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<document id="63A85DF4A2D4629003134FAA38EFA32B" ID-CLB-Dataset="163377" ID-DOI="10.11646/zootaxa.5195.3.3" ID-GBIF-Dataset="fd2040c7-2e4e-4c36-87ce-85efe9f45003" ID-ISSN="1175-5326" ID-Zenodo-Dep="7187839" ID-ZooBank="A9963C93-290B-44BA-BBDB-0C4B0299FDB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665554369870" checkinUser="plazi" docAuthor="Nakahara, Shinichi, Zilli, Alberto, Calhoun, John V., Espeland, Marianne, Padrón, Pablo Sebastián &amp; Grishin, Nick V." docDate="2022" docId="03B787FAA55F717D4E85FD56BCFF9F34" docLanguage="en" docName="zootaxa.5195.3.3.pdf" docOrigin="Zootaxa 5195 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hesperia lyrcea sensu Plotz 1882" docType="treatment" docVersion="7" lastPageNumber="250" masterDocId="FF8EFF82A55671774E12FFE5BE3A9E29" masterDocTitle="Resolving two centuries of mistaken identity: Reinterpretation of Papilio marcus Fabricius, 1787 (Lepidoptera: Nymphalidae, Hesperiidae)" masterLastPageNumber="255" masterPageNumber="241" pageNumber="250" updateTime="1699407275020" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A4DDDE776033507481E4C0F4A9F2AF83">Resolving two centuries of mistaken identity: Reinterpretation of Papilio marcus Fabricius, 1787 (Lepidoptera: Nymphalidae, Hesperiidae)</mods:title>
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<mods:name id="BDE18FCE52F49C42E4771B54B68CB37E" type="personal">
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<mods:namePart id="C89D87B66021FDD8850AFAC8C3EEA331">Nakahara, Shinichi</mods:namePart>
<mods:affiliation id="E190271AA6221D9E5A79BFC9FD588D83">Department of Organismic and Evolutionary Biology, Museum of Comparative Zoology, Harvard University, 26 Oxford St. Cambridge, MA 02318, USA</mods:affiliation>
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<mods:namePart id="677C9027CC8EB6F06A8C2832A681E08F">Zilli, Alberto</mods:namePart>
<mods:affiliation id="212CA9453EB651BBD97A0315CA32C2D2">Insects Division, Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
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<mods:namePart id="3F9FBDF42E32853BC586E4666AD3DA9C">Calhoun, John V.</mods:namePart>
<mods:affiliation id="8AA0BBC8736988BFD2B36844758E41BB">McGuire Center for Lepidoptera and Biodiversity, Florida Museum of Natural History, University of Florida, Gainesville, FL 32611, USA</mods:affiliation>
</mods:name>
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<mods:role id="1141D8F2E1FC38407A9EC57E4C360FC5">
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</mods:role>
<mods:namePart id="BE0D4355507FBEA69AC9EF4E8EB6407A">Espeland, Marianne</mods:namePart>
<mods:affiliation id="9023F588E3BB8A0D04FC9BF852718433">Leibniz Institute for the Analysis of Biodiversity Change, Zoological Research Museum Alexander Koenig, Adenauerallee 127 53113 Bonn, Germany</mods:affiliation>
</mods:name>
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<mods:role id="9A49371FF0376E3EC174B190BE5565F2">
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<mods:namePart id="6E2EBA81B534DE679918B0E4648B6FD4">Padrón, Pablo Sebastián</mods:namePart>
<mods:affiliation id="3BE56332EAB52ED7DFDFE5200894C4FF">Laboratorio de Entomología, Museo de Zoología, Universidad del Azuay, Cuenca, Ecuador</mods:affiliation>
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<mods:namePart id="608C8DA018F2F253C645E8F465A947F4">Grishin, Nick V.</mods:namePart>
<mods:affiliation id="C43EE2F7FAD097DFF711D369B95D2ED8">Howard Hughes Medical Institute, Departments of Biophysics and Biochemistry, University of Texas Southwestern Medical Center, 5323 Harry Hines Blvd., Dallas, TX 75390 - 9050, USA</mods:affiliation>
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<mods:title id="B1CA81FCA84FBCFF777DE9E7959846D0">Zootaxa</mods:title>
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<mods:date id="CEBA77CEC48C43575A3974CEE7E7181E">2022</mods:date>
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<treatment id="03B787FAA55F717D4E85FD56BCFF9F34" ID-DOI="http://doi.org/10.5281/zenodo.7190634" ID-GBIF-Taxon="202596685" ID-Zenodo-Dep="7190634" LSID="urn:lsid:plazi:treatment:03B787FAA55F717D4E85FD56BCFF9F34" httpUri="http://treatment.plazi.org/id/03B787FAA55F717D4E85FD56BCFF9F34" lastPageId="10" lastPageNumber="250" pageId="9" pageNumber="250">
<subSubSection id="C3046567A55F717E4E85FD56BFDC9CE5" box="[151,486,690,717]" pageId="9" pageNumber="250" type="nomenclature">
<paragraph id="8BA136ECA55F717E4E85FD56BFDC9CE5" blockId="9.[151,486,690,717]" box="[151,486,690,717]" pageId="9" pageNumber="250">
<heading id="D0E98180A55F717E4E85FD56BFDC9CE5" bold="true" box="[151,486,690,717]" fontSize="11" level="1" pageId="9" pageNumber="250" reason="1">
<taxonomicName id="4C1E4D6FA55F717E4E85FD56BFDC9CE5" ID-CoL="9C6QF" authority="Plotz, 1882" authorityName="sensu Plotz" authorityYear="1882" box="[151,486,690,717]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4E85FD56BFDC9CE5" bold="true" box="[151,486,690,717]" pageId="9" pageNumber="250">
<emphasis id="B96AEAFEA55F717E4E85FD56BF6F9CE4" bold="true" box="[151,341,690,717]" italics="true" pageId="9" pageNumber="250">Hesperia lyrcea</emphasis>
<bibRefCitation id="EF8F4B1DA55F717E4F4EFD56BFDC9CE5" author="Plotz, C." box="[348,486,690,717]" pageId="9" pageNumber="250" pagination="314 - 344" refId="ref12834" refString="Plotz, C. (1882) Die Hesperiinen-Gattung Hesperia Aut. Und ihre Arten. Entomologische Zeitung, Stettin, 43 (7 / 9 &amp; 10 / 12), 314 - 344 (July - September) &amp; 436 - 456 (October - December)." type="journal article" year="1882">Plötz, 1882</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C3046567A55F717E4E85FD1EBA1D9AC0" pageId="9" pageNumber="250" type="discussion">
<paragraph id="8BA136ECA55F717E4E85FD1EBA1D9AC0" blockId="9.[151,1437,763,2013]" pageId="9" pageNumber="250">
Carl Plötz described
<taxonomicName id="4C1E4D6FA55F717E4F94FD1EBC019D3C" authorityName="sensu Plotz" authorityYear="1882" box="[390,571,763,789]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4F94FD1EBC019D3C" box="[390,571,763,789]" italics="true" pageId="9" pageNumber="250">Hesperia lyrcea</emphasis>
</taxonomicName>
simultaneously in
<emphasis id="B96AEAFEA55F717E4D06FD1EBA1D9D3D" box="[788,1063,763,789]" italics="true" pageId="9" pageNumber="250">Entomologische Zeitung</emphasis>
(Stettin) in a page preceding his description of
<taxonomicName id="4C1E4D6FA55F717E4F29FCFABF809D10" baseAuthorityName="Plotz" baseAuthorityYear="1882" box="[315,442,799,825]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="aurelius">
<emphasis id="B96AEAFEA55F717E4F29FCFABF809D10" box="[315,442,799,825]" italics="true" pageId="9" pageNumber="250">H. aurelius</emphasis>
</taxonomicName>
, based on an unspecified number of specimens that were also from
<collectingCountry id="F309767CA55F717E4AD2FCFABB1F9D10" box="[1216,1317,799,825]" name="Brazil" pageId="9" pageNumber="250">Brasilien</collectingCountry>
(=
<collectingCountry id="F309767CA55F717E4B5CFCFABBAE9D10" box="[1358,1428,799,825]" name="Brazil" pageId="9" pageNumber="250">Brazil</collectingCountry>
) (
<bibRefCitation id="EF8F4B1DA55F717E4E8DFCA6BF6D9D74" author="Plotz, C." box="[159,343,834,861]" pageId="9" pageNumber="250" pagination="314 - 344" refId="ref12834" refString="Plotz, C. (1882) Die Hesperiinen-Gattung Hesperia Aut. Und ihre Arten. Entomologische Zeitung, Stettin, 43 (7 / 9 &amp; 10 / 12), 314 - 344 (July - September) &amp; 436 - 456 (October - December)." type="journal article" year="1882">Plötz 1882: 454</bibRefCitation>
). After examining illustrations prepared by Carl Plötz (see above under
<taxonomicName id="4C1E4D6FA55F717E4A6AFCA6BAE99D74" baseAuthorityName="Plotz" baseAuthorityYear="1882" box="[1144,1235,835,861]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="aurelius">
<emphasis id="B96AEAFEA55F717E4A6AFCA6BAE99D74" box="[1144,1235,835,861]" italics="true" pageId="9" pageNumber="250">aurelius</emphasis>
</taxonomicName>
),
<bibRefCitation id="EF8F4B1DA55F717E4AF8FCA6BBA69D74" author="Godman, F. D." box="[1258,1436,835,861]" pageId="9" pageNumber="250" pagination="132 - 155" refId="ref11450" refString="Godman, F. D. (1907) Notes on the American species of Hesperiidae described by Plotz. Annals and Magazine of natural History, Series 7, 20 (116), 132 - 155." type="journal article" year="1907">Godman (1907)</bibRefCitation>
synonymized
<taxonomicName id="4C1E4D6FA55F717E4F24FC82BF9A9DA8" authorityName="sensu Plotz" authorityYear="1882" box="[310,416,871,897]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4F24FC82BF9A9DA8" box="[310,416,871,897]" italics="true" pageId="9" pageNumber="250">H. lyrcea</emphasis>
</taxonomicName>
under
<emphasis id="B96AEAFEA55F717E4FE1FC82BC329DA9" box="[499,520,871,896]" italics="true" pageId="9" pageNumber="250">P.</emphasis>
“
<taxonomicName id="4C1E4D6FA55F717E4C0FFC82BC549DA9" authorityName="Fabricius" authorityYear="1787" box="[541,622,871,896]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="marcus">
<emphasis id="B96AEAFEA55F717E4C0FFC82BC549DA9" box="[541,622,871,896]" italics="true" pageId="9" pageNumber="250">marcus</emphasis>
</taxonomicName>
”.
<taxonomicName id="4C1E4D6FA55F717E4C9EFC82BD6D9DA8" authorityName="Plotz" authorityYear="1882" box="[652,855,871,897]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="aurelius">
<emphasis id="B96AEAFEA55F717E4C9EFC82BD6D9DA8" box="[652,855,871,897]" italics="true" pageId="9" pageNumber="250">Hesperia aurelius</emphasis>
</taxonomicName>
, which supposedly appeared in Carl Plötz’s plates for his new skipper species, was not commented on in
<bibRefCitation id="EF8F4B1DA55F717E4D18FC6EBDFA9D8C" author="Godman, F. D." box="[778,960,907,933]" pageId="9" pageNumber="250" pagination="132 - 155" refId="ref11450" refString="Godman, F. D. (1907) Notes on the American species of Hesperiidae described by Plotz. Annals and Magazine of natural History, Series 7, 20 (116), 132 - 155." type="journal article" year="1907">Godman (1907)</bibRefCitation>
, perhaps because
<bibRefCitation id="EF8F4B1DA55F717E4A9CFC6EBB7B9D8C" author="Godman, F. D." box="[1166,1345,907,933]" pageId="9" pageNumber="250" refId="ref11292" refString="Godman, F. D. (1900) Biologia Centrali-Americana. Insecta. Lepidoptera- Rhopalocera. 1 (168), 2 (148, 156, 157, 158, 159 &amp; 160), Dulau &amp; Co., Bernard Quaritch, London, pp. i - xlvi ([28] November 1901), pp. 457 - 460, pl. 91 (June 1899), pp. 461 - 484, pl. 92 ([28] June 1900), pp. 485 - 500, pls. 93 - 94 (August 1900), pp. 501 - 532, pls. 95 - 96 ([27] October 1900), pp. 533 - 556, pls. 97 - 98 (November 1900), pp. 557 - 588, pls. 99 - 100 (December 1900)." type="journal volume" year="1900">Godman (1900)</bibRefCitation>
already discussed its conspecificity with
<emphasis id="B96AEAFEA55F717E4C1FFC4ABC199DE1" box="[525,547,943,968]" italics="true" pageId="9" pageNumber="250">P.</emphasis>
“
<taxonomicName id="4C1E4D6FA55F717E4C25FC4ABCB29DE1" authorityName="Fabricius" authorityYear="1787" box="[567,648,943,968]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="marcus">
<emphasis id="B96AEAFEA55F717E4C25FC4ABCB29DE1" box="[567,648,943,968]" italics="true" pageId="9" pageNumber="250">marcus</emphasis>
</taxonomicName>
” as discussed above.
<bibRefCitation id="EF8F4B1DA55F717E4D91FC4ABA649DE0" author="Draudt, M. W. K." box="[899,1118,943,969]" pageId="9" pageNumber="250" pagination="913 - 952" refId="ref10588" refString="Draudt, M. W. K. (1923) B. Grypocera, breitko ¨ pfige Tagfalter. In: Seitz, A. (Ed)., Die Gross-Schmetterlinge der Erde. Vol. 5. Alfred Kernen. Stuttgart, pp. 905 - 912 (8 February [1923]) + 913 - 952 (23 May 1923) + 953 - 992 (9 July 1923) + 993 - 1000 (29 December 1923)." type="journal article" year="1923">Draudt (1923: 979)</bibRefCitation>
followed this synonymy by Godman.
<bibRefCitation id="EF8F4B1DA55F717E4F14FC36BFEF9DC4" author="Evans, W. H." box="[262,469,978,1005]" pageId="9" pageNumber="250" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans (1955: 183)</bibRefCitation>
considered
<taxonomicName id="4C1E4D6FA55F717E4C4CFC36BC999DC4" authorityName="sensu Plotz" authorityYear="1882" box="[606,675,979,1005]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4C4CFC36BC999DC4" box="[606,675,979,1005]" italics="true" pageId="9" pageNumber="250">lyrcea</emphasis>
</taxonomicName>
as a subspecies of
<taxonomicName id="4C1E4D6FA55F717E4D69FC36BB049DC4" authority="(Herrich-Schaffer, 1869)" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1869" box="[891,1342,979,1005]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="diversa">
<emphasis id="B96AEAFEA55F717E4D69FC36BA259DC4" box="[891,1055,979,1005]" italics="true" pageId="9" pageNumber="250">Vettius diversa</emphasis>
(Herrich-Schäffer, 1869)
</taxonomicName>
, a taxon initially described as
<taxonomicName id="4C1E4D6FA55F717E4F9EFC12BC7F9A38" authorityName="Herrich-Schäffer" authorityYear="1869" box="[396,581,1015,1041]" class="Insecta" family="Hesperiidae" genus="Cobalus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="diversa">
<emphasis id="B96AEAFEA55F717E4F9EFC12BC7F9A38" box="[396,581,1015,1041]" italics="true" pageId="9" pageNumber="250">Cobalus diversa</emphasis>
</taxonomicName>
.
<bibRefCitation id="EF8F4B1DA55F717E4C47FC12BCD79A38" author="Evans, W. H." box="[597,749,1014,1041]" pageId="9" pageNumber="250" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans (1955)</bibRefCitation>
refers to figures for “
<taxonomicName id="4C1E4D6FA55F717E4DF9FC12BA079A38" authorityName="Herrich-Schäffer" authorityYear="1869" box="[1003,1085,1015,1041]" class="Insecta" family="Hesperiidae" genus="Cobalus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="diversa">
<emphasis id="B96AEAFEA55F717E4DF9FC12BA079A38" box="[1003,1085,1015,1041]" italics="true" pageId="9" pageNumber="250">diversa</emphasis>
</taxonomicName>
” (i.e.,
<emphasis id="B96AEAFEA55F717E4A9CFC12BAD19A38" box="[1166,1259,1015,1041]" italics="true" pageId="9" pageNumber="250">diversus</emphasis>
) in Godman &amp; Salvin (=
<bibRefCitation id="EF8F4B1DA55F717E4F17FBFEBF9F9A1C" author="Godman, F. D." box="[261,421,1051,1077]" pageId="9" pageNumber="250" refId="ref11292" refString="Godman, F. D. (1900) Biologia Centrali-Americana. Insecta. Lepidoptera- Rhopalocera. 1 (168), 2 (148, 156, 157, 158, 159 &amp; 160), Dulau &amp; Co., Bernard Quaritch, London, pp. i - xlvi ([28] November 1901), pp. 457 - 460, pl. 91 (June 1899), pp. 461 - 484, pl. 92 ([28] June 1900), pp. 485 - 500, pls. 93 - 94 (August 1900), pp. 501 - 532, pls. 95 - 96 ([27] October 1900), pp. 533 - 556, pls. 97 - 98 (November 1900), pp. 557 - 588, pls. 99 - 100 (December 1900)." type="journal volume" year="1900">Godman 1900</bibRefCitation>
) and Seitz (=
<bibRefCitation id="EF8F4B1DA55F717E4C53FBFEBCF59A1C" author="Draudt, M. W. K." box="[577,719,1051,1077]" pageId="9" pageNumber="250" pagination="913 - 952" refId="ref10588" refString="Draudt, M. W. K. (1923) B. Grypocera, breitko ¨ pfige Tagfalter. In: Seitz, A. (Ed)., Die Gross-Schmetterlinge der Erde. Vol. 5. Alfred Kernen. Stuttgart, pp. 905 - 912 (8 February [1923]) + 913 - 952 (23 May 1923) + 953 - 992 (9 July 1923) + 993 - 1000 (29 December 1923)." type="journal article" year="1923">Draudt 1923</bibRefCitation>
) as
<taxonomicName id="4C1E4D6FA55F717E4CEEFBFEBD7B9A1C" authorityName="sensu Plotz" authorityYear="1882" box="[764,833,1051,1077]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4CEEFBFEBD7B9A1C" box="[764,833,1051,1077]" italics="true" pageId="9" pageNumber="250">lyrcea</emphasis>
</taxonomicName>
without any explanation as to these associations. This decision by
<bibRefCitation id="EF8F4B1DA55F717E4F0EFBDABF8B9A70" author="Evans, W. H." box="[284,433,1086,1113]" pageId="9" pageNumber="250" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans (1955)</bibRefCitation>
was presumably based mainly on the fact that
<taxonomicName id="4C1E4D6FA55F717E4DA6FBDABDC39A70" authorityName="sensu Plotz" authorityYear="1882" box="[948,1017,1087,1113]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4DA6FBDABDC39A70" box="[948,1017,1087,1113]" italics="true" pageId="9" pageNumber="250">lyrcea</emphasis>
</taxonomicName>
possesses dark veins on the hindwing (thus dividing yellowish ventral hindwing markings), whereas veins are not dark in
<emphasis id="B96AEAFEA55F717E4A42FB86BA5F9A55" box="[1104,1125,1123,1148]" italics="true" pageId="9" pageNumber="250">P.</emphasis>
“
<taxonomicName id="4C1E4D6FA55F717E4A6BFB86BAF09A55" authorityName="Fabricius" authorityYear="1787" box="[1145,1226,1123,1148]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="marcus">
<emphasis id="B96AEAFEA55F717E4A6BFB86BAF09A55" box="[1145,1226,1123,1148]" italics="true" pageId="9" pageNumber="250">marcus</emphasis>
</taxonomicName>
”, a character used to distinguish
<taxonomicName id="4C1E4D6FA55F717E4F25FB62BF9C9A88" authorityName="Herrich-Schäffer" authorityYear="1869" box="[311,422,1159,1185]" class="Insecta" family="Hesperiidae" genus="Cobalus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="diversa">
<emphasis id="B96AEAFEA55F717E4F25FB62BF9C9A88" box="[311,422,1159,1185]" italics="true" pageId="9" pageNumber="250">V. diversa</emphasis>
</taxonomicName>
from
<taxonomicName id="4C1E4D6FA55F717E4FFEFB62BC639A89" authorityName="Fabricius" authorityYear="1787" box="[492,601,1159,1184]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="marcus">
<emphasis id="B96AEAFEA55F717E4FFEFB62BC639A89" box="[492,601,1159,1184]" italics="true" pageId="9" pageNumber="250">V. marcus</emphasis>
</taxonomicName>
in
<bibRefCitation id="EF8F4B1DA55F717E4C6DFB62BD209A88" author="Evans, W. H." box="[639,794,1158,1185]" pageId="9" pageNumber="250" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans (1955)</bibRefCitation>
. The internal view of the valva figured for
<taxonomicName id="4C1E4D6FA55F717E4B10FB62BB7D9A88" authorityName="sensu Plotz" authorityYear="1882" box="[1282,1351,1159,1185]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4B10FB62BB7D9A88" box="[1282,1351,1159,1185]" italics="true" pageId="9" pageNumber="250">lyrcea</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8F4B1DA55F717E4B44FB62BEE99AEC" author="Evans, W. H." pageId="9" pageNumber="250" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans 1955</bibRefCitation>
: pl. 66) illustrates differences in the harpe and overall shape of the valva compared to other potential congeners figured on the same plate, including
<taxonomicName id="4C1E4D6FA55F717E4C23FB2ABCA79AC1" authorityName="Fabricius" authorityYear="1787" box="[561,669,1231,1256]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="marcus">
<emphasis id="B96AEAFEA55F717E4C23FB2ABCA79AC1" box="[561,669,1231,1256]" italics="true" pageId="9" pageNumber="250">V. marcus</emphasis>
</taxonomicName>
and other subspecies of
<taxonomicName id="4C1E4D6FA55F717E4DA0FB2ABA259AC0" authorityName="Herrich-Schäffer" authorityYear="1869" box="[946,1055,1231,1257]" class="Insecta" family="Hesperiidae" genus="Cobalus" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="diversa">
<emphasis id="B96AEAFEA55F717E4DA0FB2ABA259AC0" box="[946,1055,1231,1257]" italics="true" pageId="9" pageNumber="250">V. diversa</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3046567A55F717D4ED5FB16BCFF9F34" lastPageId="10" lastPageNumber="251" pageId="9" pageNumber="250" type="materials_examined">
<paragraph id="8BA136ECA55F717D4ED5FB16BCFF9F34" blockId="9.[151,1437,763,2013]" lastBlockId="10.[151,1437,151,285]" lastPageId="10" lastPageNumber="251" pageId="9" pageNumber="250">
A review of specimens at
<collectionCode id="ED0FAE29A55F717E4FE3FB16BC619B24" box="[497,603,1267,1293]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="250" type="Museum">NHMUK</collectionCode>
reveals
<specimenCount id="9D18FD65A55F717E4CAEFB16BD6C9B24" box="[700,854,1267,1293]" pageId="9" pageNumber="250" type="generic">32 specimens</specimenCount>
of
<taxonomicName id="4C1E4D6FA55F717E4D6CFB16BDF99B24" authorityName="sensu Plotz" authorityYear="1882" box="[894,963,1267,1293]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4D6CFB16BDF99B24" box="[894,963,1267,1293]" italics="true" pageId="9" pageNumber="250">lyrcea</emphasis>
</taxonomicName>
that show consistent differences between
<taxonomicName id="4C1E4D6FA55F717E4E85FAF2BED89B19" authorityName="Warren &amp; Turland" authorityYear="2012" box="[151,226,1303,1328]" class="Insecta" family="Hesperiidae" genus="Troyus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="genus">
<emphasis id="B96AEAFEA55F717E4E85FAF2BED89B19" box="[151,226,1303,1328]" italics="true" pageId="9" pageNumber="250">Troyus</emphasis>
</taxonomicName>
“
<taxonomicName id="4C1E4D6FA55F717E4EE6FAF2BF7F9B19" authorityName="Fabricius" authorityYear="1787" box="[244,325,1303,1328]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="marcus">
<emphasis id="B96AEAFEA55F717E4EE6FAF2BF7F9B19" box="[244,325,1303,1328]" italics="true" pageId="9" pageNumber="250">marcus</emphasis>
</taxonomicName>
” mainly by Rs, M
<subScript id="179A34A9A55F717E4C1CFACCBC2C9B1E" attach="left" box="[526,534,1321,1335]" fontSize="6" pageId="9" pageNumber="250">3</subScript>
and Cu
<subScript id="179A34A9A55F717E4C78FACCBC489B1E" attach="left" box="[618,626,1321,1335]" fontSize="6" pageId="9" pageNumber="250">1</subScript>
of the ventral hindwing being dark and thus dividing yellowish or whitish markings.
<bibRefCitation id="EF8F4B1DA55F717E4F1EFADEBF9B9B7C" author="Evans, W. H." box="[268,417,1338,1365]" pageId="9" pageNumber="250" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans (1955)</bibRefCitation>
examined
<specimenCount id="9D18FD65A55F717E4C08FADEBC8B9B7C" box="[538,689,1339,1365]" pageId="9" pageNumber="250" type="generic">34 specimens</specimenCount>
in total, 33 from
<collectingCountry id="F309767CA55F717E4D62FADEBDF79B7C" box="[880,973,1339,1365]" name="Ecuador" pageId="9" pageNumber="250">Ecuador</collectingCountry>
and a single male from
<collectingCountry id="F309767CA55F717E4AC7FADEBB369B7C" box="[1237,1292,1339,1365]" name="Peru" pageId="9" pageNumber="250">Peru</collectingCountry>
. There exists space for
<specimenCount id="9D18FD65A55F717E4F10FABABF9D9B50" box="[258,423,1375,1401]" pageId="9" pageNumber="250" type="generic">two specimens</specimenCount>
in the corresponding section of the drawer in the
<collectionCode id="ED0FAE29A55F717E4DD8FABABA0D9B50" box="[970,1079,1375,1401]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="250" type="Museum">NHMUK</collectionCode>
, thus it is reasonable to consider that these were the specimens examined by Evans to investigate the identity of
<taxonomicName id="4C1E4D6FA55F717E4A02FA66BA6F9BB4" authorityName="sensu Plotz" authorityYear="1882" box="[1040,1109,1411,1437]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4A02FA66BA6F9BB4" box="[1040,1109,1411,1437]" italics="true" pageId="9" pageNumber="250">lyrcea</emphasis>
</taxonomicName>
. Furthermore, we found
<specimenCount id="9D18FD65A55F717E4B7CFA66BF369BE8" pageId="9" pageNumber="250" type="generic">four specimens</specimenCount>
without abdomens, as well as
<specimenCount id="9D18FD65A55F717E4C71FA42BD339BE8" box="[611,777,1447,1473]" pageId="9" pageNumber="250" type="generic">two specimens</specimenCount>
with apparently intact abdomens with their genitalia glued on a card that is pinned beneath them. They were presumably dissected by humidifying the abdomens and extracting the genitalia without removing the abdomens from the specimens. In the preceding work (
<bibRefCitation id="EF8F4B1DA55F717E4AB2FA0ABB119820" author="Evans, W. H." box="[1184,1323,1519,1545]" pageId="9" pageNumber="250" refId="ref10711" refString="Evans, W. H. (1949) A Catalogue of the Hesperiidae from Europe, Asia, and Australia in the British Museum (Natural History). British Museum (Natural History), London, 502 pp. https: // doi. org / 10.5962 / bhl. title. 105941" type="book" year="1949">Evans 1949</bibRefCitation>
), Norman D. Riley stated that Evans’ intention was to provide “diagrammatic” illustrations of male genitalia, and given that any of these six individuals (summary of multiple, or none of them) might have been used to establish the identity of
<taxonomicName id="4C1E4D6FA55F717E4EA5F9BEBFDD985C" authority="sensu Evans (1955)" authorityName="sensu Evans" authorityYear="1955" box="[183,487,1626,1653]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4EA5F9BEBF79985D" box="[183,323,1627,1653]" italics="true" pageId="9" pageNumber="250">lyrcea sensu</emphasis>
<bibRefCitation id="EF8F4B1DA55F717E4F5FF9BEBFDD985C" author="Evans, W. H." box="[333,487,1626,1653]" pageId="9" pageNumber="250" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans (1955)</bibRefCitation>
</taxonomicName>
, it is impossible to draw conclusions as to how
<bibRefCitation id="EF8F4B1DA55F717E4A02F9BEBA92985C" author="Evans, W. H." box="[1040,1192,1626,1653]" pageId="9" pageNumber="250" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans (1955)</bibRefCitation>
identified this taxon. Carl Plötz’s key leading to
<taxonomicName id="4C1E4D6FA55F717E4FC2F99ABC2F98B0" authorityName="sensu Evans" authorityYear="1955" box="[464,533,1663,1689]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4FC2F99ABC2F98B0" box="[464,533,1663,1689]" italics="true" pageId="9" pageNumber="250">lyrcea</emphasis>
</taxonomicName>
is questionable as to its reliability in distinguishing this taxon from congeners, namely by stating “
<emphasis id="B96AEAFEA55F717E4F61F946BE9798C9" italics="true" pageId="9" pageNumber="250">
the pale spot beneath in cells 4 [M
<subScript id="179A34A9A55F717E4CE9F950BD3998EA" attach="left" box="[763,771,1717,1731]" fontSize="6" pageId="9" pageNumber="250">2</subScript>
-M
<subScript id="179A34A9A55F717E4D36F950BD1698EA" attach="left" box="[804,812,1717,1731]" fontSize="6" pageId="9" pageNumber="250">3</subScript>
] and 5 [M
<subScript id="179A34A9A55F717E4DBAF950BD8A98EA" attach="left" box="[936,944,1717,1731]" fontSize="6" pageId="9" pageNumber="250">1</subScript>
-M
<subScript id="179A34A9A55F717E4DC3F950BDE398EA" attach="left" box="[977,985,1717,1731]" fontSize="6" pageId="9" pageNumber="250">2</subScript>
] of the FW is sharp delimited [opposed to
</emphasis>
“
<emphasis id="B96AEAFEA55F717E4ED2F922BCFA98C8" box="[192,704,1735,1761]" italics="true" pageId="9" pageNumber="250">the lower hyaline apical spot is washed out”]</emphasis>
”. Even based solely on specimens in the drawer in the
<collectionCode id="ED0FAE29A55F717E4B39F922BBA298C8" box="[1323,1432,1735,1761]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="9" pageNumber="250" type="Museum">NHMUK</collectionCode>
, these spots in the ventral hindwing cells M
<subScript id="179A34A9A55F717E4C9CF918BCAC9922" attach="left" box="[654,662,1789,1803]" fontSize="6" pageId="9" pageNumber="250">2</subScript>
-M
<subScript id="179A34A9A55F717E4CABF918BCFB9922" attach="left" box="[697,705,1789,1803]" fontSize="6" pageId="9" pageNumber="250">3</subScript>
and M
<subScript id="179A34A9A55F717E4D05F918BD259922" attach="left" box="[791,799,1789,1803]" fontSize="6" pageId="9" pageNumber="250">1</subScript>
-M
<subScript id="179A34A9A55F717E4D53F918BD739922" attach="left" box="[833,841,1789,1803]" fontSize="6" pageId="9" pageNumber="250">2</subScript>
do not appear to show inter-specific differences in these taxa. Plötz therefore likely examined limited numbers of specimens, or perhaps single specimens, to prepare his descriptions. Despite the lack of type material and original illustrations, the most up-to-date comprehensive catalogue for
<taxonomicName id="4C1E4D6FA55F717E4F27F8B2BF819958" box="[309,443,1879,1905]" class="Insecta" family="Hesperiidae" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="family">Hesperiidae</taxonomicName>
retains the taxonomic status of
<taxonomicName id="4C1E4D6FA55F717E4D3BF8B2BD549958" authorityName="sensu Evans" authorityYear="1955" box="[809,878,1879,1905]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4D3BF8B2BD549958" box="[809,878,1879,1905]" italics="true" pageId="9" pageNumber="250">lyrcea</emphasis>
</taxonomicName>
as a subspecies of
<taxonomicName id="4C1E4D6FA55F717E4A5FF8B2BAC99958" baseAuthorityName="Herrich-Schaffer" baseAuthorityYear="1869" box="[1101,1267,1879,1905]" class="Insecta" family="Hesperiidae" genus="Vettius" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="diversa">
<emphasis id="B96AEAFEA55F717E4A5FF8B2BAC99958" box="[1101,1267,1879,1905]" italics="true" pageId="9" pageNumber="250">Vettius diversa</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF8F4B1DA55F717E4B11F8B2BBA29958" author="Mielke, O. H. H." box="[1283,1432,1879,1905]" pageId="9" pageNumber="250" pagination="3 - 11" refId="ref12282" refString="Mielke, O. H. H. (2004) Hesperiidae. In: Lamas, G. (Ed.), Checklist. Part 4 a. Hesperioidea - Papilionoidea. In: Heppner, J. B. (Ed.), Atlas of Neotropical Lepidoptera. Vol. 5 A. Association of Tropical Lepidoptera, Gainsville, pp. 3 - 11 + 25 - 86." type="book chapter" year="2004">Mielke 2004</bibRefCitation>
,
<bibRefCitation id="EF8F4B1DA55F717E4E85F89FBEE999BC" author="Mielke, O. H. H." box="[151,211,1914,1941]" pageId="9" pageNumber="250" refId="ref12356" refString="Mielke, O. H. H. (2005) Catalogue of the American Hesperioidea: Hesperiidae (Lepidoptera). Vols. 1 - 6. Sociedade Brasileira de Zoologia, Curitiba, 1536 pp." type="book" year="2005">2005</bibRefCitation>
), in accordance with
<bibRefCitation id="EF8F4B1DA55F717E4FD2F89EBC6099BC" author="Evans, W. H." box="[448,602,1914,1941]" pageId="9" pageNumber="250" refId="ref10918" refString="Evans, W. H. (1955) A catalogue of the American Hesperiidae indicating the classification and nomenclature adopted in the British Museum (Natural History). Part IV. Hesperiinae and Megathyminae. British Museum (Natural History), London, 499 pp." type="book" year="1955">Evans (1955)</bibRefCitation>
. As stated above, our attempt to locate
<typeStatus id="54A5884EA55F717E4A1CF89EBA5099BC" box="[1038,1130,1915,1941]" pageId="9" pageNumber="250" type="syntype">syntype</typeStatus>
(s) at
<collectionCode id="ED0FAE29A55F717E4AB7F89EBAD799BC" box="[1189,1261,1915,1941]" country="Germany" httpUri="http://grbio.org/cool/a2g1-8ssa" name="Zoologisches Institut und Museum Greifswald" pageId="9" pageNumber="250" type="University or college">ZIMG</collectionCode>
and
<collectionCode id="ED0FAE29A55F717E4B30F89EBB4B99BC" box="[1314,1393,1915,1941]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="9" pageNumber="250">MFNB</collectionCode>
did not yield any specimens which could represent Plötz’s type material relevant to the present study. While there appears to be no ambiguity as to Evans’ (1955) concept of
<taxonomicName id="4C1E4D6FA55F717E4CC4F826BD2199F4" authorityName="sensu Evans" authorityYear="1955" box="[726,795,1987,2013]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4CC4F826BD2199F4" box="[726,795,1987,2013]" italics="true" pageId="9" pageNumber="250">lyrcea</emphasis>
</taxonomicName>
, we are unable to apply the specific epithet
<taxonomicName id="4C1E4D6FA55F717D4B03F826BF189E98" authority="sensu Plotz (1882)" authorityName="sensu Plotz" authorityYear="1882" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" lastPageId="10" lastPageNumber="251" order="Lepidoptera" pageId="9" pageNumber="250" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55F717E4B03F826BBA699F5" box="[1297,1436,1987,2013]" italics="true" pageId="9" pageNumber="250">lyrcea sensu</emphasis>
<bibRefCitation id="EF8F4B1DA55C717D4E85FF72BF189E98" author="Plotz, C." box="[151,290,151,177]" pageId="10" pageNumber="251" pagination="314 - 344" refId="ref12834" refString="Plotz, C. (1882) Die Hesperiinen-Gattung Hesperia Aut. Und ihre Arten. Entomologische Zeitung, Stettin, 43 (7 / 9 &amp; 10 / 12), 314 - 344 (July - September) &amp; 436 - 456 (October - December)." type="journal article" year="1882">Plötz (1882)</bibRefCitation>
</taxonomicName>
mainly due to the lack of type material. In the present study, we refrain from drawing any conclusions regarding the identity of
<taxonomicName id="4C1E4D6FA55C717D4FAAFF5EBC549EFC" authorityName="sensu Plotz" authorityYear="1882" box="[440,622,187,213]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55C717D4FAAFF5EBC549EFC" box="[440,622,187,213]" italics="true" pageId="10" pageNumber="251">Hesperia lyrcea</emphasis>
</taxonomicName>
, which may or may not represent a species whose concept matches
<emphasis id="B96AEAFEA55C717D4B9AFF5EBBA79EFD" box="[1416,1437,187,212]" italics="true" pageId="10" pageNumber="251">P.</emphasis>
“
<taxonomicName id="4C1E4D6FA55C717D4EB6FF3ABECF9ED1" authorityName="Fabricius" authorityYear="1787" box="[164,245,223,248]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
<emphasis id="B96AEAFEA55C717D4EB6FF3ABECF9ED1" box="[164,245,223,248]" italics="true" pageId="10" pageNumber="251">marcus</emphasis>
</taxonomicName>
”. Under the current circumstances, we will not designate a
<typeStatus id="54A5884EA55C717D4D82FF3ABDD39ED0" box="[912,1001,223,249]" pageId="10" pageNumber="251" type="neotype">neotype</typeStatus>
for
<taxonomicName id="4C1E4D6FA55C717D4A05FF3ABA449ED0" authorityName="sensu Plotz" authorityYear="1882" box="[1047,1150,223,249]" class="Insecta" family="Hesperiidae" genus="Hesperia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="lyrcea">
<emphasis id="B96AEAFEA55C717D4A05FF3ABA449ED0" box="[1047,1150,223,249]" italics="true" pageId="10" pageNumber="251">H. lyrcea</emphasis>
</taxonomicName>
, and consequently do not regard this taxon as conspecific with
<emphasis id="B96AEAFEA55C717D4C2AFEE6BC779F35" box="[568,589,259,284]" italics="true" pageId="10" pageNumber="251">P.</emphasis>
“
<taxonomicName id="4C1E4D6FA55C717D4C72FEE6BC8B9F35" authorityName="Fabricius" authorityYear="1787" box="[608,689,259,284]" class="Insecta" family="Papilionidae" genus="Papilio" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="251" phylum="Arthropoda" rank="species" species="marcus">
<emphasis id="B96AEAFEA55C717D4C72FEE6BC8B9F35" box="[608,689,259,284]" italics="true" pageId="10" pageNumber="251">marcus</emphasis>
</taxonomicName>
”.
</paragraph>
</subSubSection>
</treatment>
</document>
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