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<document id="C5B0F1157837F94A9A28EB854F6CE14F" ID-CLB-Dataset="304119" ID-DOI="10.11646/zootaxa.5419.2.6" ID-GBIF-Dataset="247052c8-becf-49e3-8f01-a3c6a6c3f912" ID-ISSN="1175-5326" ID-Zenodo-Dep="10792134" ID-ZooBank="E7D59E77-AF1A-4DD9-809B-17552E487514" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1709795873870" checkinUser="plazi" docAuthor="Jiang, Ri-Xin &amp; Chen, Xiang-Sheng" docDate="2024" docId="03B9E05BFF9BFF8AFF0CFD76FE21079F" docLanguage="en" docName="zootaxa.5419.2.6.pdf" docOrigin="Zootaxa 5419 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5419.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macronychus xuhaoi Jiang &amp; Chen 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="279" masterDocId="FF809823FF9AFF8EFF9BFFC4FFD30066" masterDocTitle="A new species of the genus Macronychus Müller, 1806 from China (Coleoptera, Elmidae)" masterLastPageNumber="282" masterPageNumber="275" pageNumber="276" updateTime="1727957618592" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E3A0D4AF4B0C8DD7DAED1E8D657FE44B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="66AD32EF1A9BF44D68C26E9473813375">
<mods:title id="71CE28A650BA7B4140FB1CFEA092A2AC">A new species of the genus Macronychus Müller, 1806 from China (Coleoptera, Elmidae)</mods:title>
</mods:titleInfo>
<mods:name id="1D6D947ED623E25B397939EAAED31E5C" type="personal">
<mods:role id="8A4E451C1694A1FBBE9D85C0DBF8AB27">
<mods:roleTerm id="B640FE7D400A1B2B62516BB72350BCA9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C2B3CA4645893A85A9E1DC84690043FA">Jiang, Ri-Xin</mods:namePart>
<mods:nameIdentifier id="8C98BBD118B4865F4E2BC394C2B368C7" type="ORCID">0000-0002-5339-853X</mods:nameIdentifier>
<mods:affiliation id="3F2DEDD501A38FC54FDD02648A65933C">Institute of Entomology, Guizhou University, Guiyang 550025, Guizhou, P. R. China. &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources of Guizhou, Guizhou University, Guiyang, 550025, Guizhou, P. R. China. &amp; The Provincial Key Laboratory for Agricultural Pest Management of Mountainous Region, Guiyang 550025, Guizhou, P. R. China.</mods:affiliation>
</mods:name>
<mods:name id="5D948506B8AB6509F7D3F6D828014819" type="personal">
<mods:role id="995EF00AEA04F3F019C292A686FD5BB2">
<mods:roleTerm id="AE64728D178740D0B5CBF32DFD372BA8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6694466760F0F71F639B3A5900A7E2A0">Chen, Xiang-Sheng</mods:namePart>
<mods:affiliation id="DA97431297581F40C837226C69C0444E">Institute of Entomology, Guizhou University, Guiyang 550025, Guizhou, P. R. China. &amp; The Provincial Special Key Laboratory for Development and Utilization of Insect Resources of Guizhou, Guizhou University, Guiyang, 550025, Guizhou, P. R. China. &amp; The Provincial Key Laboratory for Agricultural Pest Management of Mountainous Region, Guiyang 550025, Guizhou, P. R. China.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="FE6608EA24D9B49EB5A567BD2883238A">text</mods:typeOfResource>
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<mods:title id="982D6187427C5F21E5ADF6AAF59C2072">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6A086864AFACDA6B43067DF5619A10CC">
<mods:date id="939BC2162030A3DFB06BD5EC67A02E6B">2024</mods:date>
<mods:detail id="8D6DA12503FC8ACF692C05B644DEFC88" type="pubDate">
<mods:number id="8253541D8A07C5F449DF3DAC409906F1">2024-03-06</mods:number>
</mods:detail>
<mods:detail id="FA8B7A2F39488F36225A66A8FFA97A38" type="volume">
<mods:number id="80480135F5A0895484155B885C8F0F17">5419</mods:number>
</mods:detail>
<mods:detail id="CB43C6F1DC1F1AF992C90C6834B66349" type="issue">
<mods:number id="53413CDE9E5D120A933B64825BBC352E">2</mods:number>
</mods:detail>
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<mods:start id="2C5F83572C6BC0F7C6838E348A57F91C">275</mods:start>
<mods:end id="3731B549B7AA5CEF9579A9B418D0CFA7">282</mods:end>
</mods:extent>
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<mods:location id="198B949E6849FCC38B31F15099267022">
<mods:url id="F6C6562621FFA8279D8C53E52A1E9270">http://dx.doi.org/10.11646/zootaxa.5419.2.6</mods:url>
</mods:location>
<mods:classification id="E7C0987123DA5C038ACE5277DFD90915">journal article</mods:classification>
<mods:identifier id="67918C9B097966548924891F3D1BC945" type="CLB-Dataset">304119</mods:identifier>
<mods:identifier id="77449874C1F0E597F5760E67221F160F" type="DOI">10.11646/zootaxa.5419.2.6</mods:identifier>
<mods:identifier id="A4ABC87AEA3AA7C88DBAAD314C158159" type="GBIF-Dataset">247052c8-becf-49e3-8f01-a3c6a6c3f912</mods:identifier>
<mods:identifier id="C2B9FD5B05695C344957A674108A6093" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="1A1FFC3510C3CA262846AA60FE2B4D24" type="ZooBank">E7D59E77-AF1A-4DD9-809B-17552E487514</mods:identifier>
</mods:mods>
<treatment id="03B9E05BFF9BFF8AFF0CFD76FE21079F" ID-DOI="http://doi.org/10.5281/zenodo.13885318" ID-Zenodo-Dep="13885318" LSID="urn:lsid:plazi:treatment:03B9E05BFF9BFF8AFF0CFD76FE21079F" httpUri="http://treatment.plazi.org/id/03B9E05BFF9BFF8AFF0CFD76FE21079F" lastPageId="4" lastPageNumber="279" pageId="1" pageNumber="276">
<subSubSection id="C30A02C6FF9BFF8FFF0CFD76FD1902AA" box="[151,714,690,717]" pageId="1" pageNumber="276" type="nomenclature">
<paragraph id="8BAF514DFF9BFF8FFF0CFD76FD1902AA" blockId="1.[151,714,690,753]" box="[151,714,690,717]" pageId="1" pageNumber="276">
<emphasis id="B9648D5FFF9BFF8FFF0CFD76FDC302AA" bold="true" box="[151,528,690,717]" pageId="1" pageNumber="276">
<taxonomicName id="4C102ACEFF9BFF8FFF0CFD76FE4902AB" authority="Jiang &amp; Chen, 2024" authorityName="Jiang &amp; Chen" authorityYear="2024" box="[151,410,690,717]" class="Insecta" family="Elmidae" genus="Macronychus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="276" phylum="Arthropoda" rank="species" species="xuhaoi" status="sp. nov.">
<emphasis id="B9648D5FFF9BFF8FFF0CFD76FE4902AB" bold="true" box="[151,410,690,717]" italics="true" pageId="1" pageNumber="276">Macronychus xuhaoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2573024FF9BFF8FFE3AFD77FE2D02AB" box="[417,510,691,717]" pageId="1" pageNumber="276" rank="species">sp. nov.</taxonomicNameLabel>
(
</emphasis>
许浩隆¡泥甲
<emphasis id="B9648D5FFF9BFF8FFD5BFD76FD1902AA" bold="true" box="[704,714,690,716]" pageId="1" pageNumber="276">)</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30A02C6FF9BFF8FFF0CFD13FED80297" box="[151,267,727,753]" pageId="1" pageNumber="276" type="description">
<paragraph id="8BAF514DFF9BFF8FFF0CFD13FED80297" blockId="1.[151,714,690,753]" box="[151,267,727,753]" pageId="1" pageNumber="276">
(
<figureCitation id="132B4DC8FF9BFF8FFF04FD13FED00297" box="[159,259,727,753]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1397,1421]" captionStartId-1="3.[151,250,1376,1400]" captionStartId-2="4.[151,250,1512,1536]" captionStartId-3="5.[151,250,1778,1802]" captionTargetBox-0="[163,1425,193,1362]" captionTargetBox-1="[151,1436,181,1352]" captionTargetBox-2="[151,1436,345,1489]" captionTargetBox-3="[151,1436,181,1754]" captionTargetId-0="figure-37@2.[151,1436,181,1373]" captionTargetId-1="figure-19@3.[151,1436,181,1352]" captionTargetId-2="figure-119@4.[151,1436,345,1489]" captionTargetId-3="figure-19@5.[151,1436,181,1754]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Dorsal habitus of Macronychus xuhaoi sp. nov. A. Male; B. Female." captionText-1="FIGURE 2. Diagnostic features of Macronychus xuhaoi sp. nov., male.A. Head, dorsal view; B. Antenna; C. Pronotum, dorsal view; D. Prosternal process. Note: red arrow, scutellum." captionText-2="FIGURE 3. Diagnostic features of Macronychus xuhaoi sp. nov., A–D, male; E, female. A. Metaventrite; B. Abdomen; C. Elytra; D–E. Protarsus. Note: red arrows in A: impressions of metaventrite; in B: carinae of ventrite I; in D: setal tufts of protarsus." captionText-3="FIGURE 4. Diagnostic features of Macronychus xuhaoi sp. nov., A–F, male; G–I, female. A. Aedeagus, dorsal view; B. Same, lateral view; C. Same, ventral view; D. part of aedeagus, dorsal view; E. Abdominal segment 9 and speculum gastrale; F–G. Seternite VIII; H. Ovipositor; I. stylus and apex of coxite. Note: black spot in A, setae of aedeagus." figureDoi-0="http://doi.org/10.5281/zenodo.10792136" figureDoi-1="http://doi.org/10.5281/zenodo.10792138" figureDoi-2="http://doi.org/10.5281/zenodo.10792140" figureDoi-3="http://doi.org/10.5281/zenodo.10792142" httpUri-0="https://zenodo.org/record/10792136/files/figure.png" httpUri-1="https://zenodo.org/record/10792138/files/figure.png" httpUri-2="https://zenodo.org/record/10792140/files/figure.png" httpUri-3="https://zenodo.org/record/10792142/files/figure.png" pageId="1" pageNumber="276">Figs 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30A02C6FF9BFF8FFF0CFCDBFB3D03AF" pageId="1" pageNumber="276" type="materials_examined">
<paragraph id="8BAF514DFF9BFF8FFF0CFCDBFB3D03AF" blockId="1.[151,1437,798,2013]" pageId="1" pageNumber="276">
<emphasis id="B9648D5FFF9BFF8FFF0CFCDBFEA3035E" bold="true" box="[151,368,798,825]" pageId="1" pageNumber="276">
Type material (
<specimenCount id="9D169AC4FF9BFF8FFEC8FCDAFEA3035E" box="[339,368,798,824]" count="52" pageId="1" pageNumber="276" type="male">52</specimenCount>
</emphasis>
♂♂,
<specimenCount id="9D169AC4FF9BFF8FFE2DFCDAFDD5035F" box="[438,518,798,825]" count="90" pageId="1" pageNumber="276" type="female">
<emphasis id="B9648D5FFF9BFF8FFE2DFCDAFE01035E" bold="true" box="[438,466,798,824]" pageId="1" pageNumber="276">90</emphasis>
♀♀
</specimenCount>
<emphasis id="B9648D5FFF9BFF8FFD9DFCDAFDDC035E" bold="true" box="[518,527,798,824]" pageId="1" pageNumber="276">)</emphasis>
:
<materialsCitation id="3B785B10FF9BFF8FFDBAFCDBFEE603E7" collectingDate="2017-08-14" collectingDateMax="2017-08-18" collectingDateMin="2017-08-14" collectionCode="K, GUGC" collectorName="Jian-Yue Qiu &amp; Hao Xu" country="China" county="Motuo County" elevation="2100" location="Linzhi City" pageId="1" pageNumber="276" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
<emphasis id="B9648D5FFF9BFF8FFDBAFCDBFD5D035F" bold="true" box="[545,654,799,825]" pageId="1" pageNumber="276">
<typeStatus id="54ABEFEFFF9BFF8FFDBAFCDBFD5D035F" box="[545,654,799,825]" pageId="1" pageNumber="276" type="holotype">Holotype</typeStatus>
</emphasis>
:
<collectingCountry id="F30711DDFF9BFF8FFD3BFCDBFD2E035F" box="[672,765,799,825]" name="China" pageId="1" pageNumber="276">CHINA</collectingCountry>
:
<specimenCount id="9D169AC4FF9BFF8FFC90FCDBFCF1035F" box="[779,802,799,825]" count="1" pageId="1" pageNumber="276" type="male">♂</specimenCount>
, labeled ‘China:
<collectingRegion id="49D49FAFFF9BFF8FFC71FCDBFBE8035F" box="[1002,1083,799,825]" country="China" name="Xizang" pageId="1" pageNumber="276">Xizang</collectingRegion>
(Ħǝ),
<location id="8ECF0796FF9BFF8FFB3AFCDBFAF7035F" LSID="urn:lsid:plazi:treatment:03B9E05BFF9BFF8AFF0CFD76FE21079F:8ECF0796FF9BFF8FFB3AFCDBFAF7035F" box="[1185,1316,799,825]" country="China" county="Motuo County" municipality="China" name="Linzhi City" pageId="1" pageNumber="276" stateProvince="Xizang">Linzhi City</location>
(wī市),
<collectingCounty id="62CE29C1FF9BFF8FFF0CFC87FEEE033B" box="[151,317,835,861]" pageId="1" pageNumber="276">Motuo County</collectingCounty>
(DZĸ县),
<location id="8ECF0796FF9BFF8FFE27FC87FD95033B" LSID="urn:lsid:plazi:treatment:03B9E05BFF9BFF8AFF0CFD76FE21079F:8ECF0796FF9BFF8FFE27FC87FD95033B" box="[444,582,835,861]" country="China" county="Motuo County" municipality="China" name="Bolonggong" pageId="1" pageNumber="276" stateProvince="Xizang">Bolonggong</location>
80K (波nū 80
<collectionCode id="ED01C988FF9BFF8FFC97FC87FCF1033B" box="[780,802,835,861]" pageId="1" pageNumber="276">K</collectionCode>
), H:
<quantity id="4CE8FCA8FF9BFF8FFCC6FC87FC7D033B" box="[861,942,835,861]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="276" unit="m" value="2100.0">
<elevation id="003DB67EFF9BFF8FFCC6FC87FC7D033B" box="[861,942,835,861]" metricMagnitude="3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="276" unit="m" value="2100.0">2100m</elevation>
</quantity>
,
<date id="FFAE778DFF9BFF8FFC21FC87FBAA033B" box="[954,1145,835,861]" pageId="1" pageNumber="276" value="2017-08-14" valueMax="2017-08-18" valueMin="2017-08-14">
<collectingDate id="EFEA8E65FF9BFF8FFC21FC87FBAA033B" box="[954,1145,835,861]" pageId="1" pageNumber="276" value="2017-08-14" valueMax="2017-08-18" valueMin="2017-08-14">14–18.VIII.2017</collectingDate>
</date>
,
<collectorName id="26E5349BFF9BFF8FFB1DFC87FAC8033B" box="[1158,1307,835,861]" pageId="1" pageNumber="276">Jian-Yue Qiu</collectorName>
&amp;
<collectorName id="26E5349BFF9BFF8FFAD8FC87FA4F033B" box="[1347,1436,835,861]" pageId="1" pageNumber="276">Hao Xu</collectorName>
leg.’ (
<collectionCode id="ED01C988FF9BFF8FFF4CFCA3FEFE03E7" box="[215,301,871,897]" collectionName="GUGC" pageId="1" pageNumber="276">GUGC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B785B10FF9BFF8FFEDBFCA3FE3803C3" collectingDate="2017-08-14" collectingDateMax="2017-08-18" collectingDateMin="2017-08-14" collectionCode="GUGC, MYNU" collectorName="Jian-Yue Qiu &amp; Hao Xu" country="China" county="Motuo County" elevation="2100" location="Linzhi City" pageId="1" pageNumber="276" specimenCount="141" specimenCount-female="90" specimenCount-male="51" stateProvince="Xizang" typeStatus="paratype">
<emphasis id="B9648D5FFF9BFF8FFEDBFCA3FE3103E6" bold="true" box="[320,482,870,897]" pageId="1" pageNumber="276">
<typeStatus id="54ABEFEFFF9BFF8FFEDBFCA3FE6203E7" box="[320,433,871,897]" pageId="1" pageNumber="276" type="paratype">Paratype</typeStatus>
: (
<specimenCount id="9D169AC4FF9BFF8FFE5EFCA2FDC703E7" box="[453,532,870,897]" count="51" pageId="1" pageNumber="276" type="male">51 ♂♂</specimenCount>
</emphasis>
,
<specimenCount id="9D169AC4FF9BFF8FFDB8FCA2FDA303E7" box="[547,624,870,897]" count="90" pageId="1" pageNumber="276" type="female">
<emphasis id="B9648D5FFF9BFF8FFDB8FCA2FDEC03E6" bold="true" box="[547,575,870,896]" pageId="1" pageNumber="276">90</emphasis>
♀♀
</specimenCount>
<emphasis id="B9648D5FFF9BFF8FFDEBFCA2FD7503E6" bold="true" box="[624,678,870,896]" pageId="1" pageNumber="276">): 41</emphasis>
♂♂,
<emphasis id="B9648D5FFF9BFF8FFD7DFCA2FCD103E6" bold="true" box="[742,770,870,896]" pageId="1" pageNumber="276">70</emphasis>
♀♀, with same label data as the holotype (47 ♂♂, 85 ♀♀,
<collectionCode id="ED01C988FF9BFF8FFF0CFC4FFF3903C3" box="[151,234,907,933]" collectionName="GUGC" pageId="1" pageNumber="276">GUGC</collectionCode>
; 5 ♂♂, 5 ♀♀,
<collectionCode id="ED01C988FF9BFF8FFE13FC4FFE3003C3" box="[392,483,907,933]" collectionName="MYNU" pageId="1" pageNumber="276">MYNU</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B785B10FF9BFF8FFE6EFC4EFB3903AF" collectingDate="2017-08-18" collectionCode="GUGC" collectorName="Jian-Yue Qiu &amp; Hao Xu" country="China" county="Motuo County" elevation="1000" location="Linzhi City" municipality="Motuo Town" pageId="1" pageNumber="276" specimenCount="30" specimenCount-female="20" specimenCount-male="10" stateProvince="Xizang">
<specimenCount id="9D169AC4FF9BFF8FFE6EFC4EFD9303C3" box="[501,576,906,933]" count="10" pageId="1" pageNumber="276" type="male">
<emphasis id="B9648D5FFF9BFF8FFE6EFC4EFDC203C2" bold="true" box="[501,529,906,932]" pageId="1" pageNumber="276">10</emphasis>
♂♂
</specimenCount>
,
<specimenCount id="9D169AC4FF9BFF8FFDD6FC4EFD4B03C3" box="[589,664,906,933]" count="20" pageId="1" pageNumber="276" type="female">
<emphasis id="B9648D5FFF9BFF8FFDD6FC4EFDBA03C2" bold="true" box="[589,617,906,932]" pageId="1" pageNumber="276">20</emphasis>
♀♀
</specimenCount>
,
<emphasis id="B9648D5FFF9BFF8FFD3EFC4FFCDA03C2" bold="true" box="[677,777,906,933]" pageId="1" pageNumber="276">labeled ‘</emphasis>
<collectingCountry id="F30711DDFF9BFF8FFC92FC4FFC8303C3" box="[777,848,907,933]" name="China" pageId="1" pageNumber="276">China</collectingCountry>
:
<collectingRegion id="49D49FAFFF9BFF8FFCC1FC4FFC7803C3" box="[858,939,907,933]" country="China" name="Xizang" pageId="1" pageNumber="276">Xizang</collectingRegion>
(Ħǝ),
<location id="8ECF0796FF9BFF8FFB9CFC4FFB5B03C3" LSID="urn:lsid:plazi:treatment:03B9E05BFF9BFF8AFF0CFD76FE21079F:8ECF0796FF9BFF8FFB9CFC4FFB5B03C3" box="[1031,1160,907,933]" country="China" county="Motuo County" municipality="Motuo Town" name="Linzhi City" pageId="1" pageNumber="276" stateProvince="Xizang">Linzhi City</location>
(wī市),
<collectingCounty id="62CE29C1FF9BFF8FFB61FC4FFA4F03C3" box="[1274,1436,907,933]" pageId="1" pageNumber="276">Motuo County</collectingCounty>
(DZĸ县),
<collectingMunicipality id="6BCBCB37FF9BFF8FFE97FC6BFE4F03AF" box="[268,412,943,969]" pageId="1" pageNumber="276">Motuo Town</collectingMunicipality>
(DZĸDz), H:
<quantity id="4CE8FCA8FF9BFF8FFDA0FC6BFD5803AF" box="[571,651,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="276" unit="m" value="1000.0">
<elevation id="003DB67EFF9BFF8FFDA0FC6BFD5803AF" box="[571,651,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="276" unit="m" value="1000.0">1000m</elevation>
</quantity>
,
<date id="FFAE778DFF9BFF8FFD0CFC6BFCFF03AF" box="[663,812,943,969]" pageId="1" pageNumber="276" value="2017-08-18">
<collectingDate id="EFEA8E65FF9BFF8FFD0CFC6BFCFF03AF" box="[663,812,943,969]" pageId="1" pageNumber="276" value="2017-08-18">18.VIII.2017</collectingDate>
</date>
,
<collectorName id="26E5349BFF9BFF8FFCACFC6BFC1903AF" box="[823,970,943,969]" pageId="1" pageNumber="276">Jian-Yue Qiu</collectorName>
&amp;
<collectorName id="26E5349BFF9BFF8FFC75FC6BFB9603AF" box="[1006,1093,943,969]" pageId="1" pageNumber="276">Hao Xu</collectorName>
leg.
<emphasis id="B9648D5FFF9BFF8FFBEDFC6AFBAE03AE" bold="true" box="[1142,1149,942,968]" pageId="1" pageNumber="276">’</emphasis>
(
<collectionCode id="ED01C988FF9BFF8FFB17FC6BFB3103AF" box="[1164,1250,943,969]" collectionName="GUGC" pageId="1" pageNumber="276">GUGC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A02C6FF9BFF8DFF5CFC16FCDD066F" lastPageId="3" lastPageNumber="278" pageId="1" pageNumber="276" type="description">
<paragraph id="8BAF514DFF9BFF8FFF5CFC16FAA20477" blockId="1.[151,1437,798,2013]" pageId="1" pageNumber="276">
<emphasis id="B9648D5FFF9BFF8FFF5CFC16FE8A038A" bold="true" box="[199,345,978,1004]" pageId="1" pageNumber="276">Description.</emphasis>
Male (
<figureCitation id="132B4DC8FF9BFF8FFE31FC17FDDB038B" box="[426,520,979,1005]" captionStart="FIGURE 1" captionStartId="2.[151,250,1397,1421]" captionTargetBox="[163,1425,193,1362]" captionTargetId="figure-37@2.[151,1436,181,1373]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Macronychus xuhaoi sp. nov. A. Male; B. Female." figureDoi="http://doi.org/10.5281/zenodo.10792136" httpUri="https://zenodo.org/record/10792136/files/figure.png" pageId="1" pageNumber="276">Figs 1A</figureCitation>
,
<figureCitation id="132B4DC8FF9BFF8FFD8EFC17FDF6038B" box="[533,549,979,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1376,1400]" captionTargetBox="[151,1436,181,1352]" captionTargetId="figure-19@3.[151,1436,181,1352]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Macronychus xuhaoi sp. nov., male.A. Head, dorsal view; B. Antenna; C. Pronotum, dorsal view; D. Prosternal process. Note: red arrow, scutellum." figureDoi="http://doi.org/10.5281/zenodo.10792138" httpUri="https://zenodo.org/record/10792138/files/figure.png" pageId="1" pageNumber="276">2</figureCitation>
,
<figureCitation id="132B4DC8FF9BFF8FFDA9FC17FDAB038B" box="[562,632,979,1005]" captionStart="FIGURE 3" captionStartId="4.[151,250,1512,1536]" captionTargetBox="[151,1436,345,1489]" captionTargetId="figure-119@4.[151,1436,345,1489]" captionTargetPageId="4" captionText="FIGURE 3. Diagnostic features of Macronychus xuhaoi sp. nov., A–D, male; E, female. A. Metaventrite; B. Abdomen; C. Elytra; D–E. Protarsus. Note: red arrows in A: impressions of metaventrite; in B: carinae of ventrite I; in D: setal tufts of protarsus." figureDoi="http://doi.org/10.5281/zenodo.10792140" httpUri="https://zenodo.org/record/10792140/files/figure.png" pageId="1" pageNumber="276">3A–D</figureCitation>
,
<figureCitation id="132B4DC8FF9BFF8FFD1EFC17FD18038B" box="[645,715,979,1005]" captionStart="FIGURE 4" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1436,181,1754]" captionTargetId="figure-19@5.[151,1436,181,1754]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Macronychus xuhaoi sp. nov., A–F, male; G–I, female. A. Aedeagus, dorsal view; B. Same, lateral view; C. Same, ventral view; D. part of aedeagus, dorsal view; E. Abdominal segment 9 and speculum gastrale; F–G. Seternite VIII; H. Ovipositor; I. stylus and apex of coxite. Note: black spot in A, setae of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10792142" httpUri="https://zenodo.org/record/10792142/files/figure.png" pageId="1" pageNumber="276">4A–E</figureCitation>
). Body moderately elongate (
<figureCitation id="132B4DC8FF9BFF8FFB83FC17FBA1038B" box="[1048,1138,979,1005]" captionStart="FIGURE 1" captionStartId="2.[151,250,1397,1421]" captionTargetBox="[163,1425,193,1362]" captionTargetId="figure-37@2.[151,1436,181,1373]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Macronychus xuhaoi sp. nov. A. Male; B. Female." figureDoi="http://doi.org/10.5281/zenodo.10792136" httpUri="https://zenodo.org/record/10792136/files/figure.png" pageId="1" pageNumber="276">Fig. 1A</figureCitation>
), black, legs and antennae dark brown, anterior margin of pronotum reddish brown. Dorsal surface shiny, covered with sparse short setae.
</paragraph>
<paragraph id="8BAF514DFF9BFF8FFF5CFBDFFBC70557" blockId="1.[151,1437,798,2013]" pageId="1" pageNumber="276">
Head (
<figureCitation id="132B4DC8FF9BFF8FFE8AFBDFFEBB0453" box="[273,360,1051,1077]" captionStart="FIGURE 2" captionStartId="3.[151,250,1376,1400]" captionTargetBox="[151,1436,181,1352]" captionTargetId="figure-19@3.[151,1436,181,1352]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Macronychus xuhaoi sp. nov., male.A. Head, dorsal view; B. Antenna; C. Pronotum, dorsal view; D. Prosternal process. Note: red arrow, scutellum." figureDoi="http://doi.org/10.5281/zenodo.10792138" httpUri="https://zenodo.org/record/10792138/files/figure.png" pageId="1" pageNumber="276">Fig. 2A</figureCitation>
) wider than long, disc covered with dense small punctures, and sparse short setae. Frons smooth, wrinkled near eyes, covered with sparse short and apical blunt setae. Eye large, oval. Clypeus covered with sparse punctures and setae slightly shorter than those of dorsal surface of head, apex of those setae cuspidal, distal part of clypeus microreticulated. Labrum transverse, shorter and narrower than clypeus, surface microreticulated, covered with sparse small punctures and long bristles at anterior portion, anterior margin almost straight, anterolateral angles rounded. Antenna (
<figureCitation id="132B4DC8FF9BFF8FFEEAFB0BFE1A048F" box="[369,457,1231,1257]" captionStart="FIGURE 2" captionStartId="3.[151,250,1376,1400]" captionTargetBox="[151,1436,181,1352]" captionTargetId="figure-19@3.[151,1436,181,1352]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Macronychus xuhaoi sp. nov., male.A. Head, dorsal view; B. Antenna; C. Pronotum, dorsal view; D. Prosternal process. Note: red arrow, scutellum." figureDoi="http://doi.org/10.5281/zenodo.10792138" httpUri="https://zenodo.org/record/10792138/files/figure.png" pageId="1" pageNumber="276">Fig. 2B</figureCitation>
) 7-segmented, scape about as long as wide, expanded at apex; pedicel about twice as long as wide; antennomere III about as long as pedicel, thinner and expanded near apex; IV–VI transverse, about as long as wide; VII elongate and enlarged, long oval, apex with tuft of long setae.
</paragraph>
<paragraph id="8BAF514DFF9BFF8FFF5CFAFFFACB0613" blockId="1.[151,1437,798,2013]" pageId="1" pageNumber="276">
Pronotum (
<figureCitation id="132B4DC8FF9BFF8FFEDDFAFFFE4E0533" box="[326,413,1339,1365]" captionStart="FIGURE 2" captionStartId="3.[151,250,1376,1400]" captionTargetBox="[151,1436,181,1352]" captionTargetId="figure-19@3.[151,1436,181,1352]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Macronychus xuhaoi sp. nov., male.A. Head, dorsal view; B. Antenna; C. Pronotum, dorsal view; D. Prosternal process. Note: red arrow, scutellum." figureDoi="http://doi.org/10.5281/zenodo.10792138" httpUri="https://zenodo.org/record/10792138/files/figure.png" pageId="1" pageNumber="276">Fig. 2C</figureCitation>
) longer than wide, widest in basal 1/4, narrowed from base to apex. Surface covered with sparse short setae which are of
<specimenCount id="9D169AC4FF9BFF8FFE74FA9BFD8E051F" box="[495,605,1375,1401]" count="2" pageId="1" pageNumber="276" type="generic" typeStatus="types">two types</specimenCount>
, thin and short setae with apical spiculated, and distinctly longer setae with apical blunt. Anterior margin reddish-brown, curved and with angles sub-orthogonal; lateral margins sinuated; basal margin trisinuate, posterior angles acutangular, posterior half with two large admedian prebasal gibbosities separated by mesal gap. Surface of apical half of pronotum and admedian prebasal gibbosities shiny, apical 1/6 densely micropunctured, other parts of apical half and admedian prebasal gibbosities sparsely micropunctured; transverse middle impression, mesal gap and posterior angles covered with dense small punctures and weakly microreticulate. Prosternum (
<figureCitation id="132B4DC8FF9BFF8FFEBCF9F3FEAD0637" box="[295,382,1591,1617]" captionStart="FIGURE 2" captionStartId="3.[151,250,1376,1400]" captionTargetBox="[151,1436,181,1352]" captionTargetId="figure-19@3.[151,1436,181,1352]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Macronychus xuhaoi sp. nov., male.A. Head, dorsal view; B. Antenna; C. Pronotum, dorsal view; D. Prosternal process. Note: red arrow, scutellum." figureDoi="http://doi.org/10.5281/zenodo.10792138" httpUri="https://zenodo.org/record/10792138/files/figure.png" pageId="1" pageNumber="276">Fig. 2D</figureCitation>
) disc of anterior half shiny, covered with sparse small punctures, each puncture bearing a long seta; sides of anterior half wrinkled; prosternal process (
<figureCitation id="132B4DC8FF9BFF8FFC97F99FFCB70613" box="[780,868,1627,1653]" captionStart="FIGURE 2" captionStartId="3.[151,250,1376,1400]" captionTargetBox="[151,1436,181,1352]" captionTargetId="figure-19@3.[151,1436,181,1352]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Macronychus xuhaoi sp. nov., male.A. Head, dorsal view; B. Antenna; C. Pronotum, dorsal view; D. Prosternal process. Note: red arrow, scutellum." figureDoi="http://doi.org/10.5281/zenodo.10792138" httpUri="https://zenodo.org/record/10792138/files/figure.png" pageId="1" pageNumber="276">Fig. 2D</figureCitation>
) strongly wrinkled with apex rounded.
</paragraph>
<paragraph id="8BAF514DFF9BFF8FFF5CF9BBFC2706DB" blockId="1.[151,1437,798,2013]" pageId="1" pageNumber="276">
Scutellum (
<figureCitation id="132B4DC8FF9BFF8FFED2F9BBFE7306FF" box="[329,416,1663,1689]" captionStart="FIGURE 2" captionStartId="3.[151,250,1376,1400]" captionTargetBox="[151,1436,181,1352]" captionTargetId="figure-19@3.[151,1436,181,1352]" captionTargetPageId="3" captionText="FIGURE 2. Diagnostic features of Macronychus xuhaoi sp. nov., male.A. Head, dorsal view; B. Antenna; C. Pronotum, dorsal view; D. Prosternal process. Note: red arrow, scutellum." figureDoi="http://doi.org/10.5281/zenodo.10792138" httpUri="https://zenodo.org/record/10792138/files/figure.png" pageId="1" pageNumber="276">Fig. 2C</figureCitation>
) half fusiform, about twice as long as wide, widest at middle; surface shiny, covered with several small setigerous punctures; lateral margins finely curved, apex acute.
</paragraph>
<paragraph id="8BAF514DFF9BFF8FFF5CF903FD48074F" blockId="1.[151,1437,798,2013]" pageId="1" pageNumber="276">
Elytra (
<figureCitation id="132B4DC8FF9BFF8FFE81F903FEBC0687" box="[282,367,1735,1761]" captionStart="FIGURE 3" captionStartId="4.[151,250,1512,1536]" captionTargetBox="[151,1436,345,1489]" captionTargetId="figure-119@4.[151,1436,345,1489]" captionTargetPageId="4" captionText="FIGURE 3. Diagnostic features of Macronychus xuhaoi sp. nov., A–D, male; E, female. A. Metaventrite; B. Abdomen; C. Elytra; D–E. Protarsus. Note: red arrows in A: impressions of metaventrite; in B: carinae of ventrite I; in D: setal tufts of protarsus." figureDoi="http://doi.org/10.5281/zenodo.10792140" httpUri="https://zenodo.org/record/10792140/files/figure.png" pageId="1" pageNumber="276">Fig. 3C</figureCitation>
) about twice as long as wide, parallel-sided at basal 2/3, widest near apical 1/3. Surface smooth, covered with sparse short setae, nine distinct striae composed of a row of large punctures, punctures much smaller near elytral apex. Hind wings well developed.
</paragraph>
<paragraph id="8BAF514DFF9BFF8FFF5CF8F7FCCA07F3" blockId="1.[151,1437,798,2013]" pageId="1" pageNumber="276">
Metaventrite (
<figureCitation id="132B4DC8FF9BFF8FFEF8F8F7FE6B072B" box="[355,440,1843,1869]" captionStart="FIGURE 3" captionStartId="4.[151,250,1512,1536]" captionTargetBox="[151,1436,345,1489]" captionTargetId="figure-119@4.[151,1436,345,1489]" captionTargetPageId="4" captionText="FIGURE 3. Diagnostic features of Macronychus xuhaoi sp. nov., A–D, male; E, female. A. Metaventrite; B. Abdomen; C. Elytra; D–E. Protarsus. Note: red arrows in A: impressions of metaventrite; in B: carinae of ventrite I; in D: setal tufts of protarsus." figureDoi="http://doi.org/10.5281/zenodo.10792140" httpUri="https://zenodo.org/record/10792140/files/figure.png" pageId="1" pageNumber="276">Fig. 3A</figureCitation>
) with disc smooth, covered with sparse short setae, sides with plastron setae. Median sulcus distinct, extending from posterior margin to ca. 2/3 of metasternum. Large impressions present near mesocoxae, along median sulcus and near posterior margin (
<figureCitation id="132B4DC8FF9BFF8FFD29F8BFFCDF07F3" box="[690,780,1915,1941]" captionStart="FIGURE 3" captionStartId="4.[151,250,1512,1536]" captionTargetBox="[151,1436,345,1489]" captionTargetId="figure-119@4.[151,1436,345,1489]" captionTargetPageId="4" captionText="FIGURE 3. Diagnostic features of Macronychus xuhaoi sp. nov., A–D, male; E, female. A. Metaventrite; B. Abdomen; C. Elytra; D–E. Protarsus. Note: red arrows in A: impressions of metaventrite; in B: carinae of ventrite I; in D: setal tufts of protarsus." figureDoi="http://doi.org/10.5281/zenodo.10792140" httpUri="https://zenodo.org/record/10792140/files/figure.png" pageId="1" pageNumber="276">Fig. 3A</figureCitation>
).
</paragraph>
<paragraph id="8BAF514DFF9BFF8FFF5CF85BFB8F07BB" blockId="1.[151,1437,798,2013]" pageId="1" pageNumber="276">
Legs simple, femora weakly clavate, widest subapically. Tibiae widened in apical half. Tarsi slightly shorter than length of tibiae, protarsal segments (
<figureCitation id="132B4DC8FF9BFF8FFDFDF807FD6D07BB" box="[614,702,1987,2013]" captionStart="FIGURE 3" captionStartId="4.[151,250,1512,1536]" captionTargetBox="[151,1436,345,1489]" captionTargetId="figure-119@4.[151,1436,345,1489]" captionTargetPageId="4" captionText="FIGURE 3. Diagnostic features of Macronychus xuhaoi sp. nov., A–D, male; E, female. A. Metaventrite; B. Abdomen; C. Elytra; D–E. Protarsus. Note: red arrows in A: impressions of metaventrite; in B: carinae of ventrite I; in D: setal tufts of protarsus." figureDoi="http://doi.org/10.5281/zenodo.10792140" httpUri="https://zenodo.org/record/10792140/files/figure.png" pageId="1" pageNumber="276">Fig. 3D</figureCitation>
) with ventroapical tuft of long setae.
</paragraph>
<caption id="DF6F01C5FF98FF8CFF0CFAB1FC0305EB" ID-DOI="http://doi.org/10.5281/zenodo.10792136" ID-Zenodo-Dep="10792136" box="[151,976,1397,1421]" httpUri="https://zenodo.org/record/10792136/files/figure.png" pageId="2" pageNumber="277" startId="2.[151,250,1397,1421]" targetBox="[163,1425,193,1362]" targetPageId="2" targetType="figure">
<paragraph id="8BAF514DFF98FF8CFF0CFAB1FC0305EB" blockId="2.[151,976,1397,1421]" box="[151,976,1397,1421]" pageId="2" pageNumber="277">
<emphasis id="B9648D5FFF98FF8CFF0CFAB1FEC005EB" bold="true" box="[151,275,1397,1421]" pageId="2" pageNumber="277">FIGURE 1.</emphasis>
Dorsal habitus of
<taxonomicName id="4C102ACEFF98FF8CFE55FAB1FD7105EB" authority="Jiang &amp; Chen, 2024" authorityName="Jiang &amp; Chen" authorityYear="2024" box="[462,674,1397,1421]" class="Insecta" family="Elmidae" genus="Macronychus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="277" phylum="Arthropoda" rank="species" species="xuhaoi" status="sp. nov.">
<emphasis id="B9648D5FFF98FF8CFE55FAB1FD7105EB" box="[462,674,1397,1421]" italics="true" pageId="2" pageNumber="277">Macronychus xuhaoi</emphasis>
</taxonomicName>
<emphasis id="B9648D5FFF98FF8CFD32FAB1FCCA05EB" bold="true" box="[681,793,1397,1421]" pageId="2" pageNumber="277">
<taxonomicNameLabel id="A2573024FF98FF8CFD32FAB1FD2A05EB" box="[681,761,1397,1421]" pageId="2" pageNumber="277" rank="species">sp. nov.</taxonomicNameLabel>
A.
</emphasis>
Male;
<emphasis id="B9648D5FFF98FF8CFCFBFAB1FCAB05EB" bold="true" box="[864,888,1397,1421]" pageId="2" pageNumber="277">B.</emphasis>
Female.
</paragraph>
</caption>
<paragraph id="8BAF514DFF98FF8CFF5CFA7FFD520600" blockId="2.[151,1437,1467,2034]" pageId="2" pageNumber="277">
Abdomen (
<figureCitation id="132B4DC8FF98FF8CFED3FA7FFE7305B3" box="[328,416,1467,1493]" captionStart="FIGURE 3" captionStartId="4.[151,250,1512,1536]" captionTargetBox="[151,1436,345,1489]" captionTargetId="figure-119@4.[151,1436,345,1489]" captionTargetPageId="4" captionText="FIGURE 3. Diagnostic features of Macronychus xuhaoi sp. nov., A–D, male; E, female. A. Metaventrite; B. Abdomen; C. Elytra; D–E. Protarsus. Note: red arrows in A: impressions of metaventrite; in B: carinae of ventrite I; in D: setal tufts of protarsus." figureDoi="http://doi.org/10.5281/zenodo.10792140" httpUri="https://zenodo.org/record/10792140/files/figure.png" pageId="2" pageNumber="277">Fig. 3B</figureCitation>
) with 5 visible sternites, surface with plastron setae. Ventrite I longest, about as long as ventrite II and ventrite III combined, disc without plastron setae, smooth, with sparse small punctures and short setae, carinae of ventrite I reduced, not reaching posterior margin. Ventrites II–V completely covered with plastron setae. Ventrite III and ventrite IV about same in length. Ventrite V slightly shorter than ventrite I, apical margin rounded and covered with dense long setae.
</paragraph>
<paragraph id="8BAF514DFF98FF8CFF5CF9ABFC970693" blockId="2.[151,1437,1467,2034]" pageId="2" pageNumber="277">
Aedeagus (
<figureCitation id="132B4DC8FF98FF8CFEDDF9ABFE1D06EF" box="[326,462,1647,1673]" captionStart="FIGURE 4" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1436,181,1754]" captionTargetId="figure-19@5.[151,1436,181,1754]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Macronychus xuhaoi sp. nov., A–F, male; G–I, female. A. Aedeagus, dorsal view; B. Same, lateral view; C. Same, ventral view; D. part of aedeagus, dorsal view; E. Abdominal segment 9 and speculum gastrale; F–G. Seternite VIII; H. Ovipositor; I. stylus and apex of coxite. Note: black spot in A, setae of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10792142" httpUri="https://zenodo.org/record/10792142/files/figure.png" pageId="2" pageNumber="277">Figs 4 A–D</figureCitation>
), slender, elongate, nearly symmetrical, slightly subpically constricted in lateral view; apex almost rounded, median lobe with four sclerotizations, a pair of smaller, located near base, and two larger ones, located at 1/3 of the median lobe, surface with short hair-like setae along apical 1/2; spiculum gastrale as in
<figureCitation id="132B4DC8FF98FF8CFAD8F973FA4B06B7" box="[1347,1432,1719,1745]" captionStart="FIGURE 4" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1436,181,1754]" captionTargetId="figure-19@5.[151,1436,181,1754]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Macronychus xuhaoi sp. nov., A–F, male; G–I, female. A. Aedeagus, dorsal view; B. Same, lateral view; C. Same, ventral view; D. part of aedeagus, dorsal view; E. Abdominal segment 9 and speculum gastrale; F–G. Seternite VIII; H. Ovipositor; I. stylus and apex of coxite. Note: black spot in A, setae of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10792142" httpUri="https://zenodo.org/record/10792142/files/figure.png" pageId="2" pageNumber="277">Fig. 4E</figureCitation>
. Sternite VIII (
<figureCitation id="132B4DC8FF98FF8CFEAFF91FFE5B0693" box="[308,392,1755,1781]" captionStart="FIGURE 4" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1436,181,1754]" captionTargetId="figure-19@5.[151,1436,181,1754]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Macronychus xuhaoi sp. nov., A–F, male; G–I, female. A. Aedeagus, dorsal view; B. Same, lateral view; C. Same, ventral view; D. part of aedeagus, dorsal view; E. Abdominal segment 9 and speculum gastrale; F–G. Seternite VIII; H. Ovipositor; I. stylus and apex of coxite. Note: black spot in A, setae of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10792142" httpUri="https://zenodo.org/record/10792142/files/figure.png" pageId="2" pageNumber="277">Fig. 4F</figureCitation>
) with deeply emarginate apical margin.
</paragraph>
<paragraph id="8BAF514DFF98FF8CFF5CF8C4FCDD0758" blockId="2.[151,1437,1467,2034]" pageId="2" pageNumber="277">
Measurements (n=10): CL:
<quantity id="4CE8FCA8FF98FF8CFD9AF93BFD7B077C" box="[513,680,1791,1818]" metricMagnitude="-3" metricUnit="m" metricValue="2.905" metricValueMax="3.08" metricValueMin="2.73" pageId="2" pageNumber="277" unit="mm" value="2.905" valueMax="3.08" valueMin="2.73">2.73–3.08 mm</quantity>
; HL:
<quantity id="4CE8FCA8FF98FF8CFD73F93BFC5D077C" box="[744,910,1791,1818]" metricMagnitude="-4" metricUnit="m" metricValue="3.55" metricValueMax="3.8" metricValueMin="3.3" pageId="2" pageNumber="277" unit="mm" value="0.355" valueMax="0.38" valueMin="0.33">0.33–0.38 mm</quantity>
, HW:
<quantity id="4CE8FCA8FF98FF8CFC4CF93BFBAD077C" box="[983,1150,1791,1818]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="2" pageNumber="277" unit="mm" value="0.525" valueMax="0.55" valueMin="0.5">0.50–0.55 mm</quantity>
; PL:
<quantity id="4CE8FCA8FF98FF8CFB21F93BFA8C077C" box="[1210,1375,1791,1818]" metricMagnitude="-4" metricUnit="m" metricValue="9.15" metricValueMax="9.8" metricValueMin="8.5" pageId="2" pageNumber="277" unit="mm" value="0.915" valueMax="0.98" valueMin="0.85">0.85–0.98 mm</quantity>
, PW:
<quantity id="4CE8FCA8FF98FF8CFF0CF8E7FEEE0758" box="[151,317,1827,1854]" metricMagnitude="-4" metricUnit="m" metricValue="8.75" metricValueMax="9.5" metricValueMin="8.0" pageId="2" pageNumber="277" unit="mm" value="0.875" valueMax="0.95" valueMin="0.8">0.80–0.95 mm</quantity>
; EL:
<quantity id="4CE8FCA8FF98FF8CFEE1F8E7FDCC0758" box="[378,543,1827,1854]" metricMagnitude="-3" metricUnit="m" metricValue="1.99" metricValueMax="2.1" metricValueMin="1.88" pageId="2" pageNumber="277" unit="mm" value="1.99" valueMax="2.1" valueMin="1.88">1.88–2.10 mm</quantity>
, EW:
<quantity id="4CE8FCA8FF98FF8CFDF8F8E7FCDA0758" box="[611,777,1827,1854]" metricMagnitude="-3" metricUnit="m" metricValue="1.175" metricValueMax="1.25" metricValueMin="1.1" pageId="2" pageNumber="277" unit="mm" value="1.175" valueMax="1.25" valueMin="1.1">1.10–1.25 mm</quantity>
.
</paragraph>
<paragraph id="8BAF514DFF98FF8CFF5CF883FC9C0797" blockId="2.[151,1437,1467,2034]" pageId="2" pageNumber="277">
Female (
<figureCitation id="132B4DC8FF98FF8CFEB0F883FE5B0707" box="[299,392,1863,1889]" captionStart="FIGURE 1" captionStartId="2.[151,250,1397,1421]" captionTargetBox="[163,1425,193,1362]" captionTargetId="figure-37@2.[151,1436,181,1373]" captionTargetPageId="2" captionText="FIGURE 1. Dorsal habitus of Macronychus xuhaoi sp. nov. A. Male; B. Female." figureDoi="http://doi.org/10.5281/zenodo.10792136" httpUri="https://zenodo.org/record/10792136/files/figure.png" pageId="2" pageNumber="277">Figs 1B</figureCitation>
,
<figureCitation id="132B4DC8FF98FF8CFE0CF883FE6B0707" box="[407,440,1863,1889]" captionStart="FIGURE 3" captionStartId="4.[151,250,1512,1536]" captionTargetBox="[151,1436,345,1489]" captionTargetId="figure-119@4.[151,1436,345,1489]" captionTargetPageId="4" captionText="FIGURE 3. Diagnostic features of Macronychus xuhaoi sp. nov., A–D, male; E, female. A. Metaventrite; B. Abdomen; C. Elytra; D–E. Protarsus. Note: red arrows in A: impressions of metaventrite; in B: carinae of ventrite I; in D: setal tufts of protarsus." figureDoi="http://doi.org/10.5281/zenodo.10792140" httpUri="https://zenodo.org/record/10792140/files/figure.png" pageId="2" pageNumber="277">3E</figureCitation>
,
<figureCitation id="132B4DC8FF98FF8CFE5CF883FDD50707" box="[455,518,1863,1889]" captionStart="FIGURE 4" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1436,181,1754]" captionTargetId="figure-19@5.[151,1436,181,1754]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Macronychus xuhaoi sp. nov., A–F, male; G–I, female. A. Aedeagus, dorsal view; B. Same, lateral view; C. Same, ventral view; D. part of aedeagus, dorsal view; E. Abdominal segment 9 and speculum gastrale; F–G. Seternite VIII; H. Ovipositor; I. stylus and apex of coxite. Note: black spot in A, setae of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10792142" httpUri="https://zenodo.org/record/10792142/files/figure.png" pageId="2" pageNumber="277">4G–I</figureCitation>
): externally similar to male, usually larger, protarsal segments (
<figureCitation id="132B4DC8FF98FF8CFB7AF883FAEB0707" box="[1249,1336,1863,1889]" captionStart="FIGURE 3" captionStartId="4.[151,250,1512,1536]" captionTargetBox="[151,1436,345,1489]" captionTargetId="figure-119@4.[151,1436,345,1489]" captionTargetPageId="4" captionText="FIGURE 3. Diagnostic features of Macronychus xuhaoi sp. nov., A–D, male; E, female. A. Metaventrite; B. Abdomen; C. Elytra; D–E. Protarsus. Note: red arrows in A: impressions of metaventrite; in B: carinae of ventrite I; in D: setal tufts of protarsus." figureDoi="http://doi.org/10.5281/zenodo.10792140" httpUri="https://zenodo.org/record/10792140/files/figure.png" pageId="2" pageNumber="277">Fig. 3E</figureCitation>
) lacking tuft of long setae at ventral surface. Apical margin of sternite VIII (
<figureCitation id="132B4DC8FF98FF8CFC10F8AFFC3007E3" box="[907,995,1899,1925]" captionStart="FIGURE 4" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1436,181,1754]" captionTargetId="figure-19@5.[151,1436,181,1754]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Macronychus xuhaoi sp. nov., A–F, male; G–I, female. A. Aedeagus, dorsal view; B. Same, lateral view; C. Same, ventral view; D. part of aedeagus, dorsal view; E. Abdominal segment 9 and speculum gastrale; F–G. Seternite VIII; H. Ovipositor; I. stylus and apex of coxite. Note: black spot in A, setae of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10792142" httpUri="https://zenodo.org/record/10792142/files/figure.png" pageId="2" pageNumber="277">Fig. 4G</figureCitation>
) weakly rounded. Ovipositor as in
<figureCitation id="132B4DC8FF98FF8CFAEBF8AFFF6F07CF" captionStart="FIGURE 4" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1436,181,1754]" captionTargetId="figure-19@5.[151,1436,181,1754]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Macronychus xuhaoi sp. nov., A–F, male; G–I, female. A. Aedeagus, dorsal view; B. Same, lateral view; C. Same, ventral view; D. part of aedeagus, dorsal view; E. Abdominal segment 9 and speculum gastrale; F–G. Seternite VIII; H. Ovipositor; I. stylus and apex of coxite. Note: black spot in A, setae of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10792142" httpUri="https://zenodo.org/record/10792142/files/figure.png" pageId="2" pageNumber="277">Fig. 4H</figureCitation>
, long and slender; stylus (
<figureCitation id="132B4DC8FF98FF8CFE72F84BFDEB07CF" box="[489,568,1935,1961]" captionStart="FIGURE 4" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1436,181,1754]" captionTargetId="figure-19@5.[151,1436,181,1754]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Macronychus xuhaoi sp. nov., A–F, male; G–I, female. A. Aedeagus, dorsal view; B. Same, lateral view; C. Same, ventral view; D. part of aedeagus, dorsal view; E. Abdominal segment 9 and speculum gastrale; F–G. Seternite VIII; H. Ovipositor; I. stylus and apex of coxite. Note: black spot in A, setae of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10792142" httpUri="https://zenodo.org/record/10792142/files/figure.png" pageId="2" pageNumber="277">Fig. 4I</figureCitation>
) weakly curved, apex with several small sensilla; coxite (
<figureCitation id="132B4DC8FF98FF8CFB56F84BFACF07CF" box="[1229,1308,1935,1961]" captionStart="FIGURE 4" captionStartId="5.[151,250,1778,1802]" captionTargetBox="[151,1436,181,1754]" captionTargetId="figure-19@5.[151,1436,181,1754]" captionTargetPageId="5" captionText="FIGURE 4. Diagnostic features of Macronychus xuhaoi sp. nov., A–F, male; G–I, female. A. Aedeagus, dorsal view; B. Same, lateral view; C. Same, ventral view; D. part of aedeagus, dorsal view; E. Abdominal segment 9 and speculum gastrale; F–G. Seternite VIII; H. Ovipositor; I. stylus and apex of coxite. Note: black spot in A, setae of aedeagus." figureDoi="http://doi.org/10.5281/zenodo.10792142" httpUri="https://zenodo.org/record/10792142/files/figure.png" pageId="2" pageNumber="277">Fig. 4I</figureCitation>
) with apex distinctly roundly broadened at outer margin, without sensilla; valvifer long, about twice as long as coxite, fibula with basal 1/3 distinctly expanded, finely curved at apical 1/3.
</paragraph>
<caption id="DF6F01C5FF99FF8DFF0CFAA4FD0305FA" ID-DOI="http://doi.org/10.5281/zenodo.10792138" ID-Zenodo-Dep="10792138" httpUri="https://zenodo.org/record/10792138/files/figure.png" pageId="3" pageNumber="278" startId="3.[151,250,1376,1400]" targetBox="[151,1436,181,1352]" targetPageId="3" targetType="figure">
<paragraph id="8BAF514DFF99FF8DFF0CFAA4FD0305FA" blockId="3.[151,1436,1376,1436]" pageId="3" pageNumber="278">
<emphasis id="B9648D5FFF99FF8DFF0CFAA4FEC1051E" bold="true" box="[151,274,1376,1400]" pageId="3" pageNumber="278">FIGURE 2.</emphasis>
Diagnostic features of
<taxonomicName id="4C102ACEFF99FF8DFE66FAA4FD02051E" authority="Jiang &amp; Chen, 2024" authorityName="Jiang &amp; Chen" authorityYear="2024" box="[509,721,1376,1400]" class="Insecta" family="Elmidae" genus="Macronychus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="xuhaoi" status="sp. nov.">
<emphasis id="B9648D5FFF99FF8DFE66FAA4FD02051E" box="[509,721,1376,1400]" italics="true" pageId="3" pageNumber="278">Macronychus xuhaoi</emphasis>
</taxonomicName>
<emphasis id="B9648D5FFF99FF8DFD4CFAA4FCF4051E" bold="true" box="[727,807,1376,1400]" pageId="3" pageNumber="278">
<taxonomicNameLabel id="A2573024FF99FF8DFD4CFAA4FCF4051E" box="[727,807,1376,1400]" pageId="3" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male.
<emphasis id="B9648D5FFF99FF8DFCF4FAA4FC5B051E" bold="true" box="[879,904,1376,1400]" pageId="3" pageNumber="278">A.</emphasis>
Head, dorsal view;
<emphasis id="B9648D5FFF99FF8DFBCAFAA4FBBB051E" bold="true" box="[1105,1128,1376,1400]" pageId="3" pageNumber="278">B.</emphasis>
Antenna;
<emphasis id="B9648D5FFF99FF8DFB4BFAA4FB3B051E" bold="true" box="[1232,1256,1376,1400]" pageId="3" pageNumber="278">C.</emphasis>
Pronotum, dorsal view;
<emphasis id="B9648D5FFF99FF8DFF4DFA40FF3D05FA" bold="true" box="[214,238,1412,1436]" pageId="3" pageNumber="278">D.</emphasis>
Prosternal process. Note: red arrow, scutellum.
</paragraph>
</caption>
<paragraph id="8BAF514DFF99FF8DFF5CFA0FFCDD066F" blockId="3.[151,1437,1482,2013]" pageId="3" pageNumber="278">
Measurements (n=10): CL:
<quantity id="4CE8FCA8FF99FF8DFD9AFA0EFD7B0583" box="[513,680,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="3" pageNumber="278" unit="mm" value="3.25" valueMax="3.4" valueMin="3.1">3.10–3.40 mm</quantity>
; HL:
<quantity id="4CE8FCA8FF99FF8DFD73FA0EFC5D0583" box="[744,910,1482,1509]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.3" metricValueMin="3.8" pageId="3" pageNumber="278" unit="mm" value="0.405" valueMax="0.43" valueMin="0.38">0.38–0.43 mm</quantity>
, HW:
<quantity id="4CE8FCA8FF99FF8DFC4CFA0EFBAD0583" box="[983,1150,1482,1509]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="5.8" metricValueMin="5.3" pageId="3" pageNumber="278" unit="mm" value="0.5549999999999999" valueMax="0.58" valueMin="0.53">0.53–0.58 mm</quantity>
; PL:
<quantity id="4CE8FCA8FF99FF8DFB21FA0EFA8C0583" box="[1210,1375,1482,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.05" metricValueMin="0.95" pageId="3" pageNumber="278" unit="mm" value="1.0" valueMax="1.05" valueMin="0.95">0.95–1.05 mm</quantity>
, PW: 0.90–1.00 mm; EL:
<quantity id="4CE8FCA8FF99FF8DFEE1FA2AFDCC066F" box="[378,543,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="2.35" metricValueMin="2.15" pageId="3" pageNumber="278" unit="mm" value="2.25" valueMax="2.35" valueMin="2.15">2.15–2.35 mm</quantity>
, EW:
<quantity id="4CE8FCA8FF99FF8DFDF8FA2AFCDA066F" box="[611,777,1518,1545]" metricMagnitude="-3" metricUnit="m" metricValue="1.225" metricValueMax="1.3" metricValueMin="1.15" pageId="3" pageNumber="278" unit="mm" value="1.225" valueMax="1.3" valueMin="1.15">1.15–1.30 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A02C6FF99FF8DFF5CF9D6FAF0064A" box="[199,1315,1554,1581]" pageId="3" pageNumber="278" type="distribution">
<paragraph id="8BAF514DFF99FF8DFF5CF9D6FAF0064A" blockId="3.[151,1437,1482,2013]" box="[199,1315,1554,1581]" pageId="3" pageNumber="278">
<emphasis id="B9648D5FFF99FF8DFF5CF9D6FEB2064A" bold="true" box="[199,353,1554,1580]" pageId="3" pageNumber="278">Distribution.</emphasis>
China: Only known from
<typeStatus id="54ABEFEFFF99FF8DFD12F9D7FD6A064B" box="[649,697,1555,1581]" pageId="3" pageNumber="278">type</typeStatus>
locality: Motuo County, Linzhi City,
<collectingRegion id="49D49FAFFF99FF8DFBFBF9D6FACD064A" box="[1120,1310,1554,1580]" country="China" name="Xizang" pageId="3" pageNumber="278">Xizang Province</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C30A02C6FF99FF8DFF5CF9F2FC9A0613" pageId="3" pageNumber="278" type="biology_ecology">
<paragraph id="8BAF514DFF99FF8DFF5CF9F2FC9A0613" blockId="3.[151,1437,1482,2013]" pageId="3" pageNumber="278">
<emphasis id="B9648D5FFF99FF8DFF5CF9F2FEF40636" bold="true" box="[199,295,1590,1616]" pageId="3" pageNumber="278">Biology.</emphasis>
All adults were collected by light trap near a hotel of Muotuo County (Linzhi City,
<collectingRegion id="49D49FAFFF99FF8DFB42F9F2FA4B0636" box="[1241,1432,1590,1616]" country="China" name="Xizang" pageId="3" pageNumber="278">Xizang Province</collectingRegion>
;
<figureCitation id="132B4DC8FF99FF8DFF0CF99EFF3C0613" box="[151,239,1626,1653]" captionStart="FIGURE 5" captionStartId="6.[151,250,1660,1684]" captionTargetBox="[164,1423,249,1637]" captionTargetId="figure-37@6.[164,1423,181,1637]" captionTargetPageId="6" captionText="FIGURE 5. A. The location of light trap in type locality of Macronychus xuhaoi sp. nov.; B. Water environment near the location of light trap. (both were taken by Dr. Hao Xu)." figureDoi="http://doi.org/10.5281/zenodo.10792146" httpUri="https://zenodo.org/record/10792146/files/figure.png" pageId="3" pageNumber="278">Fig. 5A</figureCitation>
), the adults might live in the nearby stream (
<figureCitation id="132B4DC8FF99FF8DFD7FF99EFCEF0613" box="[740,828,1626,1653]" captionStart="FIGURE 5" captionStartId="6.[151,250,1660,1684]" captionTargetBox="[164,1423,249,1637]" captionTargetId="figure-37@6.[164,1423,181,1637]" captionTargetPageId="6" captionText="FIGURE 5. A. The location of light trap in type locality of Macronychus xuhaoi sp. nov.; B. Water environment near the location of light trap. (both were taken by Dr. Hao Xu)." figureDoi="http://doi.org/10.5281/zenodo.10792146" httpUri="https://zenodo.org/record/10792146/files/figure.png" pageId="3" pageNumber="278">Fig. 5B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30A02C6FF99FF8DFF5CF9BAFCE506DA" pageId="3" pageNumber="278" type="etymology">
<paragraph id="8BAF514DFF99FF8DFF5CF9BAFCE506DA" blockId="3.[151,1437,1482,2013]" pageId="3" pageNumber="278">
<emphasis id="B9648D5FFF99FF8DFF5CF9BAFE9D06FE" bold="true" box="[199,334,1662,1688]" pageId="3" pageNumber="278">Etymology.</emphasis>
The species is named in honor of Dr. Hao Xu (Mianyang Normal University,
<collectingRegion id="49D49FAFFF99FF8DFB30F9BAFADB06FE" box="[1195,1288,1662,1688]" country="China" name="Sichuan" pageId="3" pageNumber="278">Sichuan</collectingRegion>
,
<collectingCountry id="F30711DDFF99FF8DFA88F9BAFA8906FE" box="[1299,1370,1662,1688]" name="China" pageId="3" pageNumber="278">China</collectingCountry>
), who collected the new species and donated it to us for this study.
</paragraph>
</subSubSection>
<subSubSection id="C30A02C6FF99FF8AFF5CF902FE21079F" lastPageId="4" lastPageNumber="279" pageId="3" pageNumber="278" type="diagnosis">
<paragraph id="8BAF514DFF99FF8DFF5CF902FB11072A" blockId="3.[151,1437,1482,2013]" pageId="3" pageNumber="278">
<emphasis id="B9648D5FFF99FF8DFF5CF902FE330686" bold="true" box="[199,480,1734,1760]" pageId="3" pageNumber="278">Comparative diagnosis.</emphasis>
<taxonomicName id="4C102ACEFF99FF8DFE7EF902FD1C0686" authority="Jiang &amp; Chen, 2024" authorityName="Jiang &amp; Chen" authorityYear="2024" box="[485,719,1734,1760]" class="Insecta" family="Elmidae" genus="Macronychus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="xuhaoi" status="sp. nov.">
<emphasis id="B9648D5FFF99FF8DFE7EF902FD1C0686" box="[485,719,1734,1760]" italics="true" pageId="3" pageNumber="278">Macronychus xuhaoi</emphasis>
</taxonomicName>
<emphasis id="B9648D5FFF99FF8DFD4EF902FCFE0686" bold="true" box="[725,813,1734,1760]" pageId="3" pageNumber="278">
<taxonomicNameLabel id="A2573024FF99FF8DFD4EF902FCFE0686" box="[725,813,1734,1760]" pageId="3" pageNumber="278" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
could be placed in the
<emphasis id="B9648D5FFF99FF8DFBB1F902FA4E0686" box="[1066,1437,1734,1760]" italics="true" pageId="3" pageNumber="278">
Macrynychus
<taxonomicName id="4C102ACEFF99FF8DFB5CF902FA4E0686" box="[1223,1437,1734,1760]" class="Insecta" family="Elmidae" genus="Macrynychus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="quadriluberculatus">quadriluberculatus</taxonomicName>
</emphasis>
species group based on the following characters: 1) median keel of pronotum flat with sides slightly declivous; 2) surface of metasternum covered with sparse macropunctures, each puncture bearing a long seta; 3) carinae of ventrite I not reaching posterior margin of ventrite; 4) femora moderately enlarged, not clavate.
</paragraph>
<paragraph id="8BAF514DFF99FF8AFF5CF892FE700127" blockId="3.[151,1437,1482,2013]" lastBlockId="4.[151,1436,151,321]" lastPageId="4" lastPageNumber="279" pageId="3" pageNumber="278">
<taxonomicName id="4C102ACEFF99FF8DFF5CF892FDED0716" authorityName="Müller" authorityYear="1806" box="[199,574,1878,1904]" class="Insecta" family="Elmidae" genus="Macrynychus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="quadrituberculatus">
<emphasis id="B9648D5FFF99FF8DFF5CF892FDED0716" box="[199,574,1878,1904]" italics="true" pageId="3" pageNumber="278">Macrynychus quadrituberculatus</emphasis>
</taxonomicName>
species group previously included only one widely-distributed species,
<taxonomicName id="4C102ACEFF99FF8DFAE5F893FEBE07F2" authorityName="Müller" authorityYear="1806" class="Insecta" family="Elmidae" genus="Macrynychus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="278" phylum="Arthropoda" rank="species" species="quadrituberculatus">
<emphasis id="B9648D5FFF99FF8DFAE5F893FEBE07F2" italics="true" pageId="3" pageNumber="278">M. quadrituberculatus</emphasis>
</taxonomicName>
. The new species can be easily distinguished from it by the following characters: 1) admedian prebasal gibbosity of pronotum much larger; 2) elytra without impression at anterior part; 3) median lobe of aedeagus weakly gradually dilated from base to near middle, with two pairs of sclerotizations, two of them much smaller and located near base, other two larger, located at 1/3 of the median lobe (vs. median lobe of aedeagus weakly get narrowed for base to middle, with only a pair of sclerotizations located at base in
<taxonomicName id="4C102ACEFF9EFF8AFBF5FF7FFAB800B3" box="[1134,1387,187,213]" class="Insecta" family="Elmidae" genus="Macrynychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="quadriluberculatus">
<emphasis id="B9648D5FFF9EFF8AFBF5FF7FFAB800B3" box="[1134,1387,187,213]" italics="true" pageId="4" pageNumber="279">M. quadriluberculatus</emphasis>
</taxonomicName>
); 4) posterior margin of female stemite VIII weakly rounded (vs. emarginated or near straight in
<taxonomicName id="4C102ACEFF9EFF8AFB0AFF1BFA58009F" box="[1169,1419,223,249]" class="Insecta" family="Elmidae" genus="Macrynychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="quadriluberculatus">
<emphasis id="B9648D5FFF9EFF8AFB0AFF1BFA58009F" box="[1169,1419,223,249]" italics="true" pageId="4" pageNumber="279">M. quadriluberculatus</emphasis>
</taxonomicName>
); 5) fibula distinctly expanded at basal 1/3, finely curved near apex (vs. only expanded at apex and straight at base in
<taxonomicName id="4C102ACEFF9EFF8AFF0CFEE3FE400127" box="[151,403,295,321]" class="Insecta" family="Elmidae" genus="Macrynychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="quadriluberculatus">
<emphasis id="B9648D5FFF9EFF8AFF0CFEE3FE400127" box="[151,403,295,321]" italics="true" pageId="4" pageNumber="279">M. quadriluberculatus</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="DF6F01C5FF9EFF8AFF0CFA2CFF28062E" ID-DOI="http://doi.org/10.5281/zenodo.10792140" ID-Zenodo-Dep="10792140" httpUri="https://zenodo.org/record/10792140/files/figure.png" pageId="4" pageNumber="279" startId="4.[151,250,1512,1536]" targetBox="[151,1436,345,1489]" targetPageId="4" targetType="figure">
<paragraph id="8BAF514DFF9EFF8AFF0CFA2CFF28062E" blockId="4.[151,1436,1512,1608]" pageId="4" pageNumber="279">
<emphasis id="B9648D5FFF9EFF8AFF0CFA2CFEC60666" bold="true" box="[151,277,1512,1536]" pageId="4" pageNumber="279">FIGURE 3.</emphasis>
Diagnostic features of
<taxonomicName id="4C102ACEFF9EFF8AFD90FA2DFD310667" authority="Jiang &amp; Chen, 2024" authorityName="Jiang &amp; Chen" authorityYear="2024" box="[523,738,1513,1537]" class="Insecta" family="Elmidae" genus="Macronychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="xuhaoi" status="sp. nov.">
<emphasis id="B9648D5FFF9EFF8AFD90FA2DFD310667" box="[523,738,1513,1537]" italics="true" pageId="4" pageNumber="279">Macronychus xuhaoi</emphasis>
</taxonomicName>
<emphasis id="B9648D5FFF9EFF8AFD71FA2DFCEE0667" bold="true" box="[746,829,1513,1537]" pageId="4" pageNumber="279">
<taxonomicNameLabel id="A2573024FF9EFF8AFD71FA2DFCEE0667" box="[746,829,1513,1537]" pageId="4" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, A–D, male; E, female.
<emphasis id="B9648D5FFF9EFF8AFBA4FA2CFB8B0666" bold="true" box="[1087,1112,1512,1536]" pageId="4" pageNumber="279">A.</emphasis>
Metaventrite;
<emphasis id="B9648D5FFF9EFF8AFB6AFA2DFADA0667" bold="true" box="[1265,1289,1513,1537]" pageId="4" pageNumber="279">B.</emphasis>
Abdomen;
<emphasis id="B9648D5FFF9EFF8AFA1FFA2CFA4F0666" bold="true" box="[1412,1436,1512,1536]" pageId="4" pageNumber="279">C.</emphasis>
Elytra;
<emphasis id="B9648D5FFF9EFF8AFF7EF9C9FECF0643" bold="true" box="[229,284,1549,1573]" pageId="4" pageNumber="279">D–E.</emphasis>
Protarsus. Note: red arrows in A: impressions of metaventrite; in B: carinae of ventrite I; in D: setal tufts of protarsus.
</paragraph>
</caption>
<paragraph id="8BAF514DFF9EFF8AFF5CF9B3FDCC070F" blockId="4.[151,1436,1655,2041]" pageId="4" pageNumber="279">
The new species is similar to
<taxonomicName id="4C102ACEFF9EFF8AFD83F9B3FB8606F7" authority="Ciampor &amp; Kodada 1998" authorityName="Ciampor &amp; Kodada" authorityYear="1998" box="[536,1109,1655,1683]" class="Insecta" family="Elmidae" genus="Macronychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="reticulatus">
<emphasis id="B9648D5FFF9EFF8AFD83F9B3FCFC06F7" box="[536,815,1655,1681]" italics="true" pageId="4" pageNumber="279">Macronychus reticulatus</emphasis>
<bibRefCitation id="EF812CBCFF9EFF8AFCADF9BCFB8606F7" author="Ciampor, F. &amp; Kodada, J." box="[822,1109,1655,1683]" pageId="4" pageNumber="279" pagination="219 - 287" refId="ref3044" refString="Ciampor, F. &amp; Kodada, J. (1998) Elmidae: I. Taxonomic revision of the genus Macronychus Muller (Coleoptera). In: Jach, M. A. &amp; Ji, L. (Eds.), Water Beetles of China. Vol. II. Zoologisch-Botanische Gesellschaft und Wiener Coleopterologenverein, Wien: pp. 219 - 287." type="book chapter" year="1998">
Čiampor &amp; Kodada
<quantity id="4CE8FCA8FF9EFF8AFB86F9B3FB8606F7" box="[1053,1109,1655,1681]" metricMagnitude="1" metricUnit="m" metricValue="5.07492" pageId="4" pageNumber="279" unit="in" value="1998.0">1998</quantity>
</bibRefCitation>
</taxonomicName>
in habitus character, like the moderately elongated body and legs.
<taxonomicName id="4C102ACEFF9EFF8AFDA8F958FD7006D3" authority="Jiang &amp; Chen, 2024" authorityName="Jiang &amp; Chen" authorityYear="2024" box="[563,675,1691,1717]" class="Insecta" family="Elmidae" genus="Macronychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="xuhaoi" status="sp. nov.">
<emphasis id="B9648D5FFF9EFF8AFDA8F958FD7006D3" box="[563,675,1691,1717]" italics="true" pageId="4" pageNumber="279">M. xuhaoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2573024FF9EFF8AFD32F95FFD2C06D3" box="[681,767,1691,1717]" pageId="4" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from the latter species by the following characters:1) body larger,&gt;
<quantity id="4CE8FCA8FF9EFF8AFE4CF97BFDE206BF" box="[471,561,1727,1754]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="279" unit="mm" value="2.5">2.5 mm</quantity>
; 2) femora slender and weakly clavated; 3) pronotum widest at basal 1/4 (vs. widest at basal 1/3); 4) median lobe of aedeagus not narrowed between base to middle in lateral view, with two pairs of sclerotizations (vs. median lob of aedeagus distinctly narrowed at basal 1/4, with one pair of sclerotizations located at base); 5) spiculum gastrale with base extended (vs. short and not extended at base); 6) apex of preterminal segment of ovipositor not bilobate.
</paragraph>
<paragraph id="8BAF514DFF9EFF8AFF5CF8B7FE21079F" blockId="4.[151,1436,1655,2041]" pageId="4" pageNumber="279">
<taxonomicName id="4C102ACEFF9EFF8AFF5CF8B7FE6207EB" authority="Jiang &amp; Chen, 2024" authorityName="Jiang &amp; Chen" authorityYear="2024" box="[199,433,1907,1933]" class="Insecta" family="Elmidae" genus="Macronychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="xuhaoi" status="sp. nov.">
<emphasis id="B9648D5FFF9EFF8AFF5CF8B7FE8D07EB" box="[199,350,1907,1933]" italics="true" pageId="4" pageNumber="279">Macronychus</emphasis>
xuhaoi
</taxonomicName>
<emphasis id="B9648D5FFF9EFF8AFE22F8B7FDC107EB" bold="true" box="[441,530,1907,1933]" pageId="4" pageNumber="279">
<taxonomicNameLabel id="A2573024FF9EFF8AFE22F8B7FDC107EB" box="[441,530,1907,1933]" pageId="4" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is also more or less similar to
<taxonomicName id="4C102ACEFF9EFF8AFCFCF8B0FB6A07EB" authority="Deleve, 1968" authorityName="Deleve" authorityYear="1968" box="[871,1209,1907,1933]" class="Insecta" family="Elmidae" genus="Macronychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="vietnamensis">
<emphasis id="B9648D5FFF9EFF8AFCFCF8B0FBCD07EB" box="[871,1054,1908,1933]" italics="true" pageId="4" pageNumber="279">M. vietnamensis</emphasis>
Delève, 1968
</taxonomicName>
and
<taxonomicName id="4C102ACEFF9EFF8AFB6BF8B0FED807D4" authority="Say, 1825" authorityName="Say" authorityYear="1825" class="Insecta" family="Elmidae" genus="Macronychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="levanidovae">
<emphasis id="B9648D5FFF9EFF8AFB6BF8B0FA4F07EB" box="[1264,1436,1907,1933]" italics="true" pageId="4" pageNumber="279">M. levanidovae</emphasis>
Say, 1825
</taxonomicName>
, which are included in
<taxonomicName id="4C102ACEFF9EFF8AFD86F85CFD6207D7" authorityName="Say" authorityYear="1825" box="[541,689,1943,1969]" class="Insecta" family="Elmidae" genus="Macronychus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="279" phylum="Arthropoda" rank="species" species="glabratus">
<emphasis id="B9648D5FFF9EFF8AFD86F85CFD6207D7" box="[541,689,1943,1969]" italics="true" pageId="4" pageNumber="279">M. glabratus</emphasis>
</taxonomicName>
species group. The new species can be distinguished from them by the following characters: 1) head and tibiae without plastron setae; 2) median lobe of aedeagus with two pairs sclerotizations (vs. with three).
</paragraph>
</subSubSection>
</treatment>
</document>
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