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<document id="48E2054D02B837F829A556C6E23B55BA" ID-CLB-Dataset="20049" ID-DOI="10.11646/zootaxa.5120.4.6" ID-GBIF-Dataset="a0421c84-a047-4499-b92f-52cab23d6e79" ID-ISSN="1175-5326" ID-Zenodo-Dep="6392907" ID-ZooBank="58CAB25D-CA92-44EE-BF1D-67DB91EB9596" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1648538015601" checkinUser="plazi" docAuthor="Martins, Luciana &amp; Tavares, Marcos" docDate="2022" docId="03B887C14B028139BEE16C55FD1291E6" docLanguage="en" docName="zootaxa.5120.4.6.pdf" docOrigin="Zootaxa 5120 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cladolella virgo Heding &amp; Panning 1954" docType="treatment" docVersion="6" lastPageNumber="568" masterDocId="FF81FFB94B0B8130BE766D38FFB09611" masterDocTitle="Additions to the morphology of the Phyllophoridae and the Sclerodactylidae. I. Type species of Phyllophorella, Selenkiella, Cladolella and Clarkiella, with the description a new species of Thorsonia (Holothuroidea: Dendrochirotida)" masterLastPageNumber="572" masterPageNumber="559" pageNumber="568" updateTime="1699318708264" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="500CFED1D9DEB1FDC36252675441A515">Additions to the morphology of the Phyllophoridae and the Sclerodactylidae. I. Type species of Phyllophorella, Selenkiella, Cladolella and Clarkiella, with the description a new species of Thorsonia (Holothuroidea: Dendrochirotida)</mods:title>
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<mods:namePart id="82A9090F55CBC00137DB6F91EA906D82">Martins, Luciana</mods:namePart>
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<mods:affiliation id="2DB9B261D5159D29E174CB9E4C8351C3">Museu de Zoologia, Universidade de São Paulo, São Paulo-SP, 04263 - 000, Brazil. &amp; martinsrluciana @ gmail. com, mdst @ usp. br; https: // orcid. org / 0000 - 0002 - 8107 - 3265</mods:affiliation>
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<mods:namePart id="E9A0C360CD0152BBA749046BD337086D">Tavares, Marcos</mods:namePart>
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<mods:affiliation id="19F00FDBC89C86CEE9CFCF4A492B748E">Museu de Zoologia, Universidade de São Paulo, São Paulo-SP, 04263 - 000, Brazil. &amp; https: // orcid. org / 0000 - 0002 - 7186 - 5787</mods:affiliation>
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<treatment id="03B887C14B028139BEE16C55FD1291E6" ID-DOI="http://doi.org/10.5281/zenodo.6392897" ID-GBIF-Taxon="194317678" ID-Zenodo-Dep="6392897" LSID="urn:lsid:plazi:treatment:03B887C14B028139BEE16C55FD1291E6" httpUri="http://treatment.plazi.org/id/03B887C14B028139BEE16C55FD1291E6" lastPageNumber="568" pageId="9" pageNumber="568">
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<paragraph id="8BAE36D74B028139BEE16C55FD249799" blockId="9.[151,660,365,428]" box="[151,660,365,392]" pageId="9" pageNumber="568">
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<taxonomicName id="4C114D544B028139BEE16C55FD249799" ID-CoL="VP92" authority="Heding &amp; Panning, 1954" authorityName="Heding &amp; Panning" authorityYear="1954" box="[151,660,365,392]" class="Holothuroidea" family="Phyllophoridae" genus="Cladolella" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="568" phylum="Echinodermata" rank="species" species="virgo">
<emphasis id="B965EAC54B028139BEE16C55FD249799" bold="true" box="[151,660,365,392]" pageId="9" pageNumber="568">
<emphasis id="B965EAC54B028139BEE16C55FEE89799" bold="true" box="[151,344,365,392]" italics="true" pageId="9" pageNumber="568">Cladolella virgo</emphasis>
<bibRefCitation id="EF804B264B028139BF166C55FD249799" author="Heding, S. G. &amp; Panning, A." box="[352,660,365,392]" pageId="9" pageNumber="568" pagination="7 - 209" refId="ref5809" refString="Heding, S. G. &amp; Panning, A. (1954) Phyllophoridae: eine Bearbeitung der Polytentaculaten Dendrochiroten Holothurien des Zoologischen Museums in Kopenhagen. Spolia Zoologica Musei Hauniensis, 13, 7 - 209." type="journal article" year="1954">Heding &amp; Panning, 1954</bibRefCitation>
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<subSubSection id="C30B655C4B028139BEE16CAAFEA697BD" box="[151,278,402,428]" pageId="9" pageNumber="568" type="description">
<paragraph id="8BAE36D74B028139BEE16CAAFEA697BD" blockId="9.[151,660,365,428]" box="[151,278,402,428]" pageId="9" pageNumber="568">
(
<figureCitation id="132A2A524B028139BEE96CAAFEBE97BD" box="[159,270,402,428]" captionStart="FIGURE 10" captionStartId="9.[151,250,1807,1831]" captionTargetBox="[151,1435,711,1787]" captionTargetId="figure-256@9.[151,1435,711,1789]" captionTargetPageId="9" captionText="FIGURE 10. Cladolella virgo Heding &amp; Panning, 1954, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from Heding &amp; Panning, 1954 and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection." figureDoi="http://doi.org/10.5281/zenodo.6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="9" pageNumber="568">Figure 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B655C4B028139BEE16CE8FA3597FA" box="[151,1413,464,491]" pageId="9" pageNumber="568" type="materials_examined">
<paragraph id="8BAE36D74B028139BEE16CE8FA3597FA" blockId="9.[151,1437,464,707]" box="[151,1413,464,491]" pageId="9" pageNumber="568">
<emphasis id="B965EAC54B028139BEE16CE8FE3197FB" bold="true" box="[151,385,464,490]" pageId="9" pageNumber="568">Material examined.</emphasis>
<materialsCitation id="3B793C8A4B028139BFF16CE8FA3597FA" ID-GBIF-Occurrence="3744207303" box="[391,1413,464,491]" collectingDate="1922-04-01" collectionCode="ZMUC" country="Indonesia" elevation="50" location="Off Doelah" pageId="9" pageNumber="568" specimenCode="HOL 256" specimenCount="1" typeStatus="holotype">
<typeStatus id="54AA88754B028139BFF16CE8FE5E97FB" box="[391,494,464,490]" pageId="9" pageNumber="568" type="holotype">Holotype</typeStatus>
:
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,
<collectingCountry id="F30676474B028139BC0D6CE8FD5597FB" box="[635,741,464,490]" name="Indonesia" pageId="9" pageNumber="568">Moluccas</collectingCountry>
,
<collectingCountry id="F30676474B028139BC996CE8FCE697FB" box="[751,854,464,490]" name="Indonesia" pageId="9" pageNumber="568">Indonesia</collectingCountry>
<quantity id="4CE99B324B028139BD2A6CE8FC2497FA" box="[860,916,464,491]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="568" unit="m" value="50.0">
<elevation id="003CD1E44B028139BD2A6CE8FC2497FA" box="[860,916,464,491]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="568" unit="m" value="50.0">50 m</elevation>
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,
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<collectingDate id="EFEBE9FF4B028139BDE96CE9FBB897FA" box="[927,1032,465,491]" pageId="9" pageNumber="568" value="1922-04-01">
1.IV.
<quantity id="4CE99B324B028139BDB86CE9FBB897FA" box="[974,1032,465,491]" metricMagnitude="0" metricUnit="m" metricValue="1.9225999999999999" pageId="9" pageNumber="568" unit="mm" value="1922.6">1922</quantity>
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, 60 mm length (
<collectionCode id="ED00AE124B028139BAC56CE9FAB197FA" box="[1203,1281,465,491]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="9" pageNumber="568">ZMUC</collectionCode>
<specimenCode id="DBB79EAC4B028139BB796CE9FAC997FA" box="[1295,1401,464,491]" collectionCode="ZMUC" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="9" pageNumber="568">HOL 256</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B655C4B028139BEB16CCCFB4F94D3" pageId="9" pageNumber="568" type="description">
<paragraph id="8BAE36D74B028139BEB16CCCFB4F94D3" blockId="9.[151,1437,464,707]" pageId="9" pageNumber="568">
<emphasis id="B965EAC54B028139BEB16CCCFEDD941F" bold="true" box="[199,365,500,526]" pageId="9" pageNumber="568">Redescription</emphasis>
. Body Ushaped; tube feet restricted to radii in double rows (
<figureCitation id="132A2A524B028139BA6B6CCCFBCD941E" box="[1053,1149,500,527]" captionStart="FIGURE 10" captionStartId="9.[151,250,1807,1831]" captionTargetBox="[151,1435,711,1787]" captionTargetId="figure-256@9.[151,1435,711,1789]" captionTargetPageId="9" captionText="FIGURE 10. Cladolella virgo Heding &amp; Panning, 1954, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from Heding &amp; Panning, 1954 and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection." figureDoi="http://doi.org/10.5281/zenodo.6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="9" pageNumber="568">Fig. 10a</figureCitation>
). Color, in ethanol, beige. 20 dendritic tentacles arranged in 2 circles (10 + 10):
<quantity id="4CE99B324B028139BC906F21FCAE9423" box="[742,798,536,563]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="9" pageNumber="568" unit="in" value="10.0">10 in</quantity>
outer circle and 10 smaller in inner circle (
<figureCitation id="132A2A524B028139BA8E6F20FAE99422" box="[1272,1369,536,563]" captionStart="FIGURE 10" captionStartId="9.[151,250,1807,1831]" captionTargetBox="[151,1435,711,1787]" captionTargetId="figure-256@9.[151,1435,711,1789]" captionTargetPageId="9" captionText="FIGURE 10. Cladolella virgo Heding &amp; Panning, 1954, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from Heding &amp; Panning, 1954 and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection." figureDoi="http://doi.org/10.5281/zenodo.6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="9" pageNumber="568">Fig. 10b</figureCitation>
). Two Polian vesicles (
<figureCitation id="132A2A524B028139BF266F04FE019446" box="[336,433,572,599]" captionStart="FIGURE 10" captionStartId="9.[151,250,1807,1831]" captionTargetBox="[151,1435,711,1787]" captionTargetId="figure-256@9.[151,1435,711,1789]" captionTargetPageId="9" captionText="FIGURE 10. Cladolella virgo Heding &amp; Panning, 1954, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from Heding &amp; Panning, 1954 and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection." figureDoi="http://doi.org/10.5281/zenodo.6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="9" pageNumber="568">Fig. 10c</figureCitation>
). Radial and interradial plates of calcareous ring fused. Radial plates subdivided, rectangular, with posterior projection and 2 AP (
<figureCitation id="132A2A524B028139BC0E6F58FD6B946A" box="[632,731,608,635]" captionStart="FIGURE 10" captionStartId="9.[151,250,1807,1831]" captionTargetBox="[151,1435,711,1787]" captionTargetId="figure-256@9.[151,1435,711,1789]" captionTargetPageId="9" captionText="FIGURE 10. Cladolella virgo Heding &amp; Panning, 1954, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from Heding &amp; Panning, 1954 and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection." figureDoi="http://doi.org/10.5281/zenodo.6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="9" pageNumber="568">Fig. 10d</figureCitation>
). Interradial plates arrow shaped. Radial and interradial plates lacking posterior processes (
<figureCitation id="132A2A524B028139BFAC6FBCFD8D948E" box="[474,573,644,671]" captionStart="FIGURE 10" captionStartId="9.[151,250,1807,1831]" captionTargetBox="[151,1435,711,1787]" captionTargetId="figure-256@9.[151,1435,711,1789]" captionTargetPageId="9" captionText="FIGURE 10. Cladolella virgo Heding &amp; Panning, 1954, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from Heding &amp; Panning, 1954 and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection." figureDoi="http://doi.org/10.5281/zenodo.6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="9" pageNumber="568">Fig. 10d</figureCitation>
). Body wall ossicles as rods, stellate at one end (with about six projections) and twelve at the other end (
<figureCitation id="132A2A524B028139BFA36F90FD8594D2" box="[469,565,680,707]" captionStart="FIGURE 10" captionStartId="9.[151,250,1807,1831]" captionTargetBox="[151,1435,711,1787]" captionTargetId="figure-256@9.[151,1435,711,1789]" captionTargetPageId="9" captionText="FIGURE 10. Cladolella virgo Heding &amp; Panning, 1954, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from Heding &amp; Panning, 1954 and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection." figureDoi="http://doi.org/10.5281/zenodo.6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="9" pageNumber="568">Fig. 10e</figureCitation>
). Tube feet with twopillared tables, arched disc, and end plate.
</paragraph>
</subSubSection>
<caption id="DF6E665F4B028139BEE16A37FBBC9161" ID-DOI="http://doi.org/10.5281/zenodo.6392930" ID-Zenodo-Dep="6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="9" pageNumber="568" startId="9.[151,250,1807,1831]" targetBox="[151,1435,711,1787]" targetPageId="9">
<paragraph id="8BAE36D74B028139BEE16A37FBBC9161" blockId="9.[151,1437,1807,1904]" pageId="9" pageNumber="568">
<emphasis id="B965EAC54B028139BEE16A37FEAF9136" bold="true" box="[151,287,1807,1831]" pageId="9" pageNumber="568">FIGURE 10.</emphasis>
<taxonomicName id="4C114D544B028139BF536A28FD7E9136" authority="Heding &amp; Panning, 1954" authorityName="Heding &amp; Panning" authorityYear="1954" box="[293,718,1807,1832]" class="Holothuroidea" family="Phyllophoridae" genus="Cladolella" kingdom="Animalia" order="Dendrochirotida" pageId="9" pageNumber="568" phylum="Echinodermata" rank="species" species="virgo">
<emphasis id="B965EAC54B028139BF536A28FE7B9136" box="[293,459,1808,1832]" italics="true" pageId="9" pageNumber="568">Cladolella virgo</emphasis>
<bibRefCitation id="EF804B264B028139BFA66A28FD7E9136" author="Heding, S. G. &amp; Panning, A." box="[464,718,1807,1832]" pageId="9" pageNumber="568" pagination="7 - 209" refId="ref5809" refString="Heding, S. G. &amp; Panning, A. (1954) Phyllophoridae: eine Bearbeitung der Polytentaculaten Dendrochiroten Holothurien des Zoologischen Museums in Kopenhagen. Spolia Zoologica Musei Hauniensis, 13, 7 - 209." type="journal article" year="1954">Heding &amp; Panning, 1954</bibRefCitation>
</taxonomicName>
, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from
<bibRefCitation id="EF804B264B028139BAD46A0CFA2D915A" author="Heding, S. G. &amp; Panning, A." box="[1186,1437,1843,1868]" pageId="9" pageNumber="568" pagination="7 - 209" refId="ref5809" refString="Heding, S. G. &amp; Panning, A. (1954) Phyllophoridae: eine Bearbeitung der Polytentaculaten Dendrochiroten Holothurien des Zoologischen Museums in Kopenhagen. Spolia Zoologica Musei Hauniensis, 13, 7 - 209." type="journal article" year="1954">Heding &amp; Panning, 1954</bibRefCitation>
and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection.
</paragraph>
</caption>
<subSubSection id="C30B655C4B028139BEE16A81FD1291E6" pageId="9" pageNumber="568" type="discussion">
<paragraph id="8BAE36D74B028139BEE16A81FD1291E6" blockId="9.[151,1436,1976,2039]" pageId="9" pageNumber="568">
<emphasis id="B965EAC54B028139BEE16A81FEBB91C2" bold="true" box="[151,267,1977,2003]" pageId="9" pageNumber="568">Remarks.</emphasis>
The opportunity of examining the
<typeStatus id="54AA88754B028139BCE56A80FD4591C3" box="[659,757,1976,2002]" pageId="9" pageNumber="568" type="holotype">holotype</typeStatus>
is taken to provide a photograph of the general aspect of the body (
<figureCitation id="132A2A524B028139BEA96AE4FE8C91E6" box="[223,316,2012,2039]" captionStart="FIGURE 10" captionStartId="9.[151,250,1807,1831]" captionTargetBox="[151,1435,711,1787]" captionTargetId="figure-256@9.[151,1435,711,1789]" captionTargetPageId="9" captionText="FIGURE 10. Cladolella virgo Heding &amp; Panning, 1954, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from Heding &amp; Panning, 1954 and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection." figureDoi="http://doi.org/10.5281/zenodo.6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="9" pageNumber="568">Fig. 10a</figureCitation>
) and calcareous ring (
<figureCitation id="132A2A524B028139BC436AE4FD2591E6" box="[565,661,2012,2039]" captionStart="FIGURE 10" captionStartId="9.[151,250,1807,1831]" captionTargetBox="[151,1435,711,1787]" captionTargetId="figure-256@9.[151,1435,711,1789]" captionTargetPageId="9" captionText="FIGURE 10. Cladolella virgo Heding &amp; Panning, 1954, (ZMUC-HOL 256). (a) specimen in dorsal view; (b) muscular system; (c) photograph of the calcareous ring in lateral view; (d) drawing of the calcareous ring (extracted from Heding &amp; Panning, 1954 and (e) rods from body wall. Abbreviations: PV, Polian vesicle; AP, anterior projection." figureDoi="http://doi.org/10.5281/zenodo.6392930" httpUri="https://zenodo.org/record/6392930/files/figure.png" pageId="9" pageNumber="568">Fig. 10c</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>