treatments-xml/data/03/B7/DE/03B7DE6C8A3BF86FFF38C0E9FB82C79B.xml
2024-06-21 12:22:17 +02:00

554 lines
79 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="D766B1D6C6DB9772A737A905BC974171" ID-CLB-Dataset="39041" ID-DOI="10.11646/zootaxa.3617.1.1" ID-GBIF-Dataset="6174b35e-3228-4d50-a6ed-844e244f8b64" ID-ISSN="1175-5326" ID-Zenodo-Dep="248150" ID-ZooBank="4DCCD152-65DA-44A3-AB19-59811384E1E7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1463638623780" checkinUser="plazi" docAuthor="Calcinai, Barbara, Bavestrello, Giorgio, Bertolino, Marco, Pica, Daniela, Wagner, Daniel &amp; Cerrano, Carlo" docDate="2013" docId="03B7DE6C8A3BF86FFF38C0E9FB82C79B" docLanguage="en" docName="zt03617p061.pdf.imf" docOrigin="Zootaxa 3617 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hymedesmia (Hymedesmia) spinata Calcinai, Bavestrello, Bertolino, Pica, Wagner &amp; Cerrano, 2013, n. sp." docType="treatment" docVersion="8" lastPageNumber="25" masterDocId="FF8EA6148A2FF877FFAFC227FFC9C52C" masterDocTitle="Sponges associated with octocorals in the Indo-Pacific, with the description of four new species" masterLastPageNumber="61" masterPageNumber="1" pageNumber="21" updateTime="1698662548627" updateUser="ExternalLinkService">
<mods:mods id="E60B5559821BC3306DDE0FD5B204F85D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1F03272F52882F3089AF9D60F69B0128">
<mods:title id="8CF398C8528EFAD7BD0663CC402BECF6">Sponges associated with octocorals in the Indo-Pacific, with the description of four new species</mods:title>
</mods:titleInfo>
<mods:name id="64346644ED77CFA6C045905F48CCE81F" type="personal">
<mods:role id="14DC90F9999FE533196AA84218BC6FF7">
<mods:roleTerm id="B21954F599B308753E02C4C95BA24207">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BCBB42249DF1A4AEC202A014EFBCB852">Calcinai, Barbara</mods:namePart>
</mods:name>
<mods:name id="722A540083AC68D95D48D8F356F1E66A" type="personal">
<mods:role id="FC219CE33E86085840E1F35D12C3B37D">
<mods:roleTerm id="40E2DC652C17C2438C4541033D8EDD91">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9EC72266B008CEA6B2280BA5E4AB4A9C">Bavestrello, Giorgio</mods:namePart>
</mods:name>
<mods:name id="916472A4FBD38D7A439F1153C0494C30" type="personal">
<mods:role id="19903704529E5737E7A5A0B7E71A82E9">
<mods:roleTerm id="C51A30E7EEF61B423C5069FD053BDED2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5DF98438ED598C9BFE32671D6ED5D26B">Bertolino, Marco</mods:namePart>
</mods:name>
<mods:name id="9C3B690D43D7798619BE147F206E2E0F" type="personal">
<mods:role id="B1030AFDDC310F1E39B9913F5FF041E3">
<mods:roleTerm id="24551F975B423603FCF924943ACDD47F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4A8DC78DFD9DE9973CFE14208ECE350A">Pica, Daniela</mods:namePart>
</mods:name>
<mods:name id="89B7607F36AE677CB63DAD21A4732F77" type="personal">
<mods:role id="C22207886CB9593FF20CE0CCA2F5F20E">
<mods:roleTerm id="70E6AEE94194A66E899BDA534F480387">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8E3FD37D54FF7183B860835998B8414D">Wagner, Daniel</mods:namePart>
</mods:name>
<mods:name id="BCF110C752D4DE4DBBDF01D46EF77DBB" type="personal">
<mods:role id="1B0A0CD66C9CA4E4ADC4D89D3EB6BE60">
<mods:roleTerm id="E7C9774C132B203B5CA37A343B2F4AA5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3D7B212CE2507BD726E3CB7A4C24B35E">Cerrano, Carlo</mods:namePart>
</mods:name>
<mods:typeOfResource id="5183855C35C5E051029E4A631144BC23">text</mods:typeOfResource>
<mods:relatedItem id="07FC73F7CBE702A3AC884AD9ECFA976F" type="host">
<mods:titleInfo id="2B90F4C92FDF465E6F1B0263721936F2">
<mods:title id="457EDFAA6246B74F129AE941E0E58995">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E4EE1B845C787B364DB24E0C996660FF">
<mods:date id="2001EF339809008A7DC7D7148A1860C6">2013</mods:date>
<mods:detail id="3C4851A7B9211B7B852D1300AB4EC97B" type="volume">
<mods:number id="B78AB0D6D7B6769C6CE6287588EC2A43">3617</mods:number>
</mods:detail>
<mods:detail id="5D083970A8D638F3CEA02AC0B54803B9" type="issue">
<mods:number id="AE9A4376F68887591B0897CCC1532F36">1</mods:number>
</mods:detail>
<mods:extent id="AC662A0E69F3935A87B809E2FF606DCC" unit="page">
<mods:start id="F1097F3DA8C93E16CF4335F0E723DA8D">1</mods:start>
<mods:end id="2B623E642C37F1BDF53EE56B6AC91BE7">61</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3A665AA904C238ABEF2EB8F3EE331827">journal article</mods:classification>
<mods:identifier id="92B3C8A5F3E1C5566ACE2755DAC4A186" type="CLB-Dataset">39041</mods:identifier>
<mods:identifier id="20BAECFFE13D126DE9105A99FB490636" type="DOI">10.11646/zootaxa.3617.1.1</mods:identifier>
<mods:identifier id="8DB32BD6B6DA7EF10CC1432EE4DCB159" type="GBIF-Dataset">6174b35e-3228-4d50-a6ed-844e244f8b64</mods:identifier>
<mods:identifier id="A8CD2F1B1CA0CE314AC67769B2E111ED" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B9315C3960FB3FD021493BE24570CCF2" type="Zenodo-Dep">248150</mods:identifier>
<mods:identifier id="A8D8DBB11997085A9A595F6B02C6D873" type="ZooBank">4DCCD152-65DA-44A3-AB19-59811384E1E7</mods:identifier>
</mods:mods>
<treatment id="03B7DE6C8A3BF86FFF38C0E9FB82C79B" ID-DOI="http://doi.org/10.5281/zenodo.6156077" ID-GBIF-Taxon="120682005" ID-Zenodo-Dep="6156077" LSID="urn:lsid:plazi:treatment:03B7DE6C8A3BF86FFF38C0E9FB82C79B" httpUri="http://treatment.plazi.org/id/03B7DE6C8A3BF86FFF38C0E9FB82C79B" lastPageId="24" lastPageNumber="25" pageId="20" pageNumber="21">
<subSubSection id="C3043CF18A3BF863FF38C0E9FE2AC625" pageId="20" pageNumber="21" type="nomenclature">
<paragraph id="8BA16F7A8A3BF863FF38C0E9FD5AC7C4" blockId="20.[151,661,614,777]" box="[151,659,718,744]" pageId="20" pageNumber="21">
<heading id="D0E9D8168A3BF863FF38C0E9FD5AC7C4" bold="true" box="[151,659,718,744]" fontSize="11" level="1" pageId="20" pageNumber="21" reason="1">
<emphasis id="B96AB3688A3BF863FF38C0E9FD5AC7C4" bold="true" box="[151,659,718,744]" pageId="20" pageNumber="21">
<taxonomicName id="4C1E14F98A3BF863FF38C0E9FD80C7C4" ID-CoL="3NH97" box="[151,585,718,744]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="20" pageNumber="21" phylum="Porifera" rank="species" species="spinata" status="sp. nov." subGenus="Hymedesmia">
<emphasis id="B96AB3688A3BF863FF38C0E9FD80C7C4" bold="true" box="[151,585,718,744]" italics="true" pageId="20" pageNumber="21">Hymedesmia (Hymedesmia) spinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2590E138A3BF863FDFFC0E9FD5AC7C4" box="[592,659,718,744]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA16F7A8A3BF863FF38C0D6FE2AC625" blockId="20.[151,661,614,777]" box="[151,483,753,777]" pageId="20" pageNumber="21">
(
<figureCitation id="132573FF8A3BF863FF30C0D6FF33C625" box="[159,250,753,777]" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">Figs. 12</figureCitation>
AJ; 13) (
<tableCitation id="C69C5AC18A3BF863FEC5C0D6FE77C625" box="[362,446,753,777]" captionStart="TABLE 5" captionStartId="22.[151,239,316,340]" captionTargetBox="[225,1354,472,642]" captionTargetPageId="22" captionText="TABLE 5. Hymedesmia (Hymedesmia) spinata n. sp. Analysis with Bhattacharyas method indicating the numbers of identified cohorts (groups) of acanthostyles, the mean length for each cohort and the number of spicules present in each cohort. S. d. Standard deviation; S. I. Separation index; separation of the groups is generally unreliable when the SI value is below 2. N. a. not applicable." httpUri="http://table.plazi.org/id/DF613FF28A39F861FF38C31BFE2BC498" pageId="20" pageNumber="21" tableUuid="DF613FF28A39F861FF38C31BFE2BC498">Tabs. 5</tableCitation>
; 6)
</paragraph>
</subSubSection>
<subSubSection id="C3043CF18A3BF86FFF38C11EFB82C79B" lastPageId="24" lastPageNumber="25" pageId="20" pageNumber="21" type="materials_examined">
<paragraph id="8BA16F7A8A3BF863FF38C11EFE52C659" blockId="20.[151,1437,825,2037]" pageId="20" pageNumber="21">
<typeStatus id="54A5D1D88A3BF863FF38C11EFF36C67D" box="[151,255,825,849]" pageId="20" pageNumber="21" type="holotype">Holotype</typeStatus>
material.
<collectionCode id="ED0FF7BF8A3BF863FEC0C11EFE09C67D" box="[367,448,825,849]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="20" pageNumber="21">MSNG</collectionCode>
56262: sample Bugor 513:
<collectingCountry id="F3092FEA8A3BF863FD58C11EFCAEC67D" box="[759,871,825,849]" name="Indonesia" pageId="20" pageNumber="21">Indonesia</collectingCountry>
, North Sulawesi, Siladen
<collectingCountry id="F3092FEA8A3BF863FB27C11EFB06C67D" box="[1160,1231,825,849]" name="Iceland" pageId="20" pageNumber="21">Island</collectingCountry>
, Siladen Barat,
<quantity id="4CE6C29F8A3BF863FA2FC11EFF66C659" metricMagnitude="1" metricUnit="m" metricValue="2.3" pageId="20" pageNumber="21" unit="m" value="23.0">23 m</quantity>
,
<date id="FFA049BA8A3BF863FF14C17AFE5EC659" box="[187,407,861,885]" pageId="20" pageNumber="21" value="2007-09-11">11 September 2007</date>
.
</paragraph>
<paragraph id="8BA16F7A8A3BF863FF68C1A5FE57C691" blockId="20.[151,1437,825,2037]" pageId="20" pageNumber="21">
<typeStatus id="54A5D1D88A3BF863FF68C1A5FEE0C6B6" box="[199,297,898,922]" pageId="20" pageNumber="21" type="paratype">Paratype</typeStatus>
material.
<collectionCode id="ED0FF7BF8A3BF863FE37C1A6FE20C6B5" box="[408,489,897,921]" country="Italy" httpUri="http://grbio.org/cool/1trn-7b59" name="Museo Civico di Storia Naturale di Genova 'Giacomo Doria'" pageId="20" pageNumber="21">MSNG</collectionCode>
56263: sample Bugor 311:
<collectingCountry id="F3092FEA8A3BF863FCB2C1A6FC44C6B5" box="[797,909,897,921]" name="Indonesia" pageId="20" pageNumber="21">Indonesia</collectingCountry>
, North Sulawesi, Siladen
<collectingCountry id="F3092FEA8A3BF863FB03C1A6FB3AC6B5" box="[1196,1267,897,921]" name="Iceland" pageId="20" pageNumber="21">Island</collectingCountry>
, Siladen Barat,
<quantity id="4CE6C29F8A3BF863FF38C182FF1BC691" box="[151,210,933,957]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="20" pageNumber="21" unit="m" value="25.0">25 m</quantity>
,
<date id="FFA049BA8A3BF863FF72C182FE52C691" box="[221,411,933,958]" pageId="20" pageNumber="21" value="2007-01-18">18 January 2007</date>
.
</paragraph>
<paragraph id="8BA16F7A8A3BF863FF68C1EEFF1FC161" blockId="20.[151,1437,825,2037]" pageId="20" pageNumber="21">
Other examined material: sample Bugor 309:
<collectingCountry id="F3092FEA8A3BF863FD61C1EEFCF7C6CD" box="[718,830,969,993]" name="Indonesia" pageId="20" pageNumber="21">Indonesia</collectingCountry>
, North Sulawesi, Siladen
<collectingCountry id="F3092FEA8A3BF863FBCCC1EEFB63C6CD" box="[1123,1194,969,993]" name="Iceland" pageId="20" pageNumber="21">Island</collectingCountry>
, Siladen Barat, depth not stated,
<date id="FFA049BA8A3BF863FEBBC1CAFE1AC129" box="[276,467,1005,1030]" pageId="20" pageNumber="21" value="2007-01-18">18 January 2007</date>
; sample
<collectingCountry id="F3092FEA8A3BF863FD98C1CAFDA3C129" box="[567,618,1005,1029]" name="India" pageId="20" pageNumber="21">Indo</collectingCountry>
23:
<collectingCountry id="F3092FEA8A3BF863FD30C1CAFCC6C129" box="[671,783,1005,1029]" name="Indonesia" pageId="20" pageNumber="21">Indonesia</collectingCountry>
, North Sulawesi, Siladen
<collectingCountry id="F3092FEA8A3BF863FB99C1CAFBB4C129" box="[1078,1149,1005,1029]" name="Iceland" pageId="20" pageNumber="21">Island</collectingCountry>
, Siladen Barat, depth not stated,
<date id="FFA049BA8A3BF863FF45C636FE6BC105" box="[234,418,1041,1065]" pageId="20" pageNumber="21" value="2008-10-08">8 October 2008</date>
; sample Bugor 410 (bis2):
<collectingCountry id="F3092FEA8A3BF863FD4DC636FC9BC105" box="[738,850,1041,1065]" name="Indonesia" pageId="20" pageNumber="21">Indonesia</collectingCountry>
, North Sulawesi, Siladen
<collectingCountry id="F3092FEA8A3BF863FB2EC636FB01C105" box="[1153,1224,1041,1065]" name="Iceland" pageId="20" pageNumber="21">Island</collectingCountry>
,
<quantity id="4CE6C29F8A3BF863FB78C636FADCC105" box="[1239,1301,1041,1065]" metricMagnitude="1" metricUnit="m" metricValue="2.6" pageId="20" pageNumber="21" unit="m" value="26.0">26 m</quantity>
,
<date id="FFA049BA8A3BF863FA8AC636FF1AC161" pageId="20" pageNumber="21" value="2007-09">September 2007</date>
.
</paragraph>
<paragraph id="8BA16F7A8A3BF863FF68C67EFE63C195" blockId="20.[151,1437,825,2037]" pageId="20" pageNumber="21">
Comparative material (
<figureCitation id="132573FF8A3BF863FE7DC67EFD81C15D" box="[466,584,1113,1137]" captionStart="FIGURE 14. A D" captionStartId="23.[151,250,1745,1768]" captionTargetBox="[174,1429,191,1717]" captionTargetId="figure@23.[174,1459,191,1719]" captionTargetPageId="23" captionText="FIGURE 14. A D, Hymedesmia (Hymedesmia) prostrata Thiele, 1903 holotype ZMB Por 3160. A, Anisotornotes and chelae; B C, Small acanthostyles; D, Isochelae; E I, Hymedesmia (Hymedesmia) dichela (Hentschel, 1911) syntype ZMB Por 4438. E, Anisotornote and close up of the extremities; F G, Acanthostyles; H, Isochela I; I, Isochela II." httpUri="https://zenodo.org/record/248164/files/figure.png" pageId="20" pageNumber="21">Fig. 14 A</figureCitation>
I):
<taxonomicName id="4C1E14F98A3BF863FDD6C67EFB01C15D" authority="Thiele, 1903" authorityName="Thiele" authorityYear="1903" box="[633,1224,1113,1137]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="20" pageNumber="21" phylum="Porifera" rank="species" species="prostrata" subGenus="Hymedesmia">
<emphasis id="B96AB3688A3BF863FDD6C67EFBE2C15D" box="[633,1067,1113,1137]" italics="true" pageId="20" pageNumber="21">Hymedesmia (Hymedesmia) prostrata</emphasis>
<bibRefCitation id="EF8F128B8A3BF863FB99C67EFB01C15D" author="Thiele" box="[1078,1224,1113,1137]" pageId="20" pageNumber="21" refString="Thiele, J. (1903) Kieselschwamme von Ternate. II. Abhandlungen herausgegeben von der Senckenbergischen naturforschenden Gesellschaft, 25, 933 - 968." type="journal article" year="1903">Thiele, 1903</bibRefCitation>
</taxonomicName>
<typeStatus id="54A5D1D88A3BF863FB7CC67EFAFCC15D" box="[1235,1333,1113,1137]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
material
<collectionCode id="ED0FF7BF8A3BF863FF38C659FF1DC1B9" box="[151,212,1150,1173]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="20" pageNumber="21">ZMB</collectionCode>
Por 3160 (only two spicule slides available);
<taxonomicName id="4C1E14F98A3BF863FD77C65AFAF4C1B9" authority="Hentschel, 1911" authorityName="Hentschel" authorityYear="1911" box="[728,1341,1149,1173]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="20" pageNumber="21" phylum="Porifera" rank="species" species="dichela" subGenus="Hymedesmia">
<emphasis id="B96AB3688A3BF863FD77C65AFBA4C1B9" box="[728,1133,1149,1173]" italics="true" pageId="20" pageNumber="21">Hymedesmia (Hymedesmia) dichela</emphasis>
(
<bibRefCitation id="EF8F128B8A3BF863FBD3C65AFAFCC1B9" author="Hentschel" box="[1148,1333,1149,1173]" pageId="20" pageNumber="21" refString="Hentschel, E. (1911) Tetraxonida. 2. In: Michaelsen, W. &amp; Hartmeyer, R. (Eds.), Die Fauna Sudwest-Australiens. Ergebnisse der Hamburger sudwest-australischen Forschungsreise 1905 Vol. 10, Fischer, Jena, pp. 279 - 393." type="journal article" year="1911">Hentschel, 1911</bibRefCitation>
)
</taxonomicName>
<typeStatus id="54A5D1D88A3BF863FAEAC659FA55C1BA" box="[1349,1436,1150,1174]" pageId="20" pageNumber="21" type="syntype">syntype</typeStatus>
material
<collectionCode id="ED0FF7BF8A3BF863FF56C685FEFFC195" box="[249,310,1186,1209]" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" pageId="20" pageNumber="21">ZMB</collectionCode>
Por 4438.
</paragraph>
<paragraph id="8BA16F7A8A3BF863FF68C6E2FED5C009" blockId="20.[151,1437,825,2037]" pageId="20" pageNumber="21">
Diagnosis.
<taxonomicName id="4C1E14F98A3BF863FEEBC6E2FDABC1F1" box="[324,610,1221,1245]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="20" pageNumber="21" phylum="Porifera" rank="species" species="spinata" status="sp. nov." subGenus="Hymedesmia">
<emphasis id="B96AB3688A3BF863FEEBC6E2FDABC1F1" box="[324,610,1221,1245]" italics="true" pageId="20" pageNumber="21">Hymedesmia (H.) spinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2590E138A3BF863FDC6C6E1FD6DC1F1" box="[617,676,1222,1245]" pageId="20" pageNumber="21" rank="species">n. sp.</taxonomicNameLabel>
is characterised by two
<typeStatus id="54A5D1D88A3BF863FC1FC6E1FC22C1F2" box="[944,1003,1222,1246]" pageId="20" pageNumber="21">types</typeStatus>
of chelae and acanthostyles in a single category, widely variable in length and in the density of spination; smaller chelae have a characteristic apical spine on each ala.
</paragraph>
<paragraph id="8BA16F7A8A3BF863FF68C716FDA8C0F5" blockId="20.[151,1437,825,2037]" pageId="20" pageNumber="21">
Description. Epibiotic on
<taxonomicName id="4C1E14F98A3BF863FE44C715FD4DC065" box="[491,644,1330,1353]" class="Anthozoa" family="Clavulariidae" genus="Carijoa" kingdom="Animalia" order="Alcyonacea" pageId="20" pageNumber="21" phylum="Cnidaria" rank="species" species="riisei">
<emphasis id="B96AB3688A3BF863FE44C715FD4DC065" box="[491,644,1330,1353]" italics="true" pageId="20" pageNumber="21">Carijoa riisei</emphasis>
</taxonomicName>
(
<figureCitation id="132573FF8A3BF863FD3CC716FD2CC065" box="[659,741,1329,1353]" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
AC), which is completely encrusted by the sponge (
<quantity id="4CE6C29F8A3BF863FA91C716FA55C065" box="[1342,1436,1329,1353]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="20" pageNumber="21" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">12 mm</quantity>
thick), leaving only the polyp openings free (
<figureCitation id="132573FF8A3BF863FD02C772FCCDC041" box="[685,772,1365,1389]" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
C). Elevated, scattered oscules are evident. The
<typeStatus id="54A5D1D88A3BF863FA95C772FA55C041" box="[1338,1436,1365,1389]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
consists of several ramified fragments of
<taxonomicName id="4C1E14F98A3BF863FDCCC75DFD74C0BD" box="[611,701,1402,1425]" class="Anthozoa" family="Clavulariidae" genus="Carijoa" kingdom="Animalia" order="Alcyonacea" pageId="20" pageNumber="21" phylum="Cnidaria" rank="species" species="riisei">
<emphasis id="B96AB3688A3BF863FDCCC75DFD74C0BD" box="[611,701,1402,1425]" italics="true" pageId="20" pageNumber="21">C. riisei</emphasis>
</taxonomicName>
up to
<quantity id="4CE6C29F8A3BF863FCADC75EFCF1C0BD" box="[770,824,1401,1425]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="20" pageNumber="21" unit="cm" value="8.0">8 cm</quantity>
long (
<figureCitation id="132573FF8A3BF863FC2FC75EFC1BC0BD" box="[896,978,1401,1425]" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
B); the
<typeStatus id="54A5D1D88A3BF863FB8CC75DFB4AC0BE" box="[1059,1155,1402,1426]" pageId="20" pageNumber="21" type="paratype">paratype</typeStatus>
covers some branches of
<taxonomicName id="4C1E14F98A3BF863FF38C7B9FF38C099" box="[151,241,1438,1461]" class="Anthozoa" family="Clavulariidae" genus="Carijoa" kingdom="Animalia" order="Alcyonacea" pageId="20" pageNumber="21" phylum="Cnidaria" rank="species" species="riisei">
<emphasis id="B96AB3688A3BF863FF38C7B9FF38C099" box="[151,241,1438,1461]" italics="true" pageId="20" pageNumber="21">C. riisei</emphasis>
</taxonomicName>
, up to
<quantity id="4CE6C29F8A3BF863FE90C7BAFE4DC099" box="[319,388,1437,1461]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="20" pageNumber="21" unit="cm" value="10.0">10 cm</quantity>
long. Consistency is soft and the surface smooth; brown
<taxonomicName id="4C1E14F98A3BF863FBA8C7B9FB85C099" box="[1031,1100,1438,1461]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="20" pageNumber="44" phylum="Porifera" rank="species" species="situ">
<emphasis id="B96AB3688A3BF863FBA8C7B9FB85C099" box="[1031,1100,1438,1461]" italics="true" pageId="20" pageNumber="21">in situ</emphasis>
</taxonomicName>
(
<figureCitation id="132573FF8A3BF863FBF3C7BAFB64C099" box="[1116,1197,1437,1461]" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
A), whitish or orange when preserved in ethanol (
<figureCitation id="132573FF8A3BF863FE62C7E6FDD7C0F5" box="[461,542,1473,1497]" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
B, C).
</paragraph>
<paragraph id="8BA16F7A8A3BF863FF68C7C2FF1DC345" blockId="20.[151,1437,825,2037]" pageId="20" pageNumber="21">
Skeleton. Ectosomal skeleton as a thin dermal membrane with scattered tornotes and chelae tangentially disposed (
<figureCitation id="132573FF8A3BF863FEA0C42EFEA1C30D" box="[271,360,1545,1569]" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
D). Choanosomal skeleton hymedesmoid; acanthostyles erect with heads on the substrate; scattered isotornotes and chelae in between. Tracts of tornotes run towards the surface connecting to ectosome (
<figureCitation id="132573FF8A3BF863FAC1C40AFF7DC345" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
E).
</paragraph>
<paragraph id="8BA16F7A8A3BF863FF68C452FCF7C2D9" blockId="20.[151,1437,825,2037]" pageId="20" pageNumber="21">
Spicules. Isotornotes straight with acerate extremities (
<figureCitation id="132573FF8A3BF863FC9AC452FC4EC3A1" box="[821,903,1653,1677]" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
F), 177.5 245
<date id="FFA049BA8A3BF863FB91C451FB46C3A1" box="[1086,1167,1653,1677]" pageId="20" pageNumber="21">x 2 5</date>
μm. Acanthostyles in a single category, very variable in size and in the distribution of spines, 52.5 237.5 x 2.5 10 μm. Smaller acanthostyles strongly spined and with the tips ending in verticils of small spines (
<figureCitation id="132573FF8A3BF863FBFAC49AFB61C3F9" box="[1109,1192,1725,1749]" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
G) are more frequent (
<figureCitation id="132573FF8A3BF863FF30C4C6FF38C3D5" box="[159,241,1761,1785]" captionStart="FIGURE 13" captionStartId="22.[151,250,1206,1229]" captionTargetBox="[193,1413,778,1174]" captionTargetId="figure@22.[151,1436,755,1175]" captionTargetPageId="22" captionText="FIGURE 13. Size / frequency distribution of acanthostyle lengths in Hymedesmia (Hymedesmia) spinata n. sp." httpUri="https://zenodo.org/record/248163/files/figure.png" pageId="20" pageNumber="21">Fig. 13</figureCitation>
); larger acanthostyles with no evident head, few spines and smooth tip (
<figureCitation id="132573FF8A3BF863FBBDC4C6FBA8C3D5" box="[1042,1121,1761,1785]" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
H) are uncommon (
<figureCitation id="132573FF8A3BF863FA94C4C6FA44C3D5" box="[1339,1421,1761,1785]" captionStart="FIGURE 13" captionStartId="22.[151,250,1206,1229]" captionTargetBox="[193,1413,778,1174]" captionTargetId="figure@22.[151,1436,755,1175]" captionTargetPageId="22" captionText="FIGURE 13. Size / frequency distribution of acanthostyle lengths in Hymedesmia (Hymedesmia) spinata n. sp." httpUri="https://zenodo.org/record/248163/files/figure.png" pageId="20" pageNumber="21">Fig. 13</figureCitation>
); numerous intermediate forms with spined and smooth tips occur. Bhattacharya analysis of the acanthostyle lengths shows the presence of three distinct size categories (
<figureCitation id="132573FF8A3BF863FD4CC50EFCFDC26D" box="[739,820,1833,1857]" captionStart="FIGURE 13" captionStartId="22.[151,250,1206,1229]" captionTargetBox="[193,1413,778,1174]" captionTargetId="figure@22.[151,1436,755,1175]" captionTargetPageId="22" captionText="FIGURE 13. Size / frequency distribution of acanthostyle lengths in Hymedesmia (Hymedesmia) spinata n. sp." httpUri="https://zenodo.org/record/248163/files/figure.png" pageId="20" pageNumber="21">Fig. 13</figureCitation>
and
<tableCitation id="C69C5AC18A3BF863FCC4C50EFC7AC26D" box="[875,947,1833,1857]" captionStart="TABLE 5" captionStartId="22.[151,239,316,340]" captionTargetBox="[225,1354,472,642]" captionTargetPageId="22" captionText="TABLE 5. Hymedesmia (Hymedesmia) spinata n. sp. Analysis with Bhattacharyas method indicating the numbers of identified cohorts (groups) of acanthostyles, the mean length for each cohort and the number of spicules present in each cohort. S. d. Standard deviation; S. I. Separation index; separation of the groups is generally unreliable when the SI value is below 2. N. a. not applicable." httpUri="http://table.plazi.org/id/DF613FF28A39F861FF38C31BFE2BC498" pageId="20" pageNumber="21" tableUuid="DF613FF28A39F861FF38C31BFE2BC498">Tab. 5</tableCitation>
). Only the first and smallest category (with a mean of 81.15 μm) is, however, represented with a considerable number of spicules. Two other groups, both with fewer spicules, cannot be considered different size classes (
<figureCitation id="132573FF8A3BF863FC97C556FC40C2A5" box="[824,905,1905,1929]" captionStart="FIGURE 13" captionStartId="22.[151,250,1206,1229]" captionTargetBox="[193,1413,778,1174]" captionTargetId="figure@22.[151,1436,755,1175]" captionTargetPageId="22" captionText="FIGURE 13. Size / frequency distribution of acanthostyle lengths in Hymedesmia (Hymedesmia) spinata n. sp." httpUri="https://zenodo.org/record/248163/files/figure.png" pageId="20" pageNumber="21">Fig. 13</figureCitation>
and
<tableCitation id="C69C5AC18A3BF863FC6DC556FBC3C2A5" box="[962,1034,1905,1929]" captionStart="TABLE 5" captionStartId="22.[151,239,316,340]" captionTargetBox="[225,1354,472,642]" captionTargetPageId="22" captionText="TABLE 5. Hymedesmia (Hymedesmia) spinata n. sp. Analysis with Bhattacharyas method indicating the numbers of identified cohorts (groups) of acanthostyles, the mean length for each cohort and the number of spicules present in each cohort. S. d. Standard deviation; S. I. Separation index; separation of the groups is generally unreliable when the SI value is below 2. N. a. not applicable." httpUri="http://table.plazi.org/id/DF613FF28A39F861FF38C31BFE2BC498" pageId="20" pageNumber="21" tableUuid="DF613FF28A39F861FF38C31BFE2BC498">Tab. 5</tableCitation>
). Arcuate chelae in two shapes and sizes: isochelae I with typical shape (
<figureCitation id="132573FF8A3BF863FDE7C5B2FD57C281" box="[584,670,1941,1965]" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">
Fig.
<date id="FFA049BA8A3BF863FD2FC5B2FD57C281" box="[640,670,1941,1965]" pageId="20" pageNumber="21" value="2012-01">12</date>
</figureCitation>
I), 22.5 27.5 μm; isochelae II, with a very thin curved shaft and inconspicuous alae (about 1/7 of the total length), 12.5 17.5 μm. A spine at the apex of each ala is visible under SEM (
<figureCitation id="132573FF8A3BF863FF4FC5FAFEFBC2D9" box="[224,306,2013,2037]" captionStart="FIGURE 12" captionStartId="21.[151,250,1851,1874]" captionTargetBox="[232,1350,182,1823]" captionTargetId="figure@21.[232,1351,179,1825]" captionTargetPageId="21" captionText="FIGURE 12. Hymedesmia (Hymedesmia) spinata n. sp. A, Sponge covering a colony of Carijoa riisei in situ (holotype); B, Holotype encrusting several fragments of C. riisei; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (F J, paratype)." httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="20" pageNumber="21">Fig. 12</figureCitation>
J). Refer to
<tableCitation id="C69C5AC18A3BF863FE1BC5FAFE33C2D9" box="[436,506,2013,2037]" captionStart="TABLE 6" captionStartId="22.[151,239,1307,1331]" captionTargetBox="[165,1415,1398,1966]" captionTargetPageId="22" captionText="TABLE 6. Spicule sizes in µm of Hymedesmia (Hymedesmia) spinata n. sp. and of the syntype of Hymedesmia (Hymedesmia) dichela (Henschel, 1911)." httpUri="http://table.plazi.org/id/DF613FF28A39F861FF38C73CFD82C07E" pageId="20" pageNumber="21" tableUuid="DF613FF28A39F861FF38C73CFD82C07E">Tab. 6</tableCitation>
for complete measurements.
</paragraph>
<caption id="DF613FF28A3AF862FF38C51CFDDAC2C7" httpUri="https://zenodo.org/record/248162/files/figure.png" pageId="21" pageNumber="22" targetBox="[232,1350,182,1823]" targetPageId="21">
<paragraph id="8BA16F7A8A3AF862FF38C51CFDDAC2C7" blockId="21.[151,1436,1851,2027]" pageId="21" pageNumber="22">
<emphasis id="B96AB3688A3AF862FF38C51CFEE8C27D" bold="true" box="[151,289,1851,1874]" pageId="21" pageNumber="22">FIGURE 12.</emphasis>
<taxonomicName id="4C1E14F98A3AF862FE86C51BFD50C27E" box="[297,665,1852,1874]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="21" pageNumber="22" phylum="Porifera" rank="species" species="spinata" status="sp. nov." subGenus="Hymedesmia">
<emphasis id="B96AB3688A3AF862FE86C51BFD50C27E" box="[297,665,1852,1874]" italics="true" pageId="21" pageNumber="22">Hymedesmia (Hymedesmia) spinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2590E138A3AF862FD0EC51AFD10C27E" box="[673,729,1853,1874]" pageId="21" pageNumber="22" rank="species">n. sp.</taxonomicNameLabel>
A, Sponge covering a colony of
<emphasis id="B96AB3688A3AF862FB9DC51AFACDC27E" box="[1074,1284,1853,1874]" italics="true" pageId="21" pageNumber="22">
<taxonomicName id="4C1E14F98A3AF862FB9DC51AFB74C27E" box="[1074,1213,1853,1874]" class="Anthozoa" family="Clavulariidae" genus="Carijoa" kingdom="Animalia" order="Alcyonacea" pageId="21" pageNumber="22" phylum="Cnidaria" rank="species" species="riisei">Carijoa riisei</taxonomicName>
<taxonomicName id="4C1E14F98A3AF862FB6BC51AFACDC27E" box="[1220,1284,1853,1874]" class="Demospongiae" family="Acarnidae" genus="Iophon" kingdom="Animalia" order="Poecilosclerida" pageId="21" pageNumber="44" phylum="Porifera" rank="species" species="situ">in situ</taxonomicName>
</emphasis>
(holotype); B, Holotype encrusting several fragments of
<taxonomicName id="4C1E14F98A3AF862FDEFC57CFD58C25D" box="[576,657,1883,1905]" class="Anthozoa" family="Clavulariidae" genus="Carijoa" kingdom="Animalia" order="Alcyonacea" pageId="21" pageNumber="22" phylum="Cnidaria" rank="species" species="riisei">
<emphasis id="B96AB3688A3AF862FDEFC57CFD58C25D" box="[576,657,1883,1905]" italics="true" pageId="21" pageNumber="22">C. riisei</emphasis>
</taxonomicName>
; C, Close up of the sponge showing the octocoral completely encrusted with only polyp openings free; D, Dermal membrane with scattered tornotes and chelae; E, Choanosomal skeleton hymedesmioid with acanthostyle heads on the substrate and scattered tornotes and chelae; F, Isotornote with close up of the acerate tip; G, Strongly spined acanthostyle and detail of the head and the spined tip; H, Acanthostyle with smooth extremity; I, Arcuate chela I; J, Arcuate chela II. (FJ, paratype).
</paragraph>
</caption>
<paragraph id="8BA16F7A8A39F861FF68C2B0FDB4C438" blockId="22.[151,1436,151,276]" pageId="22" pageNumber="23">
Remarks. As pointed out by
<bibRefCitation id="EF8F128B8A39F861FDA9C2B0FD35C583" box="[518,764,151,175]" pageId="22" pageNumber="23" refString="Goodwin, C., Jones, J., Neely, K., Brickle, P. (2011) Sponge biodiversity of the Jason Islands and Stanley, Falkland Islands with descriptions of twelve new species. Journal of the Marine Biological Association of the United Kingdom, 91 (2), 275 - 301. http: // dx. doi. org / 10.1017 / S 0025315410001542" type="journal article">Goodwin et al. (2011)</bibRefCitation>
a range in skeletal organisations has been documented from
<taxonomicName id="4C1E14F98A39F861FF38C29EFEEEC5FD" box="[151,295,185,209]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="genus">
<emphasis id="B96AB3688A39F861FF38C29EFEEEC5FD" box="[151,295,185,209]" italics="true" pageId="22" pageNumber="23">Hymedesmia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1E14F98A39F861FECEC29EFE09C5FD" box="[353,448,185,209]" class="Demospongiae" family="Hymedesmiidae" genus="Phorbas" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="genus">
<emphasis id="B96AB3688A39F861FECEC29EFE09C5FD" box="[353,448,185,209]" italics="true" pageId="22" pageNumber="23">Phorbas</emphasis>
</taxonomicName>
, the thinner specimens having a
<taxonomicName id="4C1E14F98A39F861FC95C29EFC02C5FD" box="[826,971,185,209]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="genus">
<emphasis id="B96AB3688A39F861FC95C29EFC02C5FD" box="[826,971,185,209]" italics="true" pageId="22" pageNumber="23">Hymedesmia</emphasis>
</taxonomicName>
-like skeleton and the thicker a
<taxonomicName id="4C1E14F98A39F861FA9BC29EFA5AC5FD" box="[1332,1427,185,209]" class="Demospongiae" family="Hymedesmiidae" genus="Phorbas" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="genus">
<emphasis id="B96AB3688A39F861FA9BC29EFA5AC5FD" box="[1332,1427,185,209]" italics="true" pageId="22" pageNumber="23">Phorbas</emphasis>
</taxonomicName>
- like architecture. Given the presence of acanthostyles erect on the substrate and supporting tornotes, we believe this species should be assigned to
<taxonomicName id="4C1E14F98A39F861FE49C2DBFDBEC438" box="[486,631,252,276]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="genus">
<emphasis id="B96AB3688A39F861FE49C2DBFDBEC438" box="[486,631,252,276]" italics="true" pageId="22" pageNumber="23">Hymedesmia</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF613FF28A39F861FF38C31BFE2BC498" ID-Table-UUID="DF613FF28A39F861FF38C31BFE2BC498" httpUri="http://table.plazi.org/id/DF613FF28A39F861FF38C31BFE2BC498" pageId="22" pageNumber="23" targetBox="[225,1354,472,642]" targetIsTable="true" targetPageId="22">
<paragraph id="8BA16F7A8A39F861FF38C31BFE2BC498" blockId="22.[151,1436,316,436]" pageId="22" pageNumber="23">
<emphasis id="B96AB3688A39F861FF38C31BFEC2C478" bold="true" box="[151,267,316,340]" pageId="22" pageNumber="23">TABLE 5.</emphasis>
<taxonomicName id="4C1E14F98A39F861FEBBC31AFD53C478" box="[276,666,317,340]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="species" species="spinata" status="sp. nov." subGenus="Hymedesmia">
<emphasis id="B96AB3688A39F861FEBBC31AFD53C478" box="[276,666,317,340]" italics="true" pageId="22" pageNumber="23">Hymedesmia (Hymedesmia) spinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2590E138A39F861FD0CC31AFD17C478" box="[675,734,317,340]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
Analysis with Bhattacharyas method indicating the numbers of identified cohorts (groups) of acanthostyles, the mean length for each cohort and the number of spicules present in each cohort. S. d. Standard deviation; S. I. Separation index; separation of the groups is generally unreliable when the SI value is below 2. N.a. not applicable.
</paragraph>
</caption>
<paragraph id="8BA16F7A8A39F861FF4EC3FEFA83C7AD" pageId="22" pageNumber="23">
<table id="F91E9DDA8A390788FF4EC3FFFA83C7AE" box="[225,1354,472,642]" gridcols="5" gridrows="4" pageId="22" pageNumber="23">
<tr id="352E6D388A390788FF4EC3FFFA83C4C2" box="[225,1354,472,494]" gridrow="0" pageId="22" pageNumber="23">
<th id="76FF04448A390788FF4EC3FFFEEBC4C2" box="[225,290,472,494]" gridcol="0" gridrow="0" pageId="22" pageNumber="23">Group</th>
<th id="76FF04448A390788FE78C3FFFDD5C4C2" box="[471,540,472,494]" gridcol="1" gridrow="0" pageId="22" pageNumber="23">Mean</th>
<th id="76FF04448A390788FD45C3FFFCEAC4C2" box="[746,803,472,494]" gridcol="2" gridrow="0" pageId="22" pageNumber="23">s. d.</th>
<th id="76FF04448A390788FC44C3FFFB90C4C2" box="[1003,1113,472,494]" gridcol="3" gridrow="0" pageId="22" pageNumber="23">Population</th>
<th id="76FF04448A390788FABEC3FFFA83C4C2" box="[1297,1354,472,494]" gridcol="4" gridrow="0" pageId="22" pageNumber="23">S.I.</th>
</tr>
<tr id="352E6D388A390788FF4EC02EFA83C733" box="[225,1354,521,543]" gridrow="1" pageId="22" pageNumber="23">
<th id="76FF04448A390788FF4EC02EFEEBC733" box="[225,290,521,543]" gridcol="0" gridrow="1" pageId="22" pageNumber="23">1</th>
<td id="76FF04448A390788FE78C02EFDD5C733" box="[471,540,521,543]" gridcol="1" gridrow="1" pageId="22" pageNumber="23">81.15</td>
<td id="76FF04448A390788FD45C02EFCEAC733" box="[746,803,521,543]" gridcol="2" gridrow="1" pageId="22" pageNumber="23">5.330</td>
<td id="76FF04448A390788FC44C02EFB90C733" box="[1003,1113,521,543]" gridcol="3" gridrow="1" pageId="22" pageNumber="23">99</td>
<td id="76FF04448A390788FABEC02EFA83C733" box="[1297,1354,521,543]" gridcol="4" gridrow="1" pageId="22" pageNumber="23">n.a.</td>
</tr>
<tr id="352E6D388A390788FF4EC01CFA83C77C" box="[225,1354,571,592]" gridrow="2" pageId="22" pageNumber="23">
<th id="76FF04448A390788FF4EC01CFEEBC77C" box="[225,290,571,592]" gridcol="0" gridrow="2" pageId="22" pageNumber="23">2</th>
<td id="76FF04448A390788FE78C01CFDD5C77C" box="[471,540,571,592]" gridcol="1" gridrow="2" pageId="22" pageNumber="23">129.29</td>
<td id="76FF04448A390788FD45C01CFCEAC77C" box="[746,803,571,592]" gridcol="2" gridrow="2" pageId="22" pageNumber="23">4.190</td>
<td id="76FF04448A390788FC44C01CFB90C77C" box="[1003,1113,571,592]" gridcol="3" gridrow="2" pageId="22" pageNumber="23">11</td>
<td id="76FF04448A390788FABEC01CFA83C77C" box="[1297,1354,571,592]" gridcol="4" gridrow="2" pageId="22" pageNumber="23">2.890</td>
</tr>
<tr id="352E6D388A390788FF4EC04BFA83C7AE" box="[225,1354,620,642]" gridrow="3" pageId="22" pageNumber="23">
<th id="76FF04448A390788FF4EC04BFEEBC7AE" box="[225,290,620,642]" gridcol="0" gridrow="3" pageId="22" pageNumber="23">3</th>
<td id="76FF04448A390788FE78C04BFDD5C7AE" box="[471,540,620,642]" gridcol="1" gridrow="3" pageId="22" pageNumber="23">217.50</td>
<td id="76FF04448A390788FD45C04BFCEAC7AE" box="[746,803,620,642]" gridcol="2" gridrow="3" pageId="22" pageNumber="23">4.770</td>
<td id="76FF04448A390788FC44C04BFB90C7AE" box="[1003,1113,620,642]" gridcol="3" gridrow="3" pageId="22" pageNumber="23">4</td>
<td id="76FF04448A390788FABEC04BFA83C7AE" box="[1297,1354,620,642]" gridcol="4" gridrow="3" pageId="22" pageNumber="23">3.190</td>
</tr>
</table>
</paragraph>
<caption id="DF613FF28A39F861FF38C691FB3AC1E1" box="[151,1267,1206,1229]" httpUri="https://zenodo.org/record/248163/files/figure.png" pageId="22" pageNumber="23" targetBox="[193,1413,778,1174]" targetPageId="22">
<paragraph id="8BA16F7A8A39F861FF38C691FB3AC1E1" blockId="22.[151,1267,1206,1229]" box="[151,1267,1206,1229]" pageId="22" pageNumber="23">
<emphasis id="B96AB3688A39F861FF38C691FED6C1E0" bold="true" box="[151,287,1206,1229]" pageId="22" pageNumber="23">FIGURE 13.</emphasis>
Size/frequency distribution of acanthostyle lengths in
<taxonomicName id="4C1E14F98A39F861FCE7C690FB7FC1E1" box="[840,1206,1207,1229]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="species" species="spinata" status="sp. nov." subGenus="Hymedesmia">
<emphasis id="B96AB3688A39F861FCE7C690FB7FC1E1" box="[840,1206,1207,1229]" italics="true" pageId="22" pageNumber="23">Hymedesmia (Hymedesmia) spinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2590E138A39F861FB13C69FFB3AC1E1" box="[1212,1267,1208,1229]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF613FF28A39F861FF38C73CFD82C07E" ID-Table-UUID="DF613FF28A39F861FF38C73CFD82C07E" httpUri="http://table.plazi.org/id/DF613FF28A39F861FF38C73CFD82C07E" pageId="22" pageNumber="23" targetBox="[165,1415,1398,1966]" targetIsTable="true" targetPageId="22">
<paragraph id="8BA16F7A8A39F861FF38C73CFD82C07E" blockId="22.[151,1435,1307,1362]" pageId="22" pageNumber="23">
<emphasis id="B96AB3688A39F861FF38C73CFEC6C01F" bold="true" box="[151,271,1307,1331]" pageId="22" pageNumber="23">TABLE 6.</emphasis>
Spicule sizes in µm of
<taxonomicName id="4C1E14F98A39F861FD83C73CFC7EC01E" box="[556,951,1307,1330]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="species" species="spinata" status="sp. nov." subGenus="Hymedesmia">
<emphasis id="B96AB3688A39F861FD83C73CFC7EC01E" box="[556,951,1307,1330]" italics="true" pageId="22" pageNumber="23">Hymedesmia (Hymedesmia) spinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2590E138A39F861FC6BC73CFBCAC01E" box="[964,1027,1307,1330]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
and of the syntype of
<taxonomicName id="4C1E14F98A39F861FABDC73CFD8EC07E" authority="Henschel, 1911" authorityName="Henschel" authorityYear="1911" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="species" species="dichela" subGenus="Hymedesmia">
<emphasis id="B96AB3688A39F861FABDC73CFE4EC07E" italics="true" pageId="22" pageNumber="23">Hymedesmia (Hymedesmia) dichela</emphasis>
(Henschel, 1911)
</taxonomicName>
.
</paragraph>
</caption>
<paragraph id="8BA16F7A8A39F861FDB4C7A8FAF1C281" pageId="22" pageNumber="23">
<table id="F91E9DDA8A390788FF0AC751FA4EC282" box="[165,1415,1398,1966]" gridcols="4" gridrows="7" pageId="22" pageNumber="23">
<tr id="352E6D388A390788FF0AC751FA4EC091" box="[165,1415,1398,1469]" gridrow="0" pageId="22" pageNumber="23" rowspan-0="1">
<th id="76FF04448A390788FE12C751FD07C091" box="[445,718,1398,1469]" gridcol="1" gridrow="0" pageId="22" pageNumber="23">Tornotes</th>
<th id="76FF04448A390788FD40C751FBC0C091" box="[751,1033,1398,1469]" gridcol="2" gridrow="0" pageId="22" pageNumber="23">Acanthostyles</th>
<th id="76FF04448A390788FB86C751FA4EC091" box="[1065,1415,1398,1469]" gridcol="3" gridrow="0" pageId="22" pageNumber="23">Isochelae I II</th>
</tr>
<tr id="352E6D388A390788FF0AC7FEFA4EC33D" box="[165,1415,1497,1553]" gridrow="1" pageId="22" pageNumber="23">
<th id="76FF04448A390788FF0AC7FEFE55C33D" box="[165,412,1497,1553]" gridcol="0" gridrow="1" pageId="22" pageNumber="23">
<taxonomicName id="4C1E14F98A39F861FF0AC7FDFEF6C0C3" box="[165,319,1498,1519]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="species" species="spinata" status="sp. nov." subGenus="Hymedesmia">
<emphasis id="B96AB3688A39F861FF0AC7FDFEF6C0C3" box="[165,319,1498,1519]" italics="true" pageId="22" pageNumber="23">H. (H.) spinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2590E138A39F861FEEAC7FDFEB2C0C3" box="[325,379,1498,1519]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
MSNG 56262
</th>
<td id="76FF04448A390788FE12C7FEFD07C33D" box="[445,718,1497,1553]" gridcol="1" gridrow="1" pageId="22" pageNumber="23">185 (205.5 ± 7.4) 215 x 2.5</td>
<td id="76FF04448A390788FD40C7FEFBC0C33D" box="[751,1033,1497,1553]" gridcol="2" gridrow="1" pageId="22" pageNumber="23">72.5 (122.6 ± 49) 215 x 2.5 (4.1 ± 1) 5</td>
<td id="76FF04448A390788FB86C7FEFA4EC33D" box="[1065,1415,1497,1553]" gridcol="3" gridrow="1" pageId="22" pageNumber="23">15 (15.2 ± 0.8) 25 17.5</td>
</tr>
<tr id="352E6D388A390788FF0AC40CFA4EC34F" box="[165,1415,1579,1635]" gridrow="2" pageId="22" pageNumber="23">
<th id="76FF04448A390788FF0AC40CFE55C34F" box="[165,412,1579,1635]" gridcol="0" gridrow="2" pageId="22" pageNumber="23">
<taxonomicName id="4C1E14F98A39F861FF0AC40BFEF6C36D" box="[165,319,1580,1601]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="species" species="spinata" status="sp. nov." subGenus="Hymedesmia">
<emphasis id="B96AB3688A39F861FF0AC40BFEF6C36D" box="[165,319,1580,1601]" italics="true" pageId="22" pageNumber="23">H. (H.) spinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2590E138A39F861FEEAC40BFEB2C36D" box="[325,379,1580,1601]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
MSNG 56263
</th>
<td id="76FF04448A390788FE12C40CFD07C34F" box="[445,718,1579,1635]" gridcol="1" gridrow="2" pageId="22" pageNumber="23">192.5 (206.5 ± 8.4) 222.5 x 2 (2.5 ± 0.2) 2.5</td>
<td id="76FF04448A390788FD40C40CFBC0C34F" box="[751,1033,1579,1635]" gridcol="2" gridrow="2" pageId="22" pageNumber="23">52.5 (120.4 ± 53.8) 230 x 2.5 (4.1 ± 1.1) 5</td>
<td id="76FF04448A390788FB86C40CFA4EC34F" box="[1065,1415,1579,1635]" gridcol="3" gridrow="2" pageId="22" pageNumber="23">25 (25.5 ± 15 (16.2 ± 1.3) 1.1) 27.5 17.5</td>
</tr>
<tr id="352E6D388A390788FF0AC459FA4EC39A" box="[165,1415,1662,1718]" gridrow="3" pageId="22" pageNumber="23">
<th id="76FF04448A390788FF0AC459FE55C39A" box="[165,412,1662,1718]" gridcol="0" gridrow="3" pageId="22" pageNumber="23">
<taxonomicName id="4C1E14F98A39F861FF0AC458FEF6C3B8" box="[165,319,1663,1684]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="species" species="spinata" status="sp. nov." subGenus="Hymedesmia">
<emphasis id="B96AB3688A39F861FF0AC458FEF6C3B8" box="[165,319,1663,1684]" italics="true" pageId="22" pageNumber="23">H. (H.) spinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2590E138A39F861FEEAC458FEB2C3B8" box="[325,379,1663,1684]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
Bugor 309
</th>
<td id="76FF04448A390788FE12C459FD07C39A" box="[445,718,1662,1718]" gridcol="1" gridrow="3" pageId="22" pageNumber="23">190 (211.7 ± 16.2) 245 x 2.5 (3 ± 0.8) 5</td>
<td id="76FF04448A390788FD40C459FBC0C39A" box="[751,1033,1662,1718]" gridcol="2" gridrow="3" pageId="22" pageNumber="23">77.5 (122.7 ± 49.9) 237.5 x 3.7 (4.7 ± 0.5) 5</td>
<td id="76FF04448A390788FB86C459FA4EC39A" box="[1065,1415,1662,1718]" gridcol="3" gridrow="3" pageId="22" pageNumber="23">22.5 (25.2 15 ± 1.4) 27.5</td>
</tr>
<tr id="352E6D388A390788FF0AC4F6FA4EC225" box="[165,1415,1745,1801]" gridrow="4" pageId="22" pageNumber="23">
<th id="76FF04448A390788FF0AC4F6FE55C225" box="[165,412,1745,1801]" gridcol="0" gridrow="4" pageId="22" pageNumber="23">
<taxonomicName id="4C1E14F98A39F861FF0AC4F5FEF6C3CB" box="[165,319,1746,1767]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="species" species="spinata" status="sp. nov." subGenus="Hymedesmia">
<emphasis id="B96AB3688A39F861FF0AC4F5FEF6C3CB" box="[165,319,1746,1767]" italics="true" pageId="22" pageNumber="23">H. (H.) spinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2590E138A39F861FEEAC4F5FEB2C3CB" box="[325,379,1746,1767]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
Indo 23
</th>
<td id="76FF04448A390788FE12C4F6FD07C225" box="[445,718,1745,1801]" gridcol="1" gridrow="4" pageId="22" pageNumber="23">180 (199.4 ± 10.2) 220 x 2.5</td>
<td id="76FF04448A390788FD40C4F6FBC0C225" box="[751,1033,1745,1801]" gridcol="2" gridrow="4" pageId="22" pageNumber="23">72.5 (127.3 ± 46.8) 212.5 x 5 (5.8 ± 1.4) 7.5</td>
<td id="76FF04448A390788FB86C4F6FA4EC225" box="[1065,1415,1745,1801]" gridcol="3" gridrow="4" pageId="22" pageNumber="23">20 (25.3 ± 12.5 (14 ± 1.3) 1.9) 27.5 15</td>
</tr>
<tr id="352E6D388A390788FF0AC504FA4EC277" box="[165,1415,1827,1883]" gridrow="5" pageId="22" pageNumber="23">
<th id="76FF04448A390788FF0AC504FE55C277" box="[165,412,1827,1883]" gridcol="0" gridrow="5" pageId="22" pageNumber="23">
<taxonomicName id="4C1E14F98A39F861FF0AC503FEE2C215" box="[165,299,1828,1849]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="species" species="spinata" status="sp. nov." subGenus="Hymedesmia">
<emphasis id="B96AB3688A39F861FF0AC503FEE2C215" box="[165,299,1828,1849]" italics="true" pageId="22" pageNumber="23">H. (H.) spinata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2590E138A39F861FE9FC503FE96C215" box="[304,351,1828,1849]" pageId="22" pageNumber="23" rank="species">n. sp.</taxonomicNameLabel>
Bugor 410 (bis 2)
</th>
<td id="76FF04448A390788FE12C504FD07C277" box="[445,718,1827,1883]" gridcol="1" gridrow="5" pageId="22" pageNumber="23">177.5 (192.2 ± 9.2) 205 x 2.5 (3 ± 0.6) 3.7</td>
<td id="76FF04448A390788FD40C504FBC0C277" box="[751,1033,1827,1883]" gridcol="2" gridrow="5" pageId="22" pageNumber="23">77.5 (138.4 ± 59.3) 237.5 x 5 (6.7 ± 1.9) 10</td>
<td id="76FF04448A390788FB86C504FA4EC277" box="[1065,1415,1827,1883]" gridcol="3" gridrow="5" pageId="22" pageNumber="23">22.5 (25.7 12.5 (15.3 ± 1.4) ± 1.7) 27.5 17.5</td>
</tr>
<tr id="352E6D388A390788FF0AC551FA4EC282" box="[165,1415,1910,1966]" gridrow="6" pageId="22" pageNumber="23">
<th id="76FF04448A390788FF0AC551FE55C282" box="[165,412,1910,1966]" gridcol="0" gridrow="6" pageId="22" pageNumber="23">
<taxonomicName id="4C1E14F98A39F861FF0AC550FEF6C2A0" box="[165,319,1910,1932]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="22" pageNumber="23" phylum="Porifera" rank="species" species="dichela" subGenus="Hymedesmia">
<emphasis id="B96AB3688A39F861FF0AC550FEF6C2A0" box="[165,319,1910,1932]" italics="true" pageId="22" pageNumber="23">H. (H.) dichela</emphasis>
</taxonomicName>
Syntype ZMB Por 4438
</th>
<td id="76FF04448A390788FE12C551FD07C282" box="[445,718,1910,1966]" gridcol="1" gridrow="6" pageId="22" pageNumber="23">150 (189.3 ± 18.9) 212.5 x 2 (3 ± 1.2) 5</td>
<td id="76FF04448A390788FD40C551FBC0C282" box="[751,1033,1910,1966]" gridcol="2" gridrow="6" pageId="22" pageNumber="23">65 (139.2 ± 62.3) 230 x 5 (6.1 ± 1.3) 7.5</td>
<td id="76FF04448A390788FB86C551FA4EC282" box="[1065,1415,1910,1966]" gridcol="3" gridrow="6" pageId="22" pageNumber="23">20 (21.5 ± 10 (12.5 ± 1.5) 1.8) 25 15</td>
</tr>
</table>
</paragraph>
<caption id="DF613FF28A38F860FF38C4F6FBA5C200" httpUri="https://zenodo.org/record/248164/files/figure.png" pageId="23" pageNumber="24" targetBox="[174,1429,191,1717]" targetPageId="23">
<paragraph id="8BA16F7A8A38F860FF38C4F6FBA5C200" blockId="23.[151,1437,1745,1836]" pageId="23" pageNumber="24">
<emphasis id="B96AB3688A38F860FF38C4F6FED6C3CB" bold="true" box="[151,287,1745,1768]" pageId="23" pageNumber="24">FIGURE 14.</emphasis>
AD,
<taxonomicName id="4C1E14F98A38F860FECBC4F5FCA2C3C4" authority="Thiele, 1903" authorityName="Thiele" authorityYear="1903" box="[356,875,1746,1768]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="23" pageNumber="24" phylum="Porifera" rank="species" species="prostrata" subGenus="Hymedesmia">
<emphasis id="B96AB3688A38F860FECBC4F5FD2DC3C4" box="[356,740,1746,1768]" italics="true" pageId="23" pageNumber="24">Hymedesmia (Hymedesmia) prostrata</emphasis>
Thiele, 1903
</taxonomicName>
holotype ZMB Por 3160. A, Anisotornotes and chelae; BC, Small acanthostyles; D, Isochelae; EI,
<taxonomicName id="4C1E14F98A38F860FDC1C4D4FB52C226" authority="Hentschel, 1911" authorityName="Hentschel" authorityYear="1911" box="[622,1179,1779,1802]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="23" pageNumber="24" phylum="Porifera" rank="species" species="dichela" subGenus="Hymedesmia">
<emphasis id="B96AB3688A38F860FDC1C4D4FC17C225" box="[622,990,1779,1801]" italics="true" pageId="23" pageNumber="24">Hymedesmia (Hymedesmia) dichela</emphasis>
(Hentschel, 1911)
</taxonomicName>
syntype ZMB Por 4438. E, Anisotornote and close up of the extremities; FG, Acanthostyles; H, Isochela I; I, Isochela II.
</paragraph>
</caption>
<paragraph id="8BA16F7A8A38F86FFF68C57CFD2BC48D" blockId="23.[151,1436,1883,2012]" lastBlockId="24.[151,1436,150,695]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
Sixteen species of
<taxonomicName id="4C1E14F98A38F860FE36C57BFD1DC258" box="[409,724,1884,1908]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="23" pageNumber="24" phylum="Porifera" rank="genus">
<emphasis id="B96AB3688A38F860FE36C57BFD1DC258" box="[409,724,1884,1908]" italics="true" pageId="23" pageNumber="24">Hymedesmia (Hymedesmia)</emphasis>
</taxonomicName>
have been reported from the Indo-Pacific area (van
<bibRefCitation id="EF8F128B8A38F860FA8EC57CFF1BC2BA" author="Soest" pageId="23" pageNumber="24" refString="Soest, R. W. M van, Boury-Esnault, N., Hooper, J. N. A., Rutzler, K., Voogd, N. J. de, Alvarez, B., Hajdu, E., Pisera, A. B., Vacelet, J., Manconi, R., Schoenberg, C., Janussen, D., Tabachnick, K. R. &amp; Klautau, M. (2011) World Porifera database. Available online from http: // www. marinespecies. org / porifera (accessed 2011)." type="book chapter" year="2011">Soest et al. 2011</bibRefCitation>
), 15 of which are characterised by a single category of chelae. Among these, the following have chelae sizes overlapping with our specimens:
<taxonomicName id="4C1E14F98A38F860FDB4C585FC57C295" authority="Topsent, 1906" authorityName="Topsent" authorityYear="1906" box="[539,926,1953,1977]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="23" pageNumber="24" phylum="Porifera" rank="species" species="lancifera" subGenus="Hymedesmia">
<emphasis id="B96AB3688A38F860FDB4C585FD17C295" box="[539,734,1953,1977]" italics="true" pageId="23" pageNumber="24">H. (H.) lancifera</emphasis>
(
<bibRefCitation id="EF8F128B8A38F860FD5EC586FC5FC295" author="Topsent" box="[753,918,1953,1977]" pageId="23" pageNumber="24" refString="Topsent, E. (1906) Eponges recueillies par M. Ch. Gravier dans la Mer Rouge. Bulletin du Museum national d'Histoire naturelle, 12 (7), 557 - 570." type="journal article" year="1906">Topsent, 1906</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C1E14F98A38F860FC73C585FAF4C295" authority="Thiele, 1903" authorityName="Thiele" authorityYear="1903" box="[988,1341,1953,1977]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="23" pageNumber="24" phylum="Porifera" rank="species" species="prostrata" subGenus="Hymedesmia">
<emphasis id="B96AB3688A38F860FC73C585FB68C295" box="[988,1185,1953,1977]" italics="true" pageId="23" pageNumber="24">H. (H.) prostrata</emphasis>
<bibRefCitation id="EF8F128B8A38F860FB03C586FAF4C295" author="Thiele" box="[1196,1341,1953,1977]" pageId="23" pageNumber="24" refString="Thiele, J. (1903) Kieselschwamme von Ternate. II. Abhandlungen herausgegeben von der Senckenbergischen naturforschenden Gesellschaft, 25, 933 - 968." type="journal article" year="1903">Thiele, 1903</bibRefCitation>
</taxonomicName>
both of which have chelae ~ 18 μm long,
<taxonomicName id="4C1E14F98A38F860FDB7C5E2FBBBC2F7" authority="Bergquist &amp; Fromont, 1988" authorityName="Bergquist &amp; Fromont" authorityYear="1988" box="[536,1138,1987,2012]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="23" pageNumber="24" phylum="Porifera" rank="species" species="microstrongylata" subGenus="Hymedesmia">
<emphasis id="B96AB3688A38F860FDB7C5E2FCF9C2F0" box="[536,816,1988,2012]" italics="true" pageId="23" pageNumber="24">H. (H.) microstrongylata</emphasis>
<bibRefCitation id="EF8F128B8A38F860FC97C5E4FBBBC2F7" author="Bergquist" box="[824,1138,1987,2011]" pageId="23" pageNumber="24" refString="Bergquist, P. R. &amp; Fromont, P. J. (1988) The Marine Fauna of New Zealand: Porifera, Demospongiae, Part 4 (Poecilosclerida). New Zealand Oceanographic Institute Memoir, 96, 1 - 197." type="journal article" year="1988">Bergquist &amp; Fromont, 1988</bibRefCitation>
</taxonomicName>
with chelae of 1830 μm, and
<taxonomicName id="4C1E14F98A37F86FFF65C2BEFD98C583" authority="Thomas, 1970" authorityName="Thomas" authorityYear="1970" box="[202,593,151,176]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="24" pageNumber="25" phylum="Porifera" rank="species" species="stylophora" subGenus="Hymedesmia">
<emphasis id="B96AB3688A37F86FFF65C2BEFE6BC59C" box="[202,418,152,176]" italics="true" pageId="24" pageNumber="25">H. (H.) stylophora</emphasis>
<bibRefCitation id="EF8F128B8A37F86FFE03C2B0FD98C583" author="Thomas" box="[428,593,151,175]" pageId="24" pageNumber="25" refString="Thomas, P. A. (1970) On some deep sea sponges from the Gulf of Mannar, with descriptions of three new species. Journal of the Marine Biological Association of India, 12 (1 / 2), 202 - 209." type="journal article" year="1970">Thomas, 1970</bibRefCitation>
</taxonomicName>
with chelae of 1625 μm.
<taxonomicName id="4C1E14F98A37F86FFC39C2BFFB19C59C" box="[918,1232,152,176]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="24" pageNumber="25" phylum="Porifera" rank="species" species="lancifera" subGenus="Hymedesmia">
<emphasis id="B96AB3688A37F86FFC39C2BFFB19C59C" box="[918,1232,152,176]" italics="true" pageId="24" pageNumber="25">Hymedesmia (H.) lancifera</emphasis>
</taxonomicName>
differs in having centrotylote tornotes with mucronate extremities and smaller acanthostyles (70 170 x 4.2 μm);
<taxonomicName id="4C1E14F98A37F86FFAEBC29BFE9FC5D9" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="24" pageNumber="25" phylum="Porifera" rank="species" species="microstrongylata" subGenus="Hymedesmia">
<emphasis id="B96AB3688A37F86FFAEBC29BFE9FC5D9" italics="true" pageId="24" pageNumber="25">H. (H.) microstrongylata</emphasis>
</taxonomicName>
has strongyloxeas and two categories of acanthostyles, with acanthostyle I up to 470 μm;
<taxonomicName id="4C1E14F98A37F86FFAE3C2F9FEC6C434" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="24" pageNumber="25" phylum="Porifera" rank="species" species="stylophora" subGenus="Hymedesmia">
<emphasis id="B96AB3688A37F86FFAE3C2F9FEC6C434" italics="true" pageId="24" pageNumber="25">H. (H.) stylophora</emphasis>
</taxonomicName>
has smaller acanthostyles (84117 μm) and styles or subtylostyles as dermal spicules.
<taxonomicName id="4C1E14F98A37F86FFB78C327FF37C417" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="24" pageNumber="25" phylum="Porifera" rank="species" species="prostrata" subGenus="Hymedesmia">
<emphasis id="B96AB3688A37F86FFB78C327FF37C417" italics="true" pageId="24" pageNumber="25">Hymedesmia (H.) prostrata</emphasis>
</taxonomicName>
is very close to the new species (
<figureCitation id="132573FF8A37F86FFDDBC305FD36C416" box="[628,767,290,314]" captionStart="FIGURE 14. A D" captionStartId="23.[151,250,1745,1768]" captionTargetBox="[174,1429,191,1717]" captionTargetId="figure@23.[174,1459,191,1719]" captionTargetPageId="23" captionText="FIGURE 14. A D, Hymedesmia (Hymedesmia) prostrata Thiele, 1903 holotype ZMB Por 3160. A, Anisotornotes and chelae; B C, Small acanthostyles; D, Isochelae; E I, Hymedesmia (Hymedesmia) dichela (Hentschel, 1911) syntype ZMB Por 4438. E, Anisotornote and close up of the extremities; F G, Acanthostyles; H, Isochela I; I, Isochela II." httpUri="https://zenodo.org/record/248164/files/figure.png" pageId="24" pageNumber="25">Fig. 14AD</figureCitation>
); the re-examination of the
<typeStatus id="54A5D1D88A37F86FFB9AC305FB5EC416" box="[1077,1175,290,314]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
(slides of the skeleton) confirmed that it differs mainly in having one
<typeStatus id="54A5D1D88A37F86FFD08C361FD1EC472" box="[679,727,326,350]" pageId="24" pageNumber="25">type</typeStatus>
of chelae (
<figureCitation id="132573FF8A37F86FFCF8C362FC77C471" box="[855,958,325,349]" captionStart="FIGURE 14. A D" captionStartId="23.[151,250,1745,1768]" captionTargetBox="[174,1429,191,1717]" captionTargetId="figure@23.[174,1459,191,1719]" captionTargetPageId="23" captionText="FIGURE 14. A D, Hymedesmia (Hymedesmia) prostrata Thiele, 1903 holotype ZMB Por 3160. A, Anisotornotes and chelae; B C, Small acanthostyles; D, Isochelae; E I, Hymedesmia (Hymedesmia) dichela (Hentschel, 1911) syntype ZMB Por 4438. E, Anisotornote and close up of the extremities; F G, Acanthostyles; H, Isochela I; I, Isochela II." httpUri="https://zenodo.org/record/248164/files/figure.png" pageId="24" pageNumber="25">Fig. 14D</figureCitation>
) in a large size range (10 (15.9 ± 3.9) 22.5 μm), two
<typeStatus id="54A5D1D88A37F86FFEEFC34FFEB2C4AC" box="[320,379,360,384]" pageId="24" pageNumber="25">types</typeStatus>
of acanthostyles of similar shape (187240 μm and 80117.5 μm, respectively;
<figureCitation id="132573FF8A37F86FFAA2C340FA5DC453" box="[1293,1428,359,383]" captionStart="FIGURE 14. A D" captionStartId="23.[151,250,1745,1768]" captionTargetBox="[174,1429,191,1717]" captionTargetId="figure@23.[174,1459,191,1719]" captionTargetPageId="23" captionText="FIGURE 14. A D, Hymedesmia (Hymedesmia) prostrata Thiele, 1903 holotype ZMB Por 3160. A, Anisotornotes and chelae; B C, Small acanthostyles; D, Isochelae; E I, Hymedesmia (Hymedesmia) dichela (Hentschel, 1911) syntype ZMB Por 4438. E, Anisotornote and close up of the extremities; F G, Acanthostyles; H, Isochela I; I, Isochela II." httpUri="https://zenodo.org/record/248164/files/figure.png" pageId="24" pageNumber="25">Fig. 14B, C</figureCitation>
) and anisotornotes (
<figureCitation id="132573FF8A37F86FFEC4C3ADFE18C48E" box="[363,465,394,418]" captionStart="FIGURE 14. A D" captionStartId="23.[151,250,1745,1768]" captionTargetBox="[174,1429,191,1717]" captionTargetId="figure@23.[174,1459,191,1719]" captionTargetPageId="23" captionText="FIGURE 14. A D, Hymedesmia (Hymedesmia) prostrata Thiele, 1903 holotype ZMB Por 3160. A, Anisotornotes and chelae; B C, Small acanthostyles; D, Isochelae; E I, Hymedesmia (Hymedesmia) dichela (Hentschel, 1911) syntype ZMB Por 4438. E, Anisotornote and close up of the extremities; F G, Acanthostyles; H, Isochela I; I, Isochela II." httpUri="https://zenodo.org/record/248164/files/figure.png" pageId="24" pageNumber="25">Fig. 14A</figureCitation>
) (about 187.5205 μm).
</paragraph>
<paragraph id="8BA16F7A8A37F86FFF68C389FAE9C7B9" blockId="24.[151,1436,150,695]" pageId="24" pageNumber="25">
Among the species of
<taxonomicName id="4C1E14F98A37F86FFE6FC38AFD33C4E9" box="[448,762,429,453]" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="24" pageNumber="25" phylum="Porifera" rank="genus">
<emphasis id="B96AB3688A37F86FFE6FC38AFD33C4E9" box="[448,762,429,453]" italics="true" pageId="24" pageNumber="25">Hymedesmia (Hymedesmia)</emphasis>
</taxonomicName>
from the Indo-Pacific area, only
<taxonomicName id="4C1E14F98A37F86FFBC3C389FF1EC4CB" authority="Hentschel, 1911" authorityName="Hentschel" authorityYear="1911" class="Demospongiae" family="Hymedesmiidae" genus="Hymedesmia" kingdom="Animalia" order="Poecilosclerida" pageId="24" pageNumber="25" phylum="Porifera" rank="species" species="dichela" subGenus="Hymedesmia">
<emphasis id="B96AB3688A37F86FFBC3C389FADCC4E9" box="[1132,1301,429,453]" italics="true" pageId="24" pageNumber="25">H. (H.) dichela</emphasis>
(
<bibRefCitation id="EF8F128B8A37F86FFA8CC38AFF06C4CB" author="Hentschel" pageId="24" pageNumber="25" refString="Hentschel, E. (1911) Tetraxonida. 2. In: Michaelsen, W. &amp; Hartmeyer, R. (Eds.), Die Fauna Sudwest-Australiens. Ergebnisse der Hamburger sudwest-australischen Forschungsreise 1905 Vol. 10, Fischer, Jena, pp. 279 - 393." type="journal article" year="1911">Hentschel, 1911</bibRefCitation>
)
</taxonomicName>
possesses two categories of chelae; moreover it possesses acanthostyles and anisotornotes similar in shape and size to the new species (
<figureCitation id="132573FF8A37F86FFE74C3D5FDE5C726" box="[475,556,498,522]" captionStart="FIGURE 14. A D" captionStartId="23.[151,250,1745,1768]" captionTargetBox="[174,1429,191,1717]" captionTargetId="figure@23.[174,1459,191,1719]" captionTargetPageId="23" captionText="FIGURE 14. A D, Hymedesmia (Hymedesmia) prostrata Thiele, 1903 holotype ZMB Por 3160. A, Anisotornotes and chelae; B C, Small acanthostyles; D, Isochelae; E I, Hymedesmia (Hymedesmia) dichela (Hentschel, 1911) syntype ZMB Por 4438. E, Anisotornote and close up of the extremities; F G, Acanthostyles; H, Isochela I; I, Isochela II." httpUri="https://zenodo.org/record/248164/files/figure.png" pageId="24" pageNumber="25">Fig. 14</figureCitation>
EI). Nevertheless, the examination of the
<typeStatus id="54A5D1D88A37F86FFBBCC3D4FBA3C727" box="[1043,1130,499,523]" pageId="24" pageNumber="25" type="syntype">syntype</typeStatus>
reveals that in Hentschels species the tornotes have different extremities (anisotornotes, 150 (189.3 ± 18.9) 212.5
<date id="FFA049BA8A37F86FFB35C031FB74C701" box="[1178,1213,533,557]" pageId="24" pageNumber="25">x 2</date>
(3 ± 1.2) 5 μm) (
<figureCitation id="132573FF8A37F86FFF30C010FF39C763" box="[159,240,567,591]" captionStart="FIGURE 14. A D" captionStartId="23.[151,250,1745,1768]" captionTargetBox="[174,1429,191,1717]" captionTargetId="figure@23.[174,1459,191,1719]" captionTargetPageId="23" captionText="FIGURE 14. A D, Hymedesmia (Hymedesmia) prostrata Thiele, 1903 holotype ZMB Por 3160. A, Anisotornotes and chelae; B C, Small acanthostyles; D, Isochelae; E I, Hymedesmia (Hymedesmia) dichela (Hentschel, 1911) syntype ZMB Por 4438. E, Anisotornote and close up of the extremities; F G, Acanthostyles; H, Isochela I; I, Isochela II." httpUri="https://zenodo.org/record/248164/files/figure.png" pageId="24" pageNumber="25">Fig. 14</figureCitation>
E), acanthostyles (65 (139.2 ± 62.3) 230
<date id="FFA049BA8A37F86FFD72C01FFCC9C763" box="[733,768,567,591]" pageId="24" pageNumber="25">x 5</date>
(6.1 ± 1.3) 7.5 μm) always have smooth tips, not ending in verticils of spines (
<figureCitation id="132573FF8A37F86FFE3FC07DFE2AC75E" box="[400,483,602,626]" captionStart="FIGURE 14. A D" captionStartId="23.[151,250,1745,1768]" captionTargetBox="[174,1429,191,1717]" captionTargetId="figure@23.[174,1459,191,1719]" captionTargetPageId="23" captionText="FIGURE 14. A D, Hymedesmia (Hymedesmia) prostrata Thiele, 1903 holotype ZMB Por 3160. A, Anisotornotes and chelae; B C, Small acanthostyles; D, Isochelae; E I, Hymedesmia (Hymedesmia) dichela (Hentschel, 1911) syntype ZMB Por 4438. E, Anisotornote and close up of the extremities; F G, Acanthostyles; H, Isochela I; I, Isochela II." httpUri="https://zenodo.org/record/248164/files/figure.png" pageId="24" pageNumber="25">Fig. 14</figureCitation>
F), and chelae I (20 (21.5 ± 1.8) 25 μm) and II (10 (12.5 ± 1.5) 15 μm) are similar in shape (
<figureCitation id="132573FF8A37F86FFEF8C05AFE61C7B9" box="[343,424,637,661]" captionStart="FIGURE 14. A D" captionStartId="23.[151,250,1745,1768]" captionTargetBox="[174,1429,191,1717]" captionTargetId="figure@23.[174,1459,191,1719]" captionTargetPageId="23" captionText="FIGURE 14. A D, Hymedesmia (Hymedesmia) prostrata Thiele, 1903 holotype ZMB Por 3160. A, Anisotornotes and chelae; B C, Small acanthostyles; D, Isochelae; E I, Hymedesmia (Hymedesmia) dichela (Hentschel, 1911) syntype ZMB Por 4438. E, Anisotornote and close up of the extremities; F G, Acanthostyles; H, Isochela I; I, Isochela II." httpUri="https://zenodo.org/record/248164/files/figure.png" pageId="24" pageNumber="25">Fig. 14</figureCitation>
H, I); in particular chelae II, with SEM observations, do not have apical spines.
</paragraph>
<paragraph id="8BA16F7A8A37F86FFF68C0B8FB82C79B" blockId="24.[151,1436,150,695]" box="[199,1099,671,695]" pageId="24" pageNumber="25">Etymology. Named after the presence of an apical spine on each ala of chelae II.</paragraph>
</subSubSection>
</treatment>
</document>