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<document id="3D8266EF23AAE5FEF9184B9C0C4521BF" ID-DOI="10.11646/zootaxa.3633.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5261931" ID-ZooBank="60F0956C-A1F5-44CA-9D14-8C04343AD01F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629849986103" checkinUser="felipe" docAuthor="Tedesco, Anazélia M. &amp; Aguiar, Alexandre P." docDate="2013" docId="03B987AEFFEAFF8F0AE1FD3DFEF4FF51" docLanguage="en" docName="zt03633p138.pdf" docOrigin="Zootaxa 3633 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.3633.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Toechorychus zulius Tedesco &amp; Aguiar 2013, sp. nov." docType="treatment" docVersion="3" lastPageNumber="77" masterDocId="FF80FFD6FFA0FFC30A76FFC1FF88FFE0" masterDocTitle="&lt;strong&gt; Phylogeny and Revision of &lt;em&gt; Toechorychus &lt;/ em&gt; Townes (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of thirty-five new species &lt;/ strong&gt;" masterLastPageNumber="138" masterPageNumber="1" pageNumber="75" updateTime="1699120729359" updateUser="plazi">
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<mods:title id="4E4A5E22DE0B2C5192500FB6CB701565">&lt;strong&gt; Phylogeny and Revision of &lt;em&gt; Toechorychus &lt;/ em&gt; Townes (Hymenoptera, Ichneumonidae, Cryptinae), with descriptions of thirty-five new species &lt;/ strong&gt;</mods:title>
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<mods:namePart id="C1862BC3BEAA1CCC3B1F4E1102284EDF">Tedesco, Anazélia M.</mods:namePart>
<mods:affiliation id="D5B6E1D01F9C5B2ABE6AD69F2921D447">Universidade Federal do Espírito Santo, Departamento de Ciências Biológicas, Zoologia, Av. Fernando Ferrari 514 - Goiabeiras,</mods:affiliation>
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<mods:namePart id="84E1FFE3C88B041F570001B52CFB6BE6">Aguiar, Alexandre P.</mods:namePart>
<mods:affiliation id="B958E2C1D12F1877F12E477A6FEF52B8">Universidade Federal do Espírito Santo, Departamento de Ciências Biológicas, Zoologia, Av. Fernando Ferrari 514 - Goiabeiras,</mods:affiliation>
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<mods:date id="AE0DD9521094578FDCF45DCC479F486E">2013</mods:date>
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<heading id="D0E781D4FFEAFF890AE1FD3DFDEBFCF5" bold="true" box="[151,611,763,790]" fontSize="11" level="2" pageId="74" pageNumber="75" reason="4">
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<taxonomicName id="4C104D3BFFEAFF890AE1FD3DFE71FCF5" authority="Tedesco" authorityName="Tedesco &amp; Aguiar" authorityYear="2013" box="[151,505,763,790]" class="Insecta" family="Ichneumonidae" genus="Toechorychus" kingdom="Animalia" order="Hymenoptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="zulius" status="sp. nov.">
<emphasis id="B964EAAAFFEAFF890AE1FD3DFE05FCF6" bold="true" box="[151,397,764,790]" italics="true" pageId="74" pageNumber="75">Toechorychus zulius</emphasis>
Tedesco
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, 
<taxonomicNameLabel id="A25757D1FFEAFF890870FD3AFDEBFCF5" box="[518,611,763,789]" pageId="74" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BAF36B8FFEAFF890AE1FCDFFDA9FCD6" blockId="74.[151,611,763,822]" box="[151,545,798,822]" pageId="74" pageNumber="75">
(
<figureCitation id="132B2A3DFFEAFF890AE9FCDFFF7DFCD6" box="[159,245,798,822]" captionStart="FIGURES 28–31" captionStartId="109.[151,264,1888,1911]" captionTargetBox="[151,1435,193,1867]" captionTargetId="figure-62@109.[151,1435,193,1867]" captionTargetPageId="109" captionText="FIGURES 28–31. Lateral pictures of Toechorychus. 28, T. ondensis sp. nov., holotype. 29, T. albimaculatus Taschenberg. 30, T. darienus sp. nov., holotype. 31, T. zulius sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261961" httpUri="https://zenodo.org/record/5261961/files/figure.png" pageId="74" pageNumber="75">Figs 31</figureCitation>
, 
<figureCitation id="132B2A3DFFEAFF890B77FCDFFE97FCD6" box="[257,287,798,822]" captionStart="FIGURES 45–54. 45" captionStartId="113.[151,264,1758,1781]" captionTargetBox="[151,1435,240,1737]" captionTargetId="figure-226@113.[151,1435,179,1737]" captionTargetPageId="113" captionText="FIGURES 45–54. 45, T. callangus sp. nov., holotype, ovipositor in dorsal view, showing dorsal valve with posterior V-shaped carina, a new synapomorphy discovered for the genus. 46, T. vinhaticus sp. nov., holotype, pronotum in lateral view, lateral portion of collar dorsally carinated. 47, T. morelus sp. nov., holotype, pronotum in lateral view, to show lateral portion of collar dorsally swollen. 48, T. fluminensis sp. nov., holotype, to show hind margin of metanotum without teeth or carinae. 49, T. jatainus sp. nov., holotype, to show hind margin of metanotum with two lateral carinae. 50, T. amapaeus sp. nov., holotype, head in posterior view, to show occipital carina dorsally absent. 51, T. paramaribus sp. nov., holotype, to show anterior margin of propodeum with two lateral teeth. 52, T. melgassus sp. nov., holotype, head in posterior view, to show occipital carina dorsally conspicuous. 53, T. zulius sp. nov., holotype, to show first tergite with two basolateral teeth. 54, T. nourageus sp. nov., holotype, to show first tergite without basolateral teeth." figureDoi="http://doi.org/10.5281/zenodo.5261973" httpUri="https://zenodo.org/record/5261973/files/figure.png" pageId="74" pageNumber="75">53</figureCitation>
, 
<figureCitation id="132B2A3DFFEAFF890B5DFCDFFEC2FCD6" box="[299,330,798,822]" captionStart="FIGURES 70–84" captionStartId="115.[151,264,1840,1863]" captionTargetBox="[151,1435,193,1819]" captionTargetId="figure-173@115.[151,1435,193,1819]" captionTargetPageId="115" captionText="FIGURES 70–84. Head, frontal view. 70, T. martinus sp. nov., holotype. 71, T. morelus sp. nov., holotype. 72, T. zulius sp. nov., holotype. 73, T. amazonensis sp. nov., holotype. 74, T. paramaribus sp. nov., holotype. 75, T. heredius sp. nov., holotype. 76, T. linaresius sp. nov., holotype. 77, T. marowijnus sp. nov., holotype. 78, T. darienus sp. nov., holotype. 79, T. kawus sp. nov., holotype. 80, T. jatainus sp. nov., holotype. 81, T. taperinus sp. nov., holotype. 82, T. teutonius sp. nov., holotype. 83, T. calius sp. nov., holotype. 84, T. surinamus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261981" httpUri="https://zenodo.org/record/5261981/files/figure.png" pageId="74" pageNumber="75">72</figureCitation>
, 
<figureCitation id="132B2A3DFFEAFF890B23FCDFFEFBFCD6" box="[341,371,798,822]" captionStart="FIGURES 91–96" captionStartId="117.[151,264,1674,1697]" captionTargetBox="[151,1435,193,1653]" captionTargetId="figure-84@117.[151,1435,193,1653]" captionTargetPageId="117" captionText="FIGURES 91–96. Mesoscutum, dorsal view. 91, T. marcapatus sp. nov., holotype. 92, T. darienus sp. nov., holotype. 93, T. coaracius sp. nov., holotype. 94, T. itapuensis sp. nov., holotype. 95, T. martinus sp. nov., holotype. 96, T. zulius sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261985" httpUri="https://zenodo.org/record/5261985/files/figure.png" pageId="74" pageNumber="75">96</figureCitation>
, 
<figureCitation id="132B2A3DFFEAFF890B09FCDFFE22FCD6" box="[383,426,798,822]" captionStart="FIGURE 105–118" captionStartId="119.[151,250,1884,1907]" captionTargetBox="[164,1428,179,1877]" captionTargetId="figure-155@119.[164,1428,179,1877]" captionTargetPageId="119" captionText="FIGURE 105–118. T. albimaculatus Taschenberg, fore and hind wings. FIGURES 106–118. Fore wing. 106, T. cassunungae Brauns. 107, T. stramineus Taschenberg. 108, T. ondensis sp. nov., holotype. 109, T. amapaeus sp. nov., holotype. 110, T. zulius sp. nov., holotype. 111, T. amazonensis sp. nov., holotype. 112, T. itapuensis sp. nov., holotype. 113, T. paramaribus sp. nov., holotype. 114, T. guarapuavus sp. nov., holotype. 115, T. melgassus sp. nov., holotype. 116, T. coaracius sp. nov., holotype. 117, T. marcapatus sp. nov., holotype. 118, T. barticus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261991" httpUri="https://zenodo.org/record/5261991/files/figure.png" pageId="74" pageNumber="75">110</figureCitation>
, 
<figureCitation id="132B2A3DFFEAFF890BC0FCDFFE6AFCD6" box="[438,482,798,822]" captionStart="FIGURES 157–162" captionStartId="126.[151,264,1672,1695]" captionTargetBox="[151,1435,193,1651]" captionTargetId="figure-85@126.[151,1435,193,1651]" captionTargetPageId="126" captionText="FIGURES 157–162. Propodeum, dorsal view. 157, T. kawus sp. nov., holotype. 158, T. zulius sp. nov., holotype. 159, T. taperinus sp. nov., holotype. 160, T. heredius sp. nov., holotype. 161, T. bombuscarus sp. nov., holotype. 162, T. darienus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5262009" httpUri="https://zenodo.org/record/5262009/files/figure.png" pageId="74" pageNumber="75">158</figureCitation>
, 
<figureCitation id="132B2A3DFFEAFF890B98FCDFFD91FCD6" box="[494,537,798,822]" captionStart="FIGURES 187–199" captionStartId="129.[151,264,1820,1843]" captionTargetBox="[151,1436,193,1799]" captionTargetId="figure-158@129.[151,1436,193,1799]" captionTargetPageId="129" captionText="FIGURES 187–199. Hind coxae and metasoma, dorsal view, to show color pattern. 187, T. fluminensis sp. nov., holotype. 188, T. morelus sp. nov., holotype. 189, T. itapuensis sp. nov., holotype. 190, T. calius sp. nov., holotype. 191, T. barticus sp. nov., holotype. 192, T. tumazulus sp. nov., holotype. 193, T. linaresius sp. nov., holotype. 194, T. catarinus sp. nov., holotype. 195, T. martinus sp. nov., holotype. 196, T. stramineus Taschenberg. 197, T. pirrus sp. nov., holotype. 198, T. zulius sp. nov., holotype. 199, T. marowijnus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5262019" httpUri="https://zenodo.org/record/5262019/files/figure.png" pageId="74" pageNumber="75">198</figureCitation>
)
</paragraph>
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Description. 
<typeStatus id="54AB881AFFEAFF890B51FCA7FE07FC9E" box="[295,399,870,894]" pageId="74" pageNumber="75" type="holotype">Holotype</typeStatus>
FEMALE. Fore wing length 
<quantity id="4CE89B5DFFEAFF8908ADFCA7FCCAFC9E" box="[731,834,870,894]" metricMagnitude="-3" metricUnit="m" metricValue="6.71" pageId="74" pageNumber="75" unit="mm" value="6.71">6.71 mm</quantity>
.
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<paragraph id="8BAF36B8FFEAFF890AB1FC4AFB04FACE" blockId="74.[151,1436,870,2010]" pageId="74" pageNumber="75">
<emphasis id="B964EAAAFFEAFF890AB1FC4AFE8CFC43" box="[199,260,907,931]" italics="true" pageId="74" pageNumber="75">Head</emphasis>
. Mandible 1.79 as long as basal width, moderately pilose; ventral margin slightly projected as crest; dorsal tooth distinctly longer than ventral tooth. Clypeus 1.57 as wide as high, subtriangular, minutely strigulate, apically smooth; apex 1.35 as long as base, slightly but distinctly convex; apical margin sharp, medially slightly concave. Supraclypeal area entirely strigate, with scarce punctures, moderately pilose, medially slightly prominent; between antennal foramens with three, complete, U-shaped carinae; radicle foveolate. Antenna with 26 flagellomeres; white band starting at flagellomere 5, reaching flagellomere 10; flagellum somewhat stout; subapical flagellomeres slightly flattened. Supra-antennal area with very faint but distinct median longitudinal carina, medially without longitudinal elevation, near antennal sockets striate; dorsal half medially strigulate. Paraocular area smooth. Vertex rugulose around ocelli, without sulcus; gena and vertex behind ocelli smooth, glabrous; gena in lateral view at level of dorsal portion of occipital carina narrow, ventrally wide; occipital carina stout, dorsally absent, ventrally slightly projected as crest, reaching hypostomal carina far from mandible base; hypostomal carina projected as crest; malar space 1.15 as long as basal width of mandible.
</paragraph>
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<taxonomicName id="4C104D3BFFEAFF890AB1FAFAFEB7FAB2" box="[199,319,1339,1362]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">
<emphasis id="B964EAAAFFEAFF890AB1FAFAFEB7FAB2" box="[199,319,1339,1362]" italics="true" pageId="74" pageNumber="75">Mesosoma</emphasis>
</taxonomicName>
. 
<taxonomicName id="4C104D3BFFEAFF890B3BFAFAFE4FFAB2" box="[333,455,1339,1362]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="74" pageNumber="75" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
in dorsal view 2.23 as long as wide; in lateral view, mesosoma middle width 
<quantity id="4CE89B5DFFEAFF890F47FAFBFA10FAB2" box="[1329,1432,1338,1362]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" pageId="74" pageNumber="75" unit="mm" value="0.21">0.21 mm</quantity>
. Pronotum centrally smooth, latero-ventrally rugose behind collar, margin near mesopleuron ventrally corrugated; pronotal swelling smooth; collar dorso-laterally carinated, not swollen, anteriorly smooth; epomia absent. Mesoscutum densely pilose, lobes dorsally punctulate, laterally markedly corrugated or covered with sparse coarse punctures, central lobe without longitudinal sulcus or carina; notaulus moderately impressed, posteriorly abruptly curved, U-shaped. Scutoscutellar groove faintly corrugated; scutellum 1.2 as long as wide, smooth; scutellar carina complete, posteriorly closing; postscutellum 0.29 as long as wide; hind margin of metanotum with two lateral teeth. Subalar ridge somewhat elongate; mesopleuron dorsally medially markedly strigate, ventrally smooth; mesopleural groove corrugated; epicnemial carina stout, dorsally curving forwards; dorsal end of epicnemial carina reaching 0.2 of posterior margin of pronotum, behind epicnemial carina without rounded confused-rugulose area; sternaulus medially shallow, interrupted, reaching base of mid coxa, moderately sinuate, moderately impressed, faint from anterior 0.6 to 0.9, faintly corrugated; scrobe moderately deep, forming pit with sulcus. Mesothoracic venter punctate and strigulate; median portion of posterior transverse carina of mesothoracic venter short and straight, regularly shaped, without projections. Transverse furrow at base of propodeum laterally 0.08 as long as anterior portion of propodeum deep and narrow, very faintly corrugated, almost smooth; anterior margin of propodeum with two faint lateral teeth; propodeum 1.24 as long as wide medially; anterior portion of propodeum finely strigulate, medially without longitudinal carinae; lateral longitudinal carina of propodeum indistinct; posterior portion of propodeum strigate, progressively coarse toward posterior margin, medially with longitudinal carina until midlength; anterior transverse carina of propodeum complete, stout, straight; posterior transverse carina absent; propodeal spiracle 2.00 as long as wide; pleural carina of propodeum absent, apparently present because of sculpture patterns of lower division of metapleuron and propodeum; lower division of metapleuron very faintly strigulate, sparsely pilose, juxtacoxal carina short and stout.
</paragraph>
<paragraph id="8BAF36B8FFEBFF880AB1FEC4FC52FE4C" blockId="75.[151,1436,151,1904]" pageId="75" pageNumber="76">
<emphasis id="B964EAAAFFEBFF880AB1FEC4FE83FEFC" box="[199,267,261,284]" italics="true" pageId="75" pageNumber="76">Wings</emphasis>
. Fore wing: vein 2+3Rs almost straight; ramellus absent; vein 1M+Rs entirely irregular; vein 1cu-a straight, arising near vein 1M+Rs base, angle with vein M+Cu about 90°; vein 2Cu 0.76 as long as vein 2cu-a; vein 4Rs 0.92 as long as vein 4M, slightly sinuous, almost straight; cell 1+2Rs 0.96 as high as pterostigma, rectangular; vein 2M distinctly longer than vein 3M; vein 3M distinct, 0.35 as long as vein 
<specimenCount id="9D16FD31FFEBFF880E6DFEAEFBC2FE67" box="[1051,1098,367,391]" pageId="75" pageNumber="76" type="generic">2M.</specimenCount>
Hind wing: vein 1Cu 1.37 as long as vein cu-a; vein 2-1A reaching 0.89 of distance to posterior margin.
</paragraph>
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<emphasis id="B964EAAAFFEBFF880AB1FE78FF74FE30" box="[199,252,441,464]" italics="true" pageId="75" pageNumber="76">Legs</emphasis>
. Tibia with sparse short bristles; hind coxa globose.
</paragraph>
<paragraph id="8BAF36B8FFEBFF880AB1FE1CFEA3FD64" blockId="75.[151,1436,151,1904]" pageId="75" pageNumber="76">
<emphasis id="B964EAAAFFEBFF880AB1FE1CFEB4FE14" box="[199,316,477,500]" italics="true" pageId="75" pageNumber="76">Metasoma</emphasis>
. T1 0.53 as long as hind femur, with basolateral tooth; anteriorly smooth, posteriorly coriarious; spiracle at anterior 0.55, prominent; postpetiole dorsally, behind spiracles, markedly concave, deeply depressed; T2–8 minutely coriarious; centrally almost glabrous, laterally pilose; T2 0.87 as long as wide at apex; apex of T2 2.07 as wide as base. Dorsal valve of ovipositor with nodus indistinct; apex of lower valve with series of teeth; notch absent.
</paragraph>
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<emphasis id="B964EAAAFFEBFF880AB1FD51FE8FFD48" box="[199,263,656,680]" italics="true" pageId="75" pageNumber="76">Color</emphasis>
. Head and mesosoma yellow and dark brown, metasoma orange. Head: bright yellow (243,227,126); 0.8 of mandible, ventral half of malar space, apical half of clypeus, supraclypeal area ventrally, supra-antennal area, interocellar area, vertex and gena, except complete large orbital band, and occiput, dark brown (124,082,058); scape, pedicel, and flagellum brown, blackish toward apex; f6–9 entirely white, f10 almost entirely white, f5 partially white. 
<taxonomicName id="4C104D3BFFEBFF880B38FCE1FE44FCD7" box="[334,460,800,823]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="75" pageNumber="75" phylum="Arthropoda" rank="genus">Mesosoma</taxonomicName>
: brown with whitish marks (244,231,204); propleuron brown; pronotum dark brown except collar laterally and pronotal swelling, pale yellow; mesoscutum black except longitudinal pale yellow lanceolate mark at posterior 0.5 of central lobe; remainder of mesosoma dark brown, except for the following pale yellow marks: axillary carina, tegula apically, scutellum, postscutellum, large mark extending from subalar ridge to entire dorsal margin of sternaulus except around hypoepimeron, dorsal half of hypoepimeron, upper division of metapleuron, 0.5 dorsal of lower division of metapleuron, and two lateral longitudinal marks at posterior portion of propodeum, pale yellow. Legs: mostly brown; fore and mid coxae with large dorsal pale yellow spot; fore and mid trochanters pale yellow, posteriorly brown; fore and mid femora posteriorly with brown stripe; all tibia, and all t1–t4, yellow; hind coxa dark brown with two pale yellow marks, anterior one narrow, at basal 0.7, posterior one large and dorsal, almost reaching apex of coxa, basally wide; hind first trochanter mostly brownish, posteriorly yellow. Metasoma: mostly orange (221,126,036); T1 ferruginous (183,095,052), posterior 0.2 pale yellow; T2–8 and S2–8 uniformly orange. 0.7 dorsal of lower division of metapleuron yellow.
</paragraph>
<paragraph id="8BAF36B8FFEBFF880AB1FB0EFA8CFAEC" blockId="75.[151,1436,151,1904]" pageId="75" pageNumber="76">
<emphasis id="B964EAAAFFEBFF880AB1FB0EFEBFFB08" bold="true" box="[199,311,1231,1256]" pageId="75" pageNumber="76">Variation</emphasis>
. Brown mark at supraclypeal area laterally reaching antennal foramens. Epicnemial carina dorsally yellowish. Fore and mid femora posteriorly with pale brown stripe. Posterior 0.2 of T1 bright yellow.
</paragraph>
</subSubSection>
<subSubSection id="C30A6533FFEBFF880AB1FAD6FE00F994" pageId="75" pageNumber="76" type="discussion">
<paragraph id="8BAF36B8FFEBFF880AB1FAD6FE70F9AF" blockId="75.[151,1436,151,1904]" pageId="75" pageNumber="76">
<emphasis id="B964EAAAFFEBFF880AB1FAD6FEC0FAD0" bold="true" box="[199,328,1303,1328]" pageId="75" pageNumber="76">Comments</emphasis>
. Similar to 
<taxonomicName id="4C104D3BFFEBFF880BA8FAD8FDCEFAD0" authority="Tedesco &amp; Aguiar, 2013" authorityName="Tedesco &amp; Aguiar" authorityYear="2013" box="[478,582,1304,1328]" class="Insecta" family="Ichneumonidae" genus="Toechorychus" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="kawus" status="sp. nov.">
<emphasis id="B964EAAAFFEBFF880BA8FAD8FE63FAD0" box="[478,491,1305,1328]" italics="true" pageId="75" pageNumber="76">T</emphasis>
. 
<emphasis id="B964EAAAFFEBFF880876FAD9FDCEFAD0" box="[512,582,1304,1328]" italics="true" pageId="75" pageNumber="76">kawus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757D1FFEBFF880825FAD9FD39FACF" box="[595,689,1304,1328]" pageId="75" pageNumber="76" rank="species">
<emphasis id="B964EAAAFFEBFF880825FAD9FDE5FAD0" bold="true" box="[595,621,1304,1328]" pageId="75" pageNumber="76">sp</emphasis>
. 
<emphasis id="B964EAAAFFEBFF8808F7FAD9FD23FAD0" bold="true" box="[641,683,1304,1328]" pageId="75" pageNumber="76">nov</emphasis>
.
</taxonomicNameLabel>
, from which can be separated by having ventral half of gena posteriorly brown (
<emphasis id="B964EAAAFFEBFF880B06FAFCFE0FFAB4" box="[368,391,1341,1364]" italics="true" pageId="75" pageNumber="76">vs</emphasis>
. yellow in 
<taxonomicName id="4C104D3BFFEBFF880870FAFCFDE0FAB4" authority="Tedesco &amp; Aguiar, 2013" authorityName="Tedesco &amp; Aguiar" authorityYear="2013" box="[518,616,1340,1364]" class="Insecta" family="Ichneumonidae" genus="Toechorychus" kingdom="Animalia" order="Hymenoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="kawus" status="sp. nov.">
<emphasis id="B964EAAAFFEBFF880870FAFCFD9BFAB4" box="[518,531,1341,1364]" italics="true" pageId="75" pageNumber="76">T</emphasis>
. 
<emphasis id="B964EAAAFFEBFF880854FAFDFDE0FAB4" box="[546,616,1340,1364]" italics="true" pageId="75" pageNumber="76">kawus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757D1FFEBFF880819FAFDFD40FAB3" box="[623,712,1340,1364]" pageId="75" pageNumber="76" rank="species">
<emphasis id="B964EAAAFFEBFF880819FAFDFD01FAB4" bold="true" box="[623,649,1340,1364]" pageId="75" pageNumber="76">sp</emphasis>
. 
<emphasis id="B964EAAAFFEBFF8808EEFAFDFD4AFAB4" bold="true" box="[664,706,1340,1364]" pageId="75" pageNumber="76">nov</emphasis>
.
</taxonomicNameLabel>
; 
<figureCitation id="132B2A3DFFEBFF8808AEFAFDFCA4FAB4" box="[728,812,1340,1364]" captionStart="FIGURES 70–84" captionStartId="115.[151,264,1840,1863]" captionTargetBox="[151,1435,193,1819]" captionTargetId="figure-173@115.[151,1435,193,1819]" captionTargetPageId="115" captionText="FIGURES 70–84. Head, frontal view. 70, T. martinus sp. nov., holotype. 71, T. morelus sp. nov., holotype. 72, T. zulius sp. nov., holotype. 73, T. amazonensis sp. nov., holotype. 74, T. paramaribus sp. nov., holotype. 75, T. heredius sp. nov., holotype. 76, T. linaresius sp. nov., holotype. 77, T. marowijnus sp. nov., holotype. 78, T. darienus sp. nov., holotype. 79, T. kawus sp. nov., holotype. 80, T. jatainus sp. nov., holotype. 81, T. taperinus sp. nov., holotype. 82, T. teutonius sp. nov., holotype. 83, T. calius sp. nov., holotype. 84, T. surinamus sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.5261981" httpUri="https://zenodo.org/record/5261981/files/figure.png" pageId="75" pageNumber="76">Fig. 79</figureCitation>
); propleuron completely dark brown (
<emphasis id="B964EAAAFFEBFF880EAEFAFCFB67FAB4" box="[1240,1263,1341,1364]" italics="true" pageId="75" pageNumber="76">vs</emphasis>
. dorsally black and ventrally pale yellow); collar dorso-laterally carinated, not swollen, as in 
<figureCitation id="132B2A3DFFEBFF880E47FA9EFB0EFA97" box="[1073,1158,1375,1399]" captionStart="FIGURES 45–54. 45" captionStartId="113.[151,264,1758,1781]" captionTargetBox="[151,1435,240,1737]" captionTargetId="figure-226@113.[151,1435,179,1737]" captionTargetPageId="113" captionText="FIGURES 45–54. 45, T. callangus sp. nov., holotype, ovipositor in dorsal view, showing dorsal valve with posterior V-shaped carina, a new synapomorphy discovered for the genus. 46, T. vinhaticus sp. nov., holotype, pronotum in lateral view, lateral portion of collar dorsally carinated. 47, T. morelus sp. nov., holotype, pronotum in lateral view, to show lateral portion of collar dorsally swollen. 48, T. fluminensis sp. nov., holotype, to show hind margin of metanotum without teeth or carinae. 49, T. jatainus sp. nov., holotype, to show hind margin of metanotum with two lateral carinae. 50, T. amapaeus sp. nov., holotype, head in posterior view, to show occipital carina dorsally absent. 51, T. paramaribus sp. nov., holotype, to show anterior margin of propodeum with two lateral teeth. 52, T. melgassus sp. nov., holotype, head in posterior view, to show occipital carina dorsally conspicuous. 53, T. zulius sp. nov., holotype, to show first tergite with two basolateral teeth. 54, T. nourageus sp. nov., holotype, to show first tergite without basolateral teeth." figureDoi="http://doi.org/10.5281/zenodo.5261973" httpUri="https://zenodo.org/record/5261973/files/figure.png" pageId="75" pageNumber="76">Fig. 46</figureCitation>
(
<emphasis id="B964EAAAFFEBFF880EEDFAA0FB3AFA98" box="[1179,1202,1377,1400]" italics="true" pageId="75" pageNumber="76">vs</emphasis>
. rounded, distinctly swollen, as in 
<figureCitation id="132B2A3DFFEBFF880B48FA45FE1CFA7C" box="[318,404,1412,1436]" captionStart="FIGURES 45–54. 45" captionStartId="113.[151,264,1758,1781]" captionTargetBox="[151,1435,240,1737]" captionTargetId="figure-226@113.[151,1435,179,1737]" captionTargetPageId="113" captionText="FIGURES 45–54. 45, T. callangus sp. nov., holotype, ovipositor in dorsal view, showing dorsal valve with posterior V-shaped carina, a new synapomorphy discovered for the genus. 46, T. vinhaticus sp. nov., holotype, pronotum in lateral view, lateral portion of collar dorsally carinated. 47, T. morelus sp. nov., holotype, pronotum in lateral view, to show lateral portion of collar dorsally swollen. 48, T. fluminensis sp. nov., holotype, to show hind margin of metanotum without teeth or carinae. 49, T. jatainus sp. nov., holotype, to show hind margin of metanotum with two lateral carinae. 50, T. amapaeus sp. nov., holotype, head in posterior view, to show occipital carina dorsally absent. 51, T. paramaribus sp. nov., holotype, to show anterior margin of propodeum with two lateral teeth. 52, T. melgassus sp. nov., holotype, head in posterior view, to show occipital carina dorsally conspicuous. 53, T. zulius sp. nov., holotype, to show first tergite with two basolateral teeth. 54, T. nourageus sp. nov., holotype, to show first tergite without basolateral teeth." figureDoi="http://doi.org/10.5281/zenodo.5261973" httpUri="https://zenodo.org/record/5261973/files/figure.png" pageId="75" pageNumber="76">Fig. 47</figureCitation>
); mesothoracic venter completely dark brown (
<emphasis id="B964EAAAFFEBFF8809C5FA44FC42FA7C" box="[947,970,1413,1436]" italics="true" pageId="75" pageNumber="76">vs</emphasis>
. black with yellow longitudinal stripes); lower division of metapleuron almost smooth, very faintly strigulate (
<emphasis id="B964EAAAFFEBFF8809C9FA68FC5EFA20" box="[959,982,1449,1472]" italics="true" pageId="75" pageNumber="76">vs</emphasis>
. punctate and strigulate); fore and mid femora ventrally brownish (
<emphasis id="B964EAAAFFEBFF880BA3FA0CFE64FA04" box="[469,492,1485,1508]" italics="true" pageId="75" pageNumber="76">vs</emphasis>
. entirely orange); mid coxa brown with dorsal large yellow mark (
<emphasis id="B964EAAAFFEBFF880E9FFA0CFA88FA04" box="[1257,1280,1485,1508]" italics="true" pageId="75" pageNumber="76">vs</emphasis>
. yellow, with dorsal black mark at apical 0.7); hind coxa dark brown with two yellow stripes (
<emphasis id="B964EAAAFFEBFF880E62FA30FBA3F9E8" box="[1044,1067,1521,1544]" italics="true" pageId="75" pageNumber="76">vs</emphasis>
. yellow with three black marks); spiracle of T1 prominent (
<emphasis id="B964EAAAFFEBFF880BBCF9D4FE69F9CC" box="[458,481,1557,1580]" italics="true" pageId="75" pageNumber="76">vs</emphasis>
. slightly prominent); posterior 0.2 of T1 pale yellow (
<emphasis id="B964EAAAFFEBFF880E28F9D4FBFDF9CC" box="[1118,1141,1557,1580]" italics="true" pageId="75" pageNumber="76">vs</emphasis>
. T1 medially orange and laterally pale yellow; 
<figureCitation id="132B2A3DFFEBFF880BFCF9F6FE64F9AF" box="[394,492,1591,1615]" captionStart="FIGURES 163–174" captionStartId="127.[151,264,1631,1654]" captionTargetBox="[151,1435,193,1610]" captionTargetId="figure-141@127.[151,1435,193,1610]" captionTargetPageId="127" captionText="FIGURES 163–174. Hind coxa and metasoma color patterns, dorsal view. 163, T. napus sp. nov., holotype. 164, T. teutonius sp. nov., holotype. 165, T. kawus sp. nov., holotype. 166, T. taperinus sp. nov., holotype. 167, T. darienus sp. nov., holotype. 168, T. amapaeus sp. nov., holotype. 169, T. ondensis sp. nov., holotype. 170, T. jatainus sp. nov., holotype. 171, T. albimaculatus Taschenberg. 172, T. vilhenus sp. nov., holotype. 173, T. benius sp. nov., holotype. 174, T. cassunungae Brauns." figureDoi="http://doi.org/10.5281/zenodo.5262013" httpUri="https://zenodo.org/record/5262013/files/figure.png" pageId="75" pageNumber="76">Fig. 165</figureCitation>
).
</paragraph>
<paragraph id="8BAF36B8FFEBFF880AB1F99DFE00F994" blockId="75.[151,1436,151,1904]" box="[199,392,1628,1652]" pageId="75" pageNumber="76">
<emphasis id="B964EAAAFFEBFF880AB1F99DFE8CF994" bold="true" box="[199,260,1628,1652]" pageId="75" pageNumber="76">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30A6533FFEBFF880AB1F9BEFBF7F95C" pageId="75" pageNumber="76" type="distribution">
<paragraph id="8BAF36B8FFEBFF880AB1F9BEFBF7F95C" blockId="75.[151,1436,151,1904]" pageId="75" pageNumber="76">
<emphasis id="B964EAAAFFEBFF880AB1F9BEFED1F978" bold="true" box="[199,345,1663,1688]" pageId="75" pageNumber="76">Distribution</emphasis>
. 
<collectingCountry id="F3077628FFEBFF880B1FF9BEFE54F977" box="[361,476,1663,1687]" name="Venezuela" pageId="75" pageNumber="76">Venezuela</collectingCountry>
and 
<collectingCountry id="F3077628FFEBFF880861F9BEFDD7F977" box="[535,607,1663,1687]" name="Brazil" pageId="75" pageNumber="76">Brazil</collectingCountry>
. Recorded from two localities (
<figureCitation id="132B2A3DFFEBFF8809BCF9BEFBA6F977" box="[970,1070,1663,1687]" captionStart="FIGURES 200–207" captionStartId="130.[151,264,1952,1975]" captionTargetBox="[172,1416,181,1931]" captionTargetId="figure-83@130.[172,1416,181,1931]" captionTargetPageId="130" captionText="FIGURES 200–207. Distribution records. 200, T. abactus Cresson. 201, T. albimaculatus Taschenberg. 202, T. brevicaudis Szépligeti. 203, T. cassunungae Brauns. 204, T. stramineus Taschenberg. 205, T. ondensis sp. nov., holotype. 206, T. zulius sp. nov., holotype. 207, T. amapaeus sp. nov." figureDoi="http://doi.org/10.5281/zenodo.5262021" httpUri="https://zenodo.org/record/5262021/files/figure.png" pageId="75" pageNumber="76">Fig. 206</figureCitation>
). The 
<typeStatus id="54AB881AFFEBFF880E0FF941FB21F978" box="[1145,1193,1664,1688]" pageId="75" pageNumber="76">type</typeStatus>
locality is Caxiuanã, 
<emphasis id="B964EAAAFFEBFF880AE1F965FF47F95C" box="[151,207,1700,1724]" italics="true" pageId="75" pageNumber="76">Pará</emphasis>
, 
<collectingCountry id="F3077628FFEBFF880AABF965FEA9F95C" box="[221,289,1700,1724]" name="Brazil" pageId="75" pageNumber="76">Brazil</collectingCountry>
(
<geoCoordinate id="EE24507FFFEBFF880B47F965FE11F95C" box="[305,409,1700,1724]" degrees="2" direction="south" minutes="4" orientation="latitude" pageId="75" pageNumber="76" precision="15" seconds="54" value="-2.0816667">2°4’54”S</geoCoordinate>
<geoCoordinate id="EE24507FFFEBFF880BD7F965FDADF95C" box="[417,549,1700,1724]" degrees="51" direction="west" minutes="51" orientation="longitude" pageId="75" pageNumber="76" precision="15" seconds="5" value="-51.851387">51°51’5”W</geoCoordinate>
). These records comprise a range of 7°15’ in latitude.
</paragraph>
</subSubSection>
<subSubSection id="C30A6533FFEBFF880AB1F906FE2BF93F" box="[199,419,1735,1760]" pageId="75" pageNumber="76" type="biology_ecology">
<paragraph id="8BAF36B8FFEBFF880AB1F906FE2BF93F" blockId="75.[151,1436,151,1904]" box="[199,419,1735,1760]" pageId="75" pageNumber="76">
<emphasis id="B964EAAAFFEBFF880AB1F906FE97F900" bold="true" box="[199,287,1735,1760]" pageId="75" pageNumber="76">Biology</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30A6533FFEBFF8F0AB1F92AFEF4FF51" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76" type="materials_examined">
<paragraph id="8BAF36B8FFEBFF880AB1F92AFE3FF88F" blockId="75.[151,1436,151,1904]" pageId="75" pageNumber="76">
<emphasis id="B964EAAAFFEBFF880AB1F92AFE21F8E4" bold="true" box="[199,425,1771,1796]" pageId="75" pageNumber="76">Material examined</emphasis>
. 
<specimenCount id="9D16FD31FFEBFF880BC1F92DFDAFF8E4" box="[439,551,1772,1796]" pageId="75" pageNumber="76" type="female">2 females</specimenCount>
. 
<typeStatus id="54AB881AFFEBFF880847F92DFD11F8E4" box="[561,665,1772,1796]" pageId="75" pageNumber="76" type="holotype">Holotype</typeStatus>
<materialsCitation id="3B783CE5FFEBFF8808D6F92DFD3BF8AC" accessionNumber="P05199" collectingDate="2003-11-24" collectingDateMax="2003-11-27" collectingDateMin="2003-11-24" collectionCode="UFES" collectorName="A. P. Aguiar &amp; J. Dias &amp; Right" country="Brazil" county="Para" location="Floresta Nacional de Caxiuana" municipality="Melgaco" pageId="75" pageNumber="76" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<specimenCount id="9D16FD31FFEBFF8808D6F92DFD3DF8E3" box="[672,693,1772,1795]" pageId="75" pageNumber="76" type="female">♀</specimenCount>
, 
<emphasis id="B964EAAAFFEBFF8808B5F92AFCA6F8E4" bold="true" box="[707,814,1771,1796]" pageId="75" pageNumber="76">
<collectingCountry id="F3077628FFEBFF8808B5F92AFCA6F8E4" box="[707,814,1771,1796]" name="Brazil" pageId="75" pageNumber="76">BRAZIL</collectingCountry>
</emphasis>
: 
<emphasis id="B964EAAAFFEBFF88094AF92DFCFCF8E4" box="[828,884,1772,1796]" italics="true" pageId="75" pageNumber="76">
<collectingCounty id="62CE4E34FFEBFF88094AF92DFCFCF8E4" box="[828,884,1772,1796]" pageId="75" pageNumber="76">Pará</collectingCounty>
</emphasis>
, 
<collectingMunicipality id="6BCBACC2FFEBFF8809F4F92DFC6FF8E4" box="[898,999,1772,1796]" pageId="75" pageNumber="76">Melgaço</collectingMunicipality>
, 
<location id="8ECF6063FFEBFF880984F92DFAC5F8E4" LSID="urn:lsid:plazi:treatment:03B987AEFFEAFF8F0AE1FD3DFEF4FF51:8ECF6063FFEBFF880984F92DFAC5F8E4" box="[1010,1357,1772,1796]" country="Brazil" county="Para" municipality="Melgaco" name="Floresta Nacional de Caxiuana" pageId="75" pageNumber="76">Floresta Nacional de Caxiuanã</location>
, 
<location id="8ECF6063FFEBFF880F2EF92DFEF3F8C7" LSID="urn:lsid:plazi:treatment:03B987AEFFEAFF8F0AE1FD3DFEF4FF51:8ECF6063FFEBFF880F2EF92DFEF3F8C7" country="Brazil" county="Para" municipality="Melgaco" name="Trilha Igarape Tijucaquara" pageId="75" pageNumber="76">Trilha Igarapé Tijucaquara</location>
, 
<date id="FFAE1078FFEBFF880BF1F8CEFDBAF8C7" box="[391,562,1807,1831]" pageId="75" pageNumber="76" value="2003-11-24" valueMax="2003-11-27" valueMin="2003-11-24">
<collectingDate id="EFEAE990FFEBFF880BF1F8CEFDBAF8C7" box="[391,562,1807,1831]" pageId="75" pageNumber="76" value="2003-11-24" valueMax="2003-11-27" valueMin="2003-11-24">24–27.XI.2003</collectingDate>
</date>
, YPT, 
<collectorName id="26E5536EFFEBFF8808F7F8CEFD75F8C7" box="[641,765,1807,1831]" pageId="75" pageNumber="76">A.P.Aguiar</collectorName>
&amp; 
<collectorName id="26E5536EFFEBFF880952F8CEFCE1F8C7" box="[804,873,1807,1831]" pageId="75" pageNumber="76">J.Dias</collectorName>
<emphasis id="B964EAAAFFEBFF880904F8D1FC1CF8C8" box="[882,916,1808,1832]" italics="true" pageId="75" pageNumber="76">leg</emphasis>
., P05199 [field point] (
<collectionCode id="ED01AE7DFFEBFF880ED6F8CEFB62F8C7" box="[1184,1258,1807,1831]" country="Brazil" httpUri="http://grbio.org/cool/n6nq-8nqh" name="Universidade Federal do Espirito Santo" pageId="75" pageNumber="76">UFES</collectionCode>
). 
<collectorName id="26E5536EFFEBFF880E88F8CEFAB4F8C7" box="[1278,1340,1807,1831]" pageId="75" pageNumber="76">Right</collectorName>
fore leg beyond coxa missing, otherwise in good shape
</materialsCitation>
. 
<materialsCitation id="3B783CE5FFEBFF8808B6F8F5FE3BF88F" collectingDate="1981-04-23" collectionCode="AEIC" collectorName="H. K. Townes" country="Venezuela" location="Tucuco" pageId="75" pageNumber="76" specimenCount="1" specimenCount-female="1" stateProvince="Zulia" typeStatus="paratype">
<typeStatus id="54AB881AFFEBFF8808B6F8F5FCAEF8AC" box="[704,806,1844,1868]" pageId="75" pageNumber="76" type="paratype">Paratype</typeStatus>
: 
<emphasis id="B964EAAAFFEBFF880945F8F2FC62F8AC" bold="true" box="[819,1002,1843,1868]" pageId="75" pageNumber="76">
<collectingCountry id="F3077628FFEBFF880945F8F2FC6DF8AC" box="[819,997,1843,1868]" name="Venezuela" pageId="75" pageNumber="76">VENEZUELA</collectingCountry>
:
</emphasis>
<specimenCount id="9D16FD31FFEBFF880982F8F5FBA9F8AB" box="[1012,1057,1844,1868]" pageId="75" pageNumber="76" type="female">1 ♀</specimenCount>
from 
<emphasis id="B964EAAAFFEBFF880E1CF8F5FB2DF8AC" box="[1130,1189,1844,1868]" italics="true" pageId="75" pageNumber="76">
<collectingRegion id="49D4F85AFFEBFF880E1CF8F5FB2DF8AC" box="[1130,1189,1844,1868]" country="Venezuela" name="Zulia" pageId="75" pageNumber="76">Zulia</collectingRegion>
</emphasis>
, 
<location id="8ECF6063FFEBFF880EC3F8F5FA83F8AB" LSID="urn:lsid:plazi:treatment:03B987AEFFEAFF8F0AE1FD3DFEF4FF51:8ECF6063FFEBFF880EC3F8F5FA83F8AB" box="[1205,1291,1844,1867]" country="Venezuela" name="Tucuco" pageId="75" pageNumber="76" stateProvince="Zulia">Tucuco</location>
, 
<date id="FFAE1078FFEBFF880F6FF8F5FA10F8AC" box="[1305,1432,1844,1868]" pageId="75" pageNumber="76" value="1981-04-23">
<collectingDate id="EFEAE990FFEBFF880F6FF8F5FA10F8AC" box="[1305,1432,1844,1868]" pageId="75" pageNumber="76" value="1981-04-23">23.IV.1981</collectingDate>
</date>
, 
<collectorName id="26E5536EFFEBFF880AE1F899FEABF88F" box="[151,291,1880,1903]" pageId="75" pageNumber="76">H.K.Townes</collectorName>
<emphasis id="B964EAAAFFEBFF880B5DF899FEC5F890" box="[299,333,1880,1904]" italics="true" pageId="75" pageNumber="76">leg</emphasis>
. (
<collectionCode id="ED01AE7DFFEBFF880B15F896FE22F88F" box="[355,426,1879,1903]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1008" lsid="urn:lsid:biocol.org:col:1008" name="American Entomological Institute" pageId="75" pageNumber="76">AEIC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF36B8FFECFF8F0AE1FF56FEF4FF51" blockId="76.[151,380,151,177]" box="[151,380,151,177]" pageId="76" pageNumber="77">
<emphasis id="B964EAAAFFECFF8F0AE1FF56FEF4FF51" bold="true" box="[151,380,151,177]" pageId="76" pageNumber="77">Cladistic Analyses</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>