treatments-xml/data/03/B8/87/03B887E8FFB0A607FF14AC186620FB99.xml
2024-06-21 12:22:17 +02:00

648 lines
76 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="95416A9C9792EB883CF6DAC4C28037EE" ID-CLB-Dataset="20328" ID-DOI="10.11646/zootaxa.5105.4.1" ID-GBIF-Dataset="45708bdc-2d73-4b5d-91d9-75d9f31665a2" ID-ISSN="1175-5326" ID-Zenodo-Dep="6333771" ID-ZooBank="E81605D2-5B90-446A-B209-57EEFC8EA396" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1646640338754" checkinUser="plazi" docAuthor="Huang, Hao &amp; Song, Kui" docDate="2022" docId="03B887E8FFB0A607FF14AC186620FB99" docLanguage="en" docName="zootaxa.5105.4.1.pdf" docOrigin="Zootaxa 5105 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Colias sifanica Grum-Grshimailo 1891" docType="treatment" docVersion="8" lastPageNumber="460" masterDocId="FF81FF90FFB8A60EFF83A826621BFFC0" masterDocTitle="A review of the genus Colias Fabricius, 1807 (Lepidoptera: Pieridae) from Qinghai with descriptions of a new subspecies of Colias thrasibulus Fruhstorfer, 1910" masterLastPageNumber="489" masterPageNumber="451" pageNumber="459" updateTime="1699317299447" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="E478AA90013D64BA85FF8693B7ECF9A6" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1C9FCA61F61B267EDD094880C5202D88">
<mods:title id="D1CD401FDACA87E3F1F48DA2D0775C58">A review of the genus Colias Fabricius, 1807 (Lepidoptera: Pieridae) from Qinghai with descriptions of a new subspecies of Colias thrasibulus Fruhstorfer, 1910</mods:title>
</mods:titleInfo>
<mods:name id="8F8543CE98530915A7C218EF2378EFC1" type="personal">
<mods:role id="E730C2249B5D34453E93746672612796">
<mods:roleTerm id="42A75FBABAA23CEF70BFFAA0C3F92F8A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EC7553BF9A3A251D98F9AAC7FFAC4CC7">Huang, Hao</mods:namePart>
<mods:affiliation id="3128294855BD9E4E76FD6DBA3B2AC9C3">503, Unit 1, # 1 Dongtinghu Road, Qingdao, P. R. China.</mods:affiliation>
</mods:name>
<mods:name id="E3137F97AB8F26F1B24AD03F14E705C9" type="personal">
<mods:role id="7866D2F71BEE5894CCE5FB9AE54A750B">
<mods:roleTerm id="717D30FF6E90F63FD235E2E554616D49">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B3C37C5CD9E70D92B0633103EE64524A">Song, Kui</mods:namePart>
<mods:nameIdentifier id="0B998D79975568A3D4DE8008A56314C7" type="ORCID">0000-0002-6928-7869</mods:nameIdentifier>
<mods:affiliation id="160C0EA607EAA1BE56E1C869CAE8FD75">School of Economics and Management, Qinghai Nationalities University, Bayi Road No. 3, Xining, 810007, Qinghai, P. R. China. 76342870 @ qq. com; https: // orcid. org / 0000 - 0002 - 6928 - 7869</mods:affiliation>
<mods:nameIdentifier id="695A6B87AD7D656BB529C35FF0696D57" type="email">76342870@qq.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="F1878CFD242AD26EEE26314349CB9384">text</mods:typeOfResource>
<mods:relatedItem id="1C8363CFF89472A6846C91CDA163180C" type="host">
<mods:titleInfo id="9FA775B4AF72FAF4D31AE181A926629E">
<mods:title id="5688B99D933EDB9DF0865CB878F97321">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F961114DE517ADBE3F6B1E07565B3972">
<mods:date id="2F9A97AADE27B5D0B954AAF575D0F8D1">2022</mods:date>
<mods:detail id="D128CC4F8723065EEF1A274742C38AD2" type="pubDate">
<mods:number id="FFCF5E44CBB24953B694F44ADED944DD">2022-03-07</mods:number>
</mods:detail>
<mods:detail id="1F86F8F757CDD9917EEBF6EAF5D7DA72" type="volume">
<mods:number id="9B76FFC4BD73A1B24F1F9C7EB79ED5D6">5105</mods:number>
</mods:detail>
<mods:detail id="D36595B81931282E84B4FB8DC5E327E4" type="issue">
<mods:number id="8385A856EC674C3982931B032798DF0A">4</mods:number>
</mods:detail>
<mods:extent id="76EEF58C2C4A7A16A2997F92E03C4CC8" unit="page">
<mods:start id="5B3A2E1A94F688BEE65B9C339D83C02F">451</mods:start>
<mods:end id="1A1967222835060C1622A80909FB832F">489</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="33CE06995FB171D2371BA91DB7BCF7BF">journal article</mods:classification>
<mods:identifier id="33D837D702B8CB88844038A74236892B" type="CLB-Dataset">20328</mods:identifier>
<mods:identifier id="7F5ABE85BFDCE1735F82A2EA3AA2A509" type="DOI">10.11646/zootaxa.5105.4.1</mods:identifier>
<mods:identifier id="6AEAAF8AA304037D032A89D3AEC616A4" type="GBIF-Dataset">45708bdc-2d73-4b5d-91d9-75d9f31665a2</mods:identifier>
<mods:identifier id="BBDD30DC40B81723157F1ED79DF5EEF3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="774FE23EF7C78D13A3285DCC7C8B9A4C" type="Zenodo-Dep">6333771</mods:identifier>
<mods:identifier id="275EFA52FA8F64846EDA6FBCFA43E476" type="ZooBank">E81605D2-5B90-446A-B209-57EEFC8EA396</mods:identifier>
</mods:mods>
<treatment id="03B887E8FFB0A607FF14AC186620FB99" ID-DOI="http://doi.org/10.5281/zenodo.6343664" ID-GBIF-Taxon="193892552" ID-Zenodo-Dep="6343664" LSID="urn:lsid:plazi:treatment:03B887E8FFB0A607FF14AC186620FB99" httpUri="http://treatment.plazi.org/id/03B887E8FFB0A607FF14AC186620FB99" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459">
<subSubSection id="C30B6575FFB0A606FF14AC18609BFB98" box="[151,640,1086,1113]" pageId="8" pageNumber="459" type="nomenclature">
<paragraph id="8BAE36FEFFB0A606FF14AC18609BFB98" blockId="8.[151,640,1086,1149]" box="[151,640,1086,1113]" pageId="8" pageNumber="459">
<heading id="D0E68192FFB0A606FF14AC18609BFB98" bold="true" box="[151,640,1086,1113]" fontSize="11" level="1" pageId="8" pageNumber="459" reason="1">
<taxonomicName id="4C114D7DFFB0A606FF14AC18609BFB98" ID-CoL="X2GM" authority="Grum-Grshimailo, 1891" authorityName="Grum-Grshimailo" authorityYear="1891" box="[151,640,1086,1113]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB0A606FF14AC18609BFB98" bold="true" box="[151,640,1086,1113]" pageId="8" pageNumber="459">
<emphasis id="B965EAECFFB0A606FF14AC186350FB98" bold="true" box="[151,331,1086,1113]" italics="true" pageId="8" pageNumber="459">Colias sifanica</emphasis>
<bibRefCitation id="EF804B0FFFB0A606FED0AC18609BFB98" author="Grum-Grshimailo, G. E." box="[339,640,1086,1112]" pageId="8" pageNumber="459" pagination="445 - 465" refId="ref15222" refString="Grum-Grshimailo, G. E. (1891) Lepidoptera nova in Asia Centrali novissime lecta et descripta. Horae Societatis entomologiscae rossicae, 25 (3 / 4), 445 - 465." type="journal article" year="1891">Grum-Grshimailo, 1891</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30B6575FFB0A606FF14AC44604FFBBC" box="[151,596,1122,1149]" pageId="8" pageNumber="459" type="description">
<paragraph id="8BAE36FEFFB0A606FF14AC44604FFBBC" blockId="8.[151,640,1086,1149]" box="[151,596,1122,1149]" pageId="8" pageNumber="459">
(
<figureCitation id="132A2A7BFFB0A606FF1CAC4462E6FBBC" box="[159,253,1122,1148]" captionStart="FIGURE 14" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[207,1380,181,1919]" captionTargetId="figure-19@17.[207,1380,181,1919]" captionTargetPageId="17" captionText="FIGURE 14. Habitus of Colias sifanica (with form aegidii) ♂♂ &amp; ♀ at same scale (Upper half- upper side; lower half- under- side). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.6333783" httpUri="https://zenodo.org/record/6333783/files/figure.png" pageId="8" pageNumber="459">Figs. 14</figureCitation>
,
<figureCitation id="132A2A7BFFB0A606FE8AAC44634AFBBC" box="[265,337,1122,1148]" captionStart-0="FIGURE 42" captionStart-1="FIGURE 43" captionStartId-0="24.[151,250,1943,1967]" captionStartId-1="25.[151,250,1943,1967]" captionTargetBox-0="[216,1372,181,1920]" captionTargetBox-1="[218,1370,181,1920]" captionTargetId-0="figure-21@24.[216,1372,181,1920]" captionTargetId-1="figure-19@25.[218,1370,181,1920]" captionTargetPageId-0="24" captionTargetPageId-1="25" captionText-0="FIGURE 42. ♂ genitalia and 8th tergum (with superuncus) in lateral view at same scale. Scale bar = 1 mm." captionText-1="FIGURE 43. 8th tergum in dorsal view at same scale to show superuncus. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6333797" figureDoi-1="http://doi.org/10.5281/zenodo.6333799" httpUri-0="https://zenodo.org/record/6333797/files/figure.png" httpUri-1="https://zenodo.org/record/6333799/files/figure.png" pageId="8" pageNumber="459">4243</figureCitation>
,
<figureCitation id="132A2A7BFFB0A606FEDEAC4463BEFBBD" box="[349,421,1122,1149]" captionStart-0="FIGURE 45" captionStart-1="FIGURE 46" captionStartId-0="27.[151,250,1943,1967]" captionStartId-1="28.[151,250,1943,1967]" captionTargetBox-0="[216,1372,181,1920]" captionTargetBox-1="[216,1372,181,1909]" captionTargetId-0="figure-19@27.[216,1372,181,1920]" captionTargetId-1="figure-21@28.[216,1372,181,1920]" captionTargetPageId-0="27" captionTargetPageId-1="28" captionText-0="FIGURE 45. ♂ genitalia in dorsal view at same scale. Scale bar = 1 mm." captionText-1="FIGURE 46. Ring in lateral view and scaphium in ventral view at same scale. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6333807" figureDoi-1="http://doi.org/10.5281/zenodo.6333809" httpUri-0="https://zenodo.org/record/6333807/files/figure.png" httpUri-1="https://zenodo.org/record/6333809/files/figure.png" pageId="8" pageNumber="459">4546</figureCitation>
,
<figureCitation id="132A2A7BFFB0A606FE32AC4463E1FBBC" box="[433,506,1122,1148]" captionStart-0="FIGURE 48" captionStart-1="FIGURE 49" captionStartId-0="30.[151,250,1943,1967]" captionStartId-1="31.[151,250,1943,1967]" captionTargetBox-0="[216,1372,181,1920]" captionTargetBox-1="[234,1353,181,1920]" captionTargetId-0="figure-21@30.[216,1372,181,1920]" captionTargetId-1="figure-19@31.[234,1353,181,1920]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionText-0="FIGURE 48. Phallus in lateral view and coecum-process in full-face view at same scale. Scale bar = 1 mm." captionText-1="FIGURE 49. Juxta in full-face view (nearly posterior view) at same scale. Scale bar = 1 mm. FIGURE 50. Valva in inner lateral and anterior views at same scale. Scale bar = 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6333813" figureDoi-1="http://doi.org/10.5281/zenodo.6333815" httpUri-0="https://zenodo.org/record/6333813/files/figure.png" httpUri-1="https://zenodo.org/record/6333815/files/figure.png" pageId="8" pageNumber="459">4849</figureCitation>
,
<figureCitation id="132A2A7BFFB0A606FD86AC446038FBBC" box="[517,547,1122,1148]" captionStart="FIGURES 5964" captionStartId="33.[151,264,1849,1873]" captionTargetBox="[251,1337,181,1825]" captionTargetId="figure-19@33.[251,1337,181,1825]" captionTargetPageId="33" captionText="FIGURES 5964. Biotopes of Colias species in Qinghai. 59—C. diva (Lajishan, by K. Song); 60—C. sifanica (Lajishan, by K. Song); 61—C. lada (Yushu, by K. Song); 62—C. nebulosa (Lajishan, by H. Huang); 63—C. grumi (Xidatan, Kunlun Shan, by Z.-Y. Qi); 64—C. grumi (Dachaidan, by K. Song). FIGURES 6566. Living adults of Colias species in Qinghai. 65—C. arida cakana at Wulan (by Z.-Y. Qi); 66—C. grumi at Xidatan (by Z.-Y. Qi)." figureDoi="http://doi.org/10.5281/zenodo.6333823" httpUri="https://zenodo.org/record/6333823/files/figure.png" pageId="8" pageNumber="459">60</figureCitation>
, 70)
</paragraph>
</subSubSection>
<subSubSection id="C30B6575FFB0A606FF14AC8162F6F94B" pageId="8" pageNumber="459" type="reference_group">
<paragraph id="8BAE36FEFFB0A606FF14AC816738FAF9" blockId="8.[151,1437,1191,1675]" pageId="8" pageNumber="459">
<treatmentCitationGroup id="AB0111D0FFB0A606FF14AC816738FAF9" pageId="8" pageNumber="459">
<treatmentCitation id="0AB010EFFFB0A606FF14AC816047FB7F" author="Grum-Grshimailo, G. E." box="[151,604,1191,1215]" page="447" pageId="8" pageNumber="459" year="1891">
<taxonomicName id="4C114D7DFFB0A606FF14AC816047FB7F" ID-CoL="X2GM" authority="Grum-Grshimailo, 1891: 447" authorityName="Grum-Grshimailo" authorityPageNumber="447" authorityYear="1891" box="[151,604,1191,1215]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB0A606FF14AC816334FB7F" box="[151,303,1191,1215]" italics="true" pageId="8" pageNumber="459">Colias sifanica</emphasis>
<bibRefCitation id="EF804B0FFFB0A606FEB6AC816047FB7F" author="Grum-Grshimailo, G. E." box="[309,604,1191,1215]" pageId="8" pageNumber="459" pagination="445 - 465" refId="ref15222" refString="Grum-Grshimailo, G. E. (1891) Lepidoptera nova in Asia Centrali novissime lecta et descripta. Horae Societatis entomologiscae rossicae, 25 (3 / 4), 445 - 465." type="journal article" year="1891">Grum-Grshimailo, 1891: 447</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(TL: Laji Shan and Riyue Shan:
<bibRefCitation id="EF804B0FFFB0A606FC29AC81668DFB7F" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[938,1174,1191,1215]" pageId="8" pageNumber="459" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">
Grieshuber
<emphasis id="B965EAECFFB0A606FBA3AC816648FB7F" box="[1056,1107,1191,1215]" italics="true" pageId="8" pageNumber="459">et al.</emphasis>
, 2012
</bibRefCitation>
); Verity, 1911: pl. lxx, fig. 1, cpl. I: 1,
<typeStatus id="54AA885CFFB0A606FEBEACE36381FB1D" box="[317,410,1221,1245]" pageId="8" pageNumber="459" type="lectotype">lectotype</typeStatus>
designation (
<bibRefCitation id="EF804B0FFFB0A606FDA0ACE36116FB1D" author="Grieshuber, J. &amp; Churkin, S. &amp; Worthy, R. &amp; Lvovsky, A." box="[547,781,1221,1245]" pageId="8" pageNumber="459" pagination="10 - 24" refId="ref15057" refString="Grieshuber, J., Churkin, S., Worthy, R. &amp; Lvovsky, A. (2004) The types of Colias felderi Grum-Grshimailo, 1891, Colias lada Grum-Grshimailo, 1891, and Colias sifanica Alpheraky, 1891, with additional notes on the types of Colias diva Grum- Grshimailo, 1891 (Lepidoptera: Pieridae). Helios, Moscow, 5, 10 - 24, pl. 1, figs. 1 - 7." type="journal article" year="2004">
Grieshuber
<emphasis id="B965EAECFFB0A606FD1FACE060D2FB1D" box="[668,713,1221,1245]" italics="true" pageId="8" pageNumber="459">et al</emphasis>
. 2004
</bibRefCitation>
);
<treatmentCitation id="0AB010EFFFB0A606FC9CACE361D9FB1D" author="Hoshiai, A." box="[799,962,1221,1245]" page="9" pageId="8" pageNumber="459" year="1996">
<bibRefCitation id="EF804B0FFFB0A606FC9CACE361D9FB1D" author="Hoshiai, A." box="[799,962,1221,1245]" pageId="8" pageNumber="459" pagination="8 - 10" refId="ref15351" refString="Hoshiai, A. (1996) Notes of Colias from China and surrounding area (1). In: Koiwaya, S. (Ed.), Studies of Chinese Butterflies. Vol. 3. S. Koiwaya, Tokyo, pp. 211 - 217, pls. 8 - 10. [in Japanese]" type="book chapter" year="1996">Hoshiai 1996: 9</bibRefCitation>
</treatmentCitation>
-pl. figs. 3953 for
<specimenCount id="9D17FD77FFB0A606FB0BACE066B5FB1D" box="[1160,1198,1222,1245]" pageId="8" pageNumber="459" type="male">♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFB0A606FB4CACE066EEFB1D" box="[1231,1269,1222,1245]" pageId="8" pageNumber="459" type="female">♀♀</specimenCount>
from Xiahe and Zhugqu (Zhouqu);
<bibRefCitation id="EF804B0FFFB0A606FE04ACC2600DFB3C" author="Verhulst, J." box="[391,534,1252,1276]" pageId="8" pageNumber="459" refId="ref16453" refString="Verhulst, J. (2001) Les Colias du Globe. Monograph of the genus Colias. Plates. Goecke &amp; Evers, Kelterns, 45 pp., 183 pls. [in French]" type="book" year="2001">Verhulst 2001</bibRefCitation>
: pl. 40, figs. 124 for
<specimenCount id="9D17FD77FFB0A606FD73ACC2610DFB3B" box="[752,790,1252,1275]" pageId="8" pageNumber="459" type="male">♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFB0A606FCB7ACC26140FB3B" box="[820,859,1252,1275]" pageId="8" pageNumber="459" type="female">♀♀</specimenCount>
; pl. 138 for distribution and biotope;
<treatmentCitation id="0AB010EFFFB0A606FB53ACC2677DFB3C" author="Dietz, M." box="[1232,1382,1252,1276]" page="71" pageId="8" pageNumber="459" year="2002">
<bibRefCitation id="EF804B0FFFB0A606FB53ACC2677DFB3C" author="Dietz, M." box="[1232,1382,1252,1276]" pageId="8" pageNumber="459" refId="ref14565" refString="Dietz, M. (2002) Pieridae. Colias Fabricius 1807. Species and subspecies of the Palaearctic Region. Part 1. Selbstverlag, Solms, 154 pp. [in Germany]" type="book" year="2002">Dietz 2002: 71</bibRefCitation>
</treatmentCitation>
, figs. 19 for
<specimenCount id="9D17FD77FFB0A606FE9AAD256324FADA" box="[281,319,1283,1306]" pageId="8" pageNumber="459" type="male">♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFB0A606FEE1AD256392FADA" box="[354,393,1283,1306]" pageId="8" pageNumber="459" type="female">♀♀</specimenCount>
;
<treatmentCitation id="0AB010EFFFB0A606FE15AD2560BEFADB" author="Grieshuber, J. &amp; Churkin, S. &amp; Worthy, R. &amp; Lvovsky, A." box="[406,677,1283,1307]" page="16" pageId="8" pageNumber="459" year="2004">
<bibRefCitation id="EF804B0FFFB0A606FE15AD2560BEFADB" author="Grieshuber, J. &amp; Churkin, S. &amp; Worthy, R. &amp; Lvovsky, A." box="[406,677,1283,1307]" pageId="8" pageNumber="459" pagination="10 - 24" refId="ref15057" refString="Grieshuber, J., Churkin, S., Worthy, R. &amp; Lvovsky, A. (2004) The types of Colias felderi Grum-Grshimailo, 1891, Colias lada Grum-Grshimailo, 1891, and Colias sifanica Alpheraky, 1891, with additional notes on the types of Colias diva Grum- Grshimailo, 1891 (Lepidoptera: Pieridae). Helios, Moscow, 5, 10 - 24, pl. 1, figs. 1 - 7." type="journal article" year="2004">
Grieshuber
<emphasis id="B965EAECFFB0A606FD8DAD256059FADB" box="[526,578,1283,1307]" italics="true" pageId="8" pageNumber="459">et al.</emphasis>
2004: 16
</bibRefCitation>
</treatmentCitation>
, discussion on
<typeStatus id="54AA885CFFB0A606FCC1AD2561B9FADB" box="[834,930,1283,1307]" pageId="8" pageNumber="459" type="lectotype">lectotype</typeStatus>
;
<treatmentCitation id="0AB010EFFFB0A606FC2DAD2566D1FADA" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[942,1226,1282,1307]" page="205" pageId="8" pageNumber="459" year="2012">
<bibRefCitation id="EF804B0FFFB0A606FC2DAD2566D1FADA" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[942,1226,1282,1307]" pageId="8" pageNumber="459" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">
Grieshuber
<emphasis id="B965EAECFFB0A606FBA5AD256648FADB" box="[1062,1107,1283,1307]" italics="true" pageId="8" pageNumber="459">et al</emphasis>
. 2012: 205
</bibRefCitation>
</treatmentCitation>
, discussions on type series and type locality;
<treatmentCitation id="0AB010EFFFB0A606FE3EAD07608BFAF9" author="Grieshuber, J" box="[445,656,1313,1337]" page="30" pageId="8" pageNumber="459" year="2014">
<bibRefCitation id="EF804B0FFFB0A606FE3EAD07608BFAF9" author="Grieshuber, J" box="[445,656,1313,1337]" pageId="8" pageNumber="459" refId="ref14730" refString="Grieshuber, J (2014) Guide to the butterflies of the Palearctic region. Pieridae Part II. Omnes Artes, Milano, 86 pp." type="book" year="2014">Grieshuber 2014: 30</bibRefCitation>
</treatmentCitation>
, figs.;
<bibRefCitation id="EF804B0FFFB0A606FD57AD076197FAF9" author="Grieshuber, J." box="[724,908,1313,1337]" pageId="8" pageNumber="459" refId="ref14798" refString="Grieshuber, J. (2016 a) Specimen of Colias in Museum of Zoology, the Zoological Institute of the Russian Academy of Sciences (Saint Petersburg). Available from: https: // commons. wikimedia. org / wiki / Category: Specimen _ of _ Colias _ in _ Museum _ of _ Zoology _ (Saint _ Petersburg) (accessed 23 October 2021)" type="url" year="2016">Grieshuber 2016a</bibRefCitation>
: photos of
<typeStatus id="54AA885CFFB0A606FC7FAD076642FAF9" box="[1020,1113,1313,1337]" pageId="8" pageNumber="459" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="54AA885CFFB0A606FB09AD07673BFAF9" box="[1162,1312,1313,1337]" pageId="8" pageNumber="459" type="paralectotype">paralectotypes</typeStatus>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFB0A606FF14AD666121FA55" blockId="8.[151,1437,1191,1675]" pageId="8" pageNumber="459">
<treatmentCitationGroup id="AB0111D0FFB0A606FF14AD666121FA55" pageId="8" pageNumber="459">
<treatmentCitation id="0AB010EFFFB0A606FF14AD66605DFA98" author="Bollow, C." box="[151,582,1344,1368]" page="109" pageId="8" pageNumber="459" year="1930">
<taxonomicName id="4C114D7DFFB0A606FF14AD66605DFA98" authority="Bollow, 1930: 109" authorityName="Bollow" authorityPageNumber="109" authorityYear="1930" box="[151,582,1344,1368]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="subSpecies" species="sifanica" subSpecies="herculeana">
<emphasis id="B965EAECFFB0A606FF14AD666364FA98" box="[151,383,1344,1368]" italics="true" pageId="8" pageNumber="459">C. sifanica herculeana</emphasis>
<bibRefCitation id="EF804B0FFFB0A606FE04AD66605DFA98" author="Bollow, C." box="[391,582,1344,1368]" pageId="8" pageNumber="459" pagination="107 - 112" refId="ref14383" refString="Bollow, C. (1930) Pieridae. Colias. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Abteilung I. Die Gross-Schmetterlinge des Palaearktischen Faunengebiets. Supplement zu Band 1. Die palaearktischen Tagfalter. Alfred Kernen, Stuttgart, pp. 107 - 112. [in Germany]" type="book chapter" year="1930">Bollow, 1930: 109</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(TL:
<collectingRegion id="49D5F81CFFB0A606FD07AD6660CDFA98" box="[644,726,1344,1368]" country="China" name="Qinghai" pageId="8" pageNumber="459">Qinghai</collectingRegion>
lake area:
<bibRefCitation id="EF804B0FFFB0A606FCCBAD66662FFA98" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[840,1076,1344,1368]" pageId="8" pageNumber="459" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">
Grieshuber
<emphasis id="B965EAECFFB0A606FC42AD6761EEFA98" box="[961,1013,1344,1368]" italics="true" pageId="8" pageNumber="459">et al.</emphasis>
2012
</bibRefCitation>
);
<treatmentCitation id="0AB010EFFFB0A606FBC5AD66673BFA98" author="Verhulst, J." box="[1094,1312,1344,1368]" page="185" pageId="8" pageNumber="459" year="1991">
<bibRefCitation id="EF804B0FFFB0A606FBC5AD66673BFA98" author="Verhulst, J." box="[1094,1312,1344,1368]" pageId="8" pageNumber="459" pagination="181 - 187" refId="ref16196" refString="Verhulst, J. (1991 b) Fiches specifiques des Colias F. (Colias data sheets). COLIAS SIFANICA. Lambillionea, 91 (3), 181 - 187. [in French]" type="journal article" year="1991">Verhulst, 1991b: 185</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AB010EFFFB0A606FAAFAD66637BFAB7" author="Grieshuber, J. &amp; Churkin, S. &amp; Worthy, R. &amp; Lvovsky, A." page="20" pageId="8" pageNumber="459" year="2004">
<bibRefCitation id="EF804B0FFFB0A606FAAFAD66637BFAB7" author="Grieshuber, J. &amp; Churkin, S. &amp; Worthy, R. &amp; Lvovsky, A." pageId="8" pageNumber="459" pagination="10 - 24" refId="ref15057" refString="Grieshuber, J., Churkin, S., Worthy, R. &amp; Lvovsky, A. (2004) The types of Colias felderi Grum-Grshimailo, 1891, Colias lada Grum-Grshimailo, 1891, and Colias sifanica Alpheraky, 1891, with additional notes on the types of Colias diva Grum- Grshimailo, 1891 (Lepidoptera: Pieridae). Helios, Moscow, 5, 10 - 24, pl. 1, figs. 1 - 7." type="journal article" year="2004">
Grieshuber
<emphasis id="B965EAECFFB0A606FF44AD7962E7FAB7" box="[199,252,1375,1399]" italics="true" pageId="8" pageNumber="459">et al.</emphasis>
2004: 20
</bibRefCitation>
</treatmentCitation>
, synonymy for
<taxonomicName id="4C114D7DFFB0A606FD82AD79606BFAB7" ID-CoL="X2GM" authorityName="Grum-Grshimailo" authorityYear="1891" box="[513,624,1375,1399]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB0A606FD82AD79606BFAB7" box="[513,624,1375,1399]" italics="true" pageId="8" pageNumber="459">C. sifanica</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AB010EFFFB0A606FDFCAD7961D6FAB7" author="Grieshuber, J. &amp; Lamas, G." box="[639,973,1375,1399]" page="161" pageId="8" pageNumber="459" year="2007">
<bibRefCitation id="EF804B0FFFB0A606FDFCAD7961D6FAB7" author="Grieshuber, J. &amp; Lamas, G." box="[639,973,1375,1399]" pageId="8" pageNumber="459" pagination="131 - 171" refId="ref15140" refString="Grieshuber, J. &amp; Lamas, G. (2007) A Synonymic List of the Genus Colias Fabricius, 1807 (Lepidoptera: Pieridae). Mitteilungen der Munchner Entomologischen Gesellschaft, 97, 131 - 171." type="journal article" year="2007">Grieshuber &amp; Lamas, 2007: 161</bibRefCitation>
</treatmentCitation>
, synonymic list;
<treatmentCitation id="0AB010EFFFB0A606FBFEAD796783FAB7" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[1149,1432,1375,1399]" page="117" pageId="8" pageNumber="459" year="2012">
<bibRefCitation id="EF804B0FFFB0A606FBFEAD796783FAB7" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[1149,1432,1375,1399]" pageId="8" pageNumber="459" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">Grieshuber et al, 2012: 117</bibRefCitation>
</treatmentCitation>
, discussions on
<typeStatus id="54AA885CFFB0A606FEE2AD586396FA56" box="[353,397,1406,1430]" pageId="8" pageNumber="459">type</typeStatus>
series,
<typeStatus id="54AA885CFFB0A606FE5AAD58601EFA56" box="[473,517,1406,1430]" pageId="8" pageNumber="459">type</typeStatus>
locality and taxonomic status.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFB0A606FF14ADBA6366F9D0" blockId="8.[151,1437,1191,1675]" pageId="8" pageNumber="459">
<treatmentCitationGroup id="AB0111D0FFB0A606FF14ADBA6366F9D0" pageId="8" pageNumber="459">
<taxonomicName id="4C114D7DFFB0A606FF14ADBA63EBFA74" ID-CoL="5ZLPJ" authority="Verhulst, 1990: 42" authorityName="Verhulst" authorityPageNumber="42" authorityYear="1990" box="[151,496,1436,1460]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB0A606FF14ADBA633CFA74" box="[151,295,1436,1460]" italics="true" pageId="8" pageNumber="459">Colias aegidii</emphasis>
Verhulst, 1990: 42
</taxonomicName>
, figs. 14 A for
<specimenCount id="9D17FD77FFB0A606FD19ADBA60B6FA73" box="[666,685,1436,1459]" pageId="8" pageNumber="459" type="male"></specimenCount>
genitalia, 41- pl. figs. for
<specimenCount id="9D17FD77FFB0A606FC43ADBA61FDFA73" box="[960,998,1436,1459]" pageId="8" pageNumber="459" type="male">♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFB0A606FB89ADBA662AFA73" box="[1034,1073,1436,1459]" pageId="8" pageNumber="459" type="female">♀♀</specimenCount>
;
<treatmentCitation id="0AB010EFFFB0A606FBBDADBA671EFA74" author="Hoshiai, A." box="[1086,1285,1436,1460]" page="216" pageId="8" pageNumber="459" year="1996">
<bibRefCitation id="EF804B0FFFB0A606FBBDADBA671EFA74" author="Hoshiai, A." box="[1086,1285,1436,1460]" pageId="8" pageNumber="459" pagination="8 - 10" refId="ref15351" refString="Hoshiai, A. (1996) Notes of Colias from China and surrounding area (1). In: Koiwaya, S. (Ed.), Studies of Chinese Butterflies. Vol. 3. S. Koiwaya, Tokyo, pp. 211 - 217, pls. 8 - 10. [in Japanese]" type="book chapter" year="1996">Hoshiai, 1996: 216</bibRefCitation>
</treatmentCitation>
, discussion on status;
<bibRefCitation id="EF804B0FFFB0A606FE8DAD9D63BDFA13" author="Verhulst, J." box="[270,422,1467,1491]" pageId="8" pageNumber="459" refId="ref16453" refString="Verhulst, J. (2001) Les Colias du Globe. Monograph of the genus Colias. Plates. Goecke &amp; Evers, Kelterns, 45 pp., 183 pls. [in French]" type="book" year="2001">Verhulst, 2001</bibRefCitation>
: pl. 39, figs. 124 for
<specimenCount id="9D17FD77FFB0A606FD0BAD9D60B5FA12" box="[648,686,1467,1490]" pageId="8" pageNumber="459" type="male">♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFB0A606FD4CAD9D60EDFA12" box="[719,758,1467,1490]" pageId="8" pageNumber="459" type="female">♀♀</specimenCount>
, pl. 139 for distribution and biotope;
<treatmentCitation id="0AB010EFFFB0A606FBF7AD9D670FFA13" author="Dietz, M." box="[1140,1300,1467,1491]" page="73" pageId="8" pageNumber="459" year="2002">
<bibRefCitation id="EF804B0FFFB0A606FBF7AD9D670FFA13" author="Dietz, M." box="[1140,1300,1467,1491]" pageId="8" pageNumber="459" refId="ref14565" refString="Dietz, M. (2002) Pieridae. Colias Fabricius 1807. Species and subspecies of the Palaearctic Region. Part 1. Selbstverlag, Solms, 154 pp. [in Germany]" type="book" year="2002">Dietz, 2002: 73</bibRefCitation>
</treatmentCitation>
, figs. 19 for
<specimenCount id="9D17FD77FFB0A606FF44ADFC62F6FA31" box="[199,237,1498,1521]" pageId="8" pageNumber="459" type="male">♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFB0A606FE93ADFC632CFA31" box="[272,311,1498,1521]" pageId="8" pageNumber="459" type="female">♀♀</specimenCount>
;
<treatmentCitation id="0AB010EFFFB0A606FEC6ADFF604FFA31" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[325,596,1497,1521]" page="33" pageId="8" pageNumber="459" year="2012">
<bibRefCitation id="EF804B0FFFB0A606FEC6ADFF604FFA31" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[325,596,1497,1521]" pageId="8" pageNumber="459" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">Grieshuber et al, 2012: 33</bibRefCitation>
</treatmentCitation>
, discussions on
<typeStatus id="54AA885CFFB0A606FD7FADFC6133FA32" box="[764,808,1498,1522]" pageId="8" pageNumber="459">type</typeStatus>
series,
<typeStatus id="54AA885CFFB0A606FCF4ADFC61B8FA32" box="[887,931,1498,1522]" pageId="8" pageNumber="459">type</typeStatus>
locality and taxonomic status;
<treatmentCitation id="0AB010EFFFB0A606FB66ADFF62F8F9D0" author="Grieshuber, J" page="32" pageId="8" pageNumber="459" year="2014">
<bibRefCitation id="EF804B0FFFB0A606FB66ADFF62F8F9D0" author="Grieshuber, J" pageId="8" pageNumber="459" refId="ref14730" refString="Grieshuber, J (2014) Guide to the butterflies of the Palearctic region. Pieridae Part II. Omnes Artes, Milano, 86 pp." type="book" year="2014">Grieshuber, 2014: 32</bibRefCitation>
</treatmentCitation>
, figs.
<emphasis id="B965EAECFFB0A606FEA3ADDE6366F9D0" bold="true" box="[288,381,1528,1552]" pageId="8" pageNumber="459">syn. nov.</emphasis>
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFB0A606FF14AE31671EF9EE" blockId="8.[151,1437,1191,1675]" box="[151,1285,1559,1583]" pageId="8" pageNumber="459">
<treatmentCitationGroup id="AB0111D0FFB0A606FF14AE31671EF9EE" box="[151,1285,1559,1583]" pageId="8" pageNumber="459">
<treatmentCitation id="0AB010EFFFB0A606FF14AE316072F9EF" author="Verhulst, J." box="[151,617,1559,1583]" page="181" pageId="8" pageNumber="459" year="1991">
<taxonomicName id="4C114D7DFFB0A606FF14AE316072F9EF" ID-CoL="X2GM" authority="sifanica: Verhulst, 1991 b: 181" authorityName="Verhulst" authorityPageNumber="181" authorityYear="1991" box="[151,617,1559,1583]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="subSpecies" species="sifanica" subSpecies="sifanica">
<emphasis id="B965EAECFFB0A606FF14AE31639DF9EF" box="[151,390,1559,1583]" italics="true" pageId="8" pageNumber="459">Colias sifanica sifanica</emphasis>
:
<bibRefCitation id="EF804B0FFFB0A606FE10AE316072F9EF" author="Verhulst, J." box="[403,617,1559,1583]" pageId="8" pageNumber="459" pagination="181 - 187" refId="ref16196" refString="Verhulst, J. (1991 b) Fiches specifiques des Colias F. (Colias data sheets). COLIAS SIFANICA. Lambillionea, 91 (3), 181 - 187. [in French]" type="journal article" year="1991">Verhulst, 1991b: 181</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs 12 for
<typeStatus id="54AA885CFFB0A606FD73AE316107F9EF" box="[752,796,1559,1583]" pageId="8" pageNumber="459">type</typeStatus>
specimens, pl. 1 &amp; 187-pl., figs. for
<specimenCount id="9D17FD77FFB0A606FB11AE3166A3F9EE" box="[1170,1208,1559,1582]" pageId="8" pageNumber="459" type="male">♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFB0A606FB5BAE31671BF9EE" box="[1240,1280,1559,1582]" pageId="8" pageNumber="459" type="female">♀♀</specimenCount>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BAE36FEFFB0A606FF14AE1362F6F94B" blockId="8.[151,1437,1191,1675]" pageId="8" pageNumber="459">
<treatmentCitationGroup id="AB0111D0FFB0A606FF14AE1362F6F94B" pageId="8" pageNumber="459">
<treatmentCitation id="0AB010EFFFB0A606FF14AE136113F98D" author="Chou, I. &amp; Zhang, Y. - L. &amp; Wang, Y. - L." box="[151,776,1589,1614]" page="45" pageId="8" pageNumber="459" year="2001">
<taxonomicName id="4C114D7DFFB0A606FF14AE136113F98D" authority="Chou, Zhang &amp; Wang, 2001: 45" authorityName="Chou, Zhang &amp; Wang" authorityPageNumber="45" authorityYear="2001" box="[151,776,1589,1614]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="subSpecies" species="sifanica" subSpecies="qinghaiensis">
<emphasis id="B965EAECFFB0A606FF14AE1363ACF98D" box="[151,439,1589,1613]" italics="true" pageId="8" pageNumber="459">Colias sifanica qinghaiensis</emphasis>
<bibRefCitation id="EF804B0FFFB0A606FE3DAE136113F98D" author="Chou, I. &amp; Zhang, Y. - L. &amp; Wang, Y. - L." box="[446,776,1589,1614]" pageId="8" pageNumber="459" pagination="38 - 46" refId="ref14443" refString="Chou, I., Zhang, Y. - L. &amp; Wang, Y. - L. (2001) New species, new subspecies and new record of butterflies (Lepidoptera) from China (III). Entomotaxonomia, 23, 38 - 46." type="journal article" year="2001">Chou, Zhang &amp; Wang, 2001: 45</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, 41- pl., figs. 910 for
<specimenCount id="9D17FD77FFB0A606FC70AE10661DF98D" box="[1011,1030,1590,1613]" pageId="8" pageNumber="459" type="female"></specimenCount>
<typeStatus id="54AA885CFFB0A606FB8EAE10667FF98E" box="[1037,1124,1590,1614]" pageId="8" pageNumber="459" type="paratype">paratype</typeStatus>
from Xiahe, figs. 1112 for
<specimenCount id="9D17FD77FFB0A606FA0AAE106787F98D" box="[1417,1436,1590,1613]" pageId="8" pageNumber="459" type="male"></specimenCount>
<typeStatus id="54AA885CFFB0A606FF44AE72633BF9AC" box="[199,288,1620,1644]" pageId="8" pageNumber="459" type="holotype">holotype</typeStatus>
(TL: Xining);
<treatmentCitation id="0AB010EFFFB0A606FE34AE7260D9F9AC" author="Grieshuber, J. &amp; Churkin, S. &amp; Worthy, R. &amp; Lvovsky, A." box="[439,706,1620,1644]" page="20" pageId="8" pageNumber="459" year="2004">
<bibRefCitation id="EF804B0FFFB0A606FE34AE7260D9F9AC" author="Grieshuber, J. &amp; Churkin, S. &amp; Worthy, R. &amp; Lvovsky, A." box="[439,706,1620,1644]" pageId="8" pageNumber="459" pagination="10 - 24" refId="ref15057" refString="Grieshuber, J., Churkin, S., Worthy, R. &amp; Lvovsky, A. (2004) The types of Colias felderi Grum-Grshimailo, 1891, Colias lada Grum-Grshimailo, 1891, and Colias sifanica Alpheraky, 1891, with additional notes on the types of Colias diva Grum- Grshimailo, 1891 (Lepidoptera: Pieridae). Helios, Moscow, 5, 10 - 24, pl. 1, figs. 1 - 7." type="journal article" year="2004">
Grieshuber
<emphasis id="B965EAECFFB0A606FDAEAE73607BF9AC" box="[557,608,1620,1644]" italics="true" pageId="8" pageNumber="459">et al.</emphasis>
2004: 20
</bibRefCitation>
</treatmentCitation>
, synonymy for
<taxonomicName id="4C114D7DFFB0A606FCDCAE7261D6F9AC" ID-CoL="X2GM" authorityName="Grum-Grshimailo" authorityYear="1891" box="[863,973,1620,1644]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB0A606FCDCAE7261D6F9AC" box="[863,973,1620,1644]" italics="true" pageId="8" pageNumber="459">C. sifanica</emphasis>
</taxonomicName>
;
<treatmentCitation id="0AB010EFFFB0A606FC59AE726738F9AC" author="Grieshuber, J. &amp; Lamas, G." box="[986,1315,1620,1644]" page="161" pageId="8" pageNumber="459" year="2007">
<bibRefCitation id="EF804B0FFFB0A606FC59AE726738F9AC" author="Grieshuber, J. &amp; Lamas, G." box="[986,1315,1620,1644]" pageId="8" pageNumber="459" pagination="131 - 171" refId="ref15140" refString="Grieshuber, J. &amp; Lamas, G. (2007) A Synonymic List of the Genus Colias Fabricius, 1807 (Lepidoptera: Pieridae). Mitteilungen der Munchner Entomologischen Gesellschaft, 97, 131 - 171." type="journal article" year="2007">Grieshuber &amp; Lamas, 2007: 161</bibRefCitation>
</treatmentCitation>
, synonymic list.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C30B6575FFB0A607FF14AE9F6620FB99" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459" type="materials_examined">
<paragraph id="8BAE36FEFFB0A607FF14AE9F635CFE81" blockId="8.[151,1437,1721,2035]" lastBlockId="9.[151,1437,151,1113]" lastPageId="9" lastPageNumber="460" pageId="8" pageNumber="459">
<emphasis id="B965EAECFFB0A606FF14AE9F6361F913" bold="true" box="[151,378,1721,1747]" pageId="8" pageNumber="459">Material examined</emphasis>
.
<collectingCountry id="F306766EFFB0A606FE0AAE9F63FDF913" box="[393,486,1721,1747]" name="China" pageId="8" pageNumber="459">CHINA</collectingCountry>
:
<collectingRegion id="49D5F81CFFB0A606FE72AE9F604BF913" box="[497,592,1721,1747]" country="China" name="Qinghai" pageId="8" pageNumber="459">Qinghai</collectingRegion>
:
<specimenCount id="9D17FD77FFB0A606FDD9AE9F60B3F913" box="[602,680,1721,1747]" pageId="8" pageNumber="459" type="male">87 ♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFB0A606FD4EAE9F6100F913" box="[717,795,1721,1747]" pageId="8" pageNumber="459" type="female">21 ♀♀</specimenCount>
(
<collectionCode id="ED00AE3BFFB0A606FCA9AE9F617EF913" box="[810,869,1721,1747]" pageId="8" pageNumber="459">CSK</collectionCode>
), Xiaqunsi, Pingan,
<quantity id="4CE99B1BFFB0A606FBD0AE9F66BFF913" box="[1107,1188,1721,1747]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="459" unit="m" value="2800.0">2800m</quantity>
,
<date id="FFAF103EFFB0A606FB2CAE9F674BF914" box="[1199,1360,1721,1748]" pageId="8" pageNumber="459" value="2005-06" valueMax="2005-07-31" valueMin="2005-06-00">VI.VII. 2005</date>
2019, K. Song leg.;
<specimenCount id="9D17FD77FFB0A606FEBFAEFB6397F937" box="[316,396,1757,1783]" pageId="8" pageNumber="459" type="male">24 ♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFB0A606FE35AEFB63E2F937" box="[438,505,1757,1784]" pageId="8" pageNumber="459" type="female">5 ♀♀</specimenCount>
(
<collectionCode id="ED00AE3BFFB0A606FD88AEFB605DF937" box="[523,582,1757,1783]" pageId="8" pageNumber="459">CSK</collectionCode>
), Chapu, Hualong,
<quantity id="4CE99B1BFFB0A606FCAAAEFB6161F937" box="[809,890,1757,1783]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="459" unit="m" value="2700.0">2700m</quantity>
,
<date id="FFAF103EFFB0A606FC0BAEFB6635F938" box="[904,1070,1757,1784]" pageId="8" pageNumber="459" value="2015-06" valueMax="2015-07-31" valueMin="2015-06-00">VI.VII. 2015</date>
, K. Song leg.;
<specimenCount id="9D17FD77FFB0A606FB63AEFB672BF937" box="[1248,1328,1757,1783]" pageId="8" pageNumber="459" type="male">11 ♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFB0A606FAD9AEFB6787F937" box="[1370,1436,1757,1783]" pageId="8" pageNumber="459" type="female">2 ♀♀</specimenCount>
(
<collectionCode id="ED00AE3BFFB0A606FF1CAF2762C1F8DB" box="[159,218,1793,1819]" pageId="8" pageNumber="459">CSK</collectionCode>
), Babao, Qilian,
<quantity id="4CE99B1BFFB0A606FE16AF2763FDF8DB" box="[405,486,1793,1819]" metricMagnitude="3" metricUnit="m" metricValue="2.9" pageId="8" pageNumber="459" unit="m" value="2900.0">2900m</quantity>
,
<date id="FFAF103EFFB0A606FE73AF2760B7F8DB" box="[496,684,1793,1819]" pageId="8" pageNumber="459" value="2008-07-24" valueMax="2008-07-26" valueMin="2008-07-24">2426.VII. 2008</date>
, K. Song leg.;
<collectingRegion id="49D5F81CFFB0A606FCCCAF276180F8DB" box="[847,923,1793,1819]" country="China" name="Gansu" pageId="8" pageNumber="459">Gansu</collectingRegion>
:
<specimenCount id="9D17FD77FFB0A606FC27AF2761EBF8DB" box="[932,1008,1793,1819]" pageId="8" pageNumber="459" type="male">21 ♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFB0A606FB91AF27664BF8DB" box="[1042,1104,1793,1819]" pageId="8" pageNumber="459" type="female">6 ♀♀</specimenCount>
(
<collectionCode id="ED00AE3BFFB0A606FBDDAF276685F8DB" box="[1118,1182,1793,1819]" pageId="8" pageNumber="459">CHH</collectionCode>
), Xiahe,
<quantity id="4CE99B1BFFB0A606FA82AF276783F8DB" box="[1281,1432,1793,1819]" metricMagnitude="3" metricUnit="m" metricValue="3.2" metricValueMax="3.4" metricValueMin="3.0" pageId="8" pageNumber="459" unit="m" value="3200.0" valueMax="3400.0" valueMin="3000.0">30003400m</quantity>
,
<date id="FFAF103EFFB0A606FF14AF03639AF880" box="[151,385,1829,1856]" pageId="8" pageNumber="459" value="2015-07-20" valueMax="2015-07-26" valueMin="2015-07-20">20.VII.26.VII.2015</date>
, H. Huang leg.;
<specimenCount id="9D17FD77FFB0A606FDBAAF03609EF8FF" box="[569,645,1829,1855]" pageId="8" pageNumber="459" type="male">11 ♂♂</specimenCount>
&amp;
<specimenCount id="9D17FD77FFB0A606FD2BAF0360C9F8FF" box="[680,722,1829,1855]" pageId="8" pageNumber="459" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED00AE3BFFB0A606FD62AF03613AF8FF" box="[737,801,1829,1855]" pageId="8" pageNumber="459">CHH</collectionCode>
), Xiahe,
<quantity id="4CE99B1BFFB0A606FC05AF036601F880" box="[902,1050,1829,1856]" metricMagnitude="3" metricUnit="m" metricValue="3.125" metricValueMax="3.3" metricValueMin="2.95" pageId="8" pageNumber="459" unit="m" value="3125.0" valueMax="3300.0" valueMin="2950.0">29503300m</quantity>
<date id="FFAF103EFFB0A606FBA2AF0366EFF8FF" box="[1057,1268,1829,1855]" pageId="8" pageNumber="459" value="2016-06-30" valueMax="2016-07-02" valueMin="2016-06-30">30.VI.2.VII.2016</date>
, H. Huang leg.
<emphasis id="B965EAECFFB0A606FF14AF6F632DF8A3" bold="true" box="[151,310,1864,1891]" pageId="8" pageNumber="459">
Form
<taxonomicName id="4C114D7DFFB0A606FF62AF6E6334F8A3" authorityName="Verhulst" authorityYear="1990" box="[225,303,1864,1891]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB0A606FF62AF6E6334F8A3" bold="true" box="[225,303,1864,1891]" italics="true" pageId="8" pageNumber="459">aegidii</emphasis>
</taxonomicName>
.
</emphasis>
<bibRefCitation id="EF804B0FFFB0A606FEBDAF6F63FCF8A4" author="Hoshiai, A." box="[318,487,1865,1892]" pageId="8" pageNumber="459" pagination="8 - 10" refId="ref15351" refString="Hoshiai, A. (1996) Notes of Colias from China and surrounding area (1). In: Koiwaya, S. (Ed.), Studies of Chinese Butterflies. Vol. 3. S. Koiwaya, Tokyo, pp. 211 - 217, pls. 8 - 10. [in Japanese]" type="book chapter" year="1996">Hoshiai (1996)</bibRefCitation>
figured a few specimens of
<taxonomicName id="4C114D7DFFB0A606FCA4AF6F618DF8A3" authorityName="Verhulst" authorityYear="1990" box="[807,918,1865,1891]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB0A606FCA4AF6F618DF8A3" box="[807,918,1865,1891]" italics="true" pageId="8" pageNumber="459">C. aegidii</emphasis>
</taxonomicName>
together with those of
<taxonomicName id="4C114D7DFFB0A606FB18AF6F670EF8A3" authorityName="Grum-Grshimailo" authorityYear="1891" box="[1179,1301,1865,1891]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB0A606FB18AF6F670EF8A3" box="[1179,1301,1865,1891]" italics="true" pageId="8" pageNumber="459">C. sifanica</emphasis>
</taxonomicName>
, both under the name,
<taxonomicName id="4C114D7DFFB0A606FE8FAF4B639CF847" authorityName="Grum-Grshimailo" authorityYear="1891" box="[268,391,1901,1927]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB0A606FE8FAF4B639CF847" box="[268,391,1901,1927]" italics="true" pageId="8" pageNumber="459">C. sifanica</emphasis>
</taxonomicName>
, which show continuous variations in wing-pattern. He discussed the possibility of the natural hybrid between
<taxonomicName id="4C114D7DFFB0A606FEF1AFB763F6F86B" authorityName="Grum-Grshimailo" authorityYear="1891" box="[370,493,1937,1963]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB0A606FEF1AFB763F6F86B" box="[370,493,1937,1963]" italics="true" pageId="8" pageNumber="459">C. sifanica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFB0A606FDA6AFB76173F86B" authority=", Oberthur, 1894" authorityName=", Oberthur" authorityYear="1894" box="[549,872,1937,1963]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B965EAECFFB0A606FDA6AFB760B0F86B" box="[549,683,1937,1963]" italics="true" pageId="8" pageNumber="459">C. nebulosa</emphasis>
,
<bibRefCitation id="EF804B0FFFB0A606FD39AFB76173F86B" author="Oberthur, C." box="[698,872,1937,1963]" pageId="8" pageNumber="459" pagination="1 - 41" refId="ref15957" refString="Oberthur, C. (1894) Lepidopteres d'Europe, d'Algerie, d'Asie et d'Oceanie. Etudes d'Entomologie, 19, 1 - 41, 8 pls. [in French]" type="journal article" year="1894">Oberthür, 1894</bibRefCitation>
</taxonomicName>
, and inclined to regard specimens of
<taxonomicName id="4C114D7DFFB0A606FA8DAFB76766F86B" authorityName="Verhulst" authorityYear="1990" box="[1294,1405,1937,1963]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB0A606FA8DAFB76766F86B" box="[1294,1405,1937,1963]" italics="true" pageId="8" pageNumber="459">C. aegidii</emphasis>
</taxonomicName>
as individual variations of
<taxonomicName id="4C114D7DFFB0A606FE20AF936007F80F" authorityName="Grum-Grshimailo" authorityYear="1891" box="[419,540,1973,1999]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB0A606FE20AF936007F80F" box="[419,540,1973,1999]" italics="true" pageId="8" pageNumber="459">C. sifanica</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF804B0FFFB0A606FDAAAF936136F80F" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[553,813,1973,1999]" pageId="8" pageNumber="459" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">Grieshuber et al (2012)</bibRefCitation>
stated “Although
<taxonomicName id="4C114D7DFFB0A606FC7AAF93667CF80F" authorityName="Verhulst" authorityYear="1990" box="[1017,1127,1973,1999]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB0A606FC7AAF93667CF80F" box="[1017,1127,1973,1999]" italics="true" pageId="8" pageNumber="459">C. aegidii</emphasis>
</taxonomicName>
does fly with
<taxonomicName id="4C114D7DFFB0A606FA85AF936764F80F" authorityName="Grum-Grshimailo" authorityYear="1891" box="[1286,1407,1973,1999]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB0A606FA85AF936764F80F" box="[1286,1407,1973,1999]" italics="true" pageId="8" pageNumber="459">C. sifanica</emphasis>
</taxonomicName>
, it is only found where
<taxonomicName id="4C114D7DFFB0A606FEF8AFFF63EFF833" authorityName="Grum-Grshimailo" authorityYear="1891" box="[379,500,2009,2035]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB0A606FEF8AFFF63EFF833" box="[379,500,2009,2035]" italics="true" pageId="8" pageNumber="459">C. sifanica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFB0A606FDA9AFFF6101F833" authority="nebulosa" authorityName="nebulosa Oberthur" authorityYear="1894" box="[554,794,2009,2035]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="459" phylum="Arthropoda" rank="subSpecies" species="nebulosa" subSpecies="nebulosa">
<emphasis id="B965EAECFFB0A606FDA9AFFF6101F833" box="[554,794,2009,2035]" italics="true" pageId="8" pageNumber="459">C. nebulosa nebulosa</emphasis>
</taxonomicName>
fly in close proximity to each other” and “The possibility that
<taxonomicName id="4C114D7DFFB1A607FF44A8B1630EFF71" authorityName="Verhulst" authorityYear="1990" box="[199,277,151,177]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FF44A8B1630EFF71" box="[199,277,151,177]" italics="true" pageId="9" pageNumber="460">aegidii</emphasis>
</taxonomicName>
is a natural hybrid between
<taxonomicName id="4C114D7DFFB1A607FDCEA8B160A9FF71" authorityName=", Oberthur" authorityYear="1894" box="[589,690,151,177]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B965EAECFFB1A607FDCEA8B160A9FF71" box="[589,690,151,177]" italics="true" pageId="9" pageNumber="460">nebulosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFB1A607FD64A8B1615BFF71" authorityName="Grum-Grshimailo" authorityYear="1891" box="[743,832,151,177]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB1A607FD64A8B1615BFF71" box="[743,832,151,177]" italics="true" pageId="9" pageNumber="460">sifanica</emphasis>
</taxonomicName>
cannot be excluded, but there is no direct evidence for this”. However, the senior author collected
<specimenCount id="9D17FD77FFB1A607FDFEA89D60A7FF15" box="[637,700,187,213]" pageId="9" pageNumber="460" type="male">3 ♂♂</specimenCount>
of
<taxonomicName id="4C114D7DFFB1A607FD62A89D614BFF15" authorityName="Verhulst" authorityYear="1990" box="[737,848,187,213]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FD62A89D614BFF15" box="[737,848,187,213]" italics="true" pageId="9" pageNumber="460">C. aegidii</emphasis>
</taxonomicName>
from a dry valley on north of Xiahe at
<quantity id="4CE99B1BFFB1A607FA8BA89C6787FF15" box="[1288,1436,186,213]" metricMagnitude="3" metricUnit="m" metricValue="3.125" metricValueMax="3.3" metricValueMin="2.95" pageId="9" pageNumber="460" unit="m" value="3125.0" valueMax="3300.0" valueMin="2950.0">29503300m</quantity>
together with a few
<taxonomicName id="4C114D7DFFB1A607FEFAA8F963E8FF39" authorityName="Grum-Grshimailo" authorityYear="1891" box="[377,499,223,249]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB1A607FEFAA8F963E8FF39" box="[377,499,223,249]" italics="true" pageId="9" pageNumber="460">C. sifanica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFB1A607FDA9A8F960B6FF38" authorityName="Oberthur" authorityYear="1886" box="[554,685,223,249]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="montium">
<emphasis id="B965EAECFFB1A607FDA9A8F960B6FF38" box="[554,685,223,249]" italics="true" pageId="9" pageNumber="460">C. montium</emphasis>
</taxonomicName>
; this locality is far away from the biotopes of
<taxonomicName id="4C114D7DFFB1A607FB34A8F96726FF39" authorityName=", Oberthur" authorityYear="1894" box="[1207,1341,223,249]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B965EAECFFB1A607FB34A8F96726FF39" box="[1207,1341,223,249]" italics="true" pageId="9" pageNumber="460">C. nebulosa</emphasis>
</taxonomicName>
. By this evidence and a further record from
<collectingRegion id="49D5F81CFFB1A607FDA4A925609AFEDD" box="[551,641,259,285]" country="China" name="Qinghai" pageId="9" pageNumber="460">Qinghai</collectingRegion>
(discussed below),
<taxonomicName id="4C114D7DFFB1A607FCE2A92561CAFEDD" authorityName="Verhulst" authorityYear="1990" box="[865,977,259,285]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FCE2A92561CAFEDD" box="[865,977,259,285]" italics="true" pageId="9" pageNumber="460">C. aegidii</emphasis>
</taxonomicName>
cannot be a hybrid between
<taxonomicName id="4C114D7DFFB1A607FA95A9256787FEDD" authorityName=", Oberthur" authorityYear="1894" box="[1302,1436,259,285]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B965EAECFFB1A607FA95A9256787FEDD" box="[1302,1436,259,285]" italics="true" pageId="9" pageNumber="460">C. nebulosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFB1A607FF44A901635BFE81" authorityName="Grum-Grshimailo" authorityYear="1891" box="[199,320,295,321]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB1A607FF44A901635BFE81" box="[199,320,295,321]" italics="true" pageId="9" pageNumber="460">C. sifanica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAE36FEFFB1A607FF44A96D66D0FE11" blockId="9.[151,1437,151,1113]" pageId="9" pageNumber="460">
The known specimens in literature clearly constitute a continuous series of variations from
<taxonomicName id="4C114D7DFFB1A607FB5CA96D6740FEA5" authorityName="Grum-Grshimailo" authorityYear="1891" box="[1247,1371,331,357]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB1A607FB5CA96D6740FEA5" box="[1247,1371,331,357]" italics="true" pageId="9" pageNumber="460">C. sifanica</emphasis>
</taxonomicName>
to
<taxonomicName id="4C114D7DFFB1A607FA00A96D62FEFE49" authorityName="Verhulst" authorityYear="1990" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FA00A96D62FEFE49" italics="true" pageId="9" pageNumber="460">C. aegidii</emphasis>
</taxonomicName>
without an interruption in all of the wing characters. And the following specimens are found around the boundary: 1)
<specimenCount id="9D17FD77FFB1A607FEAEA9B56375FE6D" box="[301,366,403,429]" pageId="9" pageNumber="460" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D17FD77FFB1A607FEF9A9B563A2FE6D" box="[378,441,403,429]" pageId="9" pageNumber="460" type="female">2 ♀♀</specimenCount>
(
<bibRefCitation id="EF804B0FFFB1A607FE44A9B56098FE6D" author="Hoshiai, A." box="[455,643,403,429]" pageId="9" pageNumber="460" pagination="8 - 10" refId="ref15351" refString="Hoshiai, A. (1996) Notes of Colias from China and surrounding area (1). In: Koiwaya, S. (Ed.), Studies of Chinese Butterflies. Vol. 3. S. Koiwaya, Tokyo, pp. 211 - 217, pls. 8 - 10. [in Japanese]" type="book chapter" year="1996">Hoshiai, 1996: 9</bibRefCitation>
- figs. 4041, 4850); 2)
<specimenCount id="9D17FD77FFB1A607FC1AA9B561C3FE6D" box="[921,984,403,429]" pageId="9" pageNumber="460" type="male">2 ♂♂</specimenCount>
(Verhulst, 1990: 41-pl., figs. 8 &amp; 11); 3)
<specimenCount id="9D17FD77FFB1A607FF14A99162CDFE11" box="[151,214,439,465]" pageId="9" pageNumber="460" type="female">3 ♀♀</specimenCount>
(
<bibRefCitation id="EF804B0FFFB1A607FF66A9916395FE11" author="Verhulst, J." box="[229,398,439,465]" pageId="9" pageNumber="460" refId="ref16453" refString="Verhulst, J. (2001) Les Colias du Globe. Monograph of the genus Colias. Plates. Goecke &amp; Evers, Kelterns, 45 pp., 183 pls. [in French]" type="book" year="2001">Verhulst, 2001</bibRefCitation>
: pl. 40, figs. 4, 8 &amp; 15); 4)
<specimenCount id="9D17FD77FFB1A607FD3CA99160E5FE11" box="[703,766,439,465]" pageId="9" pageNumber="460" type="male">3 ♂♂</specimenCount>
(this paper: fig. 14, Specimens I12I14).
</paragraph>
<paragraph id="8BAE36FEFFB1A607FF44A9FD61D3FD0D" blockId="9.[151,1437,151,1113]" pageId="9" pageNumber="460">
Further on, the junior author collected a male of
<taxonomicName id="4C114D7DFFB1A607FD62A9FD6154FE35" authorityName="Verhulst" authorityYear="1990" box="[737,847,475,501]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FD62A9FD6154FE35" box="[737,847,475,501]" italics="true" pageId="9" pageNumber="460">C. aegidii</emphasis>
</taxonomicName>
(
<figureCitation id="132A2A7BFFB1A607FCDEA9FD61B4FE35" box="[861,943,475,501]" captionStart="FIGURE 14" captionStartId="17.[151,250,1943,1967]" captionTargetBox="[207,1380,181,1919]" captionTargetId="figure-19@17.[207,1380,181,1919]" captionTargetPageId="17" captionText="FIGURE 14. Habitus of Colias sifanica (with form aegidii) ♂♂ &amp; ♀ at same scale (Upper half- upper side; lower half- under- side). Scale bar = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.6333783" httpUri="https://zenodo.org/record/6333783/files/figure.png" pageId="9" pageNumber="460">Fig. 14</figureCitation>
, Specimen I4) from Xiaqunsi, Pingan,
<collectingRegion id="49D5F81CFFB1A607FAEAA9FD62DCFDD9" country="China" name="Qinghai" pageId="9" pageNumber="460">Qinghai</collectingRegion>
(a locality with no
<taxonomicName id="4C114D7DFFB1A607FE24A9D96036FDD9" authorityName=", Oberthur" authorityYear="1894" box="[423,557,511,537]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B965EAECFFB1A607FE24A9D96036FDD9" box="[423,557,511,537]" italics="true" pageId="9" pageNumber="460">C. nebulosa</emphasis>
</taxonomicName>
found) among the large number of specimens of
<taxonomicName id="4C114D7DFFB1A607FBDDA9D966C2FDD9" authorityName="Grum-Grshimailo" authorityYear="1891" box="[1118,1241,511,537]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB1A607FBDDA9D966C2FDD9" box="[1118,1241,511,537]" italics="true" pageId="9" pageNumber="460">C. sifanica</emphasis>
</taxonomicName>
. He continued to collect there for 15 years and collected over
<specimenCount id="9D17FD77FFB1A607FD03AA05613EFDFD" box="[640,805,547,573]" pageId="9" pageNumber="460" type="generic">100 specimens</specimenCount>
of
<taxonomicName id="4C114D7DFFB1A607FCCBAA0561DBFDFD" authorityName="Grum-Grshimailo" authorityYear="1891" box="[840,960,547,573]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB1A607FCCBAA0561DBFDFD" box="[840,960,547,573]" italics="true" pageId="9" pageNumber="460">C. sifanica</emphasis>
</taxonomicName>
but only encountered
<taxonomicName id="4C114D7DFFB1A607FB35AA05673FFDFD" authorityName="Verhulst" authorityYear="1990" box="[1206,1316,547,573]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FB35AA05673FFDFD" box="[1206,1316,547,573]" italics="true" pageId="9" pageNumber="460">C. aegidii</emphasis>
</taxonomicName>
once. This is the first record of
<taxonomicName id="4C114D7DFFB1A607FEFFAA6163F0FDA1" authorityName="Verhulst" authorityYear="1990" box="[380,491,583,609]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FEFFAA6163F0FDA1" box="[380,491,583,609]" italics="true" pageId="9" pageNumber="460">C. aegidii</emphasis>
</taxonomicName>
either from
<collectingRegion id="49D5F81CFFB1A607FDF5AA616122FDA1" box="[630,825,583,609]" country="China" name="Qinghai" pageId="9" pageNumber="460">Qinghai province</collectingRegion>
or out of Xiahe. Verhulst (1990) erroneously recorded a male from Caka,
<collectingRegion id="49D5F81CFFB1A607FEF1AA4D63D4FD45" box="[370,463,619,645]" country="China" name="Qinghai" pageId="9" pageNumber="460">Qinghai</collectingRegion>
, which was proved to be collected actually from Xiahe (
<bibRefCitation id="EF804B0FFFB1A607FBD1AA4D6745FD45" author="Grieshuber, J. &amp; Worthy, R. &amp; Lamas, G." box="[1106,1374,619,645]" pageId="9" pageNumber="460" refId="ref15180" refString="Grieshuber, J., Worthy, R. &amp; Lamas, G. (2012) The genus Colias Fabricius, 1807. Jan Haugum's annotated catalogue of the Old World Colias. Tshikolovets Publications, Pardubice, 438 pp." type="book" year="2012">
Grieshuber
<emphasis id="B965EAECFFB1A607FB5BAA4D6703FD45" box="[1240,1304,619,645]" italics="true" pageId="9" pageNumber="460">et al.,</emphasis>
2012
</bibRefCitation>
). The co-existence of both
<taxonomicName id="4C114D7DFFB1A607FE01AAA963E7FD69" authorityName="Grum-Grshimailo" authorityYear="1891" box="[386,508,655,681]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB1A607FE01AAA963E7FD69" box="[386,508,655,681]" italics="true" pageId="9" pageNumber="460">C. sifanica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFB1A607FDB1AAA960BAFD69" authorityName="Verhulst" authorityYear="1990" box="[562,673,655,681]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FDB1AAA960BAFD69" box="[562,673,655,681]" italics="true" pageId="9" pageNumber="460">C. aegidii</emphasis>
</taxonomicName>
at Xiaqunsi, and the extreme rarity of
<taxonomicName id="4C114D7DFFB1A607FBD7AAA966D9FD69" authorityName="Verhulst" authorityYear="1990" box="[1108,1218,655,681]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FBD7AAA966D9FD69" box="[1108,1218,655,681]" italics="true" pageId="9" pageNumber="460">C. aegidii</emphasis>
</taxonomicName>
for years draw to a conclusion that
<taxonomicName id="4C114D7DFFB1A607FECBAA9563ACFD0D" authorityName="Verhulst" authorityYear="1990" box="[328,439,691,717]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FECBAA9563ACFD0D" box="[328,439,691,717]" italics="true" pageId="9" pageNumber="460">C. aegidii</emphasis>
</taxonomicName>
is merely a rare ecological form of
<taxonomicName id="4C114D7DFFB1A607FCCBAA9561D9FD0D" authorityName="Grum-Grshimailo" authorityYear="1891" box="[840,962,691,717]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB1A607FCCBAA9561D9FD0D" box="[840,962,691,717]" italics="true" pageId="9" pageNumber="460">C. sifanica</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAE36FEFFB1A607FF44AAF16706FC2D" blockId="9.[151,1437,151,1113]" pageId="9" pageNumber="460">
During a cooperation with Dr. N. Grishin, the senior author asked him to help in making a DNA analysis for the specimens of
<taxonomicName id="4C114D7DFFB1A607FEB0AADD63B6FCD5" authorityName="Grum-Grshimailo" authorityYear="1891" box="[307,429,763,789]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB1A607FEB0AADD63B6FCD5" box="[307,429,763,789]" italics="true" pageId="9" pageNumber="460">C. sifanica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFB1A607FE66AADD604FFCD5" authorityName="Verhulst" authorityYear="1990" box="[485,596,763,789]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FE66AADD604FFCD5" box="[485,596,763,789]" italics="true" pageId="9" pageNumber="460">C. aegidii</emphasis>
</taxonomicName>
collected from Xiahe. The analysis (Grishin, unpublished data) was based upon
<specimenCount id="9D17FD77FFB1A607FF55AB396344FCF9" box="[214,351,799,825]" pageId="9" pageNumber="460" type="generic">8 specimens</specimenCount>
of
<taxonomicName id="4C114D7DFFB1A607FE00AB3963E7FCF9" authorityName="Grum-Grshimailo" authorityYear="1891" box="[387,508,799,825]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB1A607FE00AB3963E7FCF9" box="[387,508,799,825]" italics="true" pageId="9" pageNumber="460">C. sifanica</emphasis>
</taxonomicName>
from Xiahe and Kukunor,
<specimenCount id="9D17FD77FFB1A607FCA4AB3961ABFCF9" box="[807,944,799,825]" pageId="9" pageNumber="460" type="generic">4 specimens</specimenCount>
of
<taxonomicName id="4C114D7DFFB1A607FC57AB396659FCF9" authorityName="Verhulst" authorityYear="1990" box="[980,1090,799,825]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FC57AB396659FCF9" box="[980,1090,799,825]" italics="true" pageId="9" pageNumber="460">C. aegidii</emphasis>
</taxonomicName>
(including the
<specimenCount id="9D17FD77FFB1A607FB69AB396732FCF9" box="[1258,1321,799,825]" pageId="9" pageNumber="460" type="male">3 ♂♂</specimenCount>
figured in this work) and
<specimenCount id="9D17FD77FFB1A607FEC3AB6563D1FC9D" box="[320,458,835,861]" pageId="9" pageNumber="460" type="generic">3 specimens</specimenCount>
of
<taxonomicName id="4C114D7DFFB1A607FE6CAB65606EFC9D" authorityName=", Oberthur" authorityYear="1894" box="[495,629,835,861]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B965EAECFFB1A607FE6CAB65606EFC9D" box="[495,629,835,861]" italics="true" pageId="9" pageNumber="460">C. nebulosa</emphasis>
</taxonomicName>
from Xiahe,
<collectingRegion id="49D5F81CFFB1A607FC89AB65617FFC9D" box="[778,868,835,861]" country="China" name="Sichuan" pageId="9" pageNumber="460">Sichuan</collectingRegion>
and
<collectingRegion id="49D5F81CFFB1A607FC19AB6561CCFC9D" box="[922,983,835,861]" country="China" name="Xizang" pageId="9" pageNumber="460">Tibet</collectingRegion>
, with
<taxonomicName id="4C114D7DFFB1A607FB98AB656675FC9D" authorityName="Grum-Grshimailo" authorityYear="1891" box="[1051,1134,835,861]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="lada">
<emphasis id="B965EAECFFB1A607FB98AB656675FC9D" box="[1051,1134,835,861]" italics="true" pageId="9" pageNumber="460">C. lada</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFB1A607FB27AB656787FC9D" authority="Fawcett, 1904" authorityName="Fawcett" authorityYear="1904" box="[1188,1436,835,861]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="nina">
<emphasis id="B965EAECFFB1A607FB27AB6566ECFC9C" box="[1188,1271,835,861]" italics="true" pageId="9" pageNumber="460">C. nina</emphasis>
Fawcett, 1904
</taxonomicName>
as out-group taxa. Three trees were got: the first was from the Z chromosome (male sex chromosome) genes, the second was on the genes from the rest of the nuclear genome and the third was from mitochondrial genome. There is no separation between
<taxonomicName id="4C114D7DFFB1A607FE31AB896030FC09" authorityName="Grum-Grshimailo" authorityYear="1891" box="[434,555,943,969]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB1A607FE31AB896030FC09" box="[434,555,943,969]" italics="true" pageId="9" pageNumber="460">C. sifanica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFB1A607FDE2AB8960CBFC09" authorityName="Verhulst" authorityYear="1990" box="[609,720,943,969]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FDE2AB8960CBFC09" box="[609,720,943,969]" italics="true" pageId="9" pageNumber="460">C. aegidii</emphasis>
</taxonomicName>
at all whilst there is clear separation between
<taxonomicName id="4C114D7DFFB1A607FB50AB896756FC09" authorityName="Grum-Grshimailo" authorityYear="1891" box="[1235,1357,943,969]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="sifanica">
<emphasis id="B965EAECFFB1A607FB50AB896756FC09" box="[1235,1357,943,969]" italics="true" pageId="9" pageNumber="460">C. sifanica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C114D7DFFB1A607FA00AB8962E7FC2D" authorityName=", Oberthur" authorityYear="1894" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="nebulosa">
<emphasis id="B965EAECFFB1A607FA00AB8962E7FC2D" italics="true" pageId="9" pageNumber="460">C. nebulosa</emphasis>
</taxonomicName>
. The result of the DNA analysis will be published separately by Dr. Grishin in the near future.
</paragraph>
<paragraph id="8BAE36FEFFB1A607FF44ABD0609FFBD1" blockId="9.[151,1437,151,1113]" box="[199,644,1014,1041]" pageId="9" pageNumber="460">
<emphasis id="B965EAECFFB1A607FF44ABD0637FFBD0" bold="true" box="[199,356,1014,1040]" pageId="9" pageNumber="460">Flight period</emphasis>
. Late June to end of July.
</paragraph>
<paragraph id="8BAE36FEFFB1A607FF44AC3C6145FBF5" blockId="9.[151,1437,151,1113]" box="[199,862,1050,1077]" pageId="9" pageNumber="460">
<emphasis id="B965EAECFFB1A607FF44AC3C6332FBF4" bold="true" box="[199,297,1050,1076]" pageId="9" pageNumber="460">Altitude</emphasis>
.
<materialsCitation id="3B793CA3FFB1A607FEB4AC3D6056FBF5" ID-GBIF-Occurrence="3698883351" box="[311,589,1051,1077]" elevation="2800" location="Qinghai" pageId="9" pageNumber="460" specimenCount="1" stateProvince="Qinghai">
<quantity id="4CE99B1BFFB1A607FEB4AC3D63D0FBF5" box="[311,459,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="9" pageNumber="460" unit="m" value="2800.0" valueMax="2900.0" valueMin="2700.0">
<elevation id="003CD1CDFFB1A607FEB4AC3D63D0FBF5" box="[311,459,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="2.8" metricValueMax="2.9" metricValueMin="2.7" pageId="9" pageNumber="460" unit="m" value="2800.0" valueMax="2900.0" valueMin="2700.0">27002900m</elevation>
</quantity>
in
<collectingRegion id="49D5F81CFFB1A607FE6CAC3D6056FBF5" box="[495,589,1051,1077]" country="China" name="Qinghai" pageId="9" pageNumber="460">Qinghai</collectingRegion>
</materialsCitation>
;
<materialsCitation id="3B793CA3FFB1A607FDDBAC3D6141FBF5" ID-GBIF-Occurrence="3698883344" box="[600,858,1051,1077]" elevation="3150" location="Gansu" pageId="9" pageNumber="460" specimenCount="1" stateProvince="Gansu">
<quantity id="4CE99B1BFFB1A607FDDBAC3D60F7FBF5" box="[600,748,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="3.15" metricValueMax="3.4" metricValueMin="2.9" pageId="9" pageNumber="460" unit="m" value="3150.0" valueMax="3400.0" valueMin="2900.0">
<elevation id="003CD1CDFFB1A607FDDBAC3D60F7FBF5" box="[600,748,1051,1077]" metricMagnitude="3" metricUnit="m" metricValue="3.15" metricValueMax="3.4" metricValueMin="2.9" pageId="9" pageNumber="460" unit="m" value="3150.0" valueMax="3400.0" valueMin="2900.0">29003400m</elevation>
</quantity>
in
<collectingRegion id="49D5F81CFFB1A607FC8CAC3D6141FBF5" box="[783,858,1051,1077]" country="China" name="Gansu" pageId="9" pageNumber="460">Gansu</collectingRegion>
</materialsCitation>
.
</paragraph>
<paragraph id="8BAE36FEFFB1A607FF44AC186620FB99" blockId="9.[151,1437,151,1113]" box="[199,1083,1086,1113]" pageId="9" pageNumber="460">
<emphasis id="B965EAECFFB1A607FF44AC1863A0FB99" bold="true" box="[199,443,1086,1113]" pageId="9" pageNumber="460">Distributional notes.</emphasis>
Form
<taxonomicName id="4C114D7DFFB1A607FD85AC19604FFB99" authorityName="Verhulst" authorityYear="1990" box="[518,596,1087,1113]" class="Insecta" family="Pieridae" genus="Colias" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="460" phylum="Arthropoda" rank="species" species="aegidii">
<emphasis id="B965EAECFFB1A607FD85AC19604FFB99" box="[518,596,1087,1113]" italics="true" pageId="9" pageNumber="460">aegidii</emphasis>
</taxonomicName>
is recorded from
<collectingRegion id="49D5F81CFFB1A607FC98AC19616EFB99" box="[795,885,1087,1113]" country="China" name="Qinghai" pageId="9" pageNumber="460">Qinghai</collectingRegion>
for the first time.
</paragraph>
</subSubSection>
</treatment>
</document>