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<document id="28298562CEA328597F64A5E9EC890268" ID-DOI="10.1590/0031-1049.2014.54.13" ID-ISSN="1807-0205" ID-Zenodo-Dep="5755768" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617966880165" checkinUser="carolina" docAuthor="Costa, Silvana Leal Nunes, Campos-Filho, Ivanklin Soares &amp; Araujo, Paula Beatriz" docDate="2014" docId="03B9878FFFE5FFBEFF4AFB2CA6A59B10" docLanguage="en" docName="PapAvZool.54.13.169-176.pdf" docOrigin="Papéis Avulsos de Zoologia (São Paulo) 54 (13)" docSource="http://dx.doi.org/10.1590/0031-1049.2014.54.13" docStyle="DocumentStyle:8E93E160036069519CAC16E940FF2271.1:PapAvZool.2014-2017.journal_article" docStyleId="8E93E160036069519CAC16E940FF2271" docStyleName="PapAvZool.2014-2017.journal_article" docStyleVersion="1" docTitle="Benthana carijos Costa &amp; Campos-Filho &amp; Araujo 2014, sp. nov." docType="treatment" docVersion="5" lastPageNumber="173" masterDocId="FF80FFF7FFE4FFBAFFA2FF9BA7519C40" masterDocTitle="New species and new records of Benthana Budde-Lund, 1908 (Isopoda: Oniscidea: Philosciidae) from southern Brazil" masterLastPageNumber="176" masterPageNumber="169" pageNumber="170" updateTime="1698903039591" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="BE85B7CBA92A5EE3292DE10599687A32">New species and new records of Benthana Budde-Lund, 1908 (Isopoda: Oniscidea: Philosciidae) from southern Brazil</mods:title>
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<mods:namePart id="901DE99035F61C71CDCC966EDA5A60FA">Costa, Silvana Leal Nunes</mods:namePart>
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<mods:namePart id="4AEB26A8597CDCEB863AA667DD1E70B9">Campos-Filho, Ivanklin Soares</mods:namePart>
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<mods:namePart id="3B95B8FA9119C10CA317189381BA12B7">Araujo, Paula Beatriz</mods:namePart>
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<mods:title id="2044300D96081D0115DA2C258DB4968F">Papéis Avulsos de Zoologia</mods:title>
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<mods:date id="073BE75D1DD528AFA74313AE9FCE80FC">2014</mods:date>
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<mods:title id="A7EF0F8876B81FBF5ABE2C83C10CBC9F">São Paulo</mods:title>
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<mods:url id="E4E7619B42C59D0127E9F89523A8CB70">http://dx.doi.org/10.1590/0031-1049.2014.54.13</mods:url>
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<mods:identifier id="4476BB44B5A84B6C3953C89F9D999670" type="DOI">10.1590/0031-1049.2014.54.13</mods:identifier>
<mods:identifier id="5FB35F65AE2C4DC6EDF11083B91012AF" type="ISSN">1807-0205</mods:identifier>
<mods:identifier id="035A89DCC5FA66EFD3C102C70CBEA2A3" type="Zenodo-Dep">5755768</mods:identifier>
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<treatment id="03B9878FFFE5FFBEFF4AFB2CA6A59B10" LSID="urn:lsid:zoobank.org:act:EE12292F-66EF-4A48-8A73-B4DD736338CA" httpUri="http://treatment.plazi.org/id/03B9878FFFE5FFBEFF4AFB2CA6A59B10" lastPageId="4" lastPageNumber="173" pageId="1" pageNumber="170">
<subSubSection id="C30A6512FFE5FFBBFF4AFB2CA6A89890" box="[232,505,1207,1232]" pageId="1" pageNumber="170" type="nomenclature">
<paragraph id="8BAF3699FFE5FFBBFF4AFB2CA6A89890" blockId="1.[232,505,1207,1264]" box="[232,505,1207,1232]" pageId="1" pageNumber="170">
<heading id="D0E781F5FFE5FFBBFF4AFB2CA6A89890" bold="true" box="[232,505,1207,1232]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="170" reason="2">
<emphasis id="B964EA8BFFE5FFBBFF4AFB2CA6A89890" bold="true" box="[232,505,1207,1232]" pageId="1" pageNumber="170">
<taxonomicName id="4C104D1AFFE5FFBBFF4AFB2CA6F19890" ID-CoL="LHPC" authority="Costa &amp; Campos-Filho &amp; Araujo, 2014" authorityName="Costa &amp; Campos-Filho &amp; Araujo" authorityYear="2014" box="[232,416,1207,1232]" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="1" pageNumber="170" phylum="Arthropoda" rank="species" species="carijos" status="sp. nov.">
<emphasis id="B964EA8BFFE5FFBBFF4AFB2CA6F19890" bold="true" box="[232,416,1207,1232]" italics="true" pageId="1" pageNumber="170">Benthana carijos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757F0FFE5FFBBFE04FB23A6A89890" box="[422,505,1208,1232]" pageId="1" pageNumber="170" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30A6512FFE5FFBEFE96FB43A6A59B10" lastPageId="4" lastPageNumber="173" pageId="1" pageNumber="170" type="description">
<paragraph id="8BAF3699FFE5FFBBFE96FB43A6FC98B0" blockId="1.[232,505,1207,1264]" box="[308,429,1240,1264]" pageId="1" pageNumber="170">
<figureCitation id="132B2A1CFFE5FFBBFE96FB43A6FC98B0" box="[308,429,1240,1264]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[132,211,1800,1819]" captionStartId-1="3.[95,174,1826,1845]" captionStartId-2="6.[132,211,1775,1794]" captionTargetBox-0="[137,1261,170,1777]" captionTargetBox-1="[94,1228,170,1802]" captionTargetBox-2="[134,680,980,1750]" captionTargetId-0="figure-1@2.[132,1266,170,1777]" captionTargetId-1="figure-1@3.[94,1228,170,1802]" captionTargetId-2="figure-525@6.[132,680,976,1751]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="6" captionText-0="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." captionText-1="FIGURE 2: Benthana carijos sp. nov. (male paratype). A: pereopod 1; B: pereopod 7; C: uropod; D: genital papilla; E: pleopod 1; F: pleopod 2; G: pleopod 5 exopod." captionText-2="FIGURE 3: Map distribution of the species of Benthana from southern Brazil. *: Benthana carijos sp. nov.; : Benthana cairensis; : Benthana longicornis; : Benthana moreirai; ·: Benthana picta; ·: Benthana serrara; : Benthana taeniata." figureDoi-0="http://doi.org/10.5281/zenodo.5755770" figureDoi-1="http://doi.org/10.5281/zenodo.5755772" figureDoi-2="http://doi.org/10.5281/zenodo.5755774" httpUri-0="https://zenodo.org/record/5755770/files/figure.png" httpUri-1="https://zenodo.org/record/5755772/files/figure.png" httpUri-2="https://zenodo.org/record/5755774/files/figure.png" pageId="1" pageNumber="170">
<emphasis id="B964EA8BFFE5FFBBFE96FB43A6FC98B0" bold="true" box="[308,429,1240,1264]" pageId="1" pageNumber="170">Figures 1-3</emphasis>
</figureCitation>
</paragraph>
<paragraph id="8BAF3699FFE5FFBBFFFCFA83A6DD9910" blockId="1.[94,643,1304,1360]" pageId="1" pageNumber="170">
ZooBank.
<uri id="FF813A9BFFE5FFBBFF6BFA83A6DD9910" pageId="1" pageNumber="170">
urn:lsid:zoobank.org:pub:
<uuid id="FFB60C4CFFE5FFBBFE6DFA83A6DD9910" pageId="1" pageNumber="170">645E1FF2-8E4F- 446F-92FE-8A565C8AA06B</uuid>
</uri>
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<paragraph id="8BAF3699FFE5FFBBFFFCFAE2A57F9AB0" blockId="1.[94,643,1400,1776]" pageId="1" pageNumber="170">
<emphasis id="B964EA8BFFE5FFBBFFFCFAE2A7BF99D0" box="[94,238,1400,1424]" italics="true" pageId="1" pageNumber="170">Type material:</emphasis>
<materialsCitation id="3B783CC4FFE5FFBBFF55FAE3A52E9A50" collectingDate="2012-06-25" collectionCode="MZUSP" collectorName="P. B. Araujo &amp; B. L. Zimmermann" country="Brazil" latitude="-27.9925" location="Anitapolis" longLatPrecision="20" longitude="-49.11333" pageId="1" pageNumber="170" specimenCode="MZUSP 30092" specimenCount="1" stateProvince="Santa Catarina" typeStatus="holotype">
<emphasis id="B964EA8BFFE5FFBBFF55FAE3A61F99D0" box="[247,334,1400,1424]" italics="true" pageId="1" pageNumber="170">
<typeStatus id="54AB883BFFE5FFBBFF55FAE3A61F99D0" box="[247,334,1400,1424]" pageId="1" pageNumber="170" type="holotype">Holotype</typeStatus>
</emphasis>
male,
<collectingCountry id="F3077609FFE5FFBBFE35FAE3A68499D0" box="[407,469,1400,1424]" name="Brazil" pageId="1" pageNumber="170">Brazil</collectingCountry>
,
<collectingRegion id="49D4F87BFFE5FFBBFE40FAE3A52E99D0" box="[482,639,1400,1424]" country="Brazil" name="Santa Catarina" pageId="1" pageNumber="170">Santa Catarina</collectingRegion>
,
<location id="8ECF6042FFE5FFBBFFFCFA03A79B99F0" LSID="urn:lsid:plazi:treatment:03B9878FFFE5FFBEFF4AFB2CA6A59B10:8ECF6042FFE5FFBBFFFCFA03A79B99F0" box="[94,202,1432,1456]" country="Brazil" latitude="-27.9925" longLatPrecision="20" longitude="-49.11333" name="Anitapolis" pageId="1" pageNumber="170" stateProvince="Santa Catarina">Anitápolis</location>
, road SC-407,
<locationDeviation id="9A1D2FBBFFE5FFBBFE22FA03A6619990" firstLocation="Anitapolis" pageId="1" pageNumber="170" secondLocation="Santa Rosa de Lima">between Anitápolis and Santa Rosa de Lima</locationDeviation>
,
<geoCoordinate id="EE24505EFFE5FFBBFE9EFA23A6E49990" box="[316,437,1464,1488]" degrees="27" direction="south" minutes="59" orientation="latitude" pageId="1" pageNumber="170" precision="15" seconds="33" value="-27.9925">27°5933”S</geoCoordinate>
,
<geoCoordinate id="EE24505EFFE5FFBBFE63FA23A5159990" box="[449,580,1464,1488]" degrees="49" direction="west" minutes="06" orientation="longitude" pageId="1" pageNumber="170" precision="15" seconds="48" value="-49.11333">49°0648”W</geoCoordinate>
, open vegetation cover, impacted area,
<date id="FFAE1059FFE5FFBBFE0BFA43A57999B0" box="[425,552,1496,1520]" pageId="1" pageNumber="170" value="2012-06-25">
<collectingDate id="EFEAE9B1FFE5FFBBFE0BFA43A57999B0" box="[425,552,1496,1520]" pageId="1" pageNumber="170" value="2012-06-25">25/VI/2012</collectingDate>
</date>
, leg.
<collectorName id="26E5534FFFE5FFBBFDF9FA43A7F59A50" pageId="1" pageNumber="170">P.B. Araujo</collectorName>
and
<collectorName id="26E5534FFFE5FFBBFF45FA63A6E29A50" box="[231,435,1528,1552]" pageId="1" pageNumber="170">B.L. Zimmermann</collectorName>
(
<specimenCode id="DBB69EE2FFE5FFBBFE6AFA63A5299A50" box="[456,632,1528,1552]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="170">MZUSP 30092</specimenCode>
)
</materialsCitation>
.
<materialsCitation id="3B783CC4FFE5FFBBFFFCF982A7FC9A10" collectingDate="2012-06-25" collectionCode="MNRJ" collectorName="P. B. Araujo &amp; B. L. Zimmermann" country="Brazil" latitude="-27.9925" location="Anitapolis" longLatPrecision="20" longitude="-49.11333" pageId="1" pageNumber="170" specimenCode="MNRJ 23311" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Santa Catarina" typeStatus="paratype">
<emphasis id="B964EA8BFFE5FFBBFFFCF982A7E89A70" box="[94,185,1561,1584]" italics="true" pageId="1" pageNumber="170">
<typeStatus id="54AB883BFFE5FFBBFFFCF982A7E89A70" box="[94,185,1561,1584]" pageId="1" pageNumber="170" type="paratype">Paratypes</typeStatus>
</emphasis>
<specimenCount id="9D16FD10FFE5FFBBFF60F983A7BD9A70" box="[194,236,1557,1584]" pageId="1" pageNumber="170" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D16FD10FFE5FFBBFF5EF983A66B9A70" box="[252,314,1557,1584]" pageId="1" pageNumber="170" type="female">2 ♀♀</specimenCount>
same data as holotype (
<specimenCode id="DBB69EE2FFE5FFBBFD9BF983A7F49A10" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="1" pageNumber="170">MNRJ 23311</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B783CC4FFE5FFBBFF14F9A3A63F9AD0" collectingDate="1979-01" collectionCode="MZUSP" country="Iceland" latitude="-25.131666" location="Cananeia" longLatPrecision="21" longitude="-47.96639" municipality="Cardoso" pageId="1" pageNumber="170" specimenCode="MZUSP 24285" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Sao Paulo" typeStatus="paratype">
<collectingRegion id="49D4F87BFFE5FFBBFF14F9A3A64D9A10" box="[182,284,1592,1616]" country="Brazil" name="Sao Paulo" pageId="1" pageNumber="170">São Paulo</collectingRegion>
:
<specimenCount id="9D16FD10FFE5FFBBFE87F9A3A60E9A10" box="[293,351,1589,1616]" pageId="1" pageNumber="170" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D16FD10FFE5FFBBFEC9F9A3A6F79A10" box="[363,422,1589,1616]" pageId="1" pageNumber="170" type="female">5 ♀♀</specimenCount>
, 3 mancas,
<location id="8ECF6042FFE5FFBBFDBEF9A3A52E9A10" LSID="urn:lsid:plazi:treatment:03B9878FFFE5FFBEFF4AFB2CA6A59B10:8ECF6042FFE5FFBBFDBEF9A3A52E9A10" box="[540,639,1592,1616]" country="Iceland" latitude="-25.131666" longLatPrecision="21" longitude="-47.96639" municipality="Cardoso" name="Cananeia" pageId="1" pageNumber="170" stateProvince="Sao Paulo">Cananéia</location>
,
<collectingCountry id="F3077609FFE5FFBBFFFDF9C3A7D69A30" box="[95,135,1624,1648]" name="Iceland" pageId="1" pageNumber="170">Ilha</collectingCountry>
do
<collectingMunicipality id="6BCBACE3FFE5FFBBFF17F9C3A65F9A30" box="[181,270,1624,1648]" pageId="1" pageNumber="170">Cardoso</collectingMunicipality>
,
<geoCoordinate id="EE24505EFFE5FFBBFEB9F9C3A6C59A30" box="[283,404,1624,1648]" degrees="25" direction="south" minutes="07" orientation="latitude" pageId="1" pageNumber="170" precision="15" seconds="54" value="-25.131666">25°0754”S</geoCoordinate>
,
<geoCoordinate id="EE24505EFFE5FFBBFE00F9C3A5759A30" box="[418,548,1624,1648]" degrees="47" direction="west" minutes="57" orientation="longitude" pageId="1" pageNumber="170" precision="15" seconds="59" value="-47.96639">47°5759”W</geoCoordinate>
, nº 110,
<date id="FFAE1059FFE5FFBBFFFDF9E3A7F79AD0" box="[95,166,1656,1680]" bridgedPair="/" pageId="1" pageNumber="170" value="1979-01">
<collectingDate id="EFEAE9B1FFE5FFBBFFFDF9E3A7F79AD0" box="[95,166,1656,1680]" bridgedPair="/" pageId="1" pageNumber="170" value="1979-01">I/1979</collectingDate>
</date>
(
<specimenCode id="DBB69EE2FFE5FFBBFF1BF9E3A6369AD0" box="[185,359,1656,1680]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="170">MZUSP 24285</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B783CC4FFE5FFBBFEDDF9E3A7FD9A90" collectionCode="MZUSP" collectorName="Cardoso" country="Iceland" location="Cananeia" pageId="1" pageNumber="170" specimenCode="MZUSP 24289" specimenCount="3" specimenCount-female="1" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D16FD10FFE5FFBBFEDDF9E3A6909AD0" box="[383,449,1653,1680]" pageId="1" pageNumber="170" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D16FD10FFE5FFBBFE77F9E3A5529AD0" box="[469,515,1653,1680]" pageId="1" pageNumber="170" type="female">1 ♀</specimenCount>
, 2 mancas,
<location id="8ECF6042FFE5FFBBFFFDF903A7939AF0" LSID="urn:lsid:plazi:treatment:03B9878FFFE5FFBEFF4AFB2CA6A59B10:8ECF6042FFE5FFBBFFFDF903A7939AF0" box="[95,194,1688,1712]" country="Iceland" name="Cananeia" pageId="1" pageNumber="170">Cananéia</location>
,
<collectingCountry id="F3077609FFE5FFBBFF6DF903A7A69AF0" box="[207,247,1688,1712]" name="Iceland" pageId="1" pageNumber="170">Ilha</collectingCountry>
do
<collectorName id="26E5534FFFE5FFBBFE81F903A62D9AF0" box="[291,380,1688,1712]" pageId="1" pageNumber="170">Cardoso</collectorName>
, nº 106, 1979 (
<specimenCode id="DBB69EE2FFE5FFBBFD8AF903A7F49A90" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="170">MZUSP 24289</specimenCode>
)
</materialsCitation>
,
<materialsCitation id="3B783CC4FFE5FFBBFF14F923A57B9AB0" collectionCode="MZUSP" collectorName="Cananeia &amp; Cardoso" country="Iceland" location="Ilha" pageId="1" pageNumber="170" specimenCode="MZUSP 24290" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D16FD10FFE5FFBBFF14F923A78C9A90" box="[182,221,1717,1744]" pageId="1" pageNumber="170" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D16FD10FFE5FFBBFF48F923A6409A90" box="[234,273,1717,1744]" pageId="1" pageNumber="170" type="female">1 ♀</specimenCount>
, 1? (without pleon),
<collectorName id="26E5534FFFE5FFBBFE4CF923A5009A90" box="[494,593,1720,1744]" pageId="1" pageNumber="170">Cananéia</collectorName>
,
<collectingCountry id="F3077609FFE5FFBBFDF9F923A5D29A90" box="[603,643,1720,1744]" name="Iceland" pageId="1" pageNumber="170">Ilha</collectingCountry>
do
<collectorName id="26E5534FFFE5FFBBFF22F943A7889AB0" box="[128,217,1752,1776]" pageId="1" pageNumber="170">Cardoso</collectorName>
, nº 057, 1979 (
<specimenCode id="DBB69EE2FFE5FFBBFED8F943A5739AB0" box="[378,546,1752,1776]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="170">MZUSP 24290</specimenCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAF3699FFE5FFBBFFFCF882A5D29B10" blockId="1.[94,643,1816,1872]" pageId="1" pageNumber="170">
<emphasis id="B964EA8BFFE5FFBBFFFCF882A7959B70" box="[94,196,1817,1840]" italics="true" pageId="1" pageNumber="170">Diagnosis:</emphasis>
Antennule with 13+2 aesthetascs; male pleopod 1 exopod elongated, with low indentation,
</paragraph>
<paragraph id="8BAF3699FFE5FFBBFD0AFF23A4529CB0" blockId="1.[680,1229,184,240]" pageId="1" pageNumber="170">not prominent dentiform protrusion, and narrow distal third.</paragraph>
<paragraph id="8BAF3699FFE5FFBBFD0AFE82A4DC9D0F" blockId="1.[680,1229,280,1872]" pageId="1" pageNumber="170">
<emphasis id="B964EA8BFFE5FFBBFD0AFE82A44E9D70" box="[680,799,281,304]" italics="true" pageId="1" pageNumber="170">Description:</emphasis>
Maximum body length
<quantity id="4CE89B7CFFE5FFBBFBBEFE83A33A9D6F" box="[1052,1131,280,304]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="1" pageNumber="170" unit="mm" value="14.0">14 mm</quantity>
, cephalothorax width 2.5 mm.
</paragraph>
<paragraph id="8BAF3699FFE5FFBBFD43FEC3A35F9E0F" blockId="1.[680,1229,280,1872]" pageId="1" pageNumber="170">Light brown color. Cephalon with irregular unpigmented spots. Antenna with unpigmented proximal portion of the fourth article and distal portion of fifth article. Pereon with two lines of unpigmented longitudinal spots paramedially, a central line of unpigmented spots from the first pereonite to the fifth pleonite; epimera with large unpigmented spots. Telson with three unpigmented spots.</paragraph>
<paragraph id="8BAF3699FFE5FFBBFD43FDC3A49098EF" blockId="1.[680,1229,280,1872]" pageId="1" pageNumber="170">
Body (
<figureCitation id="132B2A1CFFE5FFBBFC8BFDC3A42A9E30" box="[809,891,600,624]" captionStart="FIGURE 1" captionStartId="2.[132,211,1800,1819]" captionTargetBox="[137,1261,170,1777]" captionTargetId="figure-1@2.[132,1266,170,1777]" captionTargetPageId="2" captionText="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="1" pageNumber="170">Fig. 1A</figureCitation>
) convex; dorsal surface smooth and bright, bearing few long piliform scale-setae (
<figureCitation id="132B2A1CFFE5FFBBFD12FD03A4539EF0" box="[688,770,664,688]" captionStart="FIGURE 1" captionStartId="2.[132,211,1800,1819]" captionTargetBox="[137,1261,170,1777]" captionTargetId="figure-1@2.[132,1266,170,1777]" captionTargetPageId="2" captionText="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="1" pageNumber="170">Fig. 1B</figureCitation>
); pleon narrower than pereon with developed epimera; pereonites 1-4 with postero-lateral corners right-angled with rounded apices and posterior margins straight; pereonites 5-7 with postero-lateral corners gradually more acute and posterior margins gradually more arched; pleon epimera 3-5 developed, triangular, acute, and directed backwards. Cephalon (
<figureCitation id="132B2A1CFFE5FFBBFD12FCE3A4669FD0" box="[688,823,888,912]" captionStart="FIGURE 1" captionStartId="2.[132,211,1800,1819]" captionTargetBox="[137,1261,170,1777]" captionTargetId="figure-1@2.[132,1266,170,1777]" captionTargetPageId="2" captionText="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="1" pageNumber="170">Figs. 1C, D</figureCitation>
) with
<emphasis id="B964EA8BFFE5FFBBFC2BFCE3A3389FD0" box="[905,1129,888,912]" italics="true" pageId="1" pageNumber="170">linea supra-antennalis</emphasis>
sinuous;
<emphasis id="B964EA8BFFE5FFBBFD0AFC03A4629FF0" box="[680,819,920,944]" italics="true" pageId="1" pageNumber="170">linea frontalis</emphasis>
and lateral lobes lacking; clypeus and labrum similar in length; eyes with 21 ommatidia arranged in four rows. Pereonites (
<figureCitation id="132B2A1CFFE5FFBBFC59FC43A3169FB0" box="[1019,1095,984,1008]" captionStart="FIGURE 1" captionStartId="2.[132,211,1800,1819]" captionTargetBox="[137,1261,170,1777]" captionTargetId="figure-1@2.[132,1266,170,1777]" captionTargetPageId="2" captionText="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="1" pageNumber="170">Fig. 1E</figureCitation>
) with sulcus marginalis and glandular pores along the lateral margin. One line of
<emphasis id="B964EA8BFFE5FFBBFCF4FB83A4BD9870" box="[854,1004,1048,1072]" italics="true" pageId="1" pageNumber="170">noduli laterales</emphasis>
per side (
<figureCitation id="132B2A1CFFE5FFBBFBF0FB83A3C29870" box="[1106,1171,1048,1072]" captionStart="FIGURE 1" captionStartId="2.[132,211,1800,1819]" captionTargetBox="[137,1261,170,1777]" captionTargetId="figure-1@2.[132,1266,170,1777]" captionTargetPageId="2" captionText="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="1" pageNumber="170">Figs. 1</figureCitation>
E-G) with d/c coordinates reaching a maximum on pereonite 4; b/c coordinates gradually decreasing. Telson (
<figureCitation id="132B2A1CFFE5FFBBFD12FBE3A45198D0" box="[688,768,1144,1168]" captionStart="FIGURE 1" captionStartId="2.[132,211,1800,1819]" captionTargetBox="[137,1261,170,1777]" captionTargetId="figure-1@2.[132,1266,170,1777]" captionTargetPageId="2" captionText="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="1" pageNumber="170">Fig. 1H</figureCitation>
) triangular with lateral margins slightly concave and right-angled apex.
</paragraph>
<paragraph id="8BAF3699FFE5FFBBFD43FB23A35899D0" blockId="1.[680,1229,280,1872]" pageId="1" pageNumber="170">
Antennule (
<figureCitation id="132B2A1CFFE5FFBBFCC3FB23A4FA9890" box="[865,939,1208,1232]" captionStart="FIGURE 1" captionStartId="2.[132,211,1800,1819]" captionTargetBox="[137,1261,170,1777]" captionTargetId="figure-1@2.[132,1266,170,1777]" captionTargetPageId="2" captionText="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="1" pageNumber="170">Fig. 1I</figureCitation>
) triarticulate, distal article the longest, with 13 aesthetascs plus apical pair. Antenna (
<figureCitation id="132B2A1CFFE5FFBBFD55FB63A4139950" box="[759,834,1272,1296]" captionStart="FIGURE 1" captionStartId="2.[132,211,1800,1819]" captionTargetBox="[137,1261,170,1777]" captionTargetId="figure-1@2.[132,1266,170,1777]" captionTargetPageId="2" captionText="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="1" pageNumber="170">Fig. 1J</figureCitation>
) when extended posteriorly reaches fifth pereonite, flagellum triarticulate, first article longest, second and third subequal in length; apical organ short, one sixth of the length of the distal article, free sensilla inserted near the base.
</paragraph>
<paragraph id="8BAF3699FFE5FFBEFD43FA03A7E29D0F" blockId="1.[680,1229,280,1872]" lastBlockId="4.[132,681,184,783]" lastPageId="4" lastPageNumber="173" pageId="1" pageNumber="170">
Mandibles (
<figureCitation id="132B2A1CFFE5FFBBFCC7FA03A4B899F0" box="[869,1001,1432,1456]" captionStart="FIGURE 1" captionStartId="2.[132,211,1800,1819]" captionTargetBox="[137,1261,170,1777]" captionTargetId="figure-1@2.[132,1266,170,1777]" captionTargetPageId="2" captionText="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="1" pageNumber="170">Figs. 1K, L</figureCitation>
) with molar penicil consisting of at least 14 branches,
<emphasis id="B964EA8BFFE5FFBBFB88FA22A39C9990" box="[1066,1229,1464,1488]" italics="true" pageId="1" pageNumber="170">pars intermedia</emphasis>
with dense tuft of coniform setae, left mandible with three penicils, and right mandible with two penicils. Maxillule (
<figureCitation id="132B2A1CFFE5FFBBFC24F983A4B29A70" box="[902,995,1560,1584]" captionStart="FIGURE 1" captionStartId="2.[132,211,1800,1819]" captionTargetBox="[137,1261,170,1777]" captionTargetId="figure-1@2.[132,1266,170,1777]" captionTargetPageId="2" captionText="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="1" pageNumber="170">Fig. 1M</figureCitation>
) medial endite with two penicils, inserted transversely, distal portion rounded; outer endite with 4+6 teeth, five teeth of inner set pectinate (inner tooth bearing 16 denticles), one short and simple. Maxilla (
<figureCitation id="132B2A1CFFE5FFBBFB88F903A32D9AF0" box="[1066,1148,1688,1712]" captionStart="FIGURE 1" captionStartId="2.[132,211,1800,1819]" captionTargetBox="[137,1261,170,1777]" captionTargetId="figure-1@2.[132,1266,170,1777]" captionTargetPageId="2" captionText="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="1" pageNumber="170">Fig. 1N</figureCitation>
) lateral lobe twice as wide as medial lobe with distal margin sinuous, covered with trichiform setae; medial lobe rounded, covered with trichiform setae. Maxilliped (
<figureCitation id="132B2A1CFFE5FFBBFD12F883A4539B70" box="[688,770,1816,1840]" captionStart="FIGURE 1" captionStartId="2.[132,211,1800,1819]" captionTargetBox="[137,1261,170,1777]" captionTargetId="figure-1@2.[132,1266,170,1777]" captionTargetPageId="2" captionText="FIGURE 1: Benthana carijos sp. nov. (male paratype). A: habitus; B: piliform scale-seta; C: cephalon, frontal view; D: cephalon, dorsal view; E: pereon epimeron 3; F: coordinates d/c of noduli laterales; G: coordinates b/c of noduli laterales; H: telson; I: antennule; J: antenna; K: left mandible; L: right mandible; M: maxillule; N: maxilla; O: maxilliped." figureDoi="http://doi.org/10.5281/zenodo.5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="1" pageNumber="170">Fig. 1O</figureCitation>
) with rectangular base, cuticle scaled proximally, sparse piliform setae, and outer distal corner slightly prominent; endite rectangular, outer margin sinuous, distal margin rounded with two hook-like setae and medial seta overpassing distal margin, setose ventral sulcus ending in one strong tooth-like seta.
</paragraph>
<caption id="DF6F6611FFE6FFB8FF26F893A4519B0F" ID-DOI="http://doi.org/10.5281/zenodo.5755770" ID-Zenodo-Dep="5755770" httpUri="https://zenodo.org/record/5755770/files/figure.png" pageId="2" pageNumber="171" startId="2.[132,211,1800,1819]" targetBox="[137,1261,170,1777]" targetPageId="2">
<paragraph id="8BAF3699FFE6FFB8FF26F893A4519B0F" blockId="2.[132,1267,1800,1871]" pageId="2" pageNumber="171">
<emphasis id="B964EA8BFFE6FFB8FF26F893A7BA9B5B" bold="true" box="[132,235,1800,1819]" pageId="2" pageNumber="171">FIGURE 1:</emphasis>
<taxonomicName id="4C104D1AFFE6FFB8FF50F892A6279B5B" authority="Costa &amp; Campos-Filho &amp; Araujo, 2014" authorityName="Costa &amp; Campos-Filho &amp; Araujo" authorityYear="2014" box="[242,374,1801,1820]" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="2" pageNumber="171" phylum="Arthropoda" rank="species" species="carijos" status="sp. nov.">
<emphasis id="B964EA8BFFE6FFB8FF50F892A6279B5B" box="[242,374,1801,1820]" italics="true" pageId="2" pageNumber="171">Benthana carijos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757F0FFE6FFB8FEDFF892A6EF9B5C" box="[381,446,1801,1820]" pageId="2" pageNumber="171" rank="species">sp. nov.</taxonomicNameLabel>
(male paratype).
<emphasis id="B964EA8BFFE6FFB8FDF3F893A5359B5B" bold="true" box="[593,612,1800,1819]" pageId="2" pageNumber="171">A:</emphasis>
habitus;
<emphasis id="B964EA8BFFE6FFB8FD10F892A5949B5C" bold="true" box="[690,709,1801,1820]" pageId="2" pageNumber="171">B:</emphasis>
piliform scale-seta;
<emphasis id="B964EA8BFFE6FFB8FCCFF893A4D09B5B" bold="true" box="[877,897,1800,1819]" pageId="2" pageNumber="171">C:</emphasis>
cephalon, frontal view;
<emphasis id="B964EA8BFFE6FFB8FBECF892A3349B5C" bold="true" box="[1102,1125,1801,1820]" pageId="2" pageNumber="171">D:</emphasis>
cephalon, dorsal view;
<emphasis id="B964EA8BFFE6FFB8FF16F8B9A7979B75" bold="true" box="[180,198,1826,1845]" pageId="2" pageNumber="171">E:</emphasis>
pereon epimeron 3;
<emphasis id="B964EA8BFFE6FFB8FED3F8B9A6D39B75" bold="true" box="[369,386,1826,1845]" pageId="2" pageNumber="171">F:</emphasis>
coordinates d/c of
<emphasis id="B964EA8BFFE6FFB8FDBDF8B9A5CA9B75" box="[543,667,1826,1845]" italics="true" pageId="2" pageNumber="171">noduli laterales;</emphasis>
<emphasis id="B964EA8BFFE6FFB8FD02F8B9A5E49B75" bold="true" box="[672,693,1826,1845]" pageId="2" pageNumber="171">G:</emphasis>
coordinates b/c of
<emphasis id="B964EA8BFFE6FFB8FCF0F8B9A49F9B75" box="[850,974,1826,1845]" italics="true" pageId="2" pageNumber="171">noduli laterales;</emphasis>
<emphasis id="B964EA8BFFE6FFB8FC71F8B9A4B89B75" bold="true" box="[979,1001,1826,1845]" pageId="2" pageNumber="171">H:</emphasis>
telson;
<emphasis id="B964EA8BFFE6FFB8FB88F8B9A3669B75" bold="true" box="[1066,1079,1826,1845]" pageId="2" pageNumber="171">I:</emphasis>
antennule;
<emphasis id="B964EA8BFFE6FFB8FB3AF8B9A3F79B75" bold="true" box="[1176,1190,1826,1845]" pageId="2" pageNumber="171">J:</emphasis>
antenna;
<emphasis id="B964EA8BFFE6FFB8FF26F8A7A7C89B0F" bold="true" box="[132,153,1852,1871]" pageId="2" pageNumber="171">K:</emphasis>
left mandible;
<emphasis id="B964EA8BFFE6FFB8FEB4F8A7A6799B0F" bold="true" box="[278,296,1852,1871]" pageId="2" pageNumber="171">L:</emphasis>
right mandible;
<emphasis id="B964EA8BFFE6FFB8FE11F8A7A69D9B0F" bold="true" box="[435,460,1852,1871]" pageId="2" pageNumber="171">M:</emphasis>
maxillule;
<emphasis id="B964EA8BFFE6FFB8FD85F8A7A56D9B0F" bold="true" box="[551,572,1852,1871]" pageId="2" pageNumber="171">N:</emphasis>
maxilla;
<emphasis id="B964EA8BFFE6FFB8FD25F8A7A5CF9B0F" bold="true" box="[647,670,1852,1871]" pageId="2" pageNumber="171">O:</emphasis>
maxilliped.
</paragraph>
</caption>
<caption id="DF6F6611FFE7FFB9FFFDF8B9A61E9B0F" ID-DOI="http://doi.org/10.5281/zenodo.5755772" ID-Zenodo-Dep="5755772" httpUri="https://zenodo.org/record/5755772/files/figure.png" pageId="3" pageNumber="172" startId="3.[95,174,1826,1845]" targetBox="[94,1228,170,1802]" targetPageId="3">
<paragraph id="8BAF3699FFE7FFB9FFFDF8B9A61E9B0F" blockId="3.[95,1229,1826,1871]" pageId="3" pageNumber="172">
<emphasis id="B964EA8BFFE7FFB9FFFDF8B9A7929B75" bold="true" box="[95,195,1826,1845]" pageId="3" pageNumber="172">FIGURE 2:</emphasis>
<taxonomicName id="4C104D1AFFE7FFB9FF6BF8B9A61D9B75" authority="Costa &amp; Campos-Filho &amp; Araujo, 2014" authorityName="Costa &amp; Campos-Filho &amp; Araujo" authorityYear="2014" box="[201,332,1826,1845]" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="3" pageNumber="172" phylum="Arthropoda" rank="species" species="carijos" status="sp. nov.">
<emphasis id="B964EA8BFFE7FFB9FF6BF8B9A61D9B75" box="[201,332,1826,1845]" italics="true" pageId="3" pageNumber="172">Benthana carijos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757F0FFE7FFB9FEF3F8B8A6C09B76" box="[337,401,1827,1846]" pageId="3" pageNumber="172" rank="species">sp. nov.</taxonomicNameLabel>
(male paratype).
<emphasis id="B964EA8BFFE7FFB9FDBDF8B9A5639B75" bold="true" box="[543,562,1826,1845]" pageId="3" pageNumber="172">A:</emphasis>
pereopod 1;
<emphasis id="B964EA8BFFE7FFB9FD3CF8B9A5E09B75" bold="true" box="[670,689,1826,1845]" pageId="3" pageNumber="172">B:</emphasis>
pereopod 7;
<emphasis id="B964EA8BFFE7FFB9FCBCF8B9A4639B75" bold="true" box="[798,818,1826,1845]" pageId="3" pageNumber="172">C:</emphasis>
uropod;
<emphasis id="B964EA8BFFE7FFB9FCDCF8B9A4C49B75" bold="true" box="[894,917,1826,1845]" pageId="3" pageNumber="172">D:</emphasis>
genital papilla;
<emphasis id="B964EA8BFFE7FFB9FBBAF8B9A37B9B75" bold="true" box="[1048,1066,1826,1845]" pageId="3" pageNumber="172">E:</emphasis>
pleopod 1;
<emphasis id="B964EA8BFFE7FFB9FB2EF8B9A3CF9B75" bold="true" box="[1164,1182,1826,1845]" pageId="3" pageNumber="172">F:</emphasis>
pleopod 2;
<emphasis id="B964EA8BFFE7FFB9FF3BF8A7A7FF9B0F" bold="true" box="[153,174,1852,1871]" pageId="3" pageNumber="172">G:</emphasis>
pleopod 5 exopod.
</paragraph>
</caption>
<paragraph id="8BAF3699FFE0FFBEFF1FFEC3A6EB9E10" blockId="4.[132,681,184,783]" pageId="4" pageNumber="173">
Pereopods (
<figureCitation id="132B2A1CFFE0FFBEFE96FEC3A6FB9D30" box="[308,426,344,368]" captionStart="FIGURE 2" captionStartId="3.[95,174,1826,1845]" captionTargetBox="[94,1228,170,1802]" captionTargetId="figure-1@3.[94,1228,170,1802]" captionTargetPageId="3" captionText="FIGURE 2: Benthana carijos sp. nov. (male paratype). A: pereopod 1; B: pereopod 7; C: uropod; D: genital papilla; E: pleopod 1; F: pleopod 2; G: pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.5755772" httpUri="https://zenodo.org/record/5755772/files/figure.png" pageId="4" pageNumber="173">Figs. 2A, B</figureCitation>
) rather slender, piliform setae on all articles, hyaline fringe and sparse setae on merus and carpus; carpus 1 with transverse antennagrooming brush and one seta with hand-like apex; ischium 7 triangular, bearing four long setae; dactylus 1-7 with inner claw reaching distal margin of outer claw, dactylar seta long and simple, ungual seta simple, not surpassing inner claw.
</paragraph>
<paragraph id="8BAF3699FFE0FFBEFF1FFDC3A7EB9F4F" blockId="4.[132,681,184,783]" pageId="4" pageNumber="173">
Uropod (
<figureCitation id="132B2A1CFFE0FFBEFE83FDC3A6279E30" box="[289,374,600,624]" captionStart="FIGURE 2" captionStartId="3.[95,174,1826,1845]" captionTargetBox="[94,1228,170,1802]" captionTargetId="figure-1@3.[94,1228,170,1802]" captionTargetPageId="3" captionText="FIGURE 2: Benthana carijos sp. nov. (male paratype). A: pereopod 1; B: pereopod 7; C: uropod; D: genital papilla; E: pleopod 1; F: pleopod 2; G: pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.5755772" httpUri="https://zenodo.org/record/5755772/files/figure.png" pageId="4" pageNumber="173">Fig. 2C</figureCitation>
) with protopod grooved on outer margin, endopod and exopod inserted almost at the same level, with few long setae along inner and outer margins; exopod slightly longer than endopod, with six setae on apex; endopod with three setae on apex.
</paragraph>
<paragraph id="8BAF3699FFE0FFBEFF26FCA3A6559890" blockId="4.[132,682,824,1232]" pageId="4" pageNumber="173">
<emphasis id="B964EA8BFFE0FFBEFF26FCA3A7EC9F10" box="[132,189,824,848]" italics="true" pageId="4" pageNumber="173">Male:</emphasis>
Genital papilla (
<figureCitation id="132B2A1CFFE0FFBEFECDFCA3A6939F10" box="[367,450,824,848]" captionStart="FIGURE 2" captionStartId="3.[95,174,1826,1845]" captionTargetBox="[94,1228,170,1802]" captionTargetId="figure-1@3.[94,1228,170,1802]" captionTargetPageId="3" captionText="FIGURE 2: Benthana carijos sp. nov. (male paratype). A: pereopod 1; B: pereopod 7; C: uropod; D: genital papilla; E: pleopod 1; F: pleopod 2; G: pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.5755772" httpUri="https://zenodo.org/record/5755772/files/figure.png" pageId="4" pageNumber="173">Fig. 2D</figureCitation>
) with triangular frontal shield distally rounded and two subapical orifices. Pleopod 1 (
<figureCitation id="132B2A1CFFE0FFBEFF5DFCE3A61F9FD0" box="[255,334,888,912]" captionStart="FIGURE 2" captionStartId="3.[95,174,1826,1845]" captionTargetBox="[94,1228,170,1802]" captionTargetId="figure-1@3.[94,1228,170,1802]" captionTargetPageId="3" captionText="FIGURE 2: Benthana carijos sp. nov. (male paratype). A: pereopod 1; B: pereopod 7; C: uropod; D: genital papilla; E: pleopod 1; F: pleopod 2; G: pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.5755772" httpUri="https://zenodo.org/record/5755772/files/figure.png" pageId="4" pageNumber="173">Fig. 2E</figureCitation>
) exopod heart-shaped, elongated (ratio z:y 2.56), dentiform protrusion not prominent, rounded apex and distal margin with low indentation, outer lateral margin of respiratory area with six setae, and narrow distal third; endopod stout with median row of short setae on distal portion. Pleopod 2 (
<figureCitation id="132B2A1CFFE0FFBEFF6DFBA3A64A9810" box="[207,283,1080,1104]" captionStart="FIGURE 2" captionStartId="3.[95,174,1826,1845]" captionTargetBox="[94,1228,170,1802]" captionTargetId="figure-1@3.[94,1228,170,1802]" captionTargetPageId="3" captionText="FIGURE 2: Benthana carijos sp. nov. (male paratype). A: pereopod 1; B: pereopod 7; C: uropod; D: genital papilla; E: pleopod 1; F: pleopod 2; G: pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.5755772" httpUri="https://zenodo.org/record/5755772/files/figure.png" pageId="4" pageNumber="173">Fig. 2F</figureCitation>
) exopod triangular, outer margin concave with about 12 setae; endopod slender and long. Pleopod 5 exopod (
<figureCitation id="132B2A1CFFE0FFBEFEEDFBE3A6F098D0" box="[335,417,1144,1168]" captionStart="FIGURE 2" captionStartId="3.[95,174,1826,1845]" captionTargetBox="[94,1228,170,1802]" captionTargetId="figure-1@3.[94,1228,170,1802]" captionTargetPageId="3" captionText="FIGURE 2: Benthana carijos sp. nov. (male paratype). A: pereopod 1; B: pereopod 7; C: uropod; D: genital papilla; E: pleopod 1; F: pleopod 2; G: pleopod 5 exopod." figureDoi="http://doi.org/10.5281/zenodo.5755772" httpUri="https://zenodo.org/record/5755772/files/figure.png" pageId="4" pageNumber="173">Fig. 2G</figureCitation>
) triangular, outer margin slightly sinuous with four setae and transverse plumose fringe.
</paragraph>
<paragraph id="8BAF3699FFE0FFBEFF26FB63A5D69970" blockId="4.[132,680,1272,1328]" pageId="4" pageNumber="173">
<emphasis id="B964EA8BFFE0FFBEFF26FB63A7A19950" box="[132,240,1272,1296]" italics="true" pageId="4" pageNumber="173">Etymology:</emphasis>
The new species is named after the Carijós people native of southeastern and southern
<collectingCountry id="F3077609FFE0FFBEFDE6FA83A5D29970" box="[580,643,1304,1328]" name="Brazil" pageId="4" pageNumber="173">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="8BAF3699FFE0FFBEFF26FAC3A6A59B10" blockId="4.[132,681,1368,1872]" pageId="4" pageNumber="173">
<emphasis id="B964EA8BFFE0FFBEFF26FAC3A78F9930" box="[132,222,1368,1392]" italics="true" pageId="4" pageNumber="173">Remarks:</emphasis>
The male pleopod 1 exopod of
<taxonomicName id="4C104D1AFFE0FFBEFDEBFAC3A79399D0" authority="Costa &amp; Campos-Filho &amp; Araujo, 2014" authorityName="Costa &amp; Campos-Filho &amp; Araujo" authorityYear="2014" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="carijos" status="sp. nov.">
<emphasis id="B964EA8BFFE0FFBEFDEBFAC3A79399D0" italics="true" pageId="4" pageNumber="173">Benthana carijos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757F0FFE0FFBEFF6DFAE3A67599CF" box="[207,292,1400,1423]" pageId="4" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
is similar to that of
<taxonomicName id="4C104D1AFFE0FFBEFDBAFAE2A62D99F0" authority="Lemos de Castro, 1958" authorityName="Lemos de Castro" authorityYear="1958" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="bocainensis">
<emphasis id="B964EA8BFFE0FFBEFDBAFAE2A5F999D0" box="[536,680,1400,1424]" italics="true" pageId="4" pageNumber="173">B. bocainensis</emphasis>
<bibRefCitation id="EF814B68FFE0FFBEFF26FA03A62D99F0" author="LEMOS DE CASTRO, A." box="[132,380,1432,1456]" pageId="4" pageNumber="173" pagination="85 - 118" refId="ref5981" refString="LEMOS DE CASTRO, A. 1958. Revisao do ganero Benthana Budde- - Lund, 1908 (Isopoda, Oniscidae). Arquivos do Museu Nacional, 44: 85 - 118." type="journal article" year="1958">Lemos de Castro, 1958</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C104D1AFFE0FFBEFE2BFA02A7E89990" authority="Lemos de Castro, 1958" authorityName="Lemos de Castro" authorityYear="1958" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="santosi">
<emphasis id="B964EA8BFFE0FFBEFE2BFA02A6B999F0" box="[393,488,1433,1456]" italics="true" pageId="4" pageNumber="173">B. santosi</emphasis>
<bibRefCitation id="EF814B68FFE0FFBEFE53FA03A7E89990" author="LEMOS DE CASTRO, A." pageId="4" pageNumber="173" pagination="85 - 118" refId="ref5981" refString="LEMOS DE CASTRO, A. 1958. Revisao do ganero Benthana Budde- - Lund, 1908 (Isopoda, Oniscidae). Arquivos do Museu Nacional, 44: 85 - 118." type="journal article" year="1958">Lemos de Castro, 1958</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C104D1AFFE0FFBEFF4FFA22A50F9990" authority="Lemos de Castro, 1958" authorityName="Lemos de Castro" authorityYear="1958" box="[237,606,1464,1488]" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="schubarti">
<emphasis id="B964EA8BFFE0FFBEFF4FFA22A6359990" box="[237,356,1464,1488]" italics="true" pageId="4" pageNumber="173">B. schubarti</emphasis>
<bibRefCitation id="EF814B68FFE0FFBEFEC9FA23A50F9990" author="LEMOS DE CASTRO, A." box="[363,606,1464,1488]" pageId="4" pageNumber="173" pagination="85 - 118" refId="ref5981" refString="LEMOS DE CASTRO, A. 1958. Revisao do ganero Benthana Budde- - Lund, 1908 (Isopoda, Oniscidae). Arquivos do Museu Nacional, 44: 85 - 118." type="journal article" year="1958">Lemos de Castro, 1958</bibRefCitation>
</taxonomicName>
, by the not prominent dentiform protrusion and low indentation, but can be distinguished by the absence of lobe on the inner margin (
<emphasis id="B964EA8BFFE0FFBEFED1F982A6DC9A70" box="[371,397,1561,1584]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
present in
<taxonomicName id="4C104D1AFFE0FFBEFDA9F982A5C89A70" authorityName="Lemos de Castro" authorityYear="1958" box="[523,665,1560,1584]" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="bocainensis">
<emphasis id="B964EA8BFFE0FFBEFDA9F982A5C89A70" box="[523,665,1560,1584]" italics="true" pageId="4" pageNumber="173">B. bocainensis</emphasis>
</taxonomicName>
), and presence of setae on outer margin (
<emphasis id="B964EA8BFFE0FFBEFD8AF9A2A5109A10" box="[552,577,1593,1616]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
absent in
<taxonomicName id="4C104D1AFFE0FFBEFF26F9C2A7B59A30" authorityName="Lemos de Castro" authorityYear="1958" box="[132,228,1625,1648]" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="santosi">
<emphasis id="B964EA8BFFE0FFBEFF26F9C2A7B59A30" box="[132,228,1625,1648]" italics="true" pageId="4" pageNumber="173">B. santosi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C104D1AFFE0FFBEFEB9F9C2A6C29A30" authorityName="Lemos de Castro" authorityYear="1958" box="[283,403,1624,1648]" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="schubarti">
<emphasis id="B964EA8BFFE0FFBEFEB9F9C2A6C29A30" box="[283,403,1624,1648]" italics="true" pageId="4" pageNumber="173">B. schubarti</emphasis>
</taxonomicName>
).
<taxonomicName id="4C104D1AFFE0FFBEFE09F9C3A5009A30" authority="Costa &amp; Campos-Filho &amp; Araujo, 2014" authorityName="Costa &amp; Campos-Filho &amp; Araujo" authorityYear="2014" box="[427,593,1624,1648]" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="carijos" status="sp. nov.">
<emphasis id="B964EA8BFFE0FFBEFE09F9C3A5009A30" box="[427,593,1624,1648]" italics="true" pageId="4" pageNumber="173">Benthana carijos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25757F0FFE0FFBEFDFBF9C3A5F89A2F" box="[601,681,1624,1647]" pageId="4" pageNumber="173" rank="species">sp. nov.</taxonomicNameLabel>
can also be distinguished by the number of aesthetascs on the antennule (16+2
<emphasis id="B964EA8BFFE0FFBEFE20F902A6CD9AF0" box="[386,412,1689,1712]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
8+
<quantity id="4CE89B7CFFE0FFBEFE1FF903A6B69AF0" box="[445,487,1688,1712]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="173" unit="in" value="2.0">2 in</quantity>
<emphasis id="B964EA8BFFE0FFBEFE4CF902A5289AF0" box="[494,633,1688,1712]" italics="true" pageId="4" pageNumber="173">
<taxonomicName id="4C104D1AFFE0FFBEFE4CF902A5249AF0" box="[494,629,1688,1712]" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="bocainenis">B. bocainenis</taxonomicName>
,
</emphasis>
9+
<quantity id="4CE89B7CFFE0FFBEFD39F903A7C89A90" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="173" unit="in" value="2.0">2 in</quantity>
<emphasis id="B964EA8BFFE0FFBEFF3DF922A6559A90" box="[159,260,1721,1744]" italics="true" pageId="4" pageNumber="173">
<taxonomicName id="4C104D1AFFE0FFBEFF3DF922A6519A90" authorityName="Lemos de Castro" authorityYear="1958" box="[159,256,1721,1744]" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="santosi">B. santosi</taxonomicName>
,
</emphasis>
and 6+
<quantity id="4CE89B7CFFE0FFBEFEF3F923A6289A90" box="[337,377,1720,1744]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="4" pageNumber="173" unit="in" value="2.0">2 in</quantity>
<taxonomicName id="4C104D1AFFE0FFBEFE22F922A6A79A90" authorityName="Lemos de Castro" authorityYear="1958" box="[384,502,1720,1744]" class="Malacostraca" family="Philosciidae" genus="Benthana" kingdom="Animalia" order="Isopoda" pageId="4" pageNumber="173" phylum="Arthropoda" rank="species" species="schubarti">
<emphasis id="B964EA8BFFE0FFBEFE22F922A6A79A90" box="[384,502,1720,1744]" italics="true" pageId="4" pageNumber="173">B. schubarti</emphasis>
</taxonomicName>
), uropod exopod slightly longer than endopod (
<emphasis id="B964EA8BFFE0FFBEFE1AF942A6809AB0" box="[440,465,1753,1776]" italics="true" pageId="4" pageNumber="173">vs.</emphasis>
exopod twice as long as endopod in all three species), and the shape of the male pleopod 1 exopod with narrow distal third, an unique characteristic of this species.
</paragraph>
</subSubSection>
</treatment>
</document>