" box="[556,613,1204,1230]" class="Insecta" family="Tortricidae" genus="Anopinella" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="92" phylum="Arthropoda" rank="species" species="rica">Rica</taxonomicName>
</collectingCountry>
, Heredia Province, Uvita de San Rafael,
<quantity id="4C96B0D56A21460CFBFB74C4FB360DF0" box="[1121,1217,1204,1230]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="47" pageNumber="48" unit="m" value="1700.0">1700 m</quantity>
,
<date id="FFD03BF06A21460CFB4B74C4FEB20DC8" pageId="47" pageNumber="48" value="1991-03-10">10 Mar 1991</date>
, P. Hansen, r.f.
<taxonomicName id="4C6E66B36A21460CFD9774ACFDA10DCB" box="[525,598,1244,1269]" class="Magnoliopsida" family="Styracaceae" genus="Styrax" kingdom="Plantae" order="Ericales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="genus">
<emphasis id="B91AC1226A21460CFD9774ACFDA10DCB" box="[525,598,1244,1269]" italics="true" pageId="47" pageNumber="48">Styrax</emphasis>
</taxonomicName>
fruit [pupal exuvium pinned with specimen]. Deposited in UCB.
</paragraph>
<paragraph id="8BD11D306A21460CFEA2755CFC490C50" blockId="47.[264,1325,284,1550]" pageId="47" pageNumber="48">
<typeStatus id="54D5A3926A21460CFEA2755CFE690C78" box="[312,414,1324,1350]" pageId="47" pageNumber="48" type="paratype">Paratype</typeStatus>
(1Ψ).
<collectingCountry id="F3795DA06A21460CFE6B755CFD520C78" box="[497,677,1324,1350]" name="Costa Rica" pageId="47" pageNumber="48">
COSTA
<taxonomicName id="4C6E66B36A21460CFDC0755CFD520C78" box="[602,677,1324,1350]" class="Insecta" family="Tortricidae" genus="Anopinella" kingdom="Animalia" order="Lepidoptera" pageId="47" pageNumber="92" phylum="Arthropoda" rank="species" species="rica">RICA</taxonomicName>
</collectingCountry>
: Heredia Province: Uvita de San Rafael,
<quantity id="4C96B0D56A21460CFB04755CFB0A0C78" box="[1182,1277,1324,1350]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="47" pageNumber="48" unit="m" value="1700.0">1700 m</quantity>
,
<date id="FFD03BF06A21460CFA97755CFE8A0C50" pageId="47" pageNumber="48" value="1991-03-10">10 Mar 1991</date>
, P. Hansen, r.f.
<taxonomicName id="4C6E66B36A21460CFDAE7524FD890C53" box="[564,638,1364,1389]" class="Magnoliopsida" family="Styracaceae" genus="Styrax" kingdom="Plantae" order="Ericales" pageId="47" pageNumber="48" phylum="Tracheophyta" rank="genus">
<emphasis id="B91AC1226A21460CFDAE7524FD890C53" box="[564,638,1364,1389]" italics="true" pageId="47" pageNumber="48">Styrax</emphasis>
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fruit. Deposited in USNM.
</paragraph>
<paragraph id="8BD11D306A21460CFEA2750CFDAD0C80" blockId="47.[264,1325,284,1550]" pageId="47" pageNumber="48">
Biology. According to label data, both the
<typeStatus id="54D5A3926A21460CFCB3750CFC780CA8" box="[809,911,1404,1430]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
and
<typeStatus id="54D5A3926A21460CFC52750CFBDA0CA8" box="[968,1069,1404,1430]" pageId="47" pageNumber="48" type="paratype">paratype</typeStatus>
were reared from the fruit of
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<emphasis id="B91AC1226A21460CFEF875D4FE5B0C83" box="[354,428,1444,1469]" italics="true" pageId="47" pageNumber="48">Styrax</emphasis>
(Styracaceae)
</taxonomicName>
.
</paragraph>
<paragraph id="8BD11D306A21460CFEA275BCFDFA0F36" blockId="47.[264,1325,284,1550]" pageId="47" pageNumber="48">
Etymology. The specific epithet is derived from the host plant genus
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<emphasis id="B91AC1226A21460CFBC475BCFB500CDB" box="[1118,1191,1484,1509]" italics="true" pageId="47" pageNumber="48">Styrax</emphasis>
</taxonomicName>
, combined with the Latin “ivora.”
</paragraph>
</subSubSection>
</treatment>
</document>