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<document id="3532349CA7DF2DF6051E631724D130B9" ID-DOI="10.11646/zootaxa.5540.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14240496" ID-ZooBank="4DBD3A92-F14B-4C5B-95B3-2B430EC197DD" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732832094216" checkinUser="plazi" docAuthor="Wu, Jingyu &amp; Liu, Xingyue" docDate="2024" docId="03B88790FFD2FFD39EE7CE959C344C9E" docLanguage="en" docName="zootaxa.5540.1.1.pdf" docOrigin="Zootaxa 5540 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5540.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Retipenna sichuanica Yang &amp; Yang 1992" docType="treatment" docVersion="2" lastPageNumber="119" masterDocId="FF81FFE8FFA7FFA59E70CE029F264D33" masterDocTitle="Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China" masterLastPageNumber="169" masterPageNumber="1" pageNumber="118" updateTime="1732841452334" updateUser="ExternalLinkService">
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<mods:title id="F1FCD5FD1CB3BF5AA14BBF426987F952">Systematics of the green lacewing tribe Ankylopterygini Navás, 1910 (Neuroptera: Chrysopidae: Chrysopinae) from China</mods:title>
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<mods:namePart id="54709D16911C59638FBAFC0210C748C3">Wu, Jingyu</mods:namePart>
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<mods:affiliation id="B9710625D2D466B639239B59D72E2EDD">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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<mods:namePart id="51305EEE9DF8C2223E77048C84927209">Liu, Xingyue</mods:namePart>
<mods:nameIdentifier id="EACDB7726CDDE9F24099F4089D567DAB" type="ORCID">0000-0001-6677-7726</mods:nameIdentifier>
<mods:affiliation id="75294A2C42BF381C1034ACBF43E14721">Department of Entomology, College of Plant Protection, China Agricultural University, Beijing, 100193, China wujingyu 176 @ 163. com; https: // orcid. org / 0000 - 0001 - 6677 - 7726 Corresponding author: xingyue _ liu @ yahoo. com; https: // orcid. org / 0000 - 0002 - 9168 - 0659</mods:affiliation>
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<mods:title id="FF15D42683C7395DBF7A0C61186B5C59">Zootaxa</mods:title>
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<mods:date id="48825C8BAE2A42601D00BAB47E95A772">2024</mods:date>
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<mods:number id="A7927ACAB060CDE8B23F711C4A0BA461">5540</mods:number>
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<mods:identifier id="52F84BED55AA61F072E1F43C57B9210A" type="DOI">10.11646/zootaxa.5540.1.1</mods:identifier>
<mods:identifier id="2919C6A3BA94E46D9AFDE86BC0E003F3" type="ISSN">1175-5334</mods:identifier>
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<treatment id="03B88790FFD2FFD39EE7CE959C344C9E" LSID="urn:lsid:plazi:treatment:03B88790FFD2FFD39EE7CE959C344C9E" httpUri="http://treatment.plazi.org/id/03B88790FFD2FFD39EE7CE959C344C9E" lastPageId="118" lastPageNumber="119" pageId="117" pageNumber="118">
<subSubSection id="C30B650DFFD2FFD09EE7CE959DEA4D82" box="[151,716,151,178]" pageId="117" pageNumber="118" type="nomenclature">
<paragraph id="8BAE3686FFD2FFD09EE7CE959DEA4D82" blockId="117.[151,716,151,213]" box="[151,716,151,178]" pageId="117" pageNumber="118">
<emphasis id="B965EA94FFD2FFD09EE7CE959DEA4D82" bold="true" box="[151,716,151,178]" pageId="117" pageNumber="118">
[54]
<taxonomicName id="4C114D05FFD2FFD09EBFCE959DEA4D82" ID-CoL="6X3X3" authority="Yang &amp; Yang, 1992" authorityName="Yang &amp; Yang" authorityYear="1992" box="[207,716,151,178]" class="Insecta" family="Chrysopidae" genus="Retipenna" kingdom="Animalia" order="Neuroptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="sichuanica">
<emphasis id="B965EA94FFD2FFD09EBFCE959EF14D81" bold="true" box="[207,471,151,178]" italics="true" pageId="117" pageNumber="118">Retipenna sichuanica</emphasis>
Yang &amp; Yang, 1992
</taxonomicName>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFD2FFD09EE7CEB99E014DE6" box="[151,295,187,213]" pageId="117" pageNumber="118" type="description">
<paragraph id="8BAE3686FFD2FFD09EE7CEB99E014DE6" blockId="117.[151,716,151,213]" box="[151,295,187,213]" pageId="117" pageNumber="118">
(
<figureCitation id="132A2A03FFD2FFD09EEFCEB99FD04DE6" box="[159,246,187,213]" captionStart="FIGURE 64" captionStartId="117.[151,250,1436,1460]" captionTargetBox="[161,1426,315,1403]" captionTargetId="figure-49@117.[151,1436,304,1413]" captionTargetPageId="117" captionText="FIGURE 64. Retipenna sichuanica Yang &amp; Yang, 1992. A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Thorax, dorsal view; E. Male genitalia, lateral view; F. Gonarcal complex, dorsal view; G. Gonarcal complex, lateral view; H. Arcessus. Scale bar: 5 mm (A), 1 mm (B–D)." figureDoi="http://doi.org/10.5281/zenodo.14240674" httpUri="https://zenodo.org/record/14240674/files/figure.png" pageId="117" pageNumber="118">Figs 64</figureCitation>
,
<figureCitation id="132A2A03FFD2FFD09F72CEB99E394DE6" box="[258,287,187,213]" captionStart="FIGURE 88" captionStartId="147.[151,250,975,999]" captionTargetBox="[197,1391,181,952]" captionTargetId="figure-19@147.[197,1391,181,952]" captionTargetPageId="147" captionText="FIGURE 88. Distribution map of species of Retipenna from China." figureDoi="http://doi.org/10.5281/zenodo.14240722" httpUri="https://zenodo.org/record/14240722/files/figure.png" pageId="117" pageNumber="118">88</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFD2FFD09EE7CF029A144C24" box="[151,1330,255,280]" pageId="117" pageNumber="118" type="reference_group">
<paragraph id="8BAE3686FFD2FFD09EE7CF029A144C24" blockId="117.[151,1330,255,280]" box="[151,1330,255,280]" pageId="117" pageNumber="118">
<taxonomicName id="4C114D05FFD2FFD09EE7CF029D534C24" ID-CoL="6X3X3" authority="Yang &amp; Yang, 1992: 464" authorityName="Yang &amp; Yang" authorityPageNumber="464" authorityYear="1992" box="[151,629,255,280]" class="Insecta" family="Chrysopidae" genus="Retipenna" kingdom="Animalia" order="Neuroptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="sichuanica">
<emphasis id="B965EA94FFD2FFD09EE7CF029E544C2B" box="[151,370,256,280]" italics="true" pageId="117" pageNumber="118">Retipenna sichuanica</emphasis>
<materialsCitation id="3B793CDBFFD2FFD09F07CF029D534C24" box="[375,629,255,280]" country="China" location="Xinxing" municipality="Luding" pageId="117" pageNumber="118" specimenCount="1" stateProvince="Sichuan" typeStatus="holotype">Yang &amp; Yang, 1992: 464</materialsCitation>
</taxonomicName>
. Type locality:
<collectingCountry id="F3067616FFD2FFD09D60CF029C6B4C2B" box="[784,845,256,280]" name="China" pageId="117" pageNumber="118">China</collectingCountry>
(
<collectingRegion id="49D5F864FFD2FFD09D2ACF029C964C2B" box="[858,944,256,280]" country="China" name="Sichuan" pageId="117" pageNumber="118">Sichuan</collectingRegion>
,
<collectingMunicipality id="6BCAACFCFFD2FFD09DCACF029B204C2B" box="[954,1030,256,280]" pageId="117" pageNumber="118">Luding</collectingMunicipality>
,
<location id="8ECE605DFFD2FFD09A60CF029B4E4C2B" LSID="urn:lsid:plazi:treatment:03B88790FFD2FFD39EE7CE959C344C9E:8ECE605DFFD2FFD09A60CF029B4E4C2B" box="[1040,1128,256,280]" country="China" municipality="Luding" name="Xinxing" pageId="117" pageNumber="118" stateProvince="Sichuan">Xinxing</location>
).
<typeStatus id="54AA8824FFD2FFD09A08CF029BF04C2B" box="[1144,1238,256,280]" pageId="117" pageNumber="118" type="holotype">Holotype</typeStatus>
in CAU.
</paragraph>
</subSubSection>
<caption id="DF6E660EFFD2FFD09EE7CB9E9D4548CE" ID-DOI="http://doi.org/10.5281/zenodo.14240674" ID-Zenodo-Dep="14240674" httpUri="https://zenodo.org/record/14240674/files/figure.png" pageId="117" pageNumber="118" startId="117.[151,250,1436,1460]" targetBox="[161,1426,315,1403]" targetPageId="117" targetType="figure">
<paragraph id="8BAE3686FFD2FFD09EE7CB9E9D4548CE" blockId="117.[151,1437,1436,1533]" pageId="117" pageNumber="118">
<emphasis id="B965EA94FFD2FFD09EE7CB9E9E044886" bold="true" box="[151,290,1436,1461]" pageId="117" pageNumber="118">FIGURE 64.</emphasis>
<taxonomicName id="4C114D05FFD2FFD09F59CB9F9DFB4886" authority="Yang &amp; Yang, 1992" authorityName="Yang &amp; Yang" authorityYear="1992" box="[297,733,1437,1461]" class="Insecta" family="Chrysopidae" genus="Retipenna" kingdom="Animalia" order="Neuroptera" pageId="117" pageNumber="118" phylum="Arthropoda" rank="species" species="sichuanica">
<emphasis id="B965EA94FFD2FFD09F59CB9F9D204886" box="[297,518,1437,1461]" italics="true" pageId="117" pageNumber="118">Retipenna sichuanica</emphasis>
Yang &amp; Yang, 1992
</taxonomicName>
. A. Dorsal habitus; B. Head, frontal view; C. Head, lateral view; D. Thorax, dorsal view; E. Male genitalia, lateral view; F. Gonarcal complex, dorsal view; G. Gonarcal complex, lateral view; H. Arcessus. Scale bar: 5 mm (A), 1 mm (B–D).
</paragraph>
</caption>
<subSubSection id="C30B650DFFD2FFD09EE7C84D9BB04BBE" pageId="117" pageNumber="118" type="diagnosis">
<paragraph id="8BAE3686FFD2FFD09EE7C84D9BB04BBE" blockId="117.[151,1437,1615,2037]" pageId="117" pageNumber="118">
<emphasis id="B965EA94FFD2FFD09EE7C84D9E344B5A" bold="true" box="[151,274,1615,1641]" pageId="117" pageNumber="118">Diagnosis.</emphasis>
This species is characterized by the presence of the relatively large body, the head, thorax and wings without markings, and the median arch of male gonarcus with deep U-shaped invagination.
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFD2FFD39EB7C8959CD94C2E" lastPageId="118" lastPageNumber="119" pageId="117" pageNumber="118" type="description">
<paragraph id="8BAE3686FFD2FFD09EB7C8959BDB4B82" blockId="117.[151,1437,1615,2037]" box="[199,1277,1687,1714]" pageId="117" pageNumber="118">
<emphasis id="B965EA94FFD2FFD09EB7C8959E7F4B82" bold="true" box="[199,345,1687,1713]" pageId="117" pageNumber="118">Description.</emphasis>
Male. Body length
<quantity id="4CE99B63FFD2FFD09C48C8959DB84B82" box="[568,670,1687,1714]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="117" pageNumber="118" unit="mm" value="13.5">13.5 mm</quantity>
, forewing length 22.0 mm, hindwing length
<quantity id="4CE99B63FFD2FFD09AE2C8959BDF4B82" box="[1170,1273,1687,1714]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" pageId="117" pageNumber="118" unit="mm" value="18.5">18.5 mm</quantity>
.
</paragraph>
<paragraph id="8BAE3686FFD2FFD09EB7C8B99A3D4BCA" blockId="117.[151,1437,1615,2037]" pageId="117" pageNumber="118">Head without spots, vertex raised. Compound eyes prominent and black. Antennae yellow, scape and pedicel wider than flagellum. Frons, clypeus and gena not marked. Maxillary and labial palpi brownish yellow.</paragraph>
<paragraph id="8BAE3686FFD2FFD09EB7C9019A364A2E" blockId="117.[151,1437,1615,2037]" box="[199,1296,1795,1821]" pageId="117" pageNumber="118">Pronotum anterior margin shorter than posterior margin, meso- and metanotum without markings.</paragraph>
<paragraph id="8BAE3686FFD2FFD09EB7C9259B6C4A72" blockId="117.[151,1437,1615,2037]" box="[199,1098,1831,1857]" pageId="117" pageNumber="118">Legs with dense setae, without spots. Pretarsal claws brown, with basal dilation.</paragraph>
<paragraph id="8BAE3686FFD2FFD09EB7C9499BCF4A9E" blockId="117.[151,1437,1615,2037]" pageId="117" pageNumber="118">
Forewing broad, without shadings. Pterostigma indistinct. Costal area with 30 crossveins; 16 radial crossveins present.
<emphasis id="B965EA94FFD2FFD09E87C96D9E3C4ABA" box="[247,282,1903,1929]" italics="true" pageId="117" pageNumber="118">m1</emphasis>
smaller than
<emphasis id="B965EA94FFD2FFD09FC4C96D9EF04ABA" box="[436,470,1903,1929]" italics="true" pageId="117" pageNumber="118">m2</emphasis>
;
<emphasis id="B965EA94FFD2FFD09F96C9729D244ABA" box="[486,514,1904,1929]" italics="true" pageId="117" pageNumber="118">im</emphasis>
ovoid; 12 psm-psc crossveins present.
<emphasis id="B965EA94FFD2FFD09DCFC96D9CFC4ABA" box="[959,986,1903,1929]" italics="true" pageId="117" pageNumber="118">c1</emphasis>
smaller than
<emphasis id="B965EA94FFD2FFD09A05C96D9BA94ABA" box="[1141,1167,1903,1929]" italics="true" pageId="117" pageNumber="118">c2</emphasis>
,
<emphasis id="B965EA94FFD2FFD09AEEC96D9BE34ABA" box="[1182,1221,1903,1929]" italics="true" pageId="117" pageNumber="118">dcc</emphasis>
closed. A1 forked, A2 and A3 simple. Inner gradate series with 11 crossveins, outer gradate series with 12 crossveins.
</paragraph>
<paragraph id="8BAE3686FFD2FFD09EB7C9B59BCC4AC6" blockId="117.[151,1437,1615,2037]" pageId="117" pageNumber="118">Hindwing broad, without shadings. Costal area with 25 crossveins; 17 radial crossveins and 10 psm-psc crossveins present. Inner gradate series with 10 crossveins, outer gradate series with 11 crossveins.</paragraph>
<paragraph id="8BAE3686FFD1FFD39EB7CE959CD94C2E" blockId="118.[151,1436,151,429]" pageId="118" pageNumber="119">Abdomen with dense setae. Tergum VII and sternum VII subquadrate in lateral view; tergum VIII slightly shorter than tergum VII; sternum VIII fused with sternum IX and with small tubercle near tip; tergum IX fused with ectoproct. Callus cerci ovoid. Gonarcus arcuate, median arch with deep U-shaped invagination, lateral arms narrow at base and wide at tip, entoprocessus widest at tip; arcessus widest near base.</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFD1FFD39EB7CF249A6F4C72" box="[199,1353,294,321]" pageId="118" pageNumber="119" type="materials_examined">
<paragraph id="8BAE3686FFD1FFD39EB7CF249A6F4C72" blockId="118.[151,1436,151,429]" box="[199,1353,294,321]" pageId="118" pageNumber="119">
<emphasis id="B965EA94FFD1FFD39EB7CF249E104C73" bold="true" box="[199,310,294,320]" pageId="118" pageNumber="119">Material.</emphasis>
<materialsCitation id="3B793CDBFFD1FFD39F4DCF259A6F4C72" box="[317,1353,295,321]" collectingDate="1983-06-14" collectionCode="CAU" country="China" county="Luding" elevation="1800" location="Shuyong Wang" municipality="Xinxing" pageId="118" pageNumber="119" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<typeStatus id="54AA8824FFD1FFD39F4DCF259E834C72" box="[317,421,295,321]" pageId="118" pageNumber="119" type="holotype">Holotype</typeStatus>
<specimenCount id="9D17FD0FFFD1FFD39FDDCF259EE24C72" box="[429,452,295,321]" count="1" pageId="118" pageNumber="119" type="male">♂</specimenCount>
,
<collectingCountry id="F3067616FFD1FFD39FA0CF259D304C72" box="[464,534,295,321]" name="China" pageId="118" pageNumber="119">China</collectingCountry>
,
<collectingRegion id="49D5F864FFD1FFD39C50CF259D5B4C72" box="[544,637,295,321]" country="China" name="Sichuan" pageId="118" pageNumber="119">Sichuan</collectingRegion>
,
<collectingCounty id="62CF4E0AFFD1FFD39CF9CF259DFB4C72" box="[649,733,295,321]" pageId="118" pageNumber="119">Luding</collectingCounty>
,
<collectingMunicipality id="6BCAACFCFFD1FFD39C98CF259C614C72" box="[744,839,295,321]" pageId="118" pageNumber="119">Xinxing</collectingMunicipality>
,
<quantity id="4CE99B63FFD1FFD39D21CF259C8E4C72" box="[849,936,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="118" pageNumber="119" unit="m" value="1800.0">
<elevation id="003CD1B5FFD1FFD39D21CF259C8E4C72" box="[849,936,295,321]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="118" pageNumber="119" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date id="FFAF1046FFD1FFD39DC4CF259B104C72" box="[948,1078,295,321]" pageId="118" pageNumber="119" value="1983-06-14">
<collectingDate id="EFEBE9AEFFD1FFD39DC4CF259B104C72" box="[948,1078,295,321]" pageId="118" pageNumber="119" value="1983-06-14">1983.VI.14</collectingDate>
</date>
,
<location id="8ECE605DFFD1FFD39A32CF259BCB4C72" LSID="urn:lsid:plazi:treatment:03B88790FFD2FFD39EE7CE959C344C9E:8ECE605DFFD1FFD39A32CF259BCB4C72" box="[1090,1261,295,321]" country="China" county="Luding" municipality="Xinxing" name="Shuyong Wang" pageId="118" pageNumber="119" stateProvince="Sichuan">Shuyong Wang</location>
(
<collectionCode id="ED00AE43FFD1FFD39A8BCF259A1A4C72" box="[1275,1340,295,321]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="118" pageNumber="119">CAU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFD1FFD39EB7CF489D004C56" box="[199,550,330,357]" pageId="118" pageNumber="119" type="distribution">
<paragraph id="8BAE3686FFD1FFD39EB7CF489D004C56" blockId="118.[151,1436,151,429]" box="[199,550,330,357]" pageId="118" pageNumber="119">
<emphasis id="B965EA94FFD1FFD39EB7CF489E474C57" bold="true" box="[199,353,330,356]" pageId="118" pageNumber="119">Distribution.</emphasis>
<collectingCountry id="F3067616FFD1FFD39F17CF499E8C4C56" box="[359,426,331,357]" name="China" pageId="118" pageNumber="119">China</collectingCountry>
(
<collectingRegion id="49D5F864FFD1FFD39FC9CF499D3C4C56" box="[441,538,331,357]" country="China" name="Sichuan" pageId="118" pageNumber="119">Sichuan</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C30B650DFFD1FFD39EB7CF6D9C344C9E" pageId="118" pageNumber="119" type="discussion">
<paragraph id="8BAE3686FFD1FFD39EB7CF6D9C344C9E" blockId="118.[151,1436,151,429]" pageId="118" pageNumber="119">
<emphasis id="B965EA94FFD1FFD39EB7CF6D9E1D4CBA" bold="true" box="[199,315,367,393]" pageId="118" pageNumber="119">Remarks.</emphasis>
This species has no spots or markings on body, but can be distinguished by the U-shaped invagination of median arch of male gonarcus and the broad arcessus.
</paragraph>
</subSubSection>
</treatment>
</document>
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