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<document id="AC60246A9FECE83D144F3F1E2DC3472C" ID-DOI="10.1649/0010-065X-67.3.304" ID-ISSN="1938-4394" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" checkinTime="1630003085403" checkinUser="diego" docAuthor="Blanco, Jaime Ortega-, Chatzimanolis, Stylianos, Singh, Hukam &amp; Engel, Michael S." docDate="2013" docId="03B787FD8B3BFF83FD38FC6761FC90C1" docLanguage="en" docName="0010-065X-67.3.304.pdf" docOrigin="The Coleopterists Bulletin 67 (3)" docSource="http://dx.doi.org/10.1649/0010-065x-67.3.304" docStyle="DocumentStyle:0305A7CAC51C960DDA50A7292A4D3620.8:TheColeopBull.2010-2016.journal_article.0cover" docStyleId="0305A7CAC51C960DDA50A7292A4D3620" docStyleName="TheColeopBull.2010-2016.journal_article.0cover" docStyleVersion="8" docTitle="Paleosorius cambayensis Ortega-Blanco, Chatzimanolis, and Engel, new species" docType="treatment" docVersion="3" lastPageNumber="306" masterDocId="FF8EFF858B3AFF81FFDFFFFE62169461" masterDocTitle="The Oldest Fossil Of The Subfamily Osoriinae (Coleoptera: Staphylinidae), From Eocene Cambay Amber (India)" masterLastPageNumber="308" masterPageNumber="304" pageNumber="305" updateTime="1699135549677" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="FA5B7948456D39E476B0838BC56C6E46">The Oldest Fossil Of The Subfamily Osoriinae (Coleoptera: Staphylinidae), From Eocene Cambay Amber (India)</mods:title>
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<mods:name id="52D2464B399778F81576637998240AA7" type="personal">
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<mods:roleTerm id="9CABAEE1E2BC817611459B2FA4AF47AB">Author</mods:roleTerm>
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<mods:namePart id="E3044BDA426589A6380B17827003AC8C">Blanco, Jaime Ortega-</mods:namePart>
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<mods:namePart id="EA6D3F9517C3C55692022F5EA2D431C5">Chatzimanolis, Stylianos</mods:namePart>
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<mods:namePart id="6BE036D4F33D4952D2B6107F8793608B">Singh, Hukam</mods:namePart>
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<mods:namePart id="7A1ADA53436C13693EFE21CB1BEDF474">Engel, Michael S.</mods:namePart>
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<mods:title id="0214E530E0A303B057D4B5C8CE013E07">The Coleopterists Bulletin</mods:title>
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<mods:date id="409B6C469E09DC932F0BC7AE4CBB4BC5">2013</mods:date>
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<mods:number id="1AD199EDAACFEE4F1CBB8DC7DDD83FAA">2013-09-30</mods:number>
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<mods:number id="A3C1C02A775B1262D423320B42944118">67</mods:number>
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<mods:classification id="7715357872AB53077F1360AFD9463736">journal article</mods:classification>
<mods:identifier id="EB0186BE93344F46A6DB13EAA33234A1" type="DOI">10.1649/0010-065X-67.3.304</mods:identifier>
<mods:identifier id="9DAB508E06F2753EF7CEB64D982E1511" type="ISSN">1938-4394</mods:identifier>
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<treatment id="03B787FD8B3BFF83FD38FC6761FC90C1" ID-DOI="http://doi.org/10.5281/zenodo.5459133" ID-Zenodo-Dep="5459133" LSID="urn:lsid:plazi:treatment:03B787FD8B3BFF83FD38FC6761FC90C1" httpUri="http://treatment.plazi.org/id/03B787FD8B3BFF83FD38FC6761FC90C1" lastPageId="2" lastPageNumber="306" pageId="1" pageNumber="305">
<subSubSection id="C30465608B3BFF80FD38FC6761FD9785" pageId="1" pageNumber="305" type="nomenclature">
<paragraph id="8BA136EB8B3BFF80FD38FC6761FD9785" blockId="1.[743,1157,921,997]" pageId="1" pageNumber="305">
<emphasis id="B96AEAF98B3BFF80FD38FC67666B97AB" bold="true" pageId="1" pageNumber="305">
<taxonomicName id="4C1E4D688B3BFF80FD38FC6761ED97AB" ID-CoL="4C47F" authority="Ortega-Blanco, Chatzimanolis, and Engel" authorityName="Ortega-Blanco, Chatzimanolis, and Engel" class="Insecta" family="Staphylinidae" genus="Paleosorius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="305" phylum="Arthropoda" rank="species" species="cambayensis" status="sp. nov.">
<emphasis id="B96AEAF98B3BFF80FD38FC6761C897CE" bold="true" box="[743,990,921,943]" italics="true" pageId="1" pageNumber="305">Paleosorius cambayensis</emphasis>
Ortega-Blanco, Chatzimanolis, and Engel
</taxonomicName>
,
<taxonomicNameLabel id="A25957828B3BFF80FBD7FC4B666B97AB" box="[1032,1149,949,970]" pageId="1" pageNumber="305" rank="species">new species</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13252A6E8B3BFF80FC58FC3161F39785" box="[903,997,975,997]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="2.[175,207,775,794]" captionStartId-1="2.[175,207,1686,1705]" captionTargetBox-0="[151,1171,194,752]" captionTargetBox-1="[185,1156,1265,1663]" captionTargetId-0="figure-187@2.[151,1171,194,752]" captionTargetId-1="figure-168@2.[151,1171,1238,1695]" captionTargetPageId-0="2" captionTargetPageId-1="2" captionText-0="Fig. 1. Photomicrograph of holotype (SEMC-F000134) of Paleosorius cambayensis in Early Eocene (Ypresian) amber from Gujarat, India." captionText-1="Fig. 2. Illustration of holotype (SEMC-F000134) of Paleosorius cambayensis in Early Eocene (Ypresian) amber from Gujarat, India." pageId="1" pageNumber="305">Figs. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30465608B3BFF80FD04FC046660906E" box="[731,1142,1018,1039]" pageId="1" pageNumber="305" type="diagnosis">
<paragraph id="8BA136EB8B3BFF80FD04FC046660906E" blockId="1.[707,1194,1018,1729]" box="[731,1142,1018,1039]" pageId="1" pageNumber="305">
<emphasis id="B96AEAF98B3BFF80FD04FC046152906E" bold="true" box="[731,836,1018,1039]" pageId="1" pageNumber="305">Diagnosis.</emphasis>
As for the genus (
<emphasis id="B96AEAF98B3BFF80FBDCFC04667E906E" box="[1027,1128,1018,1039]" italics="true" pageId="1" pageNumber="305">vide supra</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C30465608B3BFF83FD04FBEA612297AD" lastPageId="2" lastPageNumber="306" pageId="1" pageNumber="305" type="description">
<paragraph id="8BA136EB8B3BFF83FD04FBEA612297AD" blockId="1.[707,1194,1018,1729]" lastBlockId="2.[685,1172,871,1184]" lastPageId="2" lastPageNumber="306" pageId="1" pageNumber="305">
<emphasis id="B96AEAF98B3BFF80FD04FBEA61419048" bold="true" box="[731,855,1044,1065]" pageId="1" pageNumber="305">Description.</emphasis>
As for the genus, with the following additions: total body length
<quantity id="4CE69B0E8B3BFF80FBC6FBD166649025" box="[1049,1138,1071,1092]" metricMagnitude="-3" metricUnit="m" metricValue="1.81" pageId="1" pageNumber="305" unit="mm" value="1.81">1.81 mm</quantity>
; head length
<quantity id="4CE69B0E8B3BFF80FCD9FBB76176903F" box="[774,864,1097,1118]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="305" unit="mm" value="0.26">0.26 mm</quantity>
; pronotal length
<quantity id="4CE69B0E8B3BFF80FBD8FBB76677903F" box="[1031,1121,1097,1118]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="305" unit="mm" value="0.33">0.33 mm</quantity>
; elytral length
<quantity id="4CE69B0E8B3BFF80FCDAFB9A61489018" box="[773,862,1124,1145]" metricMagnitude="-4" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="305" unit="mm" value="0.37">0.37 mm</quantity>
; head prognathous, slightly longer than wide, as wide as pronotum, with postclypeus highly angled with respect to epicranium; dorsalmost corners of postclypeus with 2 reddish dots symmetrically placed between compound eyes, reddish dots clearly defined, symmetrical, slightly anterior to compound eyes (could correspond to tentorial pits but not flat or raised, perhaps taphonomically produced); compound eyes small, separated by distance equivalent to 3 times their diameter; apparently 1 elongate seta arising dorsally close to each compound eye; ocelli absent (though corresponding area damaged); anteroclypeus and mouthparts not clearly visible; antenna filiform, not clavate, with 11 antennomeres; antennal insertion lateral, anterior to compound eyes; antennomeres increasing in size progressively; antennomeres 911 distinctly longer and wider than previous antennomeres, 11 tapered. Pronotum elongate, with rimmed anterior border but no constricted edges; mesoscutellum obscured. Elytra with trace of epipleural ridge, deformed but about as long as pronotum and without distinct sculpture. Hind wings extended, slightly bent, without clearly visible venation, apparently without pterostigma (3 or 4 thin veins at most: C, R, M
<subScript id="179A34AE8B38FF83FF73FC5862A397D5" attach="left" box="[172,181,934,948]" fontSize="6" pageId="2" pageNumber="306">4</subScript>
+Cu, 2A) or discernible folding pattern (at least as preserved), and with moderately long setae on posterior edge. Coxae globose or sub-globose (definitely not elongate); femora arched, laterally moderately flattened, longitudinally striate; protibia without ctenidium; tarsi pentamerous, with short tarsomeres except distalmost tarsomere (5) which is almost as long as combined length of preceding tarsomeres; tarsal claws simple. Abdomen elongate, with 7 terga visible dorsally; each tergum approximately 2 times wider than long, except VII (almost as long as wide) and VIII; tergum VIII distinctly reduced and rounded, as long as wide.
</paragraph>
</subSubSection>
<caption id="DF6166638B38FF83FF70FCF963969753" pageId="2" pageNumber="306" startId="2.[175,207,775,794]" targetBox="[151,1171,194,752]" targetPageId="2">
<paragraph id="8BA136EB8B38FF83FF70FCF963969753" blockId="2.[151,1172,775,818]" pageId="2" pageNumber="306">
<emphasis id="B96AEAF98B38FF83FF70FCF962FA977B" bold="true" box="[175,236,775,794]" pageId="2" pageNumber="306">Fig. 1.</emphasis>
Photomicrograph of holotype (SEMC-F000134) of
<taxonomicName id="4C1E4D688B38FF83FD1AFCF961889778" box="[709,926,775,793]" class="Insecta" family="Staphylinidae" genus="Paleosorius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="306" phylum="Arthropoda" rank="species" species="cambayensis">
<emphasis id="B96AEAF98B38FF83FD1AFCF961889778" box="[709,926,775,793]" italics="true" pageId="2" pageNumber="306">Paleosorius cambayensis</emphasis>
</taxonomicName>
in Early Eocene (Ypresian) amber from Gujarat, India.
</paragraph>
</caption>
<subSubSection id="C30465608B38FF83FD1AFC2F665F9030" pageId="2" pageNumber="306" type="materials_examined">
<paragraph id="8BA136EB8B38FF83FD1AFC2F665F9030" blockId="2.[685,1172,871,1184]" pageId="2" pageNumber="306">
<materialsCitation id="3B763CB68B38FF83FD1AFC2F665F9030" collectorName="M. S. Engel &amp; Lucknow" country="India" county="Birbal Sahni Institute of Paleobotany" latitude="21.22389" location="Eocene" longLatPrecision="21" longitude="73.075554" municipality="Basin" pageId="2" pageNumber="306" specimenCode="SEMC-F000134" specimenCount="1" typeStatus="holotype">
<emphasis id="B96AEAF98B38FF83FD1AFC2F613F9787" bold="true" box="[709,809,977,998]" pageId="2" pageNumber="306">
<typeStatus id="54A588498B38FF83FD1AFC2F61329787" box="[709,804,977,998]" pageId="2" pageNumber="306" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCode id="DBB89E908B38FF83FCE9FC2F61C19787" box="[822,983,977,998]" collectionCode="SEMC-F" pageId="2" pageNumber="306">SEMC-F000134</specimenCode>
;
<location id="8EC160308B38FF83FC3CFC2F663F9787" LSID="urn:lsid:plazi:treatment:03B787FD8B3BFF83FD38FC6761FC90C1:8EC160308B38FF83FC3CFC2F663F9787" box="[995,1065,977,998]" country="India" county="Birbal Sahni Institute of Paleobotany" latitude="21.22389" longLatPrecision="21" longitude="73.075554" municipality="Basin" name="Eocene" pageId="2" pageNumber="306">Eocene</location>
(
<location id="8EC160308B38FF83FBE8FC2F669B9787" LSID="urn:lsid:plazi:treatment:03B787FD8B3BFF83FD38FC6761FC90C1:8EC160308B38FF83FBE8FC2F669B9787" box="[1079,1165,977,998]" country="India" county="Birbal Sahni Institute of Paleobotany" latitude="21.22389" longLatPrecision="21" longitude="73.075554" municipality="Basin" name="Ypresian" pageId="2" pageNumber="306">Ypresian</location>
) amber,
<location id="8EC160308B38FF83FD22FC1261629060" LSID="urn:lsid:plazi:treatment:03B787FD8B3BFF83FD38FC6761FC90C1:8EC160308B38FF83FD22FC1261629060" box="[765,884,1004,1025]" country="India" county="Birbal Sahni Institute of Paleobotany" latitude="21.22389" longLatPrecision="21" longitude="73.075554" municipality="Basin" name="Tadkeshwar" pageId="2" pageNumber="306">Tadkeshwar</location>
mine (Surat dist.,
<collectingCountry id="F309767B8B38FF83FB9DFC1266859060" box="[1090,1171,1004,1025]" name="India" pageId="2" pageNumber="306">Cambay</collectingCountry>
<collectingMunicipality id="6BC5AC918B38FF83FD72FBF960F1907D" box="[685,743,1031,1052]" pageId="2" pageNumber="306">Basin</collectingMunicipality>
)
<geoCoordinate id="EE2A502C8B38FF83FD24FBF96164907D" box="[763,882,1030,1052]" degrees="21" direction="north" minutes="13" orientation="latitude" pageId="2" pageNumber="306" precision="15" seconds="26" value="21.22389">N21°1326″</geoCoordinate>
N,
<geoCoordinate id="EE2A502C8B38FF83FC47FBF9661A907D" box="[920,1036,1030,1052]" degrees="73" direction="east" minutes="04" orientation="longitude" pageId="2" pageNumber="306" precision="15" seconds="32" value="73.075554">E73°0432″</geoCoordinate>
, coll.
<collectorName id="26EB533D8B38FF83FBBCFBF960F19057" pageId="2" pageNumber="306">M.S. Engel</collectorName>
, 2011; to be deposited in the
<collectingCounty id="62C04E678B38FF83FBC6FBDF618C9030" pageId="2" pageNumber="306">Birbal Sahni Institute of Paleobotany</collectingCounty>
,
<collectorName id="26EB533D8B38FF83FC77FBC266159030" box="[936,1027,1084,1105]" pageId="2" pageNumber="306">Lucknow</collectorName>
,
<collectingCountry id="F309767B8B38FF83FBCEFBC266539030" box="[1041,1093,1084,1105]" name="India" pageId="2" pageNumber="306">India</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30465608B38FF83FD1AFBA861FC90C1" pageId="2" pageNumber="306" type="etymology">
<paragraph id="8BA136EB8B38FF83FD1AFBA861FC90C1" blockId="2.[685,1172,871,1184]" pageId="2" pageNumber="306">
<emphasis id="B96AEAF98B38FF83FD1AFBA8612E900A" bold="true" box="[709,824,1110,1131]" pageId="2" pageNumber="306">Etymology.</emphasis>
The specific epithet refers to the
<collectingCountry id="F309767B8B38FF83FD72FB8F60EA90E7" box="[685,764,1137,1158]" name="India" pageId="2" pageNumber="306">Cambay</collectingCountry>
Basin in
<collectingRegion id="49DAF8098B38FF83FC81FB8F61B190E7" box="[862,935,1137,1158]" country="India" name="Gujarat" pageId="2" pageNumber="306">Gujarat</collectingRegion>
,
<collectingCountry id="F309767B8B38FF83FC6CFB8F61F090E7" box="[947,998,1137,1158]" name="India" pageId="2" pageNumber="306">India</collectingCountry>
, where the amber bearing the specimen originated.
</paragraph>
</subSubSection>
</treatment>
</document>